280 lines
23 KiB
XML
280 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.960.55007" ID-GBIF-Dataset="d33b7f20-b953-459e-bf96-129919db2fc1" ID-PMC="PMC7445195" ID-Pensoft-Pub="1313-2970-960-63" ID-Pensoft-UUID="52BF27E864625C32B58BD8C17C1AA574" ID-PubMed="32884398" ID-ZooBank="53E72CF9515F47E49350F123BC2D8442" ModsDocID="1313-2970-960-63" checkinTime="1597680765858" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob & Balke, Michael" docDate="2020" docId="8B770EF32480545BB610ED488E6E7DEC" docLanguage="en" docName="ZooKeys 960: 63-78" docOrigin="ZooKeys 960" docSource="http://dx.doi.org/10.3897/zookeys.960.55007" docTitle="Exocelina tsinga Shaverdo & Surbakti & Sumoked & Balke 2020, sp. nov." docType="treatment" docUuid="F1184304-DE9D-45AA-936E-17300B34EF25" docUuidSource="ZooBank" docVersion="4" id="52BF27E864625C32B58BD8C17C1AA574" lastPageNumber="63" masterDocId="52BF27E864625C32B58BD8C17C1AA574" masterDocTitle="Two new species of the Exocelina ekari group from New Guinea with strongly modified male antennae (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="78" masterPageNumber="63" pageNumber="63" updateTime="1668169273712" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of the Exocelina ekari group from New Guinea with strongly modified male antennae (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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<mods:affiliation>Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier type="email">shaverdo@mail.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Surbakti, Suriani</mods:namePart>
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<mods:affiliation>Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sumoked, Bob</mods:namePart>
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<mods:affiliation>Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany and GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>960</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>63</mods:start>
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<mods:end>78</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.960.55007</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.960.55007</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-960-63</mods:identifier>
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<mods:identifier type="ZooBank">53E72CF9515F47E49350F123BC2D8442</mods:identifier>
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<mods:identifier type="Pensoft-UUID">52BF27E864625C32B58BD8C17C1AA574</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="166240664" LSID="urn:lsid:zoobank.org:act:F1184304-DE9D-45AA-936E-17300B34EF25" httpUri="http://treatment.plazi.org/id/8B770EF32480545BB610ED488E6E7DEC" lastPageNumber="63" pageId="0" pageNumber="63">
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<subSubSection pageId="0" pageNumber="63" type="nomenclature">
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<paragraph pageId="0" pageNumber="63">
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<taxonomicName LSID="8B770EF3-2480-545B-B610-ED488E6E7DEC" authority="Shaverdo & Surbakti & Sumoked & Balke, 2020" authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina tsinga" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="tsinga" status="sp. nov.">Exocelina tsinga</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="63">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus and colouration of Exocelina tsinga sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure5" httpUri="https://binary.pensoft.net/fig/442335" pageId="0" pageNumber="63">Figures 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Exocelina tsinga sp. nov., paratype A median lobe in ventral view B median lobe in lateral view." figureDoi="10.3897/zookeys.960.55007.figure6" httpUri="https://binary.pensoft.net/fig/442336" pageId="0" pageNumber="63">, 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Exocelina tsinga sp. nov., paratype A right paramere in external view B right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.960.55007.figure7" httpUri="https://binary.pensoft.net/fig/442337" pageId="0" pageNumber="63">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male antennae A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., paratype." figureDoi="10.3897/zookeys.960.55007.figure8" httpUri="https://binary.pensoft.net/fig/442338" pageId="0" pageNumber="63">, 8B</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Abdominal ventrite 6 A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure9" httpUri="https://binary.pensoft.net/fig/442339" pageId="0" pageNumber="63">, 9B</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Tsingogong River." figureDoi="10.3897/zookeys.960.55007.figure10" httpUri="https://binary.pensoft.net/fig/442340" pageId="0" pageNumber="63">, 10</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Small water holes on bedrock besides Tsingogong River." figureDoi="10.3897/zookeys.960.55007.figure11" httpUri="https://binary.pensoft.net/fig/442341" pageId="0" pageNumber="63">, 11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="type locality">
