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<document id="86FCE5807B1A3996674C2600C9F94244" ID-DOI="10.11646/zootaxa.5375.4.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10202772" ID-ZooBank="648DD26D-F271-4BD2-ADFB-0C5C31191FF1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1700813825895" checkinUser="plazi" docAuthor="Magnien, Philippe" docDate="2023" docId="294A879EFFD28E5037A8F974248ED8F7" docLanguage="en" docName="zootaxa.5375.4.5.pdf" docOrigin="Zootaxa 5375 (4)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.4.5/52344" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tessaratomina Stal 1865" docType="key" docVersion="5" lastPageNumber="548" masterDocId="D573FFE6FFDC8E5F373FFFFB2113DB32" masterDocTitle="New genera, species and subspecies of tessaratomine true bugs (Heteroptera: Tessaratomidae, Tessaratominae)" masterLastPageNumber="549" masterPageNumber="533" pageNumber="547" updateTime="1701864501225" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="1A744C892F7AF1B897844E49343BC91F">New genera, species and subspecies of tessaratomine true bugs (Heteroptera: Tessaratomidae, Tessaratominae)</mods:title>
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<paragraph id="A15C3688FFD28E5137A8F97423D8DD9B" blockId="14.[151,715,1679,1705]" box="[151,715,1679,1705]" pageId="14" pageNumber="547">
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<emphasis id="9397EA9AFFD28E5137A8F97423D8DD9B" bold="true" box="[151,715,1679,1705]" pageId="14" pageNumber="547">
Key to genera of the sub-tribe
<taxonomicName id="66E34D0BFFD28E51352BF97423D8DD9B" authorityName="Stal" authorityYear="1865" box="[532,715,1679,1705]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="547" phylum="Arthropoda" rank="subTribe" subTribe="Tessaratomina">Tessaratomina</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="E9F96503FFD28E5137A8F92C2313DCFB" pageId="14" pageNumber="547" type="discussion">
<paragraph id="A15C3688FFD28E5137A8F92C2313DCFB" blockId="14.[151,1437,1751,1993]" pageId="14" pageNumber="547">
The present key is an adaptation of the one given by
<bibRefCitation id="C5724B79FFD28E5135CFF9232515DDC3" author="Kumar R. &amp; Ghauri M. S. K." box="[752,1030,1751,1778]" pageId="14" pageNumber="547" pagination="1 - 32" refId="ref10342" refString="Kumar R. &amp; Ghauri M. S. K. (1970) Morphology and relationships of the Pentatomoidea (Heteroptera) 2 - World genera of Tessaratomini (Tessaratomidae), Deutsche Entomologische Zeitschrift, 17 (I - III), 1 - 32. https: // doi. org / 10.1002 / mmnd. 19700170102" type="journal article" year="1970">Kumar &amp; Ghauri (1970)</bibRefCitation>
, together with a complete diagnosis of
<taxonomicName id="66E34D0BFFD28E513788F900204FDC27" authorityName="Stal" authorityYear="1865" box="[183,348,1787,1813]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="547" phylum="Arthropoda" rank="subTribe" subTribe="Tessaratomina">Tessaratomina</taxonomicName>
. Apart from the inclusion of the two new genera, the modifications are as follows: suppression of
<taxonomicName id="66E34D0BFFD28E513789F8DB2001DC08" authorityName="Stal" authorityYear="1863" box="[182,274,1824,1850]" class="Insecta" family="Tessaratomidae" genus="Amissus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD28E513789F8DB2001DC08" box="[182,274,1824,1850]" italics="true" pageId="14" pageNumber="547">Amissus</emphasis>
</taxonomicName>
, as being not a true
<taxonomicName id="66E34D0BFFD28E5136C8F8E4224CDC0B" authority="(Magnien, 2018)" baseAuthorityName="Magnien" baseAuthorityYear="2018" box="[503,863,1823,1849]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="547" phylum="Arthropoda" rank="subTribe" subTribe="Tessaratomina">
Tessaratomina (
<bibRefCitation id="C5724B79FFD28E513597F8E42244DC0B" author="Magnien Ph." box="[680,855,1823,1849]" pageId="14" pageNumber="547" pagination="169 - 178" refId="ref10402" refString="Magnien Ph. (2018) Revision du genre Amissus Stal, 1863 (Heteroptera, Tessaratomidae). Bulletin de la Societe Entomologique de France, 123 (2), 169 - 178. https: // doi. org / 10.32475 / bsef _ 2012" type="journal article" year="2018">Magnien, 2018</bibRefCitation>
)
</taxonomicName>
, addition of
<taxonomicName id="66E34D0BFFD28E5134D1F8E42524DC0B" authorityName="Walker" authorityYear="1868" box="[1006,1079,1823,1849]" class="Insecta" family="Tessaratomidae" genus="Enada" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD28E5134D1F8E42524DC0B" box="[1006,1079,1823,1849]" italics="true" pageId="14" pageNumber="547">Enada</emphasis>
</taxonomicName>
, which has been overlooked by them, correction of concave/convex inversion in their original key, change of the character used to isolate
