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<mods:title id="D4ED4F5BE9EBE5BF86185A6081BB53B2">The genus Leucophenga (Diptera, Drosophilidae), part VII: the subpollinosa species group from China, with morphological and molecular evidence</mods:title>
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<paragraph id="0450B078FFBFFFF9FEF0FBEEFDB30448" blockId="17.[151,587,1093,1153]" box="[151,587,1093,1119]" pageId="17" pageNumber="218">
<heading id="5F180714FFBFFFF9FEF0FBEEFDB30448" bold="true" box="[151,587,1093,1119]" fontSize="11" level="1" pageId="17" pageNumber="218" reason="1">
<taxonomicName id="C3EFCBFBFFBFFFF9FEF0FBEEFDB30448" ID-CoL="8CJFD" authority="Duda, 1926" authorityName="Duda" authorityYear="1926" box="[151,587,1093,1119]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="flavicosta">
<emphasis id="369B6C6AFFBFFFF9FEF0FBEEFDB30448" bold="true" box="[151,587,1093,1119]" pageId="17" pageNumber="218">
<emphasis id="369B6C6AFFBFFFF9FEF0FBEEFE4E0448" bold="true" box="[151,438,1093,1119]" italics="true" pageId="17" pageNumber="218">Leucophenga flavicosta</emphasis>
<bibRefCitation id="607ECD89FFBFFFF9FFDAFBEEFDB30448" author="Duda" box="[445,587,1093,1119]" pageId="17" pageNumber="218" refString="Duda, O. (1926) Fauna sumatrensis. (Beitrag Nr. 26). Drosophilidae (Dipt.). Supplementa Entomologica, 14, 42 - 116." type="journal article" year="1926">Duda, 1926</bibRefCitation>
</emphasis>
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<paragraph id="0450B078FFBFFFF9FEF0FBC3FE470497" blockId="17.[151,587,1093,1153]" box="[151,447,1128,1153]" pageId="17" pageNumber="218">
(
<figureCitation id="9CD4ACFDFFBFFFF9FEF8FBC3FF150497" box="[159,237,1128,1153]" captionStart="FIGURE 2" captionStartId="8.[151,250,1530,1552]" captionTargetBox="[151,1436,193,1508]" captionTargetId="figure@8.[151,1436,193,1508]" captionTargetPageId="8" captionText="FIGURE 2. Wing of male (except for Leucophenga subpollinosa): A. L. argentina; B. L. flavicosta; C. L. formosa; D. L. nigroscutellata; E. L. subpollinosa; F. L. umbratula; G. L. aculeata sp. nov.; H. L. acuticauda sp. nov.; I. L. cultella sp. nov.; J. L. cyclophylla sp. nov.; K. L. flaviclypeata sp. nov.; L. L. fuscipedes sp. nov.; M. L. gracilenta sp. nov.; N. L. latipenis sp. nov.; O. L. magnicauda sp. nov.; P. L. rhombura sp. nov.; Q. L. rugatifolia sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/438518/files/figure.png" pageId="17" pageNumber="218">Figs. 2</figureCitation>
B, 3C, 3D, 7C, 11)
</paragraph>
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<paragraph id="0450B078FFBFFFF9FEF0FB1BFD1404D1" blockId="17.[151,1048,1200,1253]" box="[151,748,1200,1222]" pageId="17" pageNumber="218">
<treatmentCitationGroup id="24FF9756FFBFFFF9FEF0FB1BFD1404D1" box="[151,748,1200,1222]" pageId="17" pageNumber="218">
<taxonomicName id="C3EFCBFBFFBFFFF9FEF0FB1BFD1F04D1" ID-CoL="8CJFD" authority="Duda, 1926: 53" authorityName="Duda" authorityPageNumber="53" authorityYear="1926" box="[151,743,1200,1222]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="variety" species="subpollinosa" variety="flavicosta">
<emphasis id="369B6C6AFFBFFFF9FEF0FB1BFE5E04D1" box="[151,422,1200,1222]" italics="true" pageId="17" pageNumber="218">Leucophenga subpollinosa</emphasis>
var.
<emphasis id="369B6C6AFFBFFFF9FFBEFB1BFDC404D1" box="[473,572,1200,1222]" italics="true" pageId="17" pageNumber="218">flavicosta</emphasis>
,
<treatmentCitation id="854E9669FFBFFFF9FC2FFB1BFD1F04D1" author="Duda" box="[584,743,1200,1222]" page="53" pageId="17" pageNumber="218" year="1926">
<bibRefCitation id="607ECD89FFBFFFF9FC2FFB1BFD3A04D1" author="Duda" box="[584,706,1200,1222]" pageId="17" pageNumber="218" refString="Duda, O. (1926) Fauna sumatrensis. (Beitrag Nr. 26). Drosophilidae (Dipt.). Supplementa Entomologica, 14, 42 - 116." type="journal article" year="1926">Duda, 1926</bibRefCitation>
: 53
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.
