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<document id="A7C5A884AD8AB7395299E74114FCCB94" ID-CLB-Dataset="37346" ID-DOI="10.5281/zenodo.270103" ID-GBIF-Dataset="9bcf07e6-a6c2-4eb6-a757-5383dfccb217" ID-ISSN="0374-1036" ID-Zenodo-Dep="270103" ID-ZooBank="urn:lsid:zoobank.org:pub:5354869F-35B0-4035-85A0-D3C7AE4514EF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484064574471" checkinUser="plazi" docAuthor="Ryo Ogawa, Ivan Löbl &amp; Kaoru Maeto" docDate="2016" docId="2924DE4E811CFF8B4C35FA99F05BFD63" docLanguage="en" docName="ActaEntMusNatPra.56.1.195-201.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Cyparium celebense Ogawa &amp; Lobl, sp. nov." docType="treatment" docVersion="8" lastPageNumber="200" masterDocId="D51DA636811DFF8E4D51FFFCF213FF90" masterDocTitle="A new species of the genus Cyparium from northern Sulawesi, Indonesia (Coleoptera: Staphylinidae: Scaphidiinae)" masterLastPageNumber="201" masterPageNumber="195" pageNumber="196" updateTime="1698435432394" updateUser="ExternalLinkService">
<mods:mods id="17508D0FAD049A243E411F84C095765A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="56F0E798C5EB1E92B641C6FB212FF457">
<mods:title id="9A4054B2B1CF70888BB0EE8CF55F2BB5">A new species of the genus Cyparium from northern Sulawesi, Indonesia (Coleoptera: Staphylinidae: Scaphidiinae)</mods:title>
</mods:titleInfo>
<mods:name id="B33DCFA1D0BD7290E1A0B4B74B6FEC98" type="personal">
<mods:role id="3BCE4CBEF085F19796BA016175CBB643">
<mods:roleTerm id="151358AC7BBAFAA315B7D0AC1128B307">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BEBF17AD6B0BA794C01192C05D0EFBEE">Ryo Ogawa</mods:namePart>
</mods:name>
<mods:name id="5CE65F9061A249B17E7A39BED9B29A6D" type="personal">
<mods:role id="DEF1AE19F39051BB22F0CA3AD8288B8E">
<mods:roleTerm id="9EB26CE2338EB6B33075A5EB6C6F3D44">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FABB4AF14B8EE5A84CC8537FFADA9BF3">Ivan Löbl</mods:namePart>
</mods:name>
<mods:name id="E37E73AD11E0762318B039B26CD0CB61" type="personal">
<mods:role id="0ED011DE42F0F8126BA29F20F2F81379">
<mods:roleTerm id="89DBDCD245DA3143F7E1436E1D385EBB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD9E64EB285909D82C5811BFFB25E532">Kaoru Maeto</mods:namePart>
</mods:name>
<mods:typeOfResource id="0E707423CCD53837C1DABF9EE9CA443F">text</mods:typeOfResource>
<mods:relatedItem id="DB411E4F1B8E7668C17EA6BAADEB50AB" type="host">
<mods:titleInfo id="45810C2B6624ABEEFA3DCB2F7C1965BA">
<mods:title id="F6E8237713090071CC1ECB419A45D13B">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="4E83EDB314903CE5E77768A858B641D3">
<mods:date id="1ACC0641A5AB39F11F3176EA3A79C9EE">2016</mods:date>
<mods:detail id="CAAC67F544865BBFB4FA3D5A47B01F24" type="volume">
<mods:number id="9BA4A967E998449196DFC6B4AE6B3AFE">56</mods:number>
</mods:detail>
<mods:detail id="1F8D168207D9F8A7DDBE2B7A08E4675A" type="issue">
<mods:number id="909D1D26F92BA75C7C7F9F7BDA420DE6">1</mods:number>
</mods:detail>
<mods:extent id="CEABB1E118342E937808EE3315465772" unit="page">
<mods:start id="8CDA290EEF2136C3305F1CE6CA81C505">195</mods:start>
<mods:end id="2B237FDD912D77503B6E1CAC6DFD7204">201</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F37438146AD24BF6124ED1C77E739BD1">journal article</mods:classification>
<mods:identifier id="F649B88E92F8456B5A283902BA6F60A8" type="CLB-Dataset">37346</mods:identifier>
<mods:identifier id="8F009BCAEDA1C897E1EE1EEC7A0D64AC" type="DOI">10.5281/zenodo.270103</mods:identifier>
<mods:identifier id="AC51750A3EC620E8869FB03236283169" type="GBIF-Dataset">9bcf07e6-a6c2-4eb6-a757-5383dfccb217</mods:identifier>