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<paragraph pageId="0" pageNumber="63">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="63">
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Indonesia: Papua Province, Mimika Regency, Tsinga Village, Tsingogong River,
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<geoCoordinate degrees="04" direction="south" minutes="11.320" orientation="latitude" precision="1" value="-4.188667">04°11.320'S</geoCoordinate>
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,
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<geoCoordinate degrees="137" direction="east" minutes="16.364" orientation="longitude" precision="1" value="137.27274">137°16.364'E</geoCoordinate>
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, 1,306 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="type material">
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<paragraph pageId="0" pageNumber="63">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
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: male "Indonesia: Kabupaten [Regency] Mimika, Desa [Village] Tsinga, Sungai [River] Tsingogong", "1306 m, 25-30.v.2017,
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<geoCoordinate degrees="04" direction="south" minutes="11.320" orientation="latitude" precision="1" value="-4.188667">04°11.320'S</geoCoordinate>
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,
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<geoCoordinate degrees="137" direction="east" minutes="16.364" orientation="longitude" precision="1" value="137.27274">137°16.364'E</geoCoordinate>
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, B. Sumoked" (MZB).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Paratypes</emphasis>
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: 33 males, 38 females with the same label as the holotype (MZB, KSP). 6 males, 7 females "Indonesia: Kabupaten Mimika, Desa Tsinga, 1381 m, 25-30.v.2017", "
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<geoCoordinate degrees="04" direction="south" minutes="11.379" orientation="latitude" precision="1" value="-4.18965">04°11.379'S</geoCoordinate>
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,
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<geoCoordinate degrees="137" direction="east" minutes="13.456" orientation="longitude" precision="1" value="137.22427">137°13.456'E</geoCoordinate>
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, B. Sumoked" (MZB, KSP).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="description">
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<paragraph pageId="0" pageNumber="63">Description.</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Body size and form</emphasis>
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: Beetle medium-sized: TL-H 4.05-4.8 mm, TL 4.5-5.3 mm, MW 2.15-2.5 mm (holotype: TL-H 4.5 mm, TL 5 mm, MW 2.4 mm), with oblong-oval to elongate habitus (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus and colouration of Exocelina tsinga sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure5" httpUri="https://binary.pensoft.net/fig/442335" pageId="0" pageNumber="63">5</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.960.55007.figure5" httpUri="https://binary.pensoft.net/fig/442335" pageId="0" pageNumber="63" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" pageId="0" pageNumber="63">Figure 5.</emphasis>
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Habitus and colouration of
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<taxonomicName authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina tsinga" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="tsinga">
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<emphasis italics="true" pageId="0" pageNumber="63">Exocelina tsinga</emphasis>
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</taxonomicName>
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sp. nov., holotype.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Colouration</emphasis>
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: As in
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<taxonomicName lsidName="E. athesphatos" pageId="0" pageNumber="63" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">E. athesphatos</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Surface sculpture</emphasis>
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: As in
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<taxonomicName lsidName="E. athesphatos" pageId="0" pageNumber="63" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">E. athesphatos</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Structures</emphasis>
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: Pronotum with distinct, relatively narrow anteriorly lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 broadly rounded or slightly truncate.
