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<document id="F2C6EA95C782029E3D41015FACEC5C07" ID-DOI="10.3853/j.2201-4349.74.2022.1821" ID-ISSN="2201-4349" ID-Zenodo-Dep="13228969" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722893937960" checkinUser="felipe" docAuthor="Reid, Chris A. M. &amp; Runagall-Mcnaull, Aidan" docDate="2022" docId="424B87F8EC5C5065FCA0B4AEFBEEF824" docLanguage="en" docName="RecAustMus.74.5.131–150.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 74 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.74.2022.1821" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.4:RecAustMus.2019-.journal_article.0cover" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article.0cover" docStyleVersion="4" docTitle="Onthophagus longipes Paulian 1937" docType="treatment" docVersion="2" lastPageNumber="139" masterDocId="BE72FF80EC5A506DFFD1B023FFBCFF9B" masterDocTitle="A Revision of the Onthophagus pexatus Species-group, with Description of a New Brachypterous Species (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="150" masterPageNumber="131" pageNumber="137" updateTime="1722901422291" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="E905CB9AE55BFB041C9A0E31DA13BB6D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="BA536921466EB2D4D3AA4C3FD3008AB5">A Revision of the Onthophagus pexatus Species-group, with Description of a New Brachypterous Species (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="C63492994BFC72382C20E915273F5E00">Reid, Chris A. M.</mods:namePart>
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<mods:namePart id="FA5287E992D3DA407B0F7B0E623BAF97">Runagall-Mcnaull, Aidan</mods:namePart>
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<mods:title id="DA6D94E813CAF6EA30069921E0DD8588">Records of the Australian Museum</mods:title>
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<mods:date id="5D2F30F62DCEB0D096BA39046B8B16D3">2022</mods:date>
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<mods:title id="EF7F0EB8B0B0194D4AE680181216A6EB">Rec. Aust. Mus.</mods:title>
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<mods:number id="0F92D6EB48E5F0A36FDF77E48F58A1FD">2022-11-30</mods:number>
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<mods:number id="A10E87B28AE6468C385C5B1755A5C083">74</mods:number>
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<mods:number id="BCBA778F5B7F45F6EF24907C0F3ABF0A">5</mods:number>
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<mods:url id="23972901198872D558027351132F30B2">http://dx.doi.org/10.3853/j.2201-4349.74.2022.1821</mods:url>
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<mods:classification id="DF67550911869756EF49CFD5BE46BA40">journal article</mods:classification>
<mods:identifier id="186A9888F54101C7791043B9CA1B3BF8" type="DOI">10.3853/j.2201-4349.74.2022.1821</mods:identifier>
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<mods:identifier id="B16773DC8B36E51FC38B27BBBB2800BB" type="Zenodo-Dep">13228969</mods:identifier>
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<treatment id="424B87F8EC5C5065FCA0B4AEFBEEF824" ID-DOI="http://doi.org/10.5281/zenodo.13228989" ID-Zenodo-Dep="13228989" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824" httpUri="http://treatment.plazi.org/id/424B87F8EC5C5065FCA0B4AEFBEEF824" lastPageId="8" lastPageNumber="139" pageId="6" pageNumber="137">
<subSubSection id="82F86565EC5C506BFCA0B4AEFA2CFB37" box="[881,1424,1165,1196]" pageId="6" pageNumber="137" type="nomenclature">
<paragraph id="CA5D36EEEC5C506BFCA0B4AEFA2CFB37" blockId="6.[881,1424,1165,1196]" box="[881,1424,1165,1196]" pageId="6" pageNumber="137">
<heading id="91158182EC5C506BFCA0B4AEFA2CFB37" bold="true" box="[881,1424,1165,1196]" centered="true" fontSize="12" level="2" pageId="6" pageNumber="137" reason="2">
<taxonomicName id="0DE24D6DEC5C506BFCA0B4AEFA2CFB37" ID-CoL="7XNFF" authority="Paulian, 1937" authorityName="Paulian" authorityYear="1937" box="[881,1424,1165,1196]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="F896EAFCEC5C506BFCA0B4AEFA2CFB37" bold="true" box="[881,1424,1165,1196]" pageId="6" pageNumber="137">
<emphasis id="F896EAFCEC5C506BFCA0B4AEFB07FB37" bold="true" box="[881,1211,1165,1196]" italics="true" pageId="6" pageNumber="137">Onthophagus longipes</emphasis>
Paulian, 1937
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="82F86565EC5C506BFC14B4E5FA80FB7B" box="[965,1340,1222,1248]" pageId="6" pageNumber="137" type="description">
<paragraph id="CA5D36EEEC5C506BFC14B4E5FA80FB7B" blockId="6.[965,1340,1222,1248]" box="[965,1340,1222,1248]" pageId="6" pageNumber="137">
<figureCitation id="52D92A6BEC5C506BFC14B4E5FB8CFB7B" box="[965,1072,1222,1248]" captionStart="Figures 1–4" captionStartId="4.[216,293,1867,1889]" captionTargetBox="[224,1375,175,1822]" captionTargetId="figure-10@4.[167,1467,151,1828]" captionTargetPageId="4" captionText="Figures 1–4. (1, 2) Onthophagus bulga sp. nov., male, dorsal and lateral; (3, 4) O. longipes Paulian, male, dorsal and lateral." figureDoi="http://doi.org/10.5281/zenodo.13228971" httpUri="https://zenodo.org/record/13228971/files/figure.png" pageId="6" pageNumber="137">Figs 3–4</figureCitation>
,
<figureCitation id="52D92A6BEC5C506BFBECB4E5FB36FB7B" box="[1085,1162,1222,1248]" captionStart="Figures 17–20" captionStartId="9.[121,198,1260,1282]" captionTargetBox="[121,1421,144,1244]" captionTargetId="figure-10@9.[121,1421,144,1244]" captionTargetPageId="9" captionText="Figures 17–20. Onthophagus species, head, male (left), and female (right), fresh specimens. (17, 18) O. longipes Paulian; (19, 20) O. nammuldi Matthews." figureDoi="http://doi.org/10.5281/zenodo.13228983" httpUri="https://zenodo.org/record/13228983/files/figure.png" pageId="6" pageNumber="137">17–18</figureCitation>
,
<figureCitation id="52D92A6BEC5C506BFB46B4E5FB0BFB7B" box="[1175,1207,1222,1248]" captionStart="Figures 25–29" captionStartId="11.[121,198,1565,1587]" captionTargetBox="[121,748,151,1551]" captionTargetId="figure-106@11.[121,748,151,1553]" captionTargetPageId="11" captionText="Figures 25–29. Onthophagus species, venter of right metafemur. (25) O. bulga sp. nov.; (26) O. longipes Paulian; (27) O. nammuldi Matthews; (28) O. pexatus Harold; (29) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228987" httpUri="https://zenodo.org/record/13228987/files/figure.png" pageId="6" pageNumber="137">26</figureCitation>
,
<figureCitation id="52D92A6BEC5C506BFB15B4E5FB58FB7B" box="[1220,1252,1222,1248]" captionStart="Figures 30–34" captionStartId="12.[167,244,762,784]" captionTargetBox="[178,1466,144,748]" captionTargetId="figure-10@12.[166,1466,130,748]" captionTargetPageId="12" captionText="Figures 30–34. Onthophagus species, left protibia. (30) O. bulga sp. nov.; (31) O. longipes Paulian; (32) O. nammuldi Matthews; (33) O. pexatus Harold; (34) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228991" httpUri="https://zenodo.org/record/13228991/files/figure.png" pageId="6" pageNumber="137">31</figureCitation>