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<treatmentCitationGroup id="24FF9756FFBFFFF9FEF0FB64FBE004F2" box="[151,1048,1231,1253]" pageId="17" pageNumber="218">
<taxonomicName id="C3EFCBFBFFBFFFF9FEF0FB64FDC104F2" ID-CoL="8CJFD" authority="Bachli, 1971: 59" authorityName="Bachli" authorityPageNumber="59" authorityYear="1971" box="[151,569,1231,1253]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="flavicosta">
<emphasis id="369B6C6AFFBFFFF9FEF0FB64FE7004F2" box="[151,392,1231,1253]" italics="true" pageId="17" pageNumber="218">Leucophenga flavicosta</emphasis>
<treatmentCitation id="854E9669FFBFFFF9FFE9FB64FDC104F2" author="Bachli" box="[398,569,1231,1253]" page="59" pageId="17" pageNumber="218" year="1971">
<bibRefCitation id="607ECD89FFBFFFF9FFE9FB64FDEB04F2" author="Bachli" box="[398,531,1231,1253]" pageId="17" pageNumber="218" refString="Bachli, G. (1971) Leucophenga und Paraleucophenga (Diptera Brachycera), Fam. Drosophilidae. Exploration du Parc National de l'Upemba, Fascicule 71, 1 - 192 + 39 pls." type="journal article" year="1971">Bächli, 1971</bibRefCitation>
: 59
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="854E9669FFBFFFF9FC24FB64FCAA04F2" author="Lin" box="[579,850,1231,1253]" page="245" pageId="17" pageNumber="218" year="1972">
<bibRefCitation id="607ECD89FFBFFFF9FC24FB64FCE704F2" author="Lin" box="[579,799,1231,1253]" pageId="17" pageNumber="218" refString="Lin, F. J. &amp; Wheeler, M. R. (1972) The Drosophilidae of Taiwan, I. Genera Leucophenga and Paraleucophenga. The University of Texas Publication, 7213, 237 - 256." type="journal article" year="1972">Lin &amp; Wheeler, 1972</bibRefCitation>
: 245
</treatmentCitation>
;
<treatmentCitation id="854E9669FFBFFFF9FD3BFB64FBEB04F2" author="Okada" box="[860,1043,1231,1253]" page="679" pageId="17" pageNumber="218" year="1987">
<bibRefCitation id="607ECD89FFBFFFF9FD3BFB64FC1804F2" author="Okada" box="[860,992,1231,1253]" pageId="17" pageNumber="218" refString="Okada, T. (1987) The Leucophenga subpollinosa group of Japan and Southeast Asia (Diptera, Drosophilidae). Kontyu, 55 (4), 676 - 683." type="journal article" year="1987">Okada, 1987</bibRefCitation>
: 679
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.
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</paragraph>
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<paragraph id="0450B078FFBFFFF9FEF0FABAFA870562" blockId="17.[151,1437,1297,1939]" pageId="17" pageNumber="218">
<emphasis id="369B6C6AFFBFFFF9FEF0FABAFEE9053D" bold="true" box="[151,273,1297,1322]" pageId="17" pageNumber="218">Diagnosis.</emphasis>
This species differs from the other species of the
<taxonomicName id="C3EFCBFBFFBFFFF9FD25FABAFC29053E" box="[834,977,1297,1321]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="subpollinosa">
<emphasis id="369B6C6AFFBFFFF9FD25FABAFC29053E" box="[834,977,1297,1321]" italics="true" pageId="17" pageNumber="218">subpollinosa</emphasis>
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group by having the wing fuscous along R1 only (
<figureCitation id="9CD4ACFDFFBFFFF9FE9BFA9EFEB90559" box="[252,321,1333,1358]" captionStart="FIGURE 2" captionStartId="8.[151,250,1530,1552]" captionTargetBox="[151,1436,193,1508]" captionTargetId="figure@8.[151,1436,193,1508]" captionTargetPageId="8" captionText="FIGURE 2. Wing of male (except for Leucophenga subpollinosa): A. L. argentina; B. L. flavicosta; C. L. formosa; D. L. nigroscutellata; E. L. subpollinosa; F. L. umbratula; G. L. aculeata sp. nov.; H. L. acuticauda sp. nov.; I. L. cultella sp. nov.; J. L. cyclophylla sp. nov.; K. L. flaviclypeata sp. nov.; L. L. fuscipedes sp. nov.; M. L. gracilenta sp. nov.; N. L. latipenis sp. nov.; O. L. magnicauda sp. nov.; P. L. rhombura sp. nov.; Q. L. rugatifolia sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/438518/files/figure.png" pageId="17" pageNumber="218">Fig. 2</figureCitation>