<mods:identifier id="55995BD3A284A3BA0FC02567C024B13B" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="89FAFCD7BDBCA3CDBE40EFFD2FAD1D52" type="Zenodo-Dep">270103</mods:identifier>
<mods:identifier id="826C068D60B8FD3DB80019491613125E" type="ZooBank">urn:lsid:zoobank.org:pub:5354869F-35B0-4035-85A0-D3C7AE4514EF</mods:identifier>
</mods:mods>
<treatment id="2924DE4E811CFF8B4C35FA99F05BFD63" ID-DOI="http://doi.org/10.5281/zenodo.5188213" ID-GBIF-Taxon="126104327" ID-Zenodo-Dep="5188213" LSID="urn:lsid:plazi:treatment:2924DE4E811CFF8B4C35FA99F05BFD63" httpUri="http://treatment.plazi.org/id/2924DE4E811CFF8B4C35FA99F05BFD63" lastPageId="5" lastPageNumber="200" pageId="1" pageNumber="196">
<subSubSection id="E9973CD3811CFF8F4C35FA99F15DFAEE" box="[356,846,1381,1406]" pageId="1" pageNumber="196" type="nomenclature">
<paragraph id="A1326F58811CFF8F4C35FA99F15DFAEE" blockId="1.[356,846,1381,1431]" box="[356,846,1381,1406]" pageId="1" pageNumber="196">
<heading id="FA7AD834811CFF8F4C35FA99F15DFAEE" bold="true" box="[356,846,1381,1406]" centered="true" fontSize="10" level="3" pageId="1" pageNumber="196" reason="0">
<emphasis id="93F9B34A811CFF8F4C35FA99F15DFAEE" bold="true" box="[356,846,1381,1406]" pageId="1" pageNumber="196">
<taxonomicName id="668D14DB811CFF8F4C35FA99F0E2FAEE" ID-CoL="33BBX" authority="Ogawa &amp; Lobl" authorityName="Ogawa &amp; Lobl" box="[356,753,1381,1406]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="celebense" status="sp. nov.">
<emphasis id="93F9B34A811CFF8F4C35FA99F051FAEC" bold="true" box="[356,578,1381,1404]" italics="true" pageId="1" pageNumber="196">Cyparium celebense</emphasis>
Ogawa &amp; Löbl
</taxonomicName>
<taxonomicNameLabel id="88CA0E31811CFF8F4FA9FA9AF15DFAEE" box="[760,846,1382,1406]" pageId="1" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E9973CD3811CFF8F4F55FA78F0BEFA07" box="[516,685,1412,1431]" pageId="1" pageNumber="196" type="description">
<paragraph id="A1326F58811CFF8F4F55FA78F0BEFA07" blockId="1.[356,846,1381,1431]" box="[516,685,1412,1431]" pageId="1" pageNumber="196">
(
<figureCitation id="39B673DD811CFF8F4F5BFA78F02EFA07" box="[522,573,1412,1431]" captionStart="Fig. 1" captionStartId="2.[151,182,1260,1279]" captionTargetBox="[157,1115,227,1243]" captionTargetId="figure@2.[151,1126,219,1251]" captionTargetPageId="2" captionText="Fig. 1. Cyparium celebense sp. nov. A – habitus of male (left, dorsal view; right, lateral view); B – male genitalia (left, dorsal view; right, lateral view); C – antenna of female." httpUri="https://zenodo.org/record/270104/files/figure.png" pageId="1" pageNumber="196">Figs 1</figureCitation>
A–C,
<figureCitation id="39B673DD811CFF8F4F22FA78F06FFA07" box="[627,636,1412,1431]" captionStart="Fig. 3" captionStartId="3.[113,144,1284,1303]" captionTargetBox="[113,1481,476,1259]" captionTargetId="figure@3.[113,1481,476,1259]" captionTargetPageId="3" captionText="Fig. 3. Habitus and labels of four species of the genus Cyparium from Indonesia and Malaysia. A – C. bowringi Achard, 1922 (lectotype); B – C. javanum Löbl, 1990 (paratype); C – C. punctatum Pic, 1916 (holotype); D – C. variegatum Achard, 1920 (holotype)." httpUri="https://zenodo.org/record/270105/files/figure.png" pageId="1" pageNumber="196">3</figureCitation>
A–C)
</paragraph>
</subSubSection>
<subSubSection id="E9973CD3811CFF8F4D20FA51F0CAFA50" box="[113,729,1453,1473]" pageId="1" pageNumber="196" type="nomenclature">
<paragraph id="A1326F58811CFF8F4D20FA51F0CAFA50" blockId="1.[113,1089,1453,1580]" box="[113,729,1453,1473]" pageId="1" pageNumber="196">
<emphasis id="93F9B34A811CFF8F4D20FA51F2F9FA50" bold="true" box="[113,234,1453,1472]" pageId="1" pageNumber="196">Type locality.</emphasis>
Indonesia, Sulawesi Utara, Dumoga-Bone National Park.