,
<figureCitation id="52D92A6BEC5C506BFB20B4E5FAADFB7B" box="[1265,1297,1222,1248]" captionStart="Figures 37–41" captionStartId="13.[819,896,1603,1625]" captionTargetBox="[121,1422,151,1679]" captionTargetId="figure-10@13.[121,1421,151,1679]" captionTargetPageId="13" captionText="Figures 37–41. Onthophagus species, pygidium.(37) O. bulga sp. nov.; (38) O. longipes Paulian; (39) O. nammuldi Matthews; (40) O. pexatus Harold; (41) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228995" httpUri="https://zenodo.org/record/13228995/files/figure.png" pageId="6" pageNumber="137">38</figureCitation>
,
<figureCitation id="52D92A6BEC5C506BFACFB4E5FA80FB7B" box="[1310,1340,1222,1248]" captionStart="Figures 42–48" captionStartId="14.[167,244,1399,1421]" captionTargetBox="[167,1459,164,1379]" captionTargetId="figure-65@14.[166,1465,151,1381]" captionTargetPageId="14" captionText="Figures 42–48. Onthophagus species, aedeagus, lateral. (42) O. bulga sp. nov.; (43) O. longipes Paulian; (44) O. nammuldi Matthews; (45) O. squalidus Lea; (46) O. pexatus, Dairy Park, NSW; (47) O. pexatus, Wombeyan Caves, NSW; (48) O. pexatus, Woolomin, NSW." figureDoi="http://doi.org/10.5281/zenodo.13228999" httpUri="https://zenodo.org/record/13228999/files/figure.png" pageId="6" pageNumber="137">43</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="82F86565EC5C506BFCB4B4D6FBF3FAC1" pageId="6" pageNumber="137" type="reference_group">
<paragraph id="CA5D36EEEC5C506BFCB4B4D6FBF3FAC1" blockId="6.[869,1437,1269,1370]" pageId="6" pageNumber="137">
<treatmentCitationGroup id="EAF211C0EC5C506BFCB4B4D6FBF3FAC1" pageId="6" pageNumber="137">
<taxonomicName id="0DE24D6DEC5C506BFCB4B4D6FA98FA91" ID-CoL="7XNFF" authority="Paulian, 1937: 344" authorityName="Paulian" authorityPageNumber="344" authorityYear="1937" box="[869,1316,1269,1291]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="F896EAFCEC5C506BFCB4B4D6FBF2FA91" box="[869,1102,1269,1290]" italics="true" pageId="6" pageNumber="137">Onthophagus longipes</emphasis>
Paulian, 1937: 344
</taxonomicName>
;
<treatmentCitation id="4B4310FFEC5C506BFAE3B4D6FC54FABF" author="Matthews, E." page="212" pageId="6" pageNumber="137" year="1972">
<bibRefCitation id="AE734B1FEC5C506BFAE3B4D6FC54FABF" author="Matthews, E." pageId="6" pageNumber="137" pagination="3 - 330" refId="ref13173" refString="Matthews, E. 1972. A revision of the scarabaeine dung beetles of Australia. I. tribe Onthophagini. Australian Journal of Zoology. Supplementary Series 9: 3 - 330. https: // doi. org / 10.1071 / AJZS 009" type="book chapter" year="1972">Matthews, 1972: 212</bibRefCitation>
</treatmentCitation>
; Williams &amp; Williams, 1982: 44; Williams &amp; Williams, 1983: 29; Cassis &amp; Weir, 1992: 140; Monteith &amp; Kenyon, 2011: 57.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="82F86565EC5C506AFC96B54EFD0EF9A8" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137" type="materials_examined">
<paragraph id="CA5D36EEEC5C506AFC96B54EFD3CFAA5" blockId="6.[839,1467,1389,2000]" lastBlockId="7.[121,749,1229,1342]" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137">
<emphasis id="F896EAFCEC5C506BFC96B54EFB85FA1E" bold="true" box="[839,1081,1389,1413]" pageId="6" pageNumber="137">Material examined</emphasis>
(
<specimenCount id="DCE4FD67EC5C506BFB9EB54EFBCAFA1E" box="[1103,1142,1389,1413]" count="5" pageId="6" pageNumber="137" type="male">5♂</specimenCount>
,
<specimenCount id="DCE4FD67EC5C506BFB59B54DFB11FA1D" box="[1160,1197,1390,1414]" count="4" pageId="6" pageNumber="137" type="female">4♀</specimenCount>
).
<emphasis id="F896EAFCEC5C506BFB1AB54EFC0DFA38" bold="true" pageId="6" pageNumber="137">
<collectingRegion id="0826F80CEC5C506BFB1AB54EFC0DFA38" country="Australia" name="Australian Capital Territory" pageId="6" pageNumber="137">Australian Capital Territory</collectingRegion>
</emphasis>
:
<materialsCitation id="7A8A3CB3EC5C506BFC13B5A8FBEDFA5B" collectingDate="1979-03-10" collectionCode="AMS" elevation="675" latitude="-35.566666" location="Lees Ck" pageId="6" pageNumber="137" specimenCount="1" specimenCount-female="1" stateProvince="Australian Capital Territory">
<specimenCount id="DCE4FD67EC5C506BFC13B5A8FC62FA38" box="[962,990,1419,1443]" count="1" pageId="6" pageNumber="137" type="female">1♀</specimenCount>
/
<location id="CF3D6035EC5C506BFC24B5A8FBECFA38" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFC24B5A8FBECFA38" box="[1013,1104,1419,1443]" latitude="-35.566666" name="Lees Ck" pageId="6" pageNumber="137" stateProvince="Australian Capital Territory">Lees Ck</location>
[vic.
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148°86'E,
<quantity id="0D1A9B0BEC5C506BFAB5B5A8FA13FA39" box="[1380,1455,1419,1443]" metricMagnitude="2" metricUnit="m" metricValue="6.75" pageId="6" pageNumber="137" unit="m" value="675.0">
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],
<collectingMethod id="13A34EF9EC5C506BFC96B58BFC3AFA5B" box="[839,902,1448,1472]" pageId="6" pageNumber="137">pitfall</collectingMethod>
<date id="BE5C102EEC5C506BFC5AB58BFC42FA5B" box="[907,1022,1448,1472]" pageId="6" pageNumber="137" value="1979-03-10">
<collectingDate id="AE18E9C6EC5C506BFC5AB58BFC42FA5B" box="[907,1022,1448,1472]" pageId="6" pageNumber="137" value="1979-03-10">10.iii.1979</collectingDate>
</date>
(
<collectionCode id="ACF3AE2BEC5C506BFBDBB58BFBF5FA5B" box="[1034,1097,1448,1472]" pageId="6" pageNumber="137">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC5C506BFB88B58BFA3CFA60" collectingDate="1992-12-07" collectionCode="AMS" collectorName="Cathedral Rock NP &amp; D Bickel" elevation="1350" location="Barokee Swamp" pageId="6" pageNumber="137" specimenCount="1" specimenCount-female="1" stateProvince="New South Wales">
<emphasis id="F896EAFCEC5C506BFB88B58BFAA7FA5B" bold="true" box="[1113,1307,1448,1472]" pageId="6" pageNumber="137">
<collectingRegion id="0826F80CEC5C506BFB88B58BFAA7FA5B" box="[1113,1307,1448,1472]" country="Australia" name="New South Wales" pageId="6" pageNumber="137">New South Wales</collectingRegion>
</emphasis>
:
<specimenCount id="DCE4FD67EC5C506BFAF7B58BFAFFFA5B" box="[1318,1347,1448,1472]" count="1" pageId="6" pageNumber="137" type="female">1♀</specimenCount>
/
<collectorName id="67175338EC5C506BFA85B58BFC14FA46" pageId="6" pageNumber="137">Cathedral Rock NP</collectorName>
,
<location id="CF3D6035EC5C506BFC63B5E6FBD8FA45" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFC63B5E6FBD8FA45" box="[946,1124,1477,1502]" name="Barokee Swamp" pageId="6" pageNumber="137" stateProvince="New South Wales">Barokee Swamp</location>
, montane sphag. bog/ woodland
<quantity id="0D1A9B0BEC5C506BFC96B5C0FC23FA61" box="[839,927,1507,1531]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="137" unit="m" value="1350.0">
<elevation id="41CFD1DDEC5C506BFC96B5C0FC23FA61" box="[839,927,1507,1531]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="137" unit="m" value="1350.0">1350 m</elevation>