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</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Male</emphasis>
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: Antenna strongly modified (Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus and colouration of Exocelina tsinga sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure5" httpUri="https://binary.pensoft.net/fig/442335" pageId="0" pageNumber="63">5</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male antennae A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., paratype." figureDoi="10.3897/zookeys.960.55007.figure8" httpUri="https://binary.pensoft.net/fig/442338" pageId="0" pageNumber="63">8B</figureCitation>
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): antennomere 2 strongly reduced, antennomeres 3 and 4 strongly enlarged (antennomere 3 the largest), antennomeres 5 and 6 distinctly enlarged, antennomeres 7-9 stout. Pro- and mesotarsomeres 1-3 dilated. Protarsomere 4 slightly dilated, with anterolateral angle shortly expanded and with large, thick, slightly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 14 and posterior row of 6 short, thick, pointed setae (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Exocelina tsinga sp. nov., paratype A right paramere in external view B right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.960.55007.figure7" httpUri="https://binary.pensoft.net/fig/442337" pageId="0" pageNumber="63">7B</figureCitation>
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). Median lobe relatively long and slender, with slightly discontinuous outline in subapical part (mainly visible in ventral view); in lateral view, apex thin, more or less pointed and curved downwards; in ventral view, apex broad, almost truncate (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Exocelina tsinga sp. nov., paratype A median lobe in ventral view B median lobe in lateral view." figureDoi="10.3897/zookeys.960.55007.figure6" httpUri="https://binary.pensoft.net/fig/442336" pageId="0" pageNumber="63">6</figureCitation>
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). Paramere with very deep dorsal notch, separating subdistal part; subdistal part is large, broad, curved downwards and pressed closely to paramere, with fringe of seven very broad, flattened setae; proximal setae numerous, dense, thin, much more inconspicuous than subdistal (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Exocelina tsinga sp. nov., paratype A right paramere in external view B right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.960.55007.figure7" httpUri="https://binary.pensoft.net/fig/442337" pageId="0" pageNumber="63">7A</figureCitation>
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). Abdominal ventrite 6 slightly depressed medially, with 10-12 lateral striae on each side (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Abdominal ventrite 6 A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure9" httpUri="https://binary.pensoft.net/fig/442339" pageId="0" pageNumber="63">9B</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.960.55007.figure6" httpUri="https://binary.pensoft.net/fig/442336" pageId="0" pageNumber="63" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" pageId="0" pageNumber="63">Figure 6.</emphasis>
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<taxonomicName authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina tsinga" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="tsinga">
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<emphasis italics="true" pageId="0" pageNumber="63">Exocelina tsinga</emphasis>
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</taxonomicName>
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sp. nov., paratype
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<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
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median lobe in ventral view
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<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
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median lobe in lateral view.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.960.55007.figure7" httpUri="https://binary.pensoft.net/fig/442337" pageId="0" pageNumber="63" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" pageId="0" pageNumber="63">Figure 7.</emphasis>
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<taxonomicName authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina tsinga" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="tsinga">
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<emphasis italics="true" pageId="0" pageNumber="63">Exocelina tsinga</emphasis>
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</taxonomicName>
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sp. nov., paratype
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<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
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right paramere in external view
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<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
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right male protarsomeres 4-5 in ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Female</emphasis>
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: Antennae and pro- and mesotarsi not modified. Abdominal ventrite 6 without depression and lateral striae.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="affinities">
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<paragraph pageId="0" pageNumber="63">Affinities.</paragraph>
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<paragraph pageId="0" pageNumber="63">
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About the placement within the
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<taxonomicName lsidName="E. ekari" pageId="0" pageNumber="63" rank="species" species="ekari">
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<emphasis italics="true" pageId="0" pageNumber="63">E. ekari</emphasis>
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</taxonomicName>
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group, we consider the species in the same way as
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<taxonomicName lsidName="E. athesphatos" pageId="0" pageNumber="63" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">E. athesphatos</emphasis>
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</taxonomicName>
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sp. nov. Both species are very similar in external morphology (colouration, body form and sculpture, shape of male antennae) and, therefore, difficult to distinguish without detailed study (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Exocelina athesphatos sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure1" httpUri="https://binary.pensoft.net/fig/442331" pageId="0" pageNumber="63">1</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus and colouration of Exocelina tsinga sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure5" httpUri="https://binary.pensoft.net/fig/442335" pageId="0" pageNumber="63">5</figureCitation>
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). However, males antennomeres of
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<taxonomicName lsidName="E. tsinga" pageId="0" pageNumber="63" rank="species" species="tsinga">
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<emphasis italics="true" pageId="0" pageNumber="63">E. tsinga</emphasis>
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</taxonomicName>
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sp. nov. are slightly smaller than in
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<taxonomicName lsidName="E. athesphatos" pageId="0" pageNumber="63" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">E. athesphatos</emphasis>
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</taxonomicName>
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sp. nov. and have slightly different form (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male antennae A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., paratype." figureDoi="10.3897/zookeys.960.55007.figure8" httpUri="https://binary.pensoft.net/fig/442338" pageId="0" pageNumber="63">8</figureCitation>
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). The species can be easily separated by the shape of abdominal ventrite 6 (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Abdominal ventrite 6 A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure9" httpUri="https://binary.pensoft.net/fig/442339" pageId="0" pageNumber="63">9</figureCitation>
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), median lobe and paramere.