</paragraph>
</subSubSection>
<subSubSection id="E9973CD3811CFF8F4D20FA34F015F9BB" pageId="1" pageNumber="196" type="materials_examined">
<paragraph id="A1326F58811CFF8F4D20FA34F015F9BB" blockId="1.[113,1089,1453,1580]" pageId="1" pageNumber="196">
<emphasis id="93F9B34A811CFF8F4D20FA34F2E0FA4B" bold="true" box="[113,243,1480,1499]" pageId="1" pageNumber="196">Type material.</emphasis>
<materialsCitation id="11E56505811CFF8F4DA7FA35F2D5F980" ID-GBIF-Occurrence="1424516212" collectedFrom="Lowland forest" collectingDate="1985-02-24" collectingMethod="Flight interception trap" collectionCode="BMNH" country="Indonesia" location="Dumoga-Bone N. P. / Plot A, ca 200" pageId="1" pageNumber="196" specimenCode="B. M. 1985-10" specimenCount="1" stateProvince="SULAWESI UTARA" typeStatus="holotype">
<typeStatus id="7E36D1FA811CFF8F4DA7FA35F35AFA49" box="[246,329,1481,1500]" pageId="1" pageNumber="196" type="holotype">HOLOTYPE</typeStatus>
: ♁, ‘
<collectingCountry id="D99A2FC8811CFF8F4C3FFA34F3FBFA4B" box="[366,488,1480,1499]" name="Indonesia" pageId="1" pageNumber="196">INDONESIA</collectingCountry>
:
<collectingRegion id="6349A1BA811CFF8F4CBEFA34F0B1FA4C" box="[495,674,1480,1500]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="196">SULAWESI UTARA</collectingRegion>
/
<location id="A4523983811CFF8F4FE1FA35F1CBFA4B" LSID="urn:lsid:plazi:treatment:2924DE4E811CFF8B4C35FA99F05BFD63:A4523983811CFF8F4FE1FA35F1CBFA4B" box="[688,984,1480,1500]" country="Indonesia" name="Dumoga-Bone N. P. / Plot A, ca 200" pageId="1" pageNumber="196" stateProvince="SULAWESI UTARA">
Dumoga-Bone N. P. / Plot A, ca
<quantity id="6675C2BD811CFF8F4EE9FA34F1CBFA4B" box="[952,984,1480,1499]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="196" unit="m" value="200.0">200</quantity>
</location>
m /
<collectedFrom id="12912CA5811CFF8F4EA4FA34F2B1FA66" pageId="1" pageNumber="196">Lowland forest</collectedFrom>
/
<date id="D5334998811CFF8F4DE4FA1FF30FFA66" box="[181,284,1507,1527]" pageId="1" pageNumber="196" value="1985-02-24">
<collectingDate id="C577B070811CFF8F4DE4FA1FF30FFA66" box="[181,284,1507,1527]" pageId="1" pageNumber="196" value="1985-02-24">24. II. 1985</collectingDate>
</date>
//
<collectingMethod id="78CC174F811CFF8F4C64FA1FF3EEFA67" box="[309,509,1507,1527]" pageId="1" pageNumber="196">Flight interception trap</collectingMethod>
2 // R. Ent. Soc. Lond. / PROJECT WALLACE /
<specimenCode id="F12BC723811CFF8F4EE6FA18F62EFA66" box="[951,1085,1507,1527]" pageId="1" pageNumber="196">B. M. 1985–10</specimenCode>
’ (
<collectionCode id="C79CF79D811CFF8F4D26FA01F2AFF980" box="[119,188,1533,1552]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="1" pageNumber="196">BMNH</collectionCode>
).