</quantity>
yellow pans
<date id="BE5C102EEC5C506BFBEEB5C0FB0CFA60" box="[1087,1200,1507,1531]" pageId="6" pageNumber="137" value="1992-12-07">
<collectingDate id="AE18E9C6EC5C506BFBEEB5C0FB0CFA60" box="[1087,1200,1507,1531]" pageId="6" pageNumber="137" value="1992-12-07">7.xii.1992</collectingDate>
</date>
<collectorName id="67175338EC5C506BFB6DB5C0FA98FA60" box="[1212,1316,1507,1531]" pageId="6" pageNumber="137">D Bickel</collectorName>
(
<collectionCode id="ACF3AE2BEC5C506BFAE6B5C0FAC5FA60" box="[1335,1401,1507,1531]" pageId="6" pageNumber="137">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC5C506BFA41B5C0FABFF9C8" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="M Gray G Cassis" county="Enfield" elevation="1130" latitude="-31.333334" location="Survey" longLatPrecision="1216" longitude="151.9" municipality="Daisy Patch Fire Trail" pageId="6" pageNumber="137" specimenCount="1" specimenCount-female="1" stateProvince="New South Wales">
<specimenCount id="DCE4FD67EC5C506BFA41B5C0FA12FA60" box="[1424,1454,1507,1531]" count="1" pageId="6" pageNumber="137" type="female">1♀</specimenCount>
/
<collectingCounty id="233C4E62EC5C506BFC99B623FC29F983" box="[840,917,1536,1560]" pageId="6" pageNumber="137">Enfield</collectingCounty>
SF 37BM
<collectingMunicipality id="2A39AC94EC5C506BFBDDB623FB40F983" box="[1036,1276,1536,1560]" pageId="6" pageNumber="137">Daisy Patch Fire Trail</collectingMunicipality>
,
<locationDeviation id="DBEF2FCCEC5C506BFAD7B623FCDBF9AE" location="Enfield Rd" pageId="6" pageNumber="137">
<quantity id="0D1A9B0BEC5C506BFAD7B623FAEDF983" box="[1286,1361,1536,1560]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="137" unit="km" value="1.9">1.9 km</quantity>
S Enfield Rd
</locationDeviation>
<geoCoordinate id="AFD65029EC5C506BFCA3B63DFC76F9AD" box="[882,970,1566,1590]" degrees="31" direction="south" minutes="20" orientation="latitude" pageId="6" pageNumber="137" precision="925" value="-31.333334">31°20'S</geoCoordinate>
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163AS
<quantity id="0D1A9B0BEC5C506BFB71B63DFB4AF9AE" box="[1184,1270,1566,1590]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="6" pageNumber="137" unit="m" value="1130.0">
<elevation id="41CFD1DDEC5C506BFB71B63DFB4AF9AE" box="[1184,1270,1566,1590]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="6" pageNumber="137" unit="m" value="1130.0">1130 m</elevation>
</quantity>
(NPWS
<location id="CF3D6035EC5C506BFAB3B63DFA0EF9AD" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFAB3B63DFA0EF9AD" box="[1378,1458,1566,1590]" county="Enfield" latitude="-31.333334" longLatPrecision="1216" longitude="151.9" municipality="Daisy Patch Fire Trail" name="Survey" pageId="6" pageNumber="137" stateProvince="New South Wales">Survey</location>
)
<date id="BE5C102EEC5C506BFC99B618FC67F9C8" box="[840,987,1595,1619]" pageId="6" pageNumber="137" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">
<collectingDate id="AE18E9C6EC5C506BFC99B618FC67F9C8" box="[840,987,1595,1619]" pageId="6" pageNumber="137" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">4.ii–9.iv.1993</collectingDate>
</date>
<collectorName id="67175338EC5C506BFC37B618FB16F9C8" box="[998,1194,1595,1619]" pageId="6" pageNumber="137">M Gray G Cassis</collectorName>
(
<collectionCode id="ACF3AE2BEC5C506BFB6DB618FB47F9C8" box="[1212,1275,1595,1619]" pageId="6" pageNumber="137">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC5C506BFAC0B618FC2AF930" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="Kunderang Station Ck, NE &amp; M Gray G Cassis" elevation="410" latitude="-30.8" location="Survey" longLatPrecision="1220" longitude="152.1" pageId="6" pageNumber="137" specimenCount="1" specimenCount-female="1" stateProvince="New South Wales">
<specimenCount id="DCE4FD67EC5C506BFAC0B618FA92F9C8" box="[1297,1326,1595,1619]" count="1" pageId="6" pageNumber="137" type="female">1♀</specimenCount>
/
<collectorName id="67175338EC5C506BFA95B618FC54F9EB" pageId="6" pageNumber="137">Kunderang Station Ck, NE</collectorName>
facing slope above 39BR
<geoCoordinate id="AFD65029EC5C506BFB2EB67BFAEEF9EB" box="[1279,1362,1624,1648]" degrees="30" direction="south" minutes="48" orientation="latitude" pageId="6" pageNumber="137" precision="925" value="-30.8">30°48'S</geoCoordinate>
<geoCoordinate id="AFD65029EC5C506BFA89B67BFA07F9EB" box="[1368,1467,1624,1648]" degrees="152" direction="east" minutes="06" orientation="longitude" pageId="6" pageNumber="137" precision="925" value="152.1">152°06'E</geoCoordinate>
<quantity id="0D1A9B0BEC5C506BFC96B655FC33F916" box="[839,911,1654,1678]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="137" unit="m" value="410.0">
<elevation id="41CFD1DDEC5C506BFC96B655FC33F916" box="[839,911,1654,1678]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="137" unit="m" value="410.0">410 m</elevation>
</quantity>
(NPWS
<location id="CF3D6035EC5C506BFC26B655FBF9F915" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFC26B655FBF9F915" box="[1015,1093,1654,1678]" latitude="-30.8" longLatPrecision="1220" longitude="152.1" name="Survey" pageId="6" pageNumber="137" stateProvince="New South Wales">Survey</location>
)
<date id="BE5C102EEC5C506BFB87B656FB50F916" box="[1110,1260,1653,1677]" pageId="6" pageNumber="137" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">
<collectingDate id="AE18E9C6EC5C506BFB87B656FB50F916" box="[1110,1260,1653,1677]" pageId="6" pageNumber="137" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">4.ii–9.iv.1993</collectingDate>
</date>
<collectorName id="67175338EC5C506BFB27B655FA07F916" box="[1270,1467,1653,1678]" pageId="6" pageNumber="137">M Gray G Cassis</collectorName>
(
<collectionCode id="ACF3AE2BEC5C506BFC9EB6B0FC32F930" box="[847,910,1683,1707]" pageId="6" pageNumber="137">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC5C506BFC71B6B0FB48F97D" collectingDate="1999-05-13" collectionCode="AMS" collectorName="S Lassau &amp; C Lemann" county="Marengo" latitude="-30.105" location="Chaelundi Rd" longLatPrecision="73" longitude="152.412" municipality="Chimney Rd" pageId="6" pageNumber="137" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales">
<specimenCount id="DCE4FD67EC5C506BFC71B6B0FC0FF930" box="[928,947,1683,1707]" count="1" pageId="6" pageNumber="137" type="male">♂</specimenCount>
/
<collectingCounty id="233C4E62EC5C506BFC13B6B0FB9EF930" box="[962,1058,1683,1707]" pageId="6" pageNumber="137">Marengo</collectingCounty>
SF
<quantity id="0D1A9B0BEC5C506BFB9CB6B0FB2AF930" box="[1101,1174,1683,1707]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="137" unit="km" value="2.1">2.1 km</quantity>
along
<collectingMunicipality id="2A39AC94EC5C506BFB0FB6B0FAD9F930" box="[1246,1381,1683,1707]" pageId="6" pageNumber="137">Chimney Rd</collectingMunicipality>
from jn with
