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<document id="A372E28B16BDE55E4C3A19F52E19D49C" ID-DOI="10.11646/zootaxa.5375.4.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10202772" ID-ZooBank="648DD26D-F271-4BD2-ADFB-0C5C31191FF1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700813825895" checkinUser="plazi" docAuthor="Magnien, Philippe" docDate="2023" docId="294A879EFFDB8E5737A8FE0523EFDAC7" docLanguage="en" docName="zootaxa.5375.4.5.pdf" docOrigin="Zootaxa 5375 (4)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.4.5/52344" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oukhypencha ophthalmica subsp. sumaterensis Magnien 2023, ssp. nov." docType="treatment" docVersion="2" lastPageNumber="541" masterDocId="D573FFE6FFDC8E5F373FFFFB2113DB32" masterDocTitle="New genera, species and subspecies of tessaratomine true bugs (Heteroptera: Tessaratomidae, Tessaratominae)" masterLastPageNumber="549" masterPageNumber="533" pageNumber="540" updateTime="1700814248474" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8083825732A3FC4A3B9FDC8743553B0B">New genera, species and subspecies of tessaratomine true bugs (Heteroptera: Tessaratomidae, Tessaratominae)</mods:title>
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<subSubSection id="E9F96503FFDB8E5837A8FE052299D92B" box="[151,906,509,537]" pageId="7" pageNumber="540" type="nomenclature">
<paragraph id="A15C3688FFDB8E5837A8FE052299D92B" blockId="7.[151,906,509,573]" box="[151,906,509,537]" pageId="7" pageNumber="540">
<heading id="FA1481E4FFDB8E5837A8FE052299D92B" bold="true" box="[151,906,509,537]" fontSize="11" level="1" pageId="7" pageNumber="540" reason="1">
<emphasis id="9397EA9AFFDB8E5837A8FE052299D92B" bold="true" box="[151,906,509,537]" pageId="7" pageNumber="540">
<taxonomicName id="66E34D0BFFDB8E5837A8FE052209D92A" authority="(Stal, 1863)" authorityName="Magnien" authorityYear="2023" baseAuthorityName="Stal" baseAuthorityYear="1863" box="[151,794,509,537]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="540" phylum="Arthropoda" rank="subSpecies" species="ophthalmica" status="subsp. nov." subSpecies="sumaterensis">
<emphasis id="9397EA9AFFDB8E5837A8FE052396D92B" bold="true" box="[151,645,510,537]" italics="true" pageId="7" pageNumber="540">Oukhypencha ophthalmica sumaterensis</emphasis>
(
<bibRefCitation id="C5724B79FFDB8E5835AAFE062201D92A" author="Stal, K." box="[661,786,509,536]" pageId="7" pageNumber="540" pagination="571 - 603" refId="ref10541" refString="Stal, K. (1863) Hemipterorum exoticorum generum et specierum nonnullarum novarum descriptiones. Transactions of the entomological society of London, (3) 1, 571 - 603." type="journal article" year="1863">Stål, 1863</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="88A457E1FFDB8E58341EFE042299D92B" box="[801,906,511,537]" pageId="7" pageNumber="540" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E9F96503FFDB8E5837A8FDD8208BD99B" pageId="7" pageNumber="540" type="description">
<paragraph id="A15C3688FFDB8E5837A8FDD820A9D90F" blockId="7.[151,906,509,573]" box="[151,442,546,573]" pageId="7" pageNumber="540">
(
<figureCitation id="39D82A0DFFDB8E5837A0FDD82018D90F" box="[159,267,546,573]" captionStart="FIGURES 46" captionStartId="6.[151,264,1914,1938]" captionTargetBox="[157,1431,315,1885]" captionTargetId="figure-88@6.[151,1435,309,1891]" captionTargetPageId="6" captionText="FIGURES 46: habitus.—4 Oukhypencha ophthalmica ophthalmica Stål ♀ (holotype); 56: Oukhypencha ophthalmica sumaterensis n. sp.—5: ♂ (holotype)—6: ♀ (paratype). Scale 10 mm" figureDoi="http://doi.org/10.5281/zenodo.10202776" httpUri="https://zenodo.org/record/10202776/files/figure.png" pageId="7" pageNumber="540">Figs. 56</figureCitation>