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</paragraph>
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<caption doi="10.3897/zookeys.960.55007.figure8" httpUri="https://binary.pensoft.net/fig/442338" pageId="0" pageNumber="63" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" pageId="0" pageNumber="63">Figure 8.</emphasis>
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Male antennae
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<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
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<taxonomicName authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina athesphatos" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">Exocelina athesphatos</emphasis>
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</taxonomicName>
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sp. nov., paratype
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<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
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<taxonomicName lsidName="E. tsinga" pageId="0" pageNumber="63" rank="species" species="tsinga">
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<emphasis italics="true" pageId="0" pageNumber="63">E. tsinga</emphasis>
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</taxonomicName>
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sp. nov., paratype.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.960.55007.figure9" httpUri="https://binary.pensoft.net/fig/442339" pageId="0" pageNumber="63" start="Figure 9" startId="F9">
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<paragraph pageId="0" pageNumber="63">
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|
<emphasis bold="true" pageId="0" pageNumber="63">Figure 9.</emphasis>
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|
Abdominal ventrite 6
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<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
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|
<taxonomicName authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina athesphatos" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">Exocelina athesphatos</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., paratype
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<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
|
|
<taxonomicName lsidName="E. tsinga" pageId="0" pageNumber="63" rank="species" species="tsinga">
|
|
<emphasis italics="true" pageId="0" pageNumber="63">E. tsinga</emphasis>
|
|
</taxonomicName>
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|
sp. nov., holotype.
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</paragraph>
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</caption>
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|
</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="distribution">
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<paragraph pageId="0" pageNumber="63">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="63">Indonesia: Papua Province, Mimika Regency. The species is known only from the type locality.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="habitat">
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<paragraph pageId="0" pageNumber="63">Habitat.</paragraph>
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|
<paragraph pageId="0" pageNumber="63">
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|
The specimens were collected from small puddles on bedrock (Fig.
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|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Small water holes on bedrock besides Tsingogong River." figureDoi="10.3897/zookeys.960.55007.figure11" httpUri="https://binary.pensoft.net/fig/442341" pageId="0" pageNumber="63">11</figureCitation>
|
|
), besides fast flowing mountains streams (such as Tsingogong River in Fig.
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|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Tsingogong River." figureDoi="10.3897/zookeys.960.55007.figure10" httpUri="https://binary.pensoft.net/fig/442340" pageId="0" pageNumber="63">10</figureCitation>
|
|
).
|
|
</paragraph>
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|
<caption doi="10.3897/zookeys.960.55007.figure10" httpUri="https://binary.pensoft.net/fig/442340" pageId="0" pageNumber="63" start="Figure 10" startId="F10">
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|
<paragraph pageId="0" pageNumber="63">
|
|
<emphasis bold="true" pageId="0" pageNumber="63">Figure 10.</emphasis>
|
|
Tsingogong River.
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|
</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.960.55007.figure11" httpUri="https://binary.pensoft.net/fig/442341" pageId="0" pageNumber="63" start="Figure 11" startId="F11">
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|
<paragraph pageId="0" pageNumber="63">
|
|
<emphasis bold="true" pageId="0" pageNumber="63">Figure 11.</emphasis>
|
|
Small water holes on bedrock besides Tsingogong River.
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|
</paragraph>
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</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="63" type="etymology">
|
|
<paragraph pageId="0" pageNumber="63">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="63">The species is named after Tsinga Village. The name is a noun in the nominative singular standing in apposition.</paragraph>
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</subSubSection>
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</treatment>
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</document> |