<location id="CF3D6035EC5C506BFCAEB693FBA8F953" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFCAEB693FBA8F953" box="[895,1044,1712,1736]" county="Marengo" latitude="-30.105" longLatPrecision="73" longitude="152.412" municipality="Chimney Rd" name="Chaelundi Rd" pageId="6" pageNumber="137" stateProvince="New South Wales">Chaelundi Rd</location>
<geoCoordinate id="AFD65029EC5C506BFBCCB693FB3CF953" box="[1053,1152,1711,1736]" degrees="30.105" direction="south" orientation="latitude" pageId="6" pageNumber="137" precision="55" value="-30.105">30.105°S</geoCoordinate>
<geoCoordinate id="AFD65029EC5C506BFB59B693FB45F953" box="[1160,1273,1711,1736]" degrees="152.412" direction="east" orientation="longitude" pageId="6" pageNumber="137" precision="55" value="152.412">152.412°E</geoCoordinate>
<collectingMethod id="13A34EF9EC5C506BFAD3B693FA06F953" box="[1282,1466,1712,1736]" pageId="6" pageNumber="137">human dung trap</collectingMethod>
<date id="BE5C102EEC5C506BFC96B6EDFC0CF97D" box="[839,944,1742,1766]" pageId="6" pageNumber="137" value="1999-05-13">
<collectingDate id="AE18E9C6EC5C506BFC96B6EDFC0CF97D" box="[839,944,1742,1766]" pageId="6" pageNumber="137" value="1999-05-13">13.v.1999</collectingDate>
</date>
<collectorName id="67175338EC5C506BFC64B6EDFBAFF97E" box="[949,1043,1742,1766]" pageId="6" pageNumber="137">S Lassau</collectorName>
&amp;
<collectorName id="67175338EC5C506BFBE3B6EDFB23F97E" box="[1074,1183,1742,1766]" pageId="6" pageNumber="137">C Lemann</collectorName>
(
<collectionCode id="ACF3AE2BEC5C506BFB7CB6EDFB50F97D" box="[1197,1260,1742,1766]" pageId="6" pageNumber="137">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC5C506BFB2CB6EDFC62F8C0" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="M Gray G Cassis" county="New England" elevation="1350" latitude="-30.516666" location="New England" longLatPrecision="1221" longitude="152.38333" municipality="Cliffs Trail" pageId="6" pageNumber="137" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales">
<specimenCount id="DCE4FD67EC5C506BFB2CB6EDFAA7F97D" box="[1277,1307,1742,1766]" count="1" pageId="6" pageNumber="137" type="male">1♂</specimenCount>
/
<collectingCounty id="233C4E62EC5C506BFAFBB6EDFA07F97E" box="[1322,1467,1741,1765]" pageId="6" pageNumber="137">New England</collectingCounty>
NP 33AM
<collectingMunicipality id="2A39AC94EC5C506BFC68B6C8FB90F898" box="[953,1068,1771,1795]" pageId="6" pageNumber="137">Cliffs Trail</collectingMunicipality>
(top end) bordering
<location id="CF3D6035EC5C506BFAD2B6C8FA28F898" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFAD2B6C8FA28F898" box="[1283,1428,1771,1795]" county="New England" latitude="-30.516666" longLatPrecision="1221" longitude="152.38333" municipality="Cliffs Trail" name="New England" pageId="6" pageNumber="137" stateProvince="New South Wales">New England</location>
NP &amp;
<location id="CF3D6035EC5C506BFCB5B72BFC64F8BB" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFCB5B72BFC64F8BB" box="[868,984,1800,1824]" county="New England" latitude="-30.516666" longLatPrecision="1221" longitude="152.38333" municipality="Cliffs Trail" name="Styx River" pageId="6" pageNumber="137" stateProvince="New South Wales">Styx River</location>
SF, about
<locationDeviation id="DBEF2FCCEC5C506BFB9FB72BFAE2F8BB" box="[1102,1374,1800,1824]" location="Point Lookout" pageId="6" pageNumber="137">
<quantity id="0D1A9B0BEC5C506BFB9FB72BFB3AF8BB" box="[1102,1158,1800,1824]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="137" unit="km" value="3.0">3 km</quantity>
S of Point Lookout
</locationDeviation>
<geoCoordinate id="AFD65029EC5C506BFAB6B72BFA06F8BB" box="[1383,1466,1800,1824]" degrees="30" direction="south" minutes="31" orientation="latitude" pageId="6" pageNumber="137" precision="925" value="-30.516666">30°31'S</geoCoordinate>
<geoCoordinate id="AFD65029EC5C506BFC96B706FC16F8A6" box="[839,938,1829,1853]" degrees="152" direction="east" minutes="23" orientation="longitude" pageId="6" pageNumber="137" precision="925" value="152.38333">152°23'E</geoCoordinate>
<quantity id="0D1A9B0BEC5C506BFC7EB706FC43F8A6" box="[943,1023,1829,1853]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="137" unit="m" value="1350.0">
<elevation id="41CFD1DDEC5C506BFC7EB706FC43F8A6" box="[943,1023,1829,1853]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="137" unit="m" value="1350.0">1350 m</elevation>
</quantity>
(NPWS
<location id="CF3D6035EC5C506BFB8CB705FB16F8A5" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFB8CB705FB16F8A5" box="[1117,1194,1830,1854]" county="New England" latitude="-30.516666" longLatPrecision="1221" longitude="152.38333" municipality="Cliffs Trail" name="Survey" pageId="6" pageNumber="137" stateProvince="New South Wales">Survey</location>
)
<date id="BE5C102EEC5C506BFB67B706FAF6F8A6" box="[1206,1354,1829,1853]" pageId="6" pageNumber="137" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">
<collectingDate id="AE18E9C6EC5C506BFB67B706FAF6F8A6" box="[1206,1354,1829,1853]" pageId="6" pageNumber="137" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">4.ii–9.iv.1993</collectingDate>
</date>
<collectorName id="67175338EC5C506BFA9EB705FC36F8C0" pageId="6" pageNumber="137">M Gray G Cassis</collectorName>
(
<collectionCode id="ACF3AE2BEC5C506BFC46B760FC6AF8C0" box="[919,982,1859,1883]" pageId="6" pageNumber="137">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC5C506BFC37B760FC62F828" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="Stewarts Brook SF &amp; M Gray G Cassis" elevation="1250" latitude="-31.9" location="Survey" longLatPrecision="1213" longitude="151.38333" municipality="Omadale Brook Rd" pageId="6" pageNumber="137" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales">
<specimenCount id="DCE4FD67EC5C506BFC37B760FBB9F8C0" box="[998,1029,1859,1883]" count="1" pageId="6" pageNumber="137" type="male">1♂</specimenCount>
*/
<collectorName id="67175338EC5C506BFBF0B760FB59F8C0" box="[1057,1253,1859,1883]" pageId="6" pageNumber="137">Stewarts Brook SF</collectorName>
62AM
<quantity id="0D1A9B0BEC5C506BFAE2B760FAC7F8C0" box="[1331,1403,1859,1883]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="137" unit="km" value="0.7">0.7 km</quantity>
along unnamed logging track from
<collectingMunicipality id="2A39AC94EC5C506BFB54B743FAE7F8E3" box="[1157,1371,1888,1912]" pageId="6" pageNumber="137">Omadale Brook Rd</collectingMunicipality>
,
<geoCoordinate id="AFD65029EC5C506BFAB6B743FA06F8E3" box="[1383,1466,1888,1912]" degrees="31" direction="south" minutes="54" orientation="latitude" pageId="6" pageNumber="137" precision="925" value="-31.9">31°54'S</geoCoordinate>
<geoCoordinate id="AFD65029EC5C506BFC96B75EFC15F80E" box="[839,937,1917,1941]" degrees="151" direction="east" minutes="23" orientation="longitude" pageId="6" pageNumber="137" precision="925" value="151.38333">151°23'E</geoCoordinate>
<quantity id="0D1A9B0BEC5C506BFC7EB75EFC43F80E" box="[943,1023,1917,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="6" pageNumber="137" unit="m" value="1250.0">
<elevation id="41CFD1DDEC5C506BFC7EB75EFC43F80E" box="[943,1023,1917,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="6" pageNumber="137" unit="m" value="1250.0">1250 m</elevation>