, 14, 18, 21, 29)
</paragraph>
<paragraph id="A15C3688FFDB8E5837A8FD91208BD99B" blockId="7.[151,1437,618,2013]" pageId="7" pageNumber="540">
<emphasis id="9397EA9AFFDB8E5837A8FD912068D9B6" bold="true" box="[151,379,618,644]" pageId="7" pageNumber="540">Specimens studied.</emphasis>
<materialsCitation id="118B3CD5FFDB8E5836BDFD90208BD99B" collectingDate="1980-11-30" collectionCode="SMNK" collectorName="Ketambe SE &amp; Dr. Roesler" country="Indonesia" pageId="7" pageNumber="540" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Aceh" typeStatus="holotype">
<typeStatus id="7E58882AFFDB8E5836BDFD9020F9D9B7" box="[386,490,619,645]" pageId="7" pageNumber="540" type="holotype">Holotype</typeStatus>
<specimenCount id="B7E5FD01FFDB8E5836CEFD90231BD9B7" box="[497,520,619,645]" count="1" pageId="7" pageNumber="540" type="male"></specimenCount>
,
<date id="D55D1048FFDB8E58352AFD902386D9B7" box="[533,661,619,645]" pageId="7" pageNumber="540" value="1980-11-30">
<collectingDate id="C519E9A0FFDB8E58352AFD902386D9B7" box="[533,661,619,645]" pageId="7" pageNumber="540" value="1980-11-30">30 XI 1980</collectingDate>
</date>
<collectorName id="0C16535EFFDB8E5835A3FD90223AD9B7" box="[668,809,619,645]" pageId="7" pageNumber="540">Ketambe SE</collectorName>
<collectingRegion id="6327F86AFFDB8E583410FD902279D9B7" box="[815,874,619,645]" country="Indonesia" name="Aceh" pageId="7" pageNumber="540">Aceh</collectingRegion>
N
<collectingRegion id="6327F86AFFDB8E5834B3FD9022FAD9B7" box="[908,1001,619,645]" country="Indonesia" name="Sumatera Utara" pageId="7" pageNumber="540">Sumatra</collectingRegion>
<collectingCountry id="D9F47618FFDB8E5834CEFD90254DD9B7" box="[1009,1118,619,645]" name="Indonesia" pageId="7" pageNumber="540">Indonesia</collectingCountry>
<collectorName id="0C16535EFFDB8E58335AFD9025F6D9B7" box="[1125,1253,619,645]" pageId="7" pageNumber="540">Dr. Roesler</collectorName>
leg. (
<collectionCode id="C7F2AE4DFFDB8E58321BFD902464D9B7" box="[1316,1399,619,645]" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" pageId="7" pageNumber="540">SMNK</collectionCode>
)—
<typeStatus id="7E58882AFFDB8E5837A8FD7421E4D99B" box="[151,247,655,681]" pageId="7" pageNumber="540" type="paratype">paratype</typeStatus>
<specimenCount id="B7E5FD01FFDB8E5837C0FD742007D99B" box="[255,276,655,681]" count="1" pageId="7" pageNumber="540" type="female"></specimenCount>
, same data.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E9F96503FFDB8E5837F8FD492076D80B" pageId="7" pageNumber="540" type="diagnosis">
<paragraph id="A15C3688FFDB8E5837F8FD492076D80B" blockId="7.[151,1437,618,2013]" pageId="7" pageNumber="540">
<emphasis id="9397EA9AFFDB8E5837F8FD492029D9FE" bold="true" box="[199,314,690,716]" pageId="7" pageNumber="540">Diagnosis</emphasis>
. Medium sized
<taxonomicName id="66E34D0BFFDB8E5836C8FD482388D9FF" authorityName="Stal" authorityYear="1865" box="[503,667,691,717]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="540" phylum="Arthropoda" rank="subTribe" subTribe="Tessaratomina">Tessaratomina</taxonomicName>
, characterized by the following set of characters: head very small, with eyes and ocellae relatively very large, antennae long, little more than half of habitus length; connexivum bicolourous; pygophore angular, posterior ductus of spermatheca of even width aft of striated dilation which is about 60 % larger.