</materialsCitation>
<materialsCitation id="11E56505811CFF8F4D9AFA02F1A8F980" ID-GBIF-Occurrence="1424516211" box="[203,955,1530,1553]" collectedFrom="Lowland forest" collectingDate="1985-02-24" collectingMethod="Flight interception trap 1" collectionCode="BMNH" country="Indonesia" location="Flight" pageId="1" pageNumber="196" specimenCount="1" specimenCount-female="1" stateProvince="SULAWESI UTARA" typeStatus="paratype">
<typeStatus id="7E36D1FA811CFF8F4D9AFA02F330F99F" box="[203,291,1534,1553]" pageId="1" pageNumber="196" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="B78BA4D1811CFF8F4C7FFA01F35DF981" box="[302,334,1530,1553]" pageId="1" pageNumber="196" type="female">1 ♀</specimenCount>
, same data holotype but ‘
<collectingMethod id="78CC174F811CFF8F4F78FA01F110F980" box="[553,771,1533,1553]" pageId="1" pageNumber="196">Flight interception trap 1</collectingMethod>
// +31| –7’ (
<collectionCode id="C79CF79D811CFF8F4E3DFA01F1A2F980" box="[876,945,1533,1552]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="1" pageNumber="196">BMNH</collectionCode>
);
</materialsCitation>
<materialsCitation id="11E56505811CFF8F4E91FA01F015F9BB" ID-GBIF-Occurrence="1424516213" collectedFrom="Lowland forest" collectingDate="1985-02-24" collectingMethod="Flight interception trap 3" collectionCode="BMNH" country="Indonesia" location="Flight" pageId="1" pageNumber="196" specimenCount="1" specimenCount-female="1" stateProvince="SULAWESI UTARA" typeStatus="paratype">
<specimenCount id="B78BA4D1811CFF8F4E91FA01F1F2F981" box="[960,993,1530,1553]" pageId="1" pageNumber="196" type="female">1 ♀</specimenCount>
, same data above but ‘
<collectingMethod id="78CC174F811CFF8F4D9CF9E4F3B4F9BB" box="[205,423,1560,1580]" pageId="1" pageNumber="196">Flight interception trap 3</collectingMethod>
’ (
<collectionCode id="C79CF79D811CFF8F4CE6F9E4F3EFF9BB" box="[439,508,1560,1579]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="1" pageNumber="196">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="F5F23FD0811FFF8C4DC6FB10F0B3FA89" httpUri="https://zenodo.org/record/270104/files/figure.png" pageId="2" pageNumber="197" targetBox="[157,1115,227,1243]" targetPageId="2">
<paragraph id="A1326F58811FFF8C4DC6FB10F0B3FA89" blockId="2.[151,1126,1260,1306]" pageId="2" pageNumber="197">
Fig. 1.
<taxonomicName id="668D14DB811FFF8C4D84FB11F390FB6F" box="[213,387,1260,1280]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="celebense" status="sp. nov.">
<emphasis id="93F9B34A811FFF8C4D84FB11F390FB6F" box="[213,387,1260,1280]" italics="true" pageId="2" pageNumber="197">Cyparium celebense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88CA0E31811FFF8C4CD8FB11F3DEFA90" box="[393,461,1261,1280]" pageId="2" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
A
<emphasis id="93F9B34A811FFF8C4CB5FB10F3FCFB6F" bold="true" box="[484,495,1260,1279]" pageId="2" pageNumber="197">–</emphasis>
habitus of male (left, dorsal view; right, lateral view); B
<emphasis id="93F9B34A811FFF8C4E8CFB10F1FBFB6F" bold="true" box="[989,1000,1260,1279]" pageId="2" pageNumber="197">–</emphasis>
male genitalia (left, dorsal view; right, lateral view); C
<emphasis id="93F9B34A811FFF8C4CA0FAFAF3EFFA89" bold="true" box="[497,508,1286,1305]" pageId="2" pageNumber="197">–</emphasis>
antenna of female.