</quantity>
(NPWS
<location id="CF3D6035EC5C506BFB8CB75DFB16F80D" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFB8CB75DFB16F80D" box="[1117,1194,1918,1942]" latitude="-31.9" longLatPrecision="1213" longitude="151.38333" municipality="Omadale Brook Rd" name="Survey" pageId="6" pageNumber="137" stateProvince="New South Wales">Survey</location>
)
<date id="BE5C102EEC5C506BFB67B75EFAF6F80E" box="[1206,1354,1917,1941]" pageId="6" pageNumber="137" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">
<collectingDate id="AE18E9C6EC5C506BFB67B75EFAF6F80E" box="[1206,1354,1917,1941]" pageId="6" pageNumber="137" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">4.ii–9.iv.1993</collectingDate>
</date>
<collectorName id="67175338EC5C506BFA9EB75DFC37F828" pageId="6" pageNumber="137">M Gray G Cassis</collectorName>
(
<collectionCode id="ACF3AE2BEC5C506BFC46B7B8FC6AF828" box="[919,982,1947,1971]" pageId="6" pageNumber="137">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC5C506AFC36B7B8FEAEFA98" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="M Gray G Cassis" county="Stewarts Brook" elevation="1430" lastPageId="7" lastPageNumber="138" latitude="-31.933332" location="southern Green Gap" longLatPrecision="1213" longitude="151.43333" municipality="Barrington Trail" pageId="6" pageNumber="137" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales">
<specimenCount id="DCE4FD67EC5C506BFC36B7B8FBB9F828" box="[999,1029,1947,1971]" count="1" pageId="6" pageNumber="137" type="male">1♂</specimenCount>
*/
<collectingCounty id="233C4E62EC5C506BFBF0B7B8FB7FF828" box="[1057,1219,1947,1971]" pageId="6" pageNumber="137">Stewarts Brook</collectingCounty>
SF 62CM
<quantity id="0D1A9B0BEC5C506BFAE2B7B8FAC7F828" box="[1331,1403,1947,1971]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="137" unit="km" value="0.4">0.4 km</quantity>
along
<collectingMunicipality id="2A39AC94EC5C506BFC96B79BFC4FF84B" box="[839,1011,1976,2000]" pageId="6" pageNumber="137">Barrington Trail</collectingMunicipality>
from
<location id="CF3D6035EC5C506BFBE5B79BFAB6F84B" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5C506BFBE5B79BFAB6F84B" box="[1076,1290,1976,2000]" county="Stewarts Brook" latitude="-31.933332" longLatPrecision="1213" longitude="151.43333" municipality="Barrington Trail" name="southern Green Gap" pageId="6" pageNumber="137" stateProvince="New South Wales">southern Green Gap</location>
turnoff,
<geoCoordinate id="AFD65029EC5C506BFAB6B79BFA06F84B" box="[1383,1466,1976,2000]" degrees="31" direction="south" minutes="56" orientation="latitude" pageId="6" pageNumber="137" precision="925" value="-31.933332">31°56'S</geoCoordinate>
<geoCoordinate id="AFD65029EC5D506AFFA8B4EEFF67FB7E" box="[121,219,1229,1253]" degrees="151" direction="east" minutes="26" orientation="longitude" pageId="7" pageNumber="138" precision="925" value="151.43333">151°26'E</geoCoordinate>
<quantity id="0D1A9B0BEC5D506AFF30B4EDFE8DFB7E" box="[225,305,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="7" pageNumber="138" unit="m" value="1430.0">
<elevation id="41CFD1DDEC5D506AFF30B4EDFE8DFB7E" box="[225,305,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="7" pageNumber="138" unit="m" value="1430.0">1430 m</elevation>
</quantity>
(NPWS
<location id="CF3D6035EC5D506AFE5FB4EDFE67FB7D" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5D506AFE5FB4EDFE67FB7D" box="[398,475,1230,1254]" county="Stewarts Brook" latitude="-31.933332" longLatPrecision="1213" longitude="151.43333" municipality="Barrington Trail" name="Survey" pageId="7" pageNumber="138" stateProvince="New South Wales">Survey</location>
)
<date id="BE5C102EEC5D506AFE39B4EDFDC0FB7D" box="[488,636,1230,1254]" pageId="7" pageNumber="138" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">
<collectingDate id="AE18E9C6EC5D506AFE39B4EDFDC0FB7D" box="[488,636,1230,1254]" pageId="7" pageNumber="138" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">4.ii–9.iv.1993</collectingDate>
</date>
<collectorName id="67175338EC5D506AFD50B4EDFF01FA98" pageId="7" pageNumber="138">M Gray G Cassis</collectorName>
(
<collectionCode id="ACF3AE2BEC5D506AFF1AB4C8FEB6FA98" box="[203,266,1259,1283]" pageId="7" pageNumber="138">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC5D506AFECCB4C8FDC0FAA5" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="Stewarts Brook SF &amp; AG Pol Blue Ck &amp; Gray &amp; Cassis" elevation="1240" latitude="-31.916666" location="Survey" longLatPrecision="1213" longitude="151.38333" municipality="Tubbrabucca Rd" pageId="7" pageNumber="138" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales">
<specimenCount id="DCE4FD67EC5D506AFECCB4C8FE80FA98" box="[285,316,1259,1283]" count="1" pageId="7" pageNumber="138" type="male">1♂</specimenCount>
/
<collectorName id="67175338EC5D506AFE9DB4C8FDAAFA98" box="[332,534,1259,1283]" pageId="7" pageNumber="138">Stewarts Brook SF</collectorName>
62
<collectorName id="67175338EC5D506AFDE7B4C8FD55FA98" box="[566,745,1259,1283]" pageId="7" pageNumber="138">AG Pol Blue Ck</collectorName>
, above
<collectingMunicipality id="2A39AC94EC5D506AFF6EB52BFECEFABB" box="[191,370,1288,1312]" pageId="7" pageNumber="138">Tubbrabucca Rd</collectingMunicipality>
,
<geoCoordinate id="AFD65029EC5D506AFEACB52BFE6DFABB" box="[381,465,1288,1312]" degrees="31" direction="south" minutes="55" orientation="latitude" pageId="7" pageNumber="138" precision="925" value="-31.916666">31°55'S</geoCoordinate>
<geoCoordinate id="AFD65029EC5D506AFE09B52BFD86FABB" box="[472,570,1288,1312]" degrees="151" direction="east" minutes="23" orientation="longitude" pageId="7" pageNumber="138" precision="925" value="151.38333">151°23'E</geoCoordinate>
<quantity id="0D1A9B0BEC5D506AFD90B52BFD2FFABB" box="[577,659,1288,1312]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="7" pageNumber="138" unit="m" value="1240.0">
<elevation id="41CFD1DDEC5D506AFD90B52BFD2FFABB" box="[577,659,1288,1312]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="7" pageNumber="138" unit="m" value="1240.0">1240 m</elevation>
</quantity>
(NPWS
<location id="CF3D6035EC5D506AFFA8B505FF7AFAA5" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC5D506AFFA8B505FF7AFAA5" box="[121,198,1318,1342]" latitude="-31.916666" longLatPrecision="1213" longitude="151.38333" municipality="Tubbrabucca Rd" name="Survey" pageId="7" pageNumber="138" stateProvince="New South Wales">Survey</location>
)
<date id="BE5C102EEC5D506AFF05B505FEDBFAA5" box="[212,359,1318,1342]" pageId="7" pageNumber="138" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">
<collectingDate id="AE18E9C6EC5D506AFF05B505FEDBFAA5" box="[212,359,1318,1342]" pageId="7" pageNumber="138" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">4.ii–9.iv.1993</collectingDate>
</date>
M
<collectorName id="67175338EC5D506AFE5DB505FE7DFAA5" box="[396,449,1318,1342]" pageId="7" pageNumber="138">Gray</collectorName>