B), the halter yellow at base and stem, mostly dark brown at knob (
<figureCitation id="9CD4ACFDFFBFFFF9FA21FA9EFB720559" box="[1094,1162,1333,1358]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[331,1245,194,1938]" captionTargetId="figure@9.[331,1245,193,1938]" captionTargetPageId="9" captionText="FIGURE 3. Dorsal and lateral view of thorax in male (except for Leucophenga subpollinosa): A, B. L. argentina; C, D. L. flavicosta; E, F. L. formosa; G, H. L. nigroscutellata; I, J. L. subpollinosa." httpUri="https://zenodo.org/record/438519/files/figure.png" pageId="17" pageNumber="218">Fig. 3</figureCitation>
C, D), and the aedeagus slightly concaved on posteroventral 1/3, ventrolaterally with a thin process bearing a sensillum at tip (
<figureCitation id="9CD4ACFDFFBFFFF9FB6CFAF7FAA30562" box="[1291,1371,1372,1397]" captionStart="FIGURE 11" captionStartId="18.[151,250,1564,1586]" captionTargetBox="[169,1421,212,1526]" captionTargetId="figure@18.[151,1436,195,1543]" captionTargetPageId="18" captionText="FIGURE 11. Leucophenga flavicosta Duda, 1926, male terminalia in lateral view. A. Epandrium, surstylus and cercus; B. Hypandrium, paramere, gonopods and aedeagus; C. Paramere; D. Aedeagus and laterally process (lp). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/438527/files/figure.png" pageId="17" pageNumber="218">Fig. 11</figureCitation>
D).
</paragraph>
<paragraph id="0450B078FFBFFFF9FEA0FA2AFB5B063E" blockId="17.[151,1437,1297,1939]" pageId="17" pageNumber="218">
However, the only male specimen shows distinctly difference from the latest and most relevant descriptions about
<taxonomicName id="C3EFCBFBFFBFFFF9FEBBFA0EFE9D05AB" box="[220,357,1444,1468]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="flavicosta">
<emphasis id="369B6C6AFFBFFFF9FEBBFA0EFF1305AB" box="[220,235,1445,1468]" italics="true" pageId="17" pageNumber="218">L</emphasis>
.
<emphasis id="369B6C6AFFBFFFF9FE9FFA0FFE9D05AB" box="[248,357,1444,1468]" italics="true" pageId="17" pageNumber="218">flavicosta</emphasis>
</taxonomicName>
(from
<collectingCountry id="7CF8F0E8FFBFFFF9FFD5FA0FFDFF05AA" box="[434,519,1444,1469]" name="Taiwan" pageId="17" pageNumber="218">Taiwan</collectingCountry>
,
<collectingCountry id="7CF8F0E8FFBFFFF9FC76FA0FFDAC05AA" box="[529,596,1444,1469]" name="China" pageId="17" pageNumber="218">China</collectingCountry>
in
<bibRefCitation id="607ECD89FFBFFFF9FC11FA0FFC9905AA" author="Lin" box="[630,865,1444,1469]" pageId="17" pageNumber="218" refString="Lin, F. J. &amp; Wheeler, M. R. (1972) The Drosophilidae of Taiwan, I. Genera Leucophenga and Paraleucophenga. The University of Texas Publication, 7213, 237 - 256." type="journal article" year="1972">Lin &amp; Wheeler 1972</bibRefCitation>
; from
<collectingCountry id="7CF8F0E8FFBFFFF9FDCFFA0FFC1E05AA" box="[936,998,1444,1469]" name="Palau" pageId="17" pageNumber="218">Palau</collectingCountry>
in
<bibRefCitation id="607ECD89FFBFFFF9FA6EFA0FFB6A05AA" author="Okada" box="[1033,1170,1444,1469]" pageId="17" pageNumber="218" refString="Okada, T. (1987) The Leucophenga subpollinosa group of Japan and Southeast Asia (Diptera, Drosophilidae). Kontyu, 55 (4), 676 - 683." type="journal article" year="1987">Okada 1987</bibRefCitation>
) in the color patterns of abdominal tergites (