</paragraph>
</caption>
<subSubSection id="E9973CD3811FFF8B4DC6FAA3F063FD03" lastPageId="5" lastPageNumber="200" pageId="2" pageNumber="197" type="description">
<paragraph id="A1326F58811FFF8C4DC6FAA3F3C1F9A8" blockId="2.[151,1127,1375,1592]" pageId="2" pageNumber="197">
<emphasis id="93F9B34A811FFF8C4DC6FAA3F37FFAE8" bold="true" box="[151,364,1375,1400]" pageId="2" pageNumber="197">
Description.
<emphasis id="93F9B34A811FFF8C4C7BFAA3F376FAE6" bold="true" box="[298,357,1375,1398]" italics="true" pageId="2" pageNumber="197">Head</emphasis>
.
</emphasis>
Almost black, clypeus and frons dark brown to reddish brown, mouth parts yellowish-brown. Antennomeres I–VI and apical half of XI brown to yellowish-brown; VII–X dark brown to black; XI light yellowish-brown. Head with eye width about 1.2 times as interocular distance. Punctation sparse and fine. Antennomeres I–VI with few macrosetae, VII–XI covered with some macrosetae; I about 2.5 times as long as VI; VI and VII each almost as long as wide; VIII–XI each wider than long; XI about twice as long as III and 2.5 times as long as VI (
<figureCitation id="39B673DD811FFF8C4C23F9E3F3BCF9A8" box="[370,431,1567,1592]" captionStart="Fig. 1" captionStartId="2.[151,182,1260,1279]" captionTargetBox="[157,1115,227,1243]" captionTargetId="figure@2.[151,1126,219,1251]" captionTargetPageId="2" captionText="Fig. 1. Cyparium celebense sp. nov. A – habitus of male (left, dorsal view; right, lateral view); B – male genitalia (left, dorsal view; right, lateral view); C – antenna of female." httpUri="https://zenodo.org/record/270104/files/figure.png" pageId="2" pageNumber="197">Fig. 1</figureCitation>
C).
</paragraph>
<caption id="F5F23FD0811EFF8D4D20FAF8F1C5FAA2" httpUri="https://zenodo.org/record/270105/files/figure.png" pageId="3" pageNumber="198" targetBox="[113,1481,476,1259]" targetPageId="3">
<paragraph id="A1326F58811EFF8D4D20FAF8F1C5FAA2" blockId="3.[113,1482,1284,1331]" pageId="3" pageNumber="198">
Fig. 3. Habitus and labels of four species of the genus
<taxonomicName id="668D14DB811EFF8D4F12FAF9F08BFA88" box="[579,664,1285,1304]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="93F9B34A811EFF8D4F12FAF9F08BFA88" box="[579,664,1285,1304]" italics="true" pageId="3" pageNumber="198">Cyparium</emphasis>
</taxonomicName>
from Indonesia and Malaysia. A –
<emphasis id="93F9B34A811EFF8D4E95FAF9F638FA87" box="[964,1067,1284,1304]" italics="true" pageId="3" pageNumber="198">C. bowringi</emphasis>
Achard, 1922 (lectotype); B –
<emphasis id="93F9B34A811EFF8D4865FAF9F75AFA88" box="[1332,1353,1285,1304]" italics="true" pageId="3" pageNumber="198">C.</emphasis>
<taxonomicName id="668D14DB811EFF8D481CFAFAF28FFAA2" authority="Lobl, 1990" authorityName="Lobl" authorityYear="1990" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="javanum">
<emphasis id="93F9B34A811EFF8D481CFAFAF785FA88" box="[1357,1430,1286,1304]" italics="true" pageId="3" pageNumber="198">javanum</emphasis>
Löbl, 1990
</taxonomicName>
(paratype); C –
<emphasis id="93F9B34A811EFF8D4C79FADCF388FAA2" box="[296,411,1312,1331]" italics="true" pageId="3" pageNumber="198">C. punctatum</emphasis>
Pic, 1916 (holotype); D –
<emphasis id="93F9B34A811EFF8D4F2EFADCF0EAFAA2" box="[639,761,1312,1331]" italics="true" pageId="3" pageNumber="198">C. variegatum</emphasis>
Achard, 1920 (holotype).
