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<document id="1259E41DF499C313B9D07F9CFF702D46" ID-DOI="10.11646/zootaxa.991.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5046283" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625062394442" checkinUser="felipe" docAuthor="Blahnik Table Of Contents, Roger J." docDate="2005" docId="EC4BE952FFE26829FEBDCC8F89A7E932" docLanguage="en" docName="zt00991.pdf" docOrigin="Zootaxa 991 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.991.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Alterosa intervales Blahnik Table Of Contents 2005, new species" docType="treatment" docVersion="4" lastPageNumber="35" masterDocId="1072912AFFC2680BFFB5CA3E8D48EC73" masterDocTitle="Alterosa, a new caddisfly genus from Brazil (Trichoptera: Philopotamidae)" masterLastPageNumber="60" masterPageNumber="1" pageNumber="33" updateTime="1698979645207" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E13D05608B2F6BE35D2F8C8754B16FD1">Alterosa, a new caddisfly genus from Brazil (Trichoptera: Philopotamidae)</mods:title>
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<mods:namePart id="207B275DFDC70FB377637D21E7E1E8D5">Blahnik Table Of Contents, Roger J.</mods:namePart>
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<mods:title id="7B2B7CEE9DDE893528C81CFE450CCF76">Zootaxa</mods:title>
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<mods:date id="0FC3A969F40301029864343680C77CB3">2005</mods:date>
<mods:detail id="69A5C6FD496C1CF0C99DACE4D1BE4164" type="pubDate">
<mods:number id="C40155E7A488204FF542E8CC16FE9C74">2005-05-23</mods:number>
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<mods:number id="2ADF266980B6E095FD823C0579160642">991</mods:number>
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<mods:classification id="EA7B30519B73397104694285D08A0F15">journal article</mods:classification>
<mods:identifier id="435CC52413F7D6E306D64916AF8638B2" type="DOI">10.11646/zootaxa.991.1.1</mods:identifier>
<mods:identifier id="864084BA4F9A3ACEFF5BC1B7750B6882" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="91CD81B89695DE4FD256B0B07037DF28" type="Zenodo-Dep">5046283</mods:identifier>
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<treatment id="EC4BE952FFE26829FEBDCC8F89A7E932" ID-DOI="http://doi.org/10.5281/zenodo.5047060" ID-Zenodo-Dep="5047060" LSID="urn:lsid:plazi:treatment:EC4BE952FFE26829FEBDCC8F89A7E932" httpUri="http://treatment.plazi.org/id/EC4BE952FFE26829FEBDCC8F89A7E932" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
<subSubSection id="2CF80BCFFFE2682BFEBDCC8F8FC0EAB8" box="[264,648,1713,1739]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="645D5844FFE2682BFEBDCC8F8FC0EAB8" blockId="32.[264,648,1713,1774]" box="[264,648,1713,1739]" pageId="32" pageNumber="33">
<heading id="3F15EF28FFE2682BFEBDCC8F8FC0EAB8" bold="true" box="[264,648,1713,1739]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="56968456FFE2682BFEBDCC8F8FC0EAB8" bold="true" box="[264,648,1713,1739]" pageId="32" pageNumber="33">
<taxonomicName id="A3E223C7FFE2682BFEBDCC8F8CA1EAB8" authority="Blahnik Table Of Contents, 2005" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[264,489,1713,1739]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="intervales" status="sp. nov.">
<emphasis id="56968456FFE2682BFEBDCC8F8CA1EAB8" bold="true" box="[264,489,1713,1739]" italics="true" pageId="32" pageNumber="33">Alterosa intervales</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="4DA5392DFFE2682BFE4DCC8F8FC0EAB8" box="[504,648,1713,1739]" pageId="32" pageNumber="33" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2CF80BCFFFE26829FEBDCCEA8CB1E85A" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
<paragraph id="645D5844FFE2682BFEBDCCEA8CDBEA9D" blockId="32.[264,648,1713,1774]" box="[264,403,1748,1774]" pageId="32" pageNumber="33">
<figureCitation id="FCD944C1FFE2682BFEBDCCEA8CDBEA9D" box="[264,403,1748,1774]" captionStart="FIGURE 16" captionStartId="33.[264,368,1758,1782]" captionTargetBox="[321,1280,548,1725]" captionTargetId="figure-93@33.[302,1284,529,1731]" captionTargetPageId="33" captionText="FIGURE 16. Alterosa intervales, new species. Male genitalia: 16A—lateral view; 16B—dorsal view of segments IX and X; 16C—inferior appendage, dorsal view; 16D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046320" httpUri="https://zenodo.org/record/5046320/files/figure.png" pageId="32" pageNumber="33">Fig. 16A–D</figureCitation>