</paragraph>
</subSubSection>
<subSubSection id="E9F96503FFDB8E5837F8FCB925C8DC7F" pageId="7" pageNumber="540" type="description">
<paragraph id="A15C3688FFDB8E5837F8FCB92312D897" blockId="7.[151,1437,618,2013]" pageId="7" pageNumber="540">
<emphasis id="9397EA9AFFDB8E5837F8FCB9204AD86E" bold="true" box="[199,345,834,860]" pageId="7" pageNumber="540">Description.</emphasis>
Habitus (
<figureCitation id="39D82A0DFFDB8E5836F5FCB8232AD86F" box="[458,569,834,861]" captionStart="FIGURES 46" captionStartId="6.[151,264,1914,1938]" captionTargetBox="[157,1431,315,1885]" captionTargetId="figure-88@6.[151,1435,309,1891]" captionTargetPageId="6" captionText="FIGURES 46: habitus.—4 Oukhypencha ophthalmica ophthalmica Stål ♀ (holotype); 56: Oukhypencha ophthalmica sumaterensis n. sp.—5: ♂ (holotype)—6: ♀ (paratype). Scale 10 mm" figureDoi="http://doi.org/10.5281/zenodo.10202776" httpUri="https://zenodo.org/record/10202776/files/figure.png" pageId="7" pageNumber="540">Figs. 56</figureCitation>
): body of common pentatomoid shape, general hue bronze brownish above, of a lighter brownish for underside, apical half of fourth antennomere, apex of scutellum and forepart of each connexivum segment yellowish;
</paragraph>
<paragraph id="A15C3688FFDB8E5837F8FC5421EFDF23" blockId="7.[151,1437,618,2013]" pageId="7" pageNumber="540">Head. Triangular, mandibular plates surpassing the tylus, entirely punctated save for a small area around ocellae; antennae 4-segmented, long; eyes relatively very large, ocellae less so (ocular index 1.21.3, ocellar index 1.61.9).</paragraph>
<paragraph id="A15C3688FFDB8E5837F8FBE0202FDE3F" blockId="7.[151,1437,618,2013]" pageId="7" pageNumber="540">Thorax. Pronotum sexangular, antero-lateral margins slightly concave, postero-lateral margins very slightly bisinuous, humeral angle rounded; callosities conspicuous, slightly marked. Scutellum triangular, shorter than wide, its apex lanceolate. Metasternum with an elevated plate between coxae, bearing a rounded lanceolate process surpassing fore coxae, somewhat curved upwards at apex; evaporatoria small, in accordance with the description given for the tribe in Kment &amp; Vilimova (2010); legs flavescent, femora with a black ring at apex, length of apical teeth of posterior femora little more than half its diameter, tibiae furrowed outside, with a black line on on either side of furrow.</paragraph>
<paragraph id="A15C3688FFDB8E5837F8FAEC2374DE67" blockId="7.[151,1437,618,2013]" pageId="7" pageNumber="540">Abdomen. Abdominal margins straight; spiracles closer to margin than to middle of the sternites, trichobothria closer to middle, aft of transverse furrow.</paragraph>
<paragraph id="A15C3688FFDB8E5837F8FAA423EEDD63" blockId="7.[151,1437,618,2013]" pageId="7" pageNumber="540">
Male genitalia. Eighth segment visible ventrally, representing a third of visible length of genital segments, pygophore (Fig. 14) simple, its posterior margin concave, with a small central V-notch, angularly connected to lateral margins, parameres (
<figureCitation id="39D82A0DFFDB8E5836F6FA5C230ADEF3" box="[457,537,1447,1473]" captionStart="FIGURES 1518" captionStartId="10.[151,264,1797,1821]" captionTargetBox="[155,1430,348,1770]" captionTargetId="figure-115@10.[151,1435,345,1774]" captionTargetPageId="10" captionText="FIGURES 1518: male genitalia, phallus and paramere—top: phallus (lateral view)—middle: ventral processus (ventral view, right side)—bottom: paramere (dorsal and lateral view, pilosity omitted in lateral view)—15 Hypencha sp. (same specimen as Fig. 10)—16: Oukhypencha clemenceae n. sp.—17: Oukhypencha florenceae n. sp.—18: Oukhypencha ophthalmica sumaterensis n. subsp. Abbreviations: a—apophysis, c—conjunctiva, d—membranous dorsal process, lv—latero-ventral processus, mv—membranous ventral process, ph—phallobase, sd—sclerified dorsal processus, sl—sensorial lobe, v—vesica, vp—ventral processus. All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202784" httpUri="https://zenodo.org/record/10202784/files/figure.png" pageId="7" pageNumber="540">Fig. 18</figureCitation>