<taxonomicName id="66E34D0BFFD28E51327BF8B8248EDC6F" box="[1348,1437,1859,1885]" class="Insecta" family="Tessaratomidae" genus="Siphnus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD28E51327BF8B8248EDC6F" box="[1348,1437,1859,1885]" italics="true" pageId="14" pageNumber="547">Siphnus</emphasis>
</taxonomicName>
(head instead of pronotum) and addition of a secondary character to insure the correct inclusion of
<taxonomicName id="66E34D0BFFD28E5133C0F89C2095DC97" authority="Martin, 1902" authorityName="Martin" authorityYear="1902" class="Insecta" family="Tessaratomidae" genus="Embolosterna" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="547" phylum="Arthropoda" rank="species" species="vaccca">
<emphasis id="9397EA9AFFD28E5133C0F89C21F6DC97" italics="true" pageId="14" pageNumber="547">Embolosterna vaccca</emphasis>
Martin, 1902
</taxonomicName>
. For illustration of this key, readers may access pictures of all
<taxonomicName id="66E34D0BFFD28E513363F87025EEDC97" baseAuthorityName="Magnien" baseAuthorityYear="2018" box="[1116,1277,1931,1957]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="547" phylum="Arthropoda" rank="subTribe" subTribe="Tessaratomina">Tessaratomina</taxonomicName>
genera in the online catalog (
<bibRefCitation id="C5724B79FFD28E51367BF85420E7DCFB" author="Magnien, Ph." box="[324,500,1967,1993]" pageId="14" pageNumber="547" refId="ref10450" refString="Magnien, Ph. (2023) Illustrated catalog of TESSARATOMIDAE. Available from: http: // www. heteroptera. fr / tessite / (accessed 17 January 2023)" type="url" year="2023">Magnien, 2023</bibRefCitation>
).
</paragraph>
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<keyLead id="1A1295BDFFD38E5037A8FF63248FDBF8" pageId="15" pageNumber="548">1 Posterior margin of pronotum straight or concave, if slightly convex then humeral processes not spined; humeral processes normally blunt, if sharply pointed then specimens always chocolate in colour...................................... 2</keyLead>
</paragraph>
<paragraph id="A15C3688FFD38E5037A8FF28248FDA10" blockId="15.[151,1437,151,965]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FF28248FDA10" pageId="15" pageNumber="548"> Posterior margin of pronotum convex, exceptionally with a small median concavity; if posterior margin straight or slightly concave medially then humeral processes produced into long sharp pointed spines; spined specimens pale green or hazy green but never chocolate in colour............................................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A172E2DFFD38E5037A8FED1248FDA49" pageId="15" pageNumber="548">
<paragraph id="A15C3688FFD38E5037A8FED1248FDA0D" blockId="15.[151,1437,151,965]" box="[151,1436,298,320]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FED1248FDA0D" box="[151,1436,298,320]" pageId="15" pageNumber="548">2 Free projecting part of sternal processus not or only slightly flattened laterally, generally round, never absent............ 3</keyLead>
</paragraph>
<paragraph id="A15C3688FFD38E5037A8FEB3248FDA49" blockId="15.[151,1437,151,965]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FEB3248FDA49" pageId="15" pageNumber="548"> Free projecting part of sternal processus greatly flattened laterally, if free projecting part absent then raised portion of processus in contact with mesosternum always flattened laterally........................................................ 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A172E2DFFD38E5037A8FE79248FDA87" pageId="15" pageNumber="548">
<paragraph id="A15C3688FFD38E5037A8FE79248FDAA5" blockId="15.[151,1437,151,965]" box="[151,1436,386,408]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FE79248FDAA5" box="[151,1436,386,408]" pageId="15" pageNumber="548">3 Antennae length half or more than half of habitus length...................................................... 4</keyLead>
</paragraph>
<paragraph id="A15C3688FFD38E5037A8FE5B248FDA87" blockId="15.[151,1437,151,965]" box="[151,1436,415,437]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FE5B248FDA87" box="[151,1436,415,437]" pageId="15" pageNumber="548"> Antennae length less than half of habitus length............................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A172E2DFFD38E5037A8FE46248FD918" pageId="15" pageNumber="548">
<paragraph id="A15C3688FFD38E5037A8FE46248FDADD" blockId="15.[151,1437,151,965]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FE46248FDADD" pageId="15" pageNumber="548">
4 Underside dark brown,
<specimenCount id="B7E5FD01FFD38E5036F5FE4620CFDAE0" box="[458,476,445,466]" count="1" pageId="15" pageNumber="548" type="male"></specimenCount>
posterior angles of pygophore strongly produced,
<specimenCount id="B7E5FD01FFD38E5034EFFE4622F1DAE0" box="[976,994,445,466]" count="1" pageId="15" pageNumber="548" type="female"></specimenCount>
distal spermathecal duct deprived of striated dilation..............................................................................