<figureCitation id="9CD4ACFDFFBFFFF9FF1DFA63FE3805F6" box="[378,448,1480,1505]" captionStart="FIGURE 7" captionStartId="13.[151,250,1341,1363]" captionTargetBox="[172,1408,237,1297]" captionTargetId="figure@13.[149,1434,211,1310]" captionTargetPageId="13" captionText="FIGURE 7. Color patterns of abdominal tergites: A, B. Leucophenga argentina; C. L. flavicosta; D. L. formosa; E, F. L. nigroscutellata." httpUri="https://zenodo.org/record/438523/files/figure.png" pageId="17" pageNumber="218">Fig. 7</figureCitation>
C), while the aforementioned characters of wing (
<figureCitation id="9CD4ACFDFFBFFFF9FD9AFA63FBBB05F6" box="[1021,1091,1480,1505]" captionStart="FIGURE 2" captionStartId="8.[151,250,1530,1552]" captionTargetBox="[151,1436,193,1508]" captionTargetId="figure@8.[151,1436,193,1508]" captionTargetPageId="8" captionText="FIGURE 2. Wing of male (except for Leucophenga subpollinosa): A. L. argentina; B. L. flavicosta; C. L. formosa; D. L. nigroscutellata; E. L. subpollinosa; F. L. umbratula; G. L. aculeata sp. nov.; H. L. acuticauda sp. nov.; I. L. cultella sp. nov.; J. L. cyclophylla sp. nov.; K. L. flaviclypeata sp. nov.; L. L. fuscipedes sp. nov.; M. L. gracilenta sp. nov.; N. L. latipenis sp. nov.; O. L. magnicauda sp. nov.; P. L. rhombura sp. nov.; Q. L. rugatifolia sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/438518/files/figure.png" pageId="17" pageNumber="218">Fig. 2</figureCitation>
B) and halter (
<figureCitation id="9CD4ACFDFFBFFFF9FA8CFA63FAC805F6" box="[1259,1328,1480,1505]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[331,1245,194,1938]" captionTargetId="figure@9.[331,1245,193,1938]" captionTargetPageId="9" captionText="FIGURE 3. Dorsal and lateral view of thorax in male (except for Leucophenga subpollinosa): A, B. L. argentina; C, D. L. flavicosta; E, F. L. formosa; G, H. L. nigroscutellata; I, J. L. subpollinosa." httpUri="https://zenodo.org/record/438519/files/figure.png" pageId="17" pageNumber="218">Fig. 3</figureCitation>
C, D) are consistent with their descriptions (
<bibRefCitation id="607ECD89FFBFFFF9FC70FA47FCFC0612" author="Lin" box="[535,772,1516,1541]" pageId="17" pageNumber="218" refString="Lin, F. J. &amp; Wheeler, M. R. (1972) The Drosophilidae of Taiwan, I. Genera Leucophenga and Paraleucophenga. The University of Texas Publication, 7213, 237 - 256." type="journal article" year="1972">Lin &amp; Wheeler 1972</bibRefCitation>
;
<bibRefCitation id="607ECD89FFBFFFF9FD77FA47FC630612" author="Okada" box="[784,923,1516,1541]" pageId="17" pageNumber="218" refString="Okada, T. (1987) The Leucophenga subpollinosa group of Japan and Southeast Asia (Diptera, Drosophilidae). Kontyu, 55 (4), 676 - 683." type="journal article" year="1987">Okada 1987</bibRefCitation>
), which are unique in the
<taxonomicName id="C3EFCBFBFFBFFFF9FADAFA47FAB40613" box="[1213,1356,1516,1540]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="subpollinosa">
<emphasis id="369B6C6AFFBFFFF9FADAFA47FAB40613" box="[1213,1356,1516,1540]" italics="true" pageId="17" pageNumber="218">subpollinosa</emphasis>
</taxonomicName>
group. Therefore, the only male specimen was identified as
<taxonomicName id="C3EFCBFBFFBFFFF9FC81F9BAFC89063F" box="[742,881,1552,1576]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="flavicosta">
<emphasis id="369B6C6AFFBFFFF9FC81F9BAFD0D063F" box="[742,757,1553,1576]" italics="true" pageId="17" pageNumber="218">L</emphasis>
.
<emphasis id="369B6C6AFFBFFFF9FD63F9BBFC89063F" box="[772,881,1552,1576]" italics="true" pageId="17" pageNumber="218">flavicosta</emphasis>
</taxonomicName>
to avoid further confusion.