</paragraph>
</caption>
<caption id="F5F23FD08119FF8A4DC6FCC6F0F5FCF8" httpUri="https://zenodo.org/record/270106/files/figure.png" pageId="4" pageNumber="199" targetBox="[174,1097,227,809]" targetPageId="4">
<paragraph id="A1326F588119FF8A4DC6FCC6F0F5FCF8" blockId="4.[151,1127,826,872]" pageId="4" pageNumber="199">
Fig. 2. Genital characters of
<taxonomicName id="668D14DB8119FF8A4CDDFCC7F029FCDD" box="[396,570,826,846]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="celebense" status="sp. nov.">
<emphasis id="93F9B34A8119FF8A4CDDFCC7F029FCDD" box="[396,570,826,846]" italics="true" pageId="4" pageNumber="199">Cyparium celebense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88CA0E318119FF8A4F6EFCC7F090FCDE" box="[575,643,827,846]" pageId="4" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
A, B – male genitalia in lateral view (A – paramere, B – internal sac); C – female genitalia in ventral view. Scales = 0.25 mm.
</paragraph>
</caption>
<paragraph id="A1326F588119FF8A4DE4FC4BF620FA5F" blockId="4.[151,1127,951,1583]" pageId="4" pageNumber="199">
<emphasis id="93F9B34A8119FF8A4DE4FC4BF31DFC40" bold="true" box="[181,270,951,976]" pageId="4" pageNumber="199">
<emphasis id="93F9B34A8119FF8A4DE4FC4BF315FC5E" bold="true" box="[181,262,951,974]" italics="true" pageId="4" pageNumber="199">Thorax</emphasis>
.
</emphasis>
Pronotum and elytra black, without iridescent luster. Tibiae and femur dark reddish-brown; tarsus lighter than tibiae and femur. Ventral surface almost black, except for dark reddish-brown coxae (
<figureCitation id="39B673DD8119FF8A4CD8FC0BF3D4FB80" box="[393,455,1015,1040]" captionStart="Fig. 1" captionStartId="2.[151,182,1260,1279]" captionTargetBox="[157,1115,227,1243]" captionTargetId="figure@2.[151,1126,219,1251]" captionTargetPageId="2" captionText="Fig. 1. Cyparium celebense sp. nov. A – habitus of male (left, dorsal view; right, lateral view); B – male genitalia (left, dorsal view; right, lateral view); C – antenna of female." httpUri="https://zenodo.org/record/270104/files/figure.png" pageId="4" pageNumber="199">Fig. 1</figureCitation>
A). Pronotum wider than long, with anterior bead. Punctation sparse and fine, as on head. Scutellum wider than long, with exposed apex. Hypomeron and lateral portion of mesoventrite smooth. Lateral portion of metaventrite coarsely and sparsely punctate. Elytra slightly wider than long, widest at basal third to fourth, lateral margins gradually narrowed apically, minutely serrate at inner part of posterior margin, latero-posterior margin with some setae. Disc of elytra with punctation coarser than that on pronotum, with five distinct and one indistinct rows of punctures; first row extending outwards along basal margin and joined with third row. Intervals between rows of punctures finely and sparsely punctate. Sutural striae extending outwards along basal margin to form basal striae, reaching humeral area and not joined with lateral striae. Mesocoxa almost as wide as space between them. Mesepimeron about twice as long as wide. Metepisternum about six times as long as wide, without longitudinal line. Metepimeron almost as long as wide. Metacoxa almost as wide as metacoxal process. Protarsomeres I–III each 1.2 times as long as IV; V 1.5 times as long as III and twice as long as IV. Mesotarsomeres I and V each 2.0 times as long as each II–IV. Metatarsomeres I and V 1.2 times as long as III; II 1.2 times as long as III or IV.