<taxonomicName id="66E34D0BFFD38E5033DFFE20244EDAC2" authorityName="Magnien" authorityYear="2023" box="[1248,1373,475,496]" class="Insecta" family="Tessaratomidae" genus="Homoiacoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E5033DFFE20244EDAC2" box="[1248,1373,475,496]" italics="true" pageId="15" pageNumber="548">Homoiacoris</emphasis>
</taxonomicName>
<emphasis id="9397EA9AFFD38E50325BFE21248FDADD" bold="true" box="[1380,1436,474,495]" pageId="15" pageNumber="548">n. gn.</emphasis>
</keyLead>
</paragraph>
<paragraph id="A15C3688FFD38E5037A8FE03248FD918" blockId="15.[151,1437,151,965]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FE03248FD918" pageId="15" pageNumber="548">
Underside light brown,
<specimenCount id="B7E5FD01FFD38E5036F5FE0320CFD93F" box="[458,476,504,525]" count="1" pageId="15" pageNumber="548" type="male"></specimenCount>
posterior angles of pygophore not produced,
<specimenCount id="B7E5FD01FFD38E5034A4FE0322BED93F" box="[923,941,504,525]" count="1" pageId="15" pageNumber="548" type="female"></specimenCount>
distal spermathecal duct fitted with a more or less swollen longitudinally striated dilation.....................................................
<taxonomicName id="66E34D0BFFD38E5033E5FDEE244DD918" authorityName="Magnien" authorityYear="2023" box="[1242,1374,533,554]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E5033E5FDEE244DD918" box="[1242,1374,533,554]" italics="true" pageId="15" pageNumber="548">Oukhypencha</emphasis>
</taxonomicName>
<emphasis id="9397EA9AFFD38E50325BFDEE248FD918" bold="true" box="[1380,1436,533,554]" pageId="15" pageNumber="548">n. gn.</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A172E2DFFD38E5037A8FDC9248FD957" pageId="15" pageNumber="548">
<paragraph id="A15C3688FFD38E5037A8FDC9248FD97A" blockId="15.[151,1437,151,965]" box="[151,1436,562,584]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FDC9248FD97A" box="[151,1436,562,584]" pageId="15" pageNumber="548">
5 Head width great, at least one third of pronotum width...............................................
<taxonomicName id="66E34D0BFFD38E50321AFDC9248FD97A" ID-CoL="7HMG" authority="Stal" authorityName="Stal" box="[1317,1436,562,584]" class="Insecta" family="Tessaratomidae" genus="Siphnus" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E50321AFDC92462D975" box="[1317,1393,562,583]" italics="true" pageId="15" pageNumber="548">Siphnus</emphasis>
Stål
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A15C3688FFD38E5037A8FDAB248FD957" blockId="15.[151,1437,151,965]" box="[151,1436,591,613]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FDAB248FD957" box="[151,1436,591,613]" pageId="15" pageNumber="548">
Head width smaller, about one quarter of pronotum width.........................................
<taxonomicName id="66E34D0BFFD38E5033C2FDB4248FD957" ID-CoL="MXD" authority="Stal" authorityName="Stal" box="[1277,1436,591,613]" class="Insecta" family="Tessaratomidae" genus="Acidosterna" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E5033C2FDB42462D956" box="[1277,1393,591,612]" italics="true" pageId="15" pageNumber="548">Acidosterna</emphasis>
Stål
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A172E2DFFD38E5037A8FD96248FD98F" pageId="15" pageNumber="548">
<paragraph id="A15C3688FFD38E5037A8FD96248ED992" blockId="15.[151,1437,151,965]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FD96248ED992" pageId="15" pageNumber="548">
6 Length of free part of sternal processus equal to, greater than or slightly less than length in contact with sterna.............................................................................