</paragraph>
<paragraph id="645D5844FFE2682AFEBDCD1A8E71EDDD" blockId="32.[264,1324,1828,1894]" lastBlockId="33.[264,1322,284,430]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<taxonomicName id="A3E223C7FFE2682BFEBDCD1A8CADEB4E" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[264,485,1828,1853]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="intervales">
<emphasis id="56968456FFE2682BFEBDCD1A8CADEB4E" box="[264,485,1828,1853]" italics="true" pageId="32" pageNumber="33">Alterosa intervales</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="A3E223C7FFE2682BFDCECD1A8E55EB4E" authorityName="Flint" authorityYear="1971" box="[635,797,1828,1853]" class="Insecta" family="Philopotamidae" genus="Dolophilodes" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="sanctipauli">
<emphasis id="56968456FFE2682BFDCECD1A8E55EB4E" box="[635,797,1828,1853]" italics="true" pageId="32" pageNumber="33">A. sanctipauli</emphasis>
</taxonomicName>
, resembling that species in the elongate, narrow, unbranched intermediate appendages. It differs in that the setae of the intermediate appendage in
<taxonomicName id="A3E223C7FFE3682AFE07CB228F02ED46" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[434,586,284,309]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="intervales">
<emphasis id="56968456FFE3682AFE07CB228F02ED46" box="[434,586,284,309]" italics="true" pageId="33" pageNumber="34">A. intervales</emphasis>
</taxonomicName>
are very elongate and located along the apicolateral margin, while those in
<taxonomicName id="A3E223C7FFE3682AFE0DCB7A8F14ED2E" authorityName="Flint" authorityYear="1971" box="[440,604,324,349]" class="Insecta" family="Philopotamidae" genus="Dolophilodes" kingdom="Animalia" order="Trichoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="sanctipauli">
<emphasis id="56968456FFE3682AFE0DCB7A8F14ED2E" box="[440,604,324,349]" italics="true" pageId="33" pageNumber="34">A. sanctipauli</emphasis>
</taxonomicName>
are very short and terminal.
<taxonomicName id="A3E223C7FFE3682AFC0ACB7A89D6ED2E" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[959,1182,324,349]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="intervales">
<emphasis id="56968456FFE3682AFC0ACB7A89D6ED2E" box="[959,1182,324,349]" italics="true" pageId="33" pageNumber="34">Alterosa intervales</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="A3E223C7FFE3682AFEFECB528CA7EDF6" authorityName="Flint" authorityYear="1971" box="[331,495,364,390]" class="Insecta" family="Philopotamidae" genus="Dolophilodes" kingdom="Animalia" order="Trichoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="sanctipauli">
<emphasis id="56968456FFE3682AFEFECB528C15EDF6" box="[331,349,364,389]" italics="true" pageId="33" pageNumber="34">A</emphasis>
.
<emphasis id="56968456FFE3682AFED8CB528CA7EDF6" box="[365,495,364,389]" italics="true" pageId="33" pageNumber="34">sanctipauli</emphasis>
</taxonomicName>
in the shape of the apex of tergum X and presence of paired, spined longitudinal ridges found on the same structure.
</paragraph>
<caption id="309D08CCFFE3682AFEBDCCE08C09EB49" ID-DOI="http://doi.org/10.5281/zenodo.5046320" ID-Zenodo-Dep="5046320" httpUri="https://zenodo.org/record/5046320/files/figure.png" pageId="33" pageNumber="34" startId="33.[264,368,1758,1782]" targetBox="[321,1280,548,1725]" targetPageId="33">
<paragraph id="645D5844FFE3682AFEBDCCE08C09EB49" blockId="33.[264,1323,1758,1850]" pageId="33" pageNumber="34">
<emphasis id="56968456FFE3682AFEBDCCE08CD1EA85" bold="true" box="[264,409,1758,1782]" pageId="33" pageNumber="34">FIGURE 16.</emphasis>
<taxonomicName id="A3E223C7FFE3682AFE1CCCE08F3BEA86" authority="Blahnik Table Of Contents, 2005" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[425,627,1758,1781]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="intervales" status="sp. nov.">
<emphasis id="56968456FFE3682AFE1CCCE08F3BEA86" box="[425,627,1758,1781]" italics="true" pageId="33" pageNumber="34">Alterosa intervales</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="4DA5392DFFE3682AFD37CCE08E4FEA86" box="[642,775,1758,1781]" pageId="33" pageNumber="34" rank="species">new species</taxonomicNameLabel>
. Male genitalia: 16A—lateral view; 16B—dorsal view of segments IX and X; 16C—inferior appendage, dorsal view; 16D—phallic apparatus, lateral view.