</paragraph>
<paragraph id="A1326F588119FF8A4DE4FA2BF08DF9BE" blockId="4.[151,1127,951,1583]" pageId="4" pageNumber="199">
<emphasis id="93F9B34A8119FF8A4DE4FA2BF335FA60" bold="true" box="[181,294,1495,1520]" pageId="4" pageNumber="199">
<emphasis id="93F9B34A8119FF8A4DE4FA2BF30DFA7E" bold="true" box="[181,286,1495,1518]" italics="true" pageId="4" pageNumber="199">Abdomen</emphasis>
.
</emphasis>
Propygidium and pygidium dark brown to black. Propygidium and pygidium with microsculptures. Median portion of ventrite I coarsely and sparsely punctate, with punctures slightly weaker than those on metaventrite.
</paragraph>
<paragraph id="A1326F588118FF8B4DC1FF20F627FEA5" blockId="5.[113,1090,220,755]" pageId="5" pageNumber="200">
<emphasis id="93F9B34A8118FF8B4DC1FF20F2DDFF65" bold="true" box="[144,206,220,245]" pageId="5" pageNumber="200">
<emphasis id="93F9B34A8118FF8B4DC1FF20F2D4FF63" bold="true" box="[144,199,220,243]" italics="true" pageId="5" pageNumber="200">Male</emphasis>
.
</emphasis>
Pro- and mesotarsomeres I–III with tenent setae, not enlarged. Aedeagus 0.8 mm long; parameres symmetrical, slightly enlarged subapically and narrowed apically (
<figureCitation id="39B673DD8118FF8B4EDCFF00F1D9FE85" box="[909,970,252,277]" captionStart="Fig. 2" captionStartId="4.[151,182,826,845]" captionTargetBox="[174,1097,227,809]" captionTargetId="figure@4.[151,1126,219,821]" captionTargetPageId="4" captionText="Fig. 2. Genital characters of Cyparium celebense sp. nov. A, B – male genitalia in lateral view (A – paramere, B – internal sac); C – female genitalia in ventral view. Scales = 0.25 mm." httpUri="https://zenodo.org/record/270106/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
A); internal sac without sclerites, covered with fine scale-like and denticulate structures (
<figureCitation id="39B673DD8118FF8B4EF3FEE0F1F0FEA5" box="[930,995,284,309]" captionStart="Fig. 1" captionStartId="2.[151,182,1260,1279]" captionTargetBox="[157,1115,227,1243]" captionTargetId="figure@2.[151,1126,219,1251]" captionTargetPageId="2" captionText="Fig. 1. Cyparium celebense sp. nov. A – habitus of male (left, dorsal view; right, lateral view); B – male genitalia (left, dorsal view; right, lateral view); C – antenna of female." httpUri="https://zenodo.org/record/270104/files/figure.png" pageId="5" pageNumber="200">Figs 1</figureCitation>
B, 2B).
</paragraph>
<paragraph id="A1326F588118FF8B4DC1FEC0F651FEE3" blockId="5.[113,1090,220,755]" pageId="5" pageNumber="200">
<emphasis id="93F9B34A8118FF8B4DC1FEC0F2F9FEC5" bold="true" box="[144,234,316,341]" pageId="5" pageNumber="200">
<emphasis id="93F9B34A8118FF8B4DC1FEC0F2F0FEC3" bold="true" box="[144,227,316,339]" italics="true" pageId="5" pageNumber="200">Female</emphasis>
.
</emphasis>
Pro- and mesotarsomeres I–III without tenent setae, not enlarged. Ovipositor simple; bursa copulatrix sclerotized, broadly triangular (
<figureCitation id="39B673DD8118FF8B4F95FEA0F117FEE5" box="[708,772,348,373]" captionStart="Fig. 2" captionStartId="4.[151,182,826,845]" captionTargetBox="[174,1097,227,809]" captionTargetId="figure@4.[151,1126,219,821]" captionTargetPageId="4" captionText="Fig. 2. Genital characters of Cyparium celebense sp. nov. A, B – male genitalia in lateral view (A – paramere, B – internal sac); C – female genitalia in ventral view. Scales = 0.25 mm." httpUri="https://zenodo.org/record/270106/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
C). Spermatheca not detected.