) recurved, sensorial lobe marked by a brush of setae, their length about two thirds of paramere width, apophyse broadly rounded. Conjunctiva of the phallus (
<figureCitation id="39D82A0DFFDB8E5834D1FA302552DED7" box="[1006,1089,1483,1509]" captionStart="FIGURES 1518" captionStartId="10.[151,264,1797,1821]" captionTargetBox="[155,1430,348,1770]" captionTargetId="figure-115@10.[151,1435,345,1774]" captionTargetPageId="10" captionText="FIGURES 1518: male genitalia, phallus and paramere—top: phallus (lateral view)—middle: ventral processus (ventral view, right side)—bottom: paramere (dorsal and lateral view, pilosity omitted in lateral view)—15 Hypencha sp. (same specimen as Fig. 10)—16: Oukhypencha clemenceae n. sp.—17: Oukhypencha florenceae n. sp.—18: Oukhypencha ophthalmica sumaterensis n. subsp. Abbreviations: a—apophysis, c—conjunctiva, d—membranous dorsal process, lv—latero-ventral processus, mv—membranous ventral process, ph—phallobase, sd—sclerified dorsal processus, sl—sensorial lobe, v—vesica, vp—ventral processus. All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202784" httpUri="https://zenodo.org/record/10202784/files/figure.png" pageId="7" pageNumber="540">Fig. 18</figureCitation>
) with three pairs of processes, as follow: distally, a pair of membranous processes, latero-dorsal, and two pairs of strongly sclerotized ventral processes, one central, bifid, its biggest arm free, hook-shaped, its smaller one only weekly sclerotized, free only at apex, the other latero-ventral, hook-shaped, very large.
</paragraph>
<paragraph id="A15C3688FFDB8E5837F8F9A02409DDD3" blockId="7.[151,1437,618,2013]" pageId="7" pageNumber="540">
Female genitalia (
<figureCitation id="39D82A0DFFDB8E5836A5F9A020E0DD47" box="[410,499,1627,1653]" captionStart="FIGURES 1923" captionStartId="11.[151,264,1312,1336]" captionTargetBox="[155,1431,183,1289]" captionTargetId="figure-15@11.[151,1435,181,1289]" captionTargetPageId="11" captionText="FIGURES 1923: Oukhypencha, female genitalia, evaporatoria.—1922: female genitalia.—23: evaporatoria.—19, 23: Oukhypencha clemenceae n. sp.—20: Oukhypencha ophthalmica ophthalmica—21: Oukhypencha ophthalmica sumaterensis— 22: Oukhypencha stali. Abbreviations: ad—anterior duct, ev—evaporatoria field, ip—intermediate part, o—ostiole, pd—posterior duct, pw—posterior wall of vagina, r—receptacle, rs—ring sclerite, sd—striated dilation All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202786" httpUri="https://zenodo.org/record/10202786/files/figure.png" pageId="7" pageNumber="540">Fig. 21</figureCitation>
). Ring sclerites existing; spermatheca of common
<typeStatus id="7E58882AFFDB8E583373F9A0256FDD47" box="[1100,1148,1627,1653]" pageId="7" pageNumber="540">type</typeStatus>
in
<taxonomicName id="66E34D0BFFDB8E583396F9A0245ADD47" authorityName="Stal" authorityYear="1865" box="[1193,1353,1627,1653]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="540" phylum="Arthropoda" rank="tribe" tribe="Tessaratomini">Tessaratomini</taxonomicName>
; apical receptacle (distal bulb) spherical with a basal “neck”, intermediate part (pump apparatus) with two well developed flanges; spermathecal duct bipartite, anterior duct slender, posterior duct much larger, with an anterior striated dilation about 60 % wider than rest, rear part about two times longer, of even diameter on all its length.