</paragraph>
</caption>
<paragraph id="645D5844FFE06829FE8DCB228866ED28" blockId="34.[264,1326,284,1345]" pageId="34" pageNumber="35">
Adult. Color overall, dark brown; legs, palps, and antennae slightly paler, wings irregularly mottled with small light brown spots. Male forewing
<quantity id="A31AF5A1FFE06829FC7ECB7F8901ED28" box="[971,1097,321,347]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.3" metricValueMin="7.0" pageId="34" pageNumber="35" unit="mm" value="7.15" valueMax="7.3" valueMin="7.0">7–7.3 mm</quantity>
.; female
<quantity id="A31AF5A1FFE06829FB1ACB7F8862ED28" box="[1199,1322,321,347]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.0" metricValueMin="6.8" pageId="34" pageNumber="35" unit="mm" value="6.9" valueMax="7.0" valueMin="6.8">6.8–7 mm</quantity>
.
</paragraph>
<paragraph id="645D5844FFE06829FE8DCB598CB1E85A" blockId="34.[264,1326,284,1345]" pageId="34" pageNumber="35">Male genitalia. Tergum VIII with posteromesal margin deeply emarginate, emargination V­shaped and extending more than half way to anterior margin. Sternum IX with anterolateral margin forming a rounded projection in dorsal half; posteroventral margin greatly produced, forming broadly rounded expansion; tergum IX greatly reduced, membranous or fused to base of tergum X. Tergum X with basolateral margins rounded, tapering apically; dorsally with mesally paired, longitudinal rows of short spines, converging apically; apex sensillate, rounded as viewed dorsally, subtruncate in lateral view; tergum textured basolaterally, with scabrous cuticle. Intermediate appendage heavily sclerotized, elongate, extending past preanal appendages, rod­like, curved near base; apex acute, with linear, preapical row of about four elongate spine­like setae. Preanal appendage narrow, constricted basally, moderately elongate, not greatly modified, with scant setae and numerous minute seta­like spines; apex rounded, with distinct terminal seta. Inferior appendages elongate, linear, flattened on mesal surface; 1st article, in lateral view, narrow, about 3x longer than wide, tapering apically; 2nd article subequal in length to 1st article, nearly uniform in width, only slightly narrower at base than apex of 1st article; apex rounded, with small pad of short, stiff apicomesal setae. Phallobase tubular, short, scarcely curved; endotheca with numerous fine spines in several distinct, paired tracts near base, apex elongate, without spines.</paragraph>
</subSubSection>
<subSubSection id="2CF80BCFFFE06829FE8DCE0889A7E932" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="645D5844FFE06829FE8DCE088F3CE8D2" blockId="34.[264,1326,284,1345]" pageId="34" pageNumber="35">
<materialsCitation id="D48A5219FFE06829FE8DCE088F3CE8D2" collectingDate="2002-10-01" collectionCode="MZUSP" collectorName="Blahnik &amp; Prather" country="Brazil" elevation="800" latitude="-24.275517" location="Agua Comprida" longLatPrecision="1" longitude="-48.417366" municipality="Parque Estadual Intervales" pageId="34" pageNumber="35" specimenCode="UMSP000088786" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="56968456FFE06829FE8DCE088E6BE823" bold="true" box="[312,803,1078,1104]" pageId="34" pageNumber="35">
<typeStatus id="BB59E6E6FFE06829FE8DCE088CE2E823" box="[312,426,1078,1104]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="1CF518D4FFE06829FDB2CE088F34E823" box="[519,636,1078,1104]" name="Brazil" pageId="34" pageNumber="35">BRAZIL</collectingCountry>
:
<collectingRegion id="A62696A6FFE06829FD2CCE088E56E823" box="[665,798,1078,1104]" country="Brazil" name="Sao Paulo" pageId="34" pageNumber="35">São Paulo</collectingRegion>
:
</emphasis>
<collectingMunicipality id="8439C23EFFE06829FC8CCE098935E822" box="[825,1149,1079,1105]" pageId="34" pageNumber="35">Parque Estadual Intervales</collectingMunicipality>
, small trib. to
<location id="613D0E9FFFE06829FEBDCE618C86E80A" LSID="urn:lsid:plazi:treatment:EC4BE952FFE26829FEBDCC8F89A7E932:613D0E9FFFE06829FEBDCE618C86E80A" box="[264,462,1119,1145]" country="Brazil" latitude="-24.275517" longLatPrecision="1" longitude="-48.417366" municipality="Parque Estadual Intervales" name="Agua Comprida" pageId="34" pageNumber="35" stateProvince="Sao Paulo">Agua Comprida</location>
,