<taxonomicName id="66E34D0BFFD38E5033B6FD71248ED992" ID-CoL="52CC" authority="Amyot &amp; Serville" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[1161,1437,650,672]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E5033B6FD7125F9D9AD" box="[1161,1258,650,671]" italics="true" pageId="15" pageNumber="548">Hypencha</emphasis>
Amyot &amp; Serville
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A15C3688FFD38E5037A8FD53248FD98F" blockId="15.[151,1437,151,965]" box="[151,1436,679,701]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FD53248FD98F" box="[151,1436,679,701]" pageId="15" pageNumber="548"> Free part of sternal processus reduced, always less than length in contact......................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A172E2DFFD38E5037A8FD3E248FD800" pageId="15" pageNumber="548">
<paragraph id="A15C3688FFD38E5037A8FD3E248ED9CA" blockId="15.[151,1437,151,965]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FD3E248ED9CA" pageId="15" pageNumber="548">
7 Postero-lateral margins of segment 7 produced backwards into long flat and apically furcate processes; scutellum long, extending well onto membrane of forewing and gradually tapering apically..........................
<taxonomicName id="66E34D0BFFD38E5033B3FD18248ED9CA" ID-CoL="5VH6" authority="Amyot &amp; Serville" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[1164,1437,738,760]" class="Insecta" family="Tessaratomidae" genus="Mucanum" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E5033B3FD1825FFD9CA" box="[1164,1260,739,760]" italics="true" pageId="15" pageNumber="548">Mucanum</emphasis>
Amyot &amp; Serville
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A15C3688FFD38E5037A8FCFB248FD800" blockId="15.[151,1437,151,965]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FCFB248FD800" pageId="15" pageNumber="548"> Postero-lateral margins of segment 7 not or slightly produced, the latter never furcate apically, scutellum not or slightly extending onto membrane, never gradually tapering posteriorly................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A172E2DFFD38E5037A8FCC1248FD85F" pageId="15" pageNumber="548">
<paragraph id="A15C3688FFD38E5037A8FCC1248FD862" blockId="15.[151,1437,151,965]" box="[151,1436,826,848]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FCC1248FD862" box="[151,1436,826,848]" pageId="15" pageNumber="548">
8 Humeral processus conspicuous, if not then apex of scutellum with a large yellow spot................
<taxonomicName id="66E34D0BFFD38E5033D5FCC1248FD862" ID-CoL="6345D" authority="Stal" authorityName="Stal" box="[1258,1436,826,848]" class="Insecta" family="Tessaratomidae" genus="Embolosterna" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E5033D5FCC12462D87D" box="[1258,1393,826,847]" italics="true" pageId="15" pageNumber="548">Embolosterna</emphasis>
Stål
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A15C3688FFD38E5037A8FCA3248FD85F" blockId="15.[151,1437,151,965]" box="[151,1436,855,877]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FCA3248FD85F" box="[151,1436,855,877]" pageId="15" pageNumber="548">
Humerus rounded, not produced, apex of scutellum concolorous...............................
<taxonomicName id="66E34D0BFFD38E5033F7FCA3248FD85F" ID-CoL="63V9C" authority="Berthold" authorityName="Berthold" authorityYear="1827" box="[1224,1436,855,877]" class="Insecta" family="Tessaratomidae" genus="Tessaratoma" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E5033F7FCA32451D85F" box="[1224,1346,856,877]" italics="true" pageId="15" pageNumber="548">Tessaratoma</emphasis>
Berthold
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A172E2DFFD38E5037A8FC8E248ED8F7" pageId="15" pageNumber="548">
<paragraph id="A15C3688FFD38E5037A8FC8E248FD89A" blockId="15.[151,1437,151,965]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FC8E248FD89A" pageId="15" pageNumber="548">
9 Width of pronotum at least equal and often much greater than that of abdomen, taken at level of 4th abdominal segment...........................................................................................
<taxonomicName id="66E34D0BFFD38E5033D1FC69248FD89A" ID-CoL="75V2" authority="Dallas" authorityName="Dallas" authorityYear="1851" box="[1262,1436,914,936]" class="Insecta" family="Tessaratomidae" genus="Pygoplatys" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E5033D1FC69244AD895" box="[1262,1369,914,935]" italics="true" pageId="15" pageNumber="548">Pygoplatys</emphasis>
Dallas
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A15C3688FFD38E5037A8FC4B248ED8F7" blockId="15.[151,1437,151,965]" box="[151,1437,943,965]" pageId="15" pageNumber="548">
<keyLead id="1A1295BDFFD38E5037A8FC4B248ED8F7" box="[151,1437,943,965]" pageId="15" pageNumber="548">
Width of pronotum at most 90 % of that of abdomen...............................................
<taxonomicName id="66E34D0BFFD38E50322CFC54248ED8F7" ID-CoL="6346T" authority="Walker" authorityName="Walker" authorityYear="1868" box="[1299,1437,943,965]" class="Insecta" family="Tessaratomidae" genus="Enada" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="9397EA9AFFD38E50322CFC542441D8F6" box="[1299,1362,943,964]" italics="true" pageId="15" pageNumber="548">Enada</emphasis>
Walker
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>