</paragraph>
<paragraph id="A1326F588118FF8B4DC1FE81F063FD03" blockId="5.[113,1090,220,755]" pageId="5" pageNumber="200">
<emphasis id="93F9B34A8118FF8B4DC1FE81F322FE04" bold="true" box="[144,305,381,404]" italics="true" pageId="5" pageNumber="200">Measurements</emphasis>
(n = 3). Length (PL+EL): 2.13–2.36 mm. PW: 1.31–1.42 mm. EW: 1.44– 1.67 mm. HW: 0.56–0.60 mm. ID: 0.16–0.18 mm. PL/PW: 0.64–0.67, EL/EW: 0.84–0.89. Approximate ratio of each antennal segment in length (width) (n = 1) = 1.5 (0.6): 1.1 (0.5): 1.0 (0.4): 0.8 (0.4): 0.9 (0.5): 0.6 (0.6): 0.9 (1.0): 0.8 (1.1): 0.8 (1.3): 0.9 (1.4): 1.6 (1.4).
<emphasis id="93F9B34A8118FF8B4D20FE00F37FFD83" bold="true" box="[113,364,508,531]" pageId="5" pageNumber="200">Differential diagnosis.</emphasis>
This species is very similar to
<taxonomicName id="668D14DB8118FF8B4FEAFE01F616FD85" authority="Lobl, 1990" authorityName="Lobl" authorityYear="1990" box="[699,1029,508,533]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="javanum">
<emphasis id="93F9B34A8118FF8B4FEAFE01F19AFD84" box="[699,905,509,532]" italics="true" pageId="5" pageNumber="200">Cyparium javanum</emphasis>
<bibRefCitation id="C51C12A98118FF8B4EC1FE00F616FD85" author="LOBL I" box="[912,1029,508,533]" journalOrPublisher="Elytron" pageId="5" pageNumber="200" pagination="125 - 129" part="4" refString="LOBL I. 1990: Cyparium javanum sp. n., a new Scaphidiidae (Coleoptera) from Indonesia. Elytron 4: 125 - 129." title="Cyparium javanum sp. n., a new Scaphidiidae (Coleoptera) from Indonesia" type="journal article" year="1990">Löbl, 1990</bibRefCitation>
</taxonomicName>
from Java, Indonesia, in particular by its colour pattern, the small body size, and the antennomere XI yellowish-brown, but may be easily distinguished from
<emphasis id="93F9B34A8118FF8B4FB2FDC1F14EFDC4" box="[739,861,573,596]" italics="true" pageId="5" pageNumber="200">
C.
<taxonomicName id="668D14DB8118FF8B4E50FDC1F14EFDC4" box="[769,861,573,596]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="javanum">javanum</taxonomicName>
</emphasis>
by the lateral portion of the metaventrite and the median portion of the abdominal ventrite I coarsely punctate, and the apical portion of parameres hardly widened.
</paragraph>
</subSubSection>
<subSubSection id="E9973CD38118FF8B4D20FD61F395FD43" pageId="5" pageNumber="200" type="etymology">
<paragraph id="A1326F588118FF8B4D20FD61F395FD43" blockId="5.[113,1090,220,755]" pageId="5" pageNumber="200">
<emphasis id="93F9B34A8118FF8B4D20FD61F2E1FD25" bold="true" box="[113,242,669,693]" pageId="5" pageNumber="200">Etymology.</emphasis>
The species epithet (
<emphasis id="93F9B34A8118FF8B4C81FD60F095FD24" box="[464,646,668,692]" italics="true" pageId="5" pageNumber="200">celebensis, -is, -e</emphasis>
) is an adjective derived from Celebes, the former name of Sulawesi.
</paragraph>
</subSubSection>
<subSubSection id="E9973CD38118FF8B4D20FD20F05BFD63" box="[113,584,732,755]" pageId="5" pageNumber="200" type="distribution">
<paragraph id="A1326F588118FF8B4D20FD20F05BFD63" blockId="5.[113,1090,220,755]" box="[113,584,732,755]" pageId="5" pageNumber="200">
<emphasis id="93F9B34A8118FF8B4D20FD20F317FD63" bold="true" box="[113,260,732,755]" pageId="5" pageNumber="200">Distribution.</emphasis>
Indonesia: northern Sulawesi.
</paragraph>
</subSubSection>
</treatment>
</document>
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