</paragraph>
</subSubSection>
<subSubSection id="4CF5E3F3FFBFFFFAFEA0F99FFEB4075D" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218" type="description">
<paragraph id="0450B078FFBFFFF9FEA0F99FFF2C0784" blockId="17.[151,1437,1297,1939]" pageId="17" pageNumber="218">
<emphasis id="369B6C6AFFBFFFF9FEA0F99FFEA0065B" bold="true" box="[199,344,1588,1613]" pageId="17" pageNumber="218">Description.</emphasis>
Ocellar triangle brown, with ca. 2 setae above ocellar setae. Frons brownish yellow, with three indistinct, brownish, longitudinal stripes medially and laterally. Pedicel and first flagellomere brownish yellow. Clypeus brownish yellow medially, brownish laterally. Palpus brownish yellow. Mesonotum brownish yellow, with three brown, longitudinal stripes medially and sublaterally (
<figureCitation id="9CD4ACFDFFBFFFF9FD08F90BFC4206AE" box="[879,954,1696,1721]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[331,1245,194,1938]" captionTargetId="figure@9.[331,1245,193,1938]" captionTargetPageId="9" captionText="FIGURE 3. Dorsal and lateral view of thorax in male (except for Leucophenga subpollinosa): A, B. L. argentina; C, D. L. flavicosta; E, F. L. formosa; G, H. L. nigroscutellata; I, J. L. subpollinosa." httpUri="https://zenodo.org/record/438519/files/figure.png" pageId="17" pageNumber="218">Fig. 3</figureCitation>
C). Postpronotal lobe yellow (
<figureCitation id="9CD4ACFDFFBFFFF9FB4AF90BFA8F06AE" box="[1325,1399,1696,1721]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[331,1245,194,1938]" captionTargetId="figure@9.[331,1245,193,1938]" captionTargetPageId="9" captionText="FIGURE 3. Dorsal and lateral view of thorax in male (except for Leucophenga subpollinosa): A, B. L. argentina; C, D. L. flavicosta; E, F. L. formosa; G, H. L. nigroscutellata; I, J. L. subpollinosa." httpUri="https://zenodo.org/record/438519/files/figure.png" pageId="17" pageNumber="218">Fig. 3</figureCitation>
D). Acrostichal setulae in ca. 1012 irregular rows (
<figureCitation id="9CD4ACFDFFBFFFF9FC9AF96FFCB206CA" box="[765,842,1732,1757]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[331,1245,194,1938]" captionTargetId="figure@9.[331,1245,193,1938]" captionTargetPageId="9" captionText="FIGURE 3. Dorsal and lateral view of thorax in male (except for Leucophenga subpollinosa): A, B. L. argentina; C, D. L. flavicosta; E, F. L. formosa; G, H. L. nigroscutellata; I, J. L. subpollinosa." httpUri="https://zenodo.org/record/438519/files/figure.png" pageId="17" pageNumber="218">Fig. 3</figureCitation>
C). Scutellum entirely brown (
<figureCitation id="9CD4ACFDFFBFFFF9FAABF96FFAE106CA" box="[1228,1305,1732,1757]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[331,1245,194,1938]" captionTargetId="figure@9.[331,1245,193,1938]" captionTargetPageId="9" captionText="FIGURE 3. Dorsal and lateral view of thorax in male (except for Leucophenga subpollinosa): A, B. L. argentina; C, D. L. flavicosta; E, F. L. formosa; G, H. L. nigroscutellata; I, J. L. subpollinosa." httpUri="https://zenodo.org/record/438519/files/figure.png" pageId="17" pageNumber="218">Fig. 3</figureCitation>
C). Pleura, katepisternum and meron mostly brown above (
<figureCitation id="9CD4ACFDFFBFFFF9FCCCF943FD150716" box="[683,749,1768,1793]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[331,1245,194,1938]" captionTargetId="figure@9.[331,1245,193,1938]" captionTargetPageId="9" captionText="FIGURE 3. Dorsal and lateral view of thorax in male (except for Leucophenga subpollinosa): A, B. L. argentina; C, D. L. flavicosta; E, F. L. formosa; G, H. L. nigroscutellata; I, J. L. subpollinosa." httpUri="https://zenodo.org/record/438519/files/figure.png" pageId="17" pageNumber="218">Fig. 3</figureCitation>