G
<collectorName id="67175338EC5D506AFE33B505FD9AFAA5" box="[482,550,1318,1342]" pageId="7" pageNumber="138">Cassis</collectorName>
(
<collectionCode id="ACF3AE2BEC5D506AFDE4B505FDC8FAA5" box="[565,628,1318,1342]" pageId="7" pageNumber="138">AMS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="9E9D6666EC5D506AFEF1B4AEFB5BFB38" ID-DOI="http://doi.org/10.5281/zenodo.13228977" ID-Zenodo-Dep="13228977" box="[288,1255,1165,1187]" httpUri="https://zenodo.org/record/13228977/files/figure.png" pageId="7" pageNumber="138" startId="7.[288,365,1165,1187]" targetBox="[219,1322,167,1151]" targetPageId="7" targetType="figure">
<paragraph id="CA5D36EEEC5D506AFEF1B4AEFB5BFB38" blockId="7.[288,1255,1165,1187]" box="[288,1255,1165,1187]" pageId="7" pageNumber="138">
<emphasis id="F896EAFCEC5D506AFEF1B4AEFE11FB39" bold="true" box="[288,429,1165,1187]" pageId="7" pageNumber="138">Figures 11, 12</emphasis>
. Lateral margin pronotum. (
<emphasis id="F896EAFCEC5D506AFD15B4AEFD66FB39" box="[708,730,1165,1186]" italics="true" pageId="7" pageNumber="138">11</emphasis>
)
<taxonomicName id="0DE24D6DEC5D506AFD39B4AEFC1BFB39" authorityName="Reid &amp; Runagall-Mcnaull" authorityYear="2022" box="[744,935,1165,1186]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="bulga" status="sp. nov.">
<emphasis id="F896EAFCEC5D506AFD39B4AEFC1BFB39" box="[744,935,1165,1186]" italics="true" pageId="7" pageNumber="138">Onthophagus bulga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3A55787EC5D506AFC7CB4ADFC46FB38" box="[941,1018,1166,1187]" pageId="7" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
; (
<emphasis id="F896EAFCEC5D506AFBDAB4AEFB9FFB39" box="[1035,1059,1165,1186]" italics="true" pageId="7" pageNumber="138">12</emphasis>
)
<taxonomicName id="0DE24D6DEC5D506AFBE0B4AEFB5BFB38" authority="Harold." authorityName="Harold." box="[1073,1255,1165,1187]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="pexatus">
<emphasis id="F896EAFCEC5D506AFBE0B4AEFB2BFB38" box="[1073,1175,1165,1187]" italics="true" pageId="7" pageNumber="138">O. pexatus</emphasis>
Harold.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="CA5D36EEEC5D506AFFA8B56EFD0EF9A8" blockId="7.[121,749,1357,1587]" pageId="7" pageNumber="138">
<emphasis id="F896EAFCEC5D506AFFA8B56EFEE6FAFD" bold="true" box="[121,346,1357,1382]" pageId="7" pageNumber="138">Additional records</emphasis>
(not included in
<bibRefCitation id="AE734B1FEC5D506AFDF6B56DFD5CFAFD" author="Matthews, E." box="[551,736,1358,1382]" pageId="7" pageNumber="138" pagination="3 - 330" refId="ref13173" refString="Matthews, E. 1972. A revision of the scarabaeine dung beetles of Australia. I. tribe Onthophagini. Australian Journal of Zoology. Supplementary Series 9: 3 - 330. https: // doi. org / 10.1071 / AJZS 009" type="book chapter" year="1972">Matthews, 1972</bibRefCitation>
).
<emphasis id="F896EAFCEC5D506AFFA8B548FE7EFA18" bold="true" box="[121,450,1387,1411]" pageId="7" pageNumber="138">
<collectingRegion id="0826F80CEC5D506AFFA8B548FE7EFA18" box="[121,450,1387,1411]" country="Australia" name="Australian Capital Territory" pageId="7" pageNumber="138">Australian Capital Territory</collectingRegion>
</emphasis>
: 2/ 3k &amp; 6k NE Piccadilly Circus (
<collectionCode id="ACF3AE2BEC5D506AFF02B5ABFEA0FA3B" box="[211,284,1416,1440]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
);
<emphasis id="F896EAFCEC5D506AFEE2B5ABFDB9FA3B" bold="true" box="[307,517,1416,1440]" pageId="7" pageNumber="138">
<collectingRegion id="0826F80CEC5D506AFEE2B5ABFDB9FA3B" box="[307,517,1416,1440]" country="Australia" name="New South Wales" pageId="7" pageNumber="138">New South Wales</collectingRegion>
</emphasis>
: 3/ Moppy Lookout (
<collectionCode id="ACF3AE2BEC5D506AFF50B585FF75FA25" box="[129,201,1446,1470]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
); 1/ Mt Gibraltar (
<collectionCode id="ACF3AE2BEC5D506AFE4CB585FE59FA25" box="[413,485,1446,1470]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
); 7/ Robertson (
<collectionCode id="ACF3AE2BEC5D506AFD4CB585FD5CFA25" box="[669,736,1446,1470]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="7" pageNumber="138" type="Museum">QMB</collectionCode>
); 2/ Thungutti Campground (
<collectionCode id="ACF3AE2BEC5D506AFE67B5E0FDBCFA40" box="[438,512,1475,1499]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
);
<emphasis id="F896EAFCEC5D506AFDC6B5E0FDCBFA40" bold="true" box="[535,631,1475,1499]" pageId="7" pageNumber="138">
<collectingRegion id="0826F80CEC5D506AFDC6B5E0FDCBFA40" box="[535,631,1475,1499]" country="Australia" name="Victoria" pageId="7" pageNumber="138">Victoria</collectingRegion>
</emphasis>
: 2/ Ghost Point (
<collectionCode id="ACF3AE2BEC5D506AFF13B5C3FEB5FA63" box="[194,265,1504,1528]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
); 1/ 2.2k NE Warburton (
<collectionCode id="ACF3AE2BEC5D506AFDF0B5C3FDD4FA63" box="[545,616,1504,1528]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
); 1/ 6k ESE Tanjil Bren (
<collectionCode id="ACF3AE2BEC5D506AFED1B5DDFEFBF98D" box="[256,327,1534,1558]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
); 1/ Tarra-Bulga NP (
<collectionCode id="ACF3AE2BEC5D506AFDFDB5DDFDCFF98D" box="[556,627,1534,1558]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
); 5/ Wilson Promontory (
<collectionCode id="ACF3AE2BEC5D506AFED7B638FEF7F9A8" box="[262,331,1563,1587]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
,
<collectionCode id="ACF3AE2BEC5D506AFE87B638FE1FF9A8" box="[342,419,1563,1587]" pageId="7" pageNumber="138">QDAF</collectionCode>
); 2/ Yanks Folly (
<collectionCode id="ACF3AE2BEC5D506AFD8EB638FD1AF9A8" box="[607,678,1563,1587]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="138">ANIC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="82F86565EC5D5065FFA8B660FBEEF824" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138" type="description">
<paragraph id="CA5D36EEEC5D506AFFA8B660FE4CF8B3" blockId="7.[121,749,1603,1832]" pageId="7" pageNumber="138">
<emphasis id="F896EAFCEC5D506AFFA8B660FEA7F9C0" bold="true" box="[121,283,1603,1627]" pageId="7" pageNumber="138">Redescription</emphasis>
. Mature specimens with shiny shallowly microsculptured black head and pronotum, duller and densely microsculptured grey-black elytra, with dull microsculptured striae, and dull densely microsculptured grey-black pygidium; freshly emerged specimens usually teneral, with reddish elytral apices, venter, pygidium and legs; antennae reddish-brown, with brown clubs. Length, male
<quantity id="0D1A9B0BEC5D506AFF62B733FE9FF8B3" box="[179,291,1808,1832]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="7" pageNumber="138" unit="mm" value="5.25" valueMax="6.0" valueMin="4.5">4.5–6 mm</quantity>
; female
<quantity id="0D1A9B0BEC5D506AFEACB733FE51F8B3" box="[381,493,1808,1832]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="7" pageNumber="138" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">4–5.5 mm</quantity>
.