</paragraph>
<paragraph id="A15C3688FFDB8E5837F8F9102037DC1B" blockId="7.[151,1437,618,2013]" pageId="7" pageNumber="540">
Measurements (mm). Male (
<typeStatus id="7E58882AFFDB8E583539F9102379DC37" box="[518,618,1771,1797]" pageId="7" pageNumber="540" type="holotype">holotype</typeStatus>
)—length 19.8; width 10.7; antennal articles length (A1: 10, A2: 3.6, A3: 3.6, A4: 4.0)
</paragraph>
<paragraph id="A15C3688FFDB8E5837F8F8C825C8DC7F" blockId="7.[151,1437,618,2013]" box="[199,1243,1842,1869]" pageId="7" pageNumber="540">Female—length 23,4; width 10.7; antennal articles length (A1: 1.0, A2: 3.5, A3: 3.7, A4: 4.1)</paragraph>
</subSubSection>
<subSubSection id="E9F96503FFDB8E5837F8F8AC2513DC43" box="[199,1024,1879,1905]" pageId="7" pageNumber="540" type="etymology">
<paragraph id="A15C3688FFDB8E5837F8F8AC2513DC43" blockId="7.[151,1437,618,2013]" box="[199,1024,1879,1905]" pageId="7" pageNumber="540">
<emphasis id="9397EA9AFFDB8E5837F8F8AC205BDC43" bold="true" box="[199,328,1879,1905]" pageId="7" pageNumber="540">Etymology</emphasis>
. The name of this subspecies refers to its geographical origin.
</paragraph>
</subSubSection>
<subSubSection id="E9F96503FFDB8E5837F8F8812340DCA7" box="[199,595,1914,1941]" pageId="7" pageNumber="540" type="distribution">
<paragraph id="A15C3688FFDB8E5837F8F8812340DCA7" blockId="7.[151,1437,618,2013]" box="[199,595,1914,1941]" pageId="7" pageNumber="540">
<emphasis id="9397EA9AFFDB8E5837F8F881204ADCA6" bold="true" box="[199,345,1914,1940]" pageId="7" pageNumber="540">Distribution</emphasis>
.
<collectingCountry id="D9F47618FFDB8E583658F88020C7DCA7" box="[359,468,1915,1941]" name="Indonesia" pageId="7" pageNumber="540">Indonesia</collectingCountry>
(
<collectingRegion id="6327F86AFFDB8E5836DCF8802354DCA7" box="[483,583,1915,1941]" country="Indonesia" name="Sumatera Utara" pageId="7" pageNumber="540">Sumatra</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="E9F96503FFDB8E5737F8F86523EFDAC7" lastPageId="8" lastPageNumber="541" pageId="7" pageNumber="540" type="discussion">
<paragraph id="A15C3688FFDB8E5737F8F8652288DBE7" blockId="7.[151,1437,618,2013]" lastBlockId="8.[151,1436,151,501]" lastPageId="8" lastPageNumber="541" pageId="7" pageNumber="540">
<emphasis id="9397EA9AFFDB8E5837F8F865205FDC8A" bold="true" box="[199,332,1950,1976]" pageId="7" pageNumber="540">Discussion.</emphasis>
This taxon is represented by only
<specimenCount id="B7E5FD01FFDB8E5835E8F8642292DC8B" box="[727,897,1951,1977]" count="2" pageId="7" pageNumber="540" type="generic">two specimens</specimenCount>
, collected together in
<collectingRegion id="6327F86AFFDB8E583343F86425CEDC8B" box="[1148,1245,1951,1977]" country="Indonesia" name="Sumatera Utara" pageId="7" pageNumber="540">Sumatra</collectingRegion>
. It is to be noted that the male
<typeStatus id="7E58882AFFDB8E583606F8382088DCEF" box="[313,411,1987,2013]" pageId="7" pageNumber="540" type="holotype">holotype</typeStatus>
is slightly teratologic, with a malformation of left outer part of the abdomen, including pygophore which is somewhat assymetric, its left side smaller than the other. It does not seem to have affected the genitalia, parameres and phallus being almost perfectly symmetrical.