D). Wing (
<figureCitation id="9CD4ACFDFFBFFFF9FD03F943FC500716" box="[868,936,1768,1793]" captionStart="FIGURE 2" captionStartId="8.[151,250,1530,1552]" captionTargetBox="[151,1436,193,1508]" captionTargetId="figure@8.[151,1436,193,1508]" captionTargetPageId="8" captionText="FIGURE 2. Wing of male (except for Leucophenga subpollinosa): A. L. argentina; B. L. flavicosta; C. L. formosa; D. L. nigroscutellata; E. L. subpollinosa; F. L. umbratula; G. L. aculeata sp. nov.; H. L. acuticauda sp. nov.; I. L. cultella sp. nov.; J. L. cyclophylla sp. nov.; K. L. flaviclypeata sp. nov.; L. L. fuscipedes sp. nov.; M. L. gracilenta sp. nov.; N. L. latipenis sp. nov.; O. L. magnicauda sp. nov.; P. L. rhombura sp. nov.; Q. L. rugatifolia sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/438518/files/figure.png" pageId="17" pageNumber="218">Fig. 2</figureCitation>
B): Costal vein between R2+3 and R4+5 distally with ca. 5 peg-like spinules on ventral surface. Abdominal tergites dark brown (
<figureCitation id="9CD4ACFDFFBFFFF9FA51F8A4FB840730" box="[1078,1148,1807,1832]" captionStart="FIGURE 7" captionStartId="13.[151,250,1341,1363]" captionTargetBox="[172,1408,237,1297]" captionTargetId="figure@13.[149,1434,211,1310]" captionTargetPageId="13" captionText="FIGURE 7. Color patterns of abdominal tergites: A, B. Leucophenga argentina; C. L. flavicosta; D. L. formosa; E, F. L. nigroscutellata." httpUri="https://zenodo.org/record/438523/files/figure.png" pageId="17" pageNumber="218">Fig. 7</figureCitation>
C), with a pair of yellow patches submedially between 2nd to 3rd tergites. Male terminalia: Epandrium with ca. 1 seta near posterior margin per side (
<figureCitation id="9CD4ACFDFFBFFFF9FF67F8FCFEA90778" box="[256,337,1879,1904]" captionStart="FIGURE 11" captionStartId="18.[151,250,1564,1586]" captionTargetBox="[169,1421,212,1526]" captionTargetId="figure@18.[151,1436,195,1543]" captionTargetPageId="18" captionText="FIGURE 11. Leucophenga flavicosta Duda, 1926, male terminalia in lateral view. A. Epandrium, surstylus and cercus; B. Hypandrium, paramere, gonopods and aedeagus; C. Paramere; D. Aedeagus and laterally process (lp). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/438527/files/figure.png" pageId="17" pageNumber="218">Fig. 11</figureCitation>
A). Surstylus rounded apically (
<figureCitation id="9CD4ACFDFFBFFFF9FCDBF8FCFCF40778" box="[700,780,1879,1904]" captionStart="FIGURE 11" captionStartId="18.[151,250,1564,1586]" captionTargetBox="[169,1421,212,1526]" captionTargetId="figure@18.[151,1436,195,1543]" captionTargetPageId="18" captionText="FIGURE 11. Leucophenga flavicosta Duda, 1926, male terminalia in lateral view. A. Epandrium, surstylus and cercus; B. Hypandrium, paramere, gonopods and aedeagus; C. Paramere; D. Aedeagus and laterally process (lp). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/438527/files/figure.png" pageId="17" pageNumber="218">Fig. 11</figureCitation>
A). Paramere with ca. 7 sensillum submediodorsally (
<figureCitation id="9CD4ACFDFFBFFFF9FB08F8FCFF4A0784" captionStart="FIGURE 11" captionStartId="18.[151,250,1564,1586]" captionTargetBox="[169,1421,212,1526]" captionTargetId="figure@18.[151,1436,195,1543]" captionTargetPageId="18" captionText="FIGURE 11. Leucophenga flavicosta Duda, 1926, male terminalia in lateral view. A. Epandrium, surstylus and cercus; B. Hypandrium, paramere, gonopods and aedeagus; C. Paramere; D. Aedeagus and laterally process (lp). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/438527/files/figure.png" pageId="17" pageNumber="218">Fig. 11</figureCitation>
C).