</paragraph>
<paragraph id="CA5D36EEEC5D506AFFA8B71BFAB2FAE3" blockId="7.[121,749,1848,1990]" lastBlockId="7.[793,1422,1230,1990]" pageId="7" pageNumber="138">
<emphasis id="F896EAFCEC5D506AFFA8B71BFF0FF8CB" bold="true" box="[121,179,1848,1872]" pageId="7" pageNumber="138">Male</emphasis>
. Head (
<figureCitation id="52D92A6BEC5D506AFED7B71BFEE9F8CB" box="[262,341,1848,1872]" captionStart="Figures 17–20" captionStartId="9.[121,198,1260,1282]" captionTargetBox="[121,1421,144,1244]" captionTargetId="figure-10@9.[121,1421,144,1244]" captionTargetPageId="9" captionText="Figures 17–20. Onthophagus species, head, male (left), and female (right), fresh specimens. (17, 18) O. longipes Paulian; (19, 20) O. nammuldi Matthews." figureDoi="http://doi.org/10.5281/zenodo.13228983" httpUri="https://zenodo.org/record/13228983/files/figure.png" pageId="7" pageNumber="138">Fig. 17</figureCitation>
). Surface smooth, shiny but shallowly microreticulate, with dense large punctures at base becoming smaller and sparser towards apex, minute stubble on genae and longer semi-erect setae on sides of vertex. Clypeal apical margin uptilted, arcuately excavate, each side of excavation triangularly produced, side margins slightly rounded; clypeal suture entirely effaced at frontal portion, genal portions present and feebly raised; frons flat, without elevations, slightly medially depressed; eyes narrow, 5–7 facet rows in width, separated by 15–20 eye widths, canthus narrowly complete; mentum shallowly excavate at apex.
</paragraph>
<paragraph id="CA5D36EEEC5D5065FCE9B5A3FEE4F974" blockId="7.[793,1422,1230,1990]" lastBlockId="8.[166,795,1223,1981]" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138">
Thorax (
<figureCitation id="52D92A6BEC5D506AFC49B5A3FC42FA03" box="[920,1022,1408,1432]" captionStart="Figures 1–4" captionStartId="4.[216,293,1867,1889]" captionTargetBox="[224,1375,175,1822]" captionTargetId="figure-10@4.[167,1467,151,1828]" captionTargetPageId="4" captionText="Figures 1–4. (1, 2) Onthophagus bulga sp. nov., male, dorsal and lateral; (3, 4) O. longipes Paulian, male, dorsal and lateral." figureDoi="http://doi.org/10.5281/zenodo.13228971" httpUri="https://zenodo.org/record/13228971/files/figure.png" pageId="7" pageNumber="138">Figs 3–4</figureCitation>
,
<figureCitation id="52D92A6BEC5D506AFBDCB5A3FB96FA03" box="[1037,1066,1408,1432]" captionStart="Figures 25–29" captionStartId="11.[121,198,1565,1587]" captionTargetBox="[121,748,151,1551]" captionTargetId="figure-106@11.[121,748,151,1553]" captionTargetPageId="11" captionText="Figures 25–29. Onthophagus species, venter of right metafemur. (25) O. bulga sp. nov.; (26) O. longipes Paulian; (27) O. nammuldi Matthews; (28) O. pexatus Harold; (29) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228987" httpUri="https://zenodo.org/record/13228987/files/figure.png" pageId="7" pageNumber="138">26</figureCitation>
,
<figureCitation id="52D92A6BEC5D506AFBE8B5A3FBEBFA03" box="[1081,1111,1408,1432]" captionStart="Figures 30–34" captionStartId="12.[167,244,762,784]" captionTargetBox="[178,1466,144,748]" captionTargetId="figure-10@12.[166,1466,130,748]" captionTargetPageId="12" captionText="Figures 30–34. Onthophagus species, left protibia. (30) O. bulga sp. nov.; (31) O. longipes Paulian; (32) O. nammuldi Matthews; (33) O. pexatus Harold; (34) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228991" httpUri="https://zenodo.org/record/13228991/files/figure.png" pageId="7" pageNumber="138">31</figureCitation>
). Pronotum. Broad in large males (as wide as elytra), narrower and flatter in small males. Moderately convex, slightly tumid at middle of anterior slope in larger specimens; surface smooth, entirely strongly and closely punctured, punctures shallowly ocellate and often slightly elongate, intervals shiny in fresh material but microreticulate, less than half diameter of punctures, and covered in dense short semi-erect to almost recumbent setae; anterior angles 80–90°; anterior margined, sides margined and crenulate with curved short setae between crenulations, base with thin raised edge which may be partly effaced; pronotal hypomeron dull and densely microsculptured, strongly punctured on outer half, punctures separated by 1–2 diameters, with elevated rims, increasing in diameter towards sides, outermost punctures with long semi-erect setae. Elytra. Semi-ovate in dorsal view, with prominent humeri and preapical swelling, longer than pronotum along midline. Duller than pronotum but weakly shiny, striae and suture similar, densely microreticulate; intervals 1–6 flat or slightly uneven in teneral specimens, without obvious small tubercles or elevations, 7th interval may have a few small round shiny tubercles, outer interval with a few lunate punctures; intervals 1–7 with fairly dense erect short scale-like setae, not in distinct rows, outermost interval more densely setose; strial punctures usually distinct, slightly broader and deeper than striae; epipleura with single row of curved setae. Metaventrite medially shiny, anteriorly and laterally (lateral to mesocoxae) dull and microreticulate, with scattered semi-recumbent setae, anterior smoothly convex, midline grooved; wing fully developed, folded twice in repose. Legs. Protibiae elongated and narrowed, inner apical angle with compact dense tuft of yellow setae about twice length of apical spur; distal face of apical tooth with loose sparse tuft of yellow setae about as long as apical spur, 4th (smallest) lateral tooth at about midpoint; metafemoral ventral surface rough, with short elongate ridges (elevated edges of punctures), mostly dull and microreticulate, with moderately large but shallow close punctures separated by 0.5–2 diameters.
</paragraph>
<caption id="9E9D6666EC525065FF7FB451FA0FFB13" ID-DOI="http://doi.org/10.5281/zenodo.13228981" ID-Zenodo-Dep="13228981" box="[174,1459,1138,1160]" httpUri="https://zenodo.org/record/13228981/files/figure.png" pageId="8" pageNumber="139" startId="8.[174,251,1138,1160]" targetBox="[167,1467,151,1119]" targetPageId="8" targetType="figure">
<paragraph id="CA5D36EEEC525065FF7FB451FA0FFB13" blockId="8.[174,1459,1138,1160]" box="[174,1459,1138,1160]" pageId="8" pageNumber="139">
<emphasis id="F896EAFCEC525065FF7FB451FE82FB1C" bold="true" box="[174,318,1138,1160]" pageId="8" pageNumber="139">Figures 13–16</emphasis>
.
<taxonomicName id="0DE24D6DEC525065FE9EB451FDB2FB1C" authorityName="Reid &amp; Runagall-Mcnaull" authorityYear="2022" box="[335,526,1138,1159]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="bulga" status="sp. nov.">
<emphasis id="F896EAFCEC525065FE9EB451FDB2FB1C" box="[335,526,1138,1159]" italics="true" pageId="8" pageNumber="139">Onthophagus bulga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3A55787EC525065FDC2B451FDDCFB1C" box="[531,608,1138,1159]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
, head, male (left) and female (right). (
<emphasis id="F896EAFCEC525065FC1FB451FBB6FB1C" box="[974,1034,1138,1159]" italics="true" pageId="8" pageNumber="139">13, 14</emphasis>
) fresh specimens; (
<emphasis id="F896EAFCEC525065FB16B451FABFFB1C" box="[1223,1283,1138,1159]" italics="true" pageId="8" pageNumber="139">15, 16</emphasis>
) worn specimens.