</paragraph>
<paragraph id="A15C3688FFD48E5737F8FF24201FDA73" blockId="8.[151,1436,151,501]" pageId="8" pageNumber="541">
As expressed here above, it was chosen to represent Ståls species because of its female specimens being closest to Ståls type. This can only be verified when males from
<collectingCountry id="D9F47618FFD48E57341CFEF82294DA2F" box="[803,903,259,285]" name="Thailand" pageId="8" pageNumber="541">Thailand</collectingCountry>
will be available to compare to the
<typeStatus id="7E58882AFFD48E573224FEF8246EDA2F" box="[1307,1405,259,285]" pageId="8" pageNumber="541" type="holotype">holotype</typeStatus>
of this taxon.
</paragraph>
<paragraph id="A15C3688FFD48E5737F8FEB023EFDAC7" blockId="8.[151,1436,151,501]" pageId="8" pageNumber="541">
The female
<typeStatus id="7E58882AFFD48E57366EFEB020A2DA57" box="[337,433,331,357]" pageId="8" pageNumber="541" type="paratype">paratype</typeStatus>
is very similar to
<taxonomicName id="66E34D0BFFD48E5735BAFEB02225DA57" authorityName="Stal" authorityYear="1863" box="[645,822,331,357]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="541" phylum="Arthropoda" rank="species" sensu="stricto" species="ophthalmica">
<emphasis id="9397EA9AFFD48E5735BAFEB02225DA57" box="[645,822,331,357]" italics="true" pageId="8" pageNumber="541">O. ophthalmica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88A457E1FFD48E573401FEB0226EDA57" box="[830,893,331,357]" pageId="8" pageNumber="541" sensu="stricto">
<emphasis id="9397EA9AFFD48E573401FEB02264DA56" box="[830,887,331,356]" italics="true" pageId="8" pageNumber="541">s. str</emphasis>
.
</taxonomicNameLabel>
, but not identical. Their habitus are very close, but O.
<emphasis id="9397EA9AFFD48E5737D5FE942068DABA" box="[234,379,367,392]" italics="true" pageId="8" pageNumber="541">sumaterensis</emphasis>
has anterior margins of pronotum somewhat concave, whereas
<taxonomicName id="66E34D0BFFD48E573361FE94241CDABB" authorityName="Stal" authorityYear="1863" box="[1118,1295,367,393]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="548" phylum="Arthropoda" rank="species" species="ophthalmica">
O.
<emphasis id="9397EA9AFFD48E5733BCFE94241CDABB" box="[1155,1295,367,393]" italics="true" pageId="8" pageNumber="541">ophthalmica</emphasis>
</taxonomicName>
has straight ones. Spermatheca are also similar, but O.
<emphasis id="9397EA9AFFD48E57354DFE682210DA9E" box="[626,771,403,428]" italics="true" pageId="8" pageNumber="541">sumaterensis</emphasis>
has receptacle neck and anterior ductus shorter and striated dilation much bigger than
<taxonomicName id="66E34D0BFFD48E5736F5FE4C236CDAE3" authorityName="Stal" authorityYear="1863" box="[458,639,439,465]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="548" phylum="Arthropoda" rank="species" species="ophthalmica">
O.
<emphasis id="9397EA9AFFD48E5736D0FE4C236CDAE3" box="[495,639,439,465]" italics="true" pageId="8" pageNumber="541">ophthalmica</emphasis>
</taxonomicName>
<emphasis id="9397EA9AFFD48E573540FE4C2390DAE3" box="[639,643,439,465]" italics="true" pageId="8" pageNumber="541">.</emphasis>
Added to the impossibility of comparing male features, it led to the decision of choosing a subspecies status for this taxon.
</paragraph>
</subSubSection>
</treatment>
</document>