</paragraph>
<caption id="5090E0F0FFBCFFFAFEF0F9B7FA860646" httpUri="https://zenodo.org/record/438527/files/figure.png" pageId="18" pageNumber="219" targetBox="[169,1421,212,1526]" targetPageId="18">
<paragraph id="0450B078FFBCFFFAFEF0F9B7FA860646" blockId="18.[151,1436,1564,1617]" pageId="18" pageNumber="219">
<emphasis id="369B6C6AFFBCFFFAFEF0F9B7FED90625" bold="true" box="[151,289,1564,1586]" pageId="18" pageNumber="219">FIGURE 11.</emphasis>
<taxonomicName id="C3EFCBFBFFBCFFFAFF4DF9B7FD5B0625" authority="Duda, 1926" authorityName="Duda" authorityYear="1926" box="[298,675,1564,1586]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="flavicosta">
<emphasis id="369B6C6AFFBCFFFAFF4DF9B7FDE70625" box="[298,543,1564,1586]" italics="true" pageId="18" pageNumber="219">Leucophenga flavicosta</emphasis>
Duda, 1926
</taxonomicName>
, male terminalia in lateral view. A. Epandrium, surstylus and cercus; B. Hypandrium, paramere, gonopods and aedeagus; C. Paramere; D. Aedeagus and laterally process (lp). Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="0450B078FFBCFFFAFEA0F9D6FE2A0715" blockId="18.[151,1436,1661,1938]" pageId="18" pageNumber="219">
<emphasis id="369B6C6AFFBCFFFAFEA0F9D6FE850681" bold="true" box="[199,381,1661,1686]" pageId="18" pageNumber="219">Measurements.</emphasis>
BL =
<quantity id="C3171D9DFFBCFFFAFFA1F9D6FDD40681" box="[454,556,1661,1686]" metricMagnitude="-3" metricUnit="m" metricValue="2.53" pageId="18" pageNumber="219" unit="mm" value="2.53">2.53 mm</quantity>
, THL =
<quantity id="C3171D9DFFBCFFFAFCEBF9D5FD0A0681" box="[652,754,1662,1686]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="18" pageNumber="219" unit="mm" value="1.07">1.07 mm</quantity>
, WL =
<quantity id="C3171D9DFFBCFFFAFD2FF9D5FC560681" box="[840,942,1662,1686]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="18" pageNumber="219" unit="mm" value="2.1">2.10 mm</quantity>
, WW =
<quantity id="C3171D9DFFBCFFFAFA6AF9D5FB8B0681" box="[1037,1139,1662,1686]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="18" pageNumber="219" unit="mm" value="0.97">0.97 mm</quantity>
, arb = 6/3, avd = 0.83, adf = 1.71, flw = 1.86, FW/HW = 0.40, ch/o = 0.02, prorb = 0.59, rcorb = 0.59, vb = 0.33, dcl = 0.46, presctl = 0.54, sctl = 1.42, sterno = 0.87, orbito = 1.80, dcp = 0.32, sctlp = 1.09, C = 1.89, 4c = 1.31,
<date id="705196B8FFBCFFFAFA15F96DFB7606C9" box="[1138,1166,1734,1758]" pageId="18" pageNumber="219">4v</date>
= 2.14,
<date id="705196B8FFBCFFFAFA88F96EFAF306C9" box="[1263,1291,1733,1758]" pageId="18" pageNumber="219">5x</date>
= 1.54, ac = 2.38, M = 0.69, C3F = 0.89.
</paragraph>
<paragraph id="0450B078FFBCFFFAFEA0F8A6FEB4075D" blockId="18.[151,1436,1661,1938]" pageId="18" pageNumber="219">
<materialsCitation id="B487BA25FFBCFFFAFEA0F8A6FEB4075D" ID-GBIF-Occurrence="1450621017" collectingDate="2003-08-16" collectionCode="SCAU" collectorName="JJ Gao." country="China" county="Specimens" elevation="470" location="Ziyang" municipality="Haoping" pageId="18" pageNumber="219" specimenCount="1" specimenCount-male="1" stateProvince="Shaanxi">
<emphasis id="369B6C6AFFBCFFFAFEA0F8A6FE3F0731" bold="true" box="[199,455,1805,1830]" pageId="18" pageNumber="219">
<collectingCounty id="ED31C8F4FFBCFFFAFEA0F8A6FEBD0731" box="[199,325,1805,1830]" pageId="18" pageNumber="219">Specimens</collectingCounty>
examined.