</paragraph>
</caption>
<paragraph id="CA5D36EEEC525065FF14B6D6FEC1F826" blockId="8.[166,795,1223,1981]" pageId="8" pageNumber="139">
Abdomen (
<figureCitation id="52D92A6BEC525065FEE6B6D6FE3EF896" box="[311,386,1781,1805]" captionStart="Figures 37–41" captionStartId="13.[819,896,1603,1625]" captionTargetBox="[121,1422,151,1679]" captionTargetId="figure-10@13.[121,1421,151,1679]" captionTargetPageId="13" captionText="Figures 37–41. Onthophagus species, pygidium.(37) O. bulga sp. nov.; (38) O. longipes Paulian; (39) O. nammuldi Matthews; (40) O. pexatus Harold; (41) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228995" httpUri="https://zenodo.org/record/13228995/files/figure.png" pageId="8" pageNumber="139">Fig. 38</figureCitation>
). Ventrites dull, densely microreticulate, obscuring puncturation, each with single transverse row of semi-recumbent pale setae; pygidium flat or almost so, densely and evenly microreticulate except shiny raised margins; punctures distinct or obscured by microsculpture, sparse, separated by 1–3 diameters; setae short, but thin, length 7–10× width.
</paragraph>
<paragraph id="CA5D36EEEC525065FCB4B4E4FBD4FA81" blockId="8.[839,1466,1223,1306]" pageId="8" pageNumber="139">
Aedeagus (
<figureCitation id="52D92A6BEC525065FC0DB4E4FB96FB7B" box="[988,1066,1223,1248]" captionStart="Figures 42–48" captionStartId="14.[167,244,1399,1421]" captionTargetBox="[167,1459,164,1379]" captionTargetId="figure-65@14.[166,1465,151,1381]" captionTargetPageId="14" captionText="Figures 42–48. Onthophagus species, aedeagus, lateral. (42) O. bulga sp. nov.; (43) O. longipes Paulian; (44) O. nammuldi Matthews; (45) O. squalidus Lea; (46) O. pexatus, Dairy Park, NSW; (47) O. pexatus, Wombeyan Caves, NSW; (48) O. pexatus, Woolomin, NSW." figureDoi="http://doi.org/10.5281/zenodo.13228999" httpUri="https://zenodo.org/record/13228999/files/figure.png" pageId="8" pageNumber="139">Fig. 43</figureCitation>
). In lateral view, parameres “crested”, with an angular apico-dorsal elevation and prominent but short apico-ventral “beak”.
</paragraph>
<paragraph id="CA5D36EEEC525065FC96B513FAB0F98E" blockId="8.[839,1467,1327,1557]" pageId="8" pageNumber="139">
<emphasis id="F896EAFCEC525065FC96B513FC24FAD3" bold="true" box="[839,920,1328,1352]" pageId="8" pageNumber="139">Female</emphasis>
. Head (
<figureCitation id="52D92A6BEC525065FC36B50CFB8EFAD3" box="[999,1074,1327,1352]" captionStart="Figures 17–20" captionStartId="9.[121,198,1260,1282]" captionTargetBox="[121,1421,144,1244]" captionTargetId="figure-10@9.[121,1421,144,1244]" captionTargetPageId="9" captionText="Figures 17–20. Onthophagus species, head, male (left), and female (right), fresh specimens. (17, 18) O. longipes Paulian; (19, 20) O. nammuldi Matthews." figureDoi="http://doi.org/10.5281/zenodo.13228983" httpUri="https://zenodo.org/record/13228983/files/figure.png" pageId="8" pageNumber="139">Fig. 18</figureCitation>
). Sides more rounded, surface rougher than male, shiny, smooth at base becoming transversely rugose on clypeus, with dense large punctures throughout; median depression of vertex deeper than male, with sides slightly elevated; clypeal suture slightly elevated in frontal portion, sometimes effaced before reaching genal sutures, straight; pronotum slightly narrower than elytra; protibiae short and straight, with broad lateral teeth.
</paragraph>
<paragraph id="CA5D36EEEC525065FC96B609FABAF894" blockId="8.[839,1467,1578,1983]" pageId="8" pageNumber="139">
<emphasis id="F896EAFCEC525065FC96B609FC3BF9D9" bold="true" box="[839,903,1578,1602]" pageId="8" pageNumber="139">Notes</emphasis>
.
<materialsCitation id="7A8A3CB3EC525065FC45B609FABEF894" county="Illawarra" location="Matthews'" municipality="The" pageId="8" pageNumber="139" specimenCount="19" specimenCount-male="19" stateProvince="Victoria" typeStatus="holotype">
Matthews’ redescription was based on
<specimenCount id="DCE4FD67EC525065FAFEB609FA32F9D9" box="[1327,1422,1578,1602]" count="19" pageId="8" pageNumber="139" type="male">19 males</specimenCount>
and females, including the male
<typeStatus id="1559884CEC525065FB50B664FB62F9C4" box="[1153,1246,1607,1631]" pageId="8" pageNumber="139" type="holotype">holotype</typeStatus>
from
<collectingRegion id="0826F80CEC525065FAF4B664FAC2F9C4" box="[1317,1406,1607,1631]" country="Australia" name="Victoria" pageId="8" pageNumber="139">Victoria</collectingRegion>
, with most specimens in
<collectingRegion id="0826F80CEC525065FBC4B646FBD7F9E6" box="[1045,1131,1637,1661]" country="Australia" name="Victoria" pageId="8" pageNumber="139">Victoria</collectingRegion>
and the northernmost from the
<collectingCounty id="233C4E62EC525065FC96B6A1FC16F901" box="[839,938,1666,1690]" pageId="8" pageNumber="139">Illawarra</collectingCounty>
,
<collectingRegion id="0826F80CEC525065FC64B6A0FBCBF901" box="[949,1143,1666,1690]" country="Australia" name="New South Wales" pageId="8" pageNumber="139">New South Wales</collectingRegion>
.
<collectingMunicipality id="2A39AC94EC525065FB50B6A1FB16F901" box="[1153,1194,1666,1690]" pageId="8" pageNumber="139">The</collectingMunicipality>
specimens recorded here extend the distribution of this species north to the Dorrigo Plateau. Our material conforms to
<location id="CF3D6035EC525065FB18B69EFA85F94E" LSID="urn:lsid:plazi:treatment:424B87F8EC5C5065FCA0B4AEFBEEF824:CF3D6035EC525065FB18B69EFA85F94E" box="[1225,1337,1725,1749]" county="Illawarra" municipality="The" name="Matthews'" pageId="8" pageNumber="139" stateProvince="Victoria">Matthews’</location>
description except that the eyes are slightly more variable in size and the striae are usually distinctly punctured
</materialsCitation>
.
</paragraph>
<paragraph id="CA5D36EEEC525065FCB4B736FBEEF824" blockId="8.[839,1467,1578,1983]" pageId="8" pageNumber="139">
All sites recorded here are in closed forest, which appears to be its preferred habitat (
<bibRefCitation id="AE734B1FEC525065FBA4B711FA90F8D1" author="Matthews, E." box="[1141,1324,1842,1866]" pageId="8" pageNumber="139" pagination="3 - 330" refId="ref13173" refString="Matthews, E. 1972. A revision of the scarabaeine dung beetles of Australia. I. tribe Onthophagini. Australian Journal of Zoology. Supplementary Series 9: 3 - 330. https: // doi. org / 10.1071 / AJZS 009" type="book chapter" year="1972">Matthews (1972</bibRefCitation>
; Williams &amp; Williams, 1982), except Barokee Swamp, which is open woodland around a grassy bog at high elevation. The species is rare at its known sites in
<collectingRegion id="0826F80CEC525065FBADB7A8FAFEF839" box="[1148,1346,1930,1954]" country="Australia" name="New South Wales" pageId="8" pageNumber="139">New South Wales</collectingRegion>
, almost all records being singletons.
</paragraph>
</subSubSection>
</treatment>
</document>
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