<geoCoordinate id="01D63E83FFE06829FE6ACE618F39E80A" box="[479,625,1119,1145]" degrees="24" direction="south" minutes="16.531" orientation="latitude" pageId="34" pageNumber="35" precision="1" value="-24.275517">24°16.531'S</geoCoordinate>
,
<geoCoordinate id="01D63E83FFE06829FD31CE618E56E80A" box="[644,798,1119,1145]" degrees="48" direction="west" minutes="25.042" orientation="longitude" pageId="34" pageNumber="35" precision="1" value="-48.417366">48°25.042'W</geoCoordinate>
, el
<quantity id="A31AF5A1FFE06829FCE6CE618EEFE80A" box="[851,935,1119,1145]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="34" pageNumber="35" unit="m" value="800.0">
<elevation id="EFCFBF77FFE06829FCE6CE618EEFE80A" box="[851,935,1119,1145]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="34" pageNumber="35" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="105C7E84FFE06829FC0CCE618979E80A" box="[953,1073,1119,1145]" pageId="34" pageNumber="35" value="2002-10-01">
<collectingDate id="0018876CFFE06829FC0CCE618979E80A" box="[953,1073,1119,1145]" pageId="34" pageNumber="35" value="2002-10-01">01.x.2002</collectingDate>
</date>
,
<collectorName id="C9173D92FFE06829FBF6CE6189E8E80A" box="[1091,1184,1119,1145]" pageId="34" pageNumber="35">Blahnik</collectorName>
&amp;
<collectorName id="C9173D92FFE06829FB67CE618861E80A" box="[1234,1321,1119,1145]" pageId="34" pageNumber="35">Prather</collectorName>
, (
<specimenCode id="3444F03FFFE06829FEA5CEB98CAFE8D2" box="[272,487,1159,1185]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="34" pageNumber="35" type="University or college">UMSP000088786</specimenCode>
) (
<collectionCode id="02F3C081FFE06829FE4ACEB98F20E8D2" box="[511,616,1159,1185]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="35">MZUSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="645D5844FFE06829FE8DCE908F15E882" blockId="34.[264,1326,284,1345]" pageId="34" pageNumber="35">
<materialsCitation id="D48A5219FFE06829FE8DCE908F11E882" collectingDate="2002-10-01" collectionCode="UMSP, MZUSP" collectorName="Blahnik &amp; Prather" country="Brazil" elevation="800" latitude="-24.275517" location="Agua Comprida" longLatPrecision="1" longitude="-48.417366" municipality="Parque Estadual Intervales" pageId="34" pageNumber="35" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<emphasis id="56968456FFE06829FE8DCE908FA7E8BB" bold="true" box="[312,751,1198,1224]" pageId="34" pageNumber="35">
<typeStatus id="BB59E6E6FFE06829FE8DCE908CF2E8BB" box="[312,442,1198,1224]" pageId="34" pageNumber="35" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="1CF518D4FFE06829FE60CE908F02E8BB" box="[469,586,1198,1224]" name="Brazil" pageId="34" pageNumber="35">BRAZIL</collectingCountry>
:
<collectingRegion id="A62696A6FFE06829FDD0CE908FA2E8BB" box="[613,746,1198,1224]" country="Brazil" name="Sao Paulo" pageId="34" pageNumber="35">São Paulo</collectingRegion>
:
</emphasis>
—
<specimenCount id="72E493CDFFE06829FC9DCE918E31E8BA" box="[808,889,1199,1225]" pageId="34" pageNumber="35" type="male">1 male</specimenCount>
,
<specimenCount id="72E493CDFFE06829FC3CCE918948E8BA" box="[905,1024,1199,1225]" pageId="34" pageNumber="35" type="female">3 females</specimenCount>
, same collection data as holotype (
<collectionCode id="02F3C081FFE06829FECBCEE98C99E882" box="[382,465,1239,1265]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="34" pageNumber="35" type="University or college">UMSP</collectionCode>
) (
<collectionCode id="02F3C081FFE06829FE5DCEE98F19E882" box="[488,593,1239,1265]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="35">MZUSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="645D5844FFE06829FE8DCEC189A7E932" blockId="34.[264,1326,284,1345]" pageId="34" pageNumber="35">
Etymology. This species is named
<taxonomicName id="A3E223C7FFE06829FD60CEC18E00E96B" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[725,840,1279,1304]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="intervales">
<emphasis id="56968456FFE06829FD60CEC18E00E96B" box="[725,840,1279,1304]" italics="true" pageId="34" pageNumber="35">intervales</emphasis>
</taxonomicName>
, used as a noun in apposition, for Parque Estadual Intervales, the very beautiful park where the
<typeStatus id="BB59E6E6FFE06829FC37CF198EFDE932" box="[898,949,1319,1345]" pageId="34" pageNumber="35">type</typeStatus>
specimens were collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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