</emphasis>
<collectingCountry id="7CF8F0E8FFBCFFFAFFA8F8A5FDD40731" box="[463,556,1806,1830]" name="China" pageId="18" pageNumber="219">CHINA</collectingCountry>
:
<specimenCount id="12E97BF1FFBCFFFAFC5FF8A5FDA30731" box="[568,603,1806,1830]" pageId="18" pageNumber="219" type="male">1♂</specimenCount>
(
<collectionCode id="62FE28BDFFBCFFFAFC0CF8A5FD420731" box="[619,698,1806,1830]" pageId="18" pageNumber="219">SCAU</collectionCode>
, No. 128964),
<collectingMunicipality id="E4342A02FFBCFFFAFD03F8A6FC300731" box="[868,968,1805,1830]" pageId="18" pageNumber="219">Haoping</collectingMunicipality>
,
<location id="0130E6A3FFBCFFFAFDB3F8A6FBDE0731" LSID="urn:lsid:plazi:treatment:8C46016EFFBFFFFAFEF0FBEEFEB60785:0130E6A3FFBCFFFAFDB3F8A6FBDE0731" box="[980,1062,1805,1830]" country="China" county="Specimens" municipality="Haoping" name="Ziyang" pageId="18" pageNumber="219" stateProvince="Shaanxi">Ziyang</location>
,
<collectingRegion id="C62B7E9AFFBCFFFAFA54F8A6FB680731" box="[1075,1168,1805,1830]" country="China" name="Shaanxi" pageId="18" pageNumber="219">Shaanxi</collectingRegion>
,
<quantity id="C3171D9DFFBCFFFAFAFBF8A5FB270731" box="[1180,1247,1806,1830]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="18" pageNumber="219" unit="m" value="470.0">
<elevation id="8FC2574BFFBCFFFAFAFBF8A5FB270731" box="[1180,1247,1806,1830]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="18" pageNumber="219" unit="m" value="470.0">470m</elevation>
</quantity>
,
<date id="705196B8FFBCFFFAFA8CF8A6FA8D0731" box="[1259,1397,1805,1830]" pageId="18" pageNumber="219" value="2003-08-16">
<collectingDate id="60156F50FFBCFFFAFA8CF8A6FA8D0731" box="[1259,1397,1805,1830]" pageId="18" pageNumber="219" value="2003-08-16">16.viii.2003</collectingDate>
</date>
, ex tussock,
<collectorName id="A91AD5AEFFBCFFFAFE9EF899FEB4075D" box="[249,332,1842,1866]" pageId="18" pageNumber="219">JJ Gao.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4CF5E3F3FFBCFFFAFEA0F8FEFEB60785" pageId="18" pageNumber="219" type="distribution">
<paragraph id="0450B078FFBCFFFAFEA0F8FEFEB60785" blockId="18.[151,1436,1661,1938]" pageId="18" pageNumber="219">
<emphasis id="369B6C6AFFBCFFFAFEA0F8FEFE980779" bold="true" box="[199,352,1877,1902]" pageId="18" pageNumber="219">Distribution.</emphasis>
<collectingCountry id="7CF8F0E8FFBCFFFAFF0EF8FEFE540779" box="[361,428,1877,1902]" name="China" pageId="18" pageNumber="219">China</collectingCountry>
(
<collectingRegion id="C62B7E9AFFBCFFFAFFDBF8FEFDE20779" box="[444,538,1877,1902]" country="China" name="Shaanxi" pageId="18" pageNumber="219">Shaanxi</collectingRegion>
,
<collectingRegion id="C62B7E9AFFBCFFFAFC4FF8FEFD7E0779" box="[552,646,1877,1902]" country="China" name="Taiwan)" pageId="18" pageNumber="219">
<collectingCountry id="7CF8F0E8FFBCFFFAFC4FF8FEFD860779" box="[552,638,1877,1902]" name="Taiwan" pageId="18" pageNumber="219">Taiwan</collectingCountry>
)
</collectingRegion>
,
<collectingCountry id="7CF8F0E8FFBCFFFAFCF4F8FEFD370779" box="[659,719,1877,1902]" name="India" pageId="18" pageNumber="219">India</collectingCountry>
,
<collectingCountry id="7CF8F0E8FFBCFFFAFCBBF8FEFCE50779" box="[732,797,1877,1902]" name="Palau" pageId="18" pageNumber="219">Palau</collectingCountry>
,
<collectingCountry id="7CF8F0E8FFBCFFFAFD4DF8FEFC530779" box="[810,939,1877,1902]" name="Micronesia" pageId="18" pageNumber="219">Micronesia</collectingCountry>
, Borneo, New
<collectingCountry id="7CF8F0E8FFBCFFFAFA30F8FEFB500779" box="[1111,1192,1877,1902]" name="Guinea-Bissau" pageId="18" pageNumber="219">Guinea</collectingCountry>
(Madang),
<collectingCountry id="7CF8F0E8FFBCFFFAFB48F8FEFA640779" box="[1327,1436,1877,1902]" name="Indonesia" pageId="18" pageNumber="219">Indonesia</collectingCountry>
(
<collectingRegion id="C62B7E9AFFBCFFFAFEF8F8D1FF2D0785" box="[159,213,1914,1938]" country="Indonesia" name="Jawa Timur" pageId="18" pageNumber="219">Java</collectingRegion>
,
<collectingRegion id="C62B7E9AFFBCFFFAFE87F8D1FEBA0785" box="[224,322,1914,1938]" country="Indonesia" name="Sumatera Utara" pageId="18" pageNumber="219">Sumatra</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>