treatments-xml/data/22/B3/AB/22B3ABC733915CC7A3062A1197061F58.xml

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<mods:title id="BC3D13EDF8A05D64C4CB96E9FF4BEAFD">Diversity underfoot of agromyzids (Agromyzidae, Diptera) mining thalli of liverworts and hornworts</mods:title>
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<mods:namePart id="CC5F25C1853D8E2473F44111179312FF">Kato, Makoto</mods:namePart>
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<mods:namePart id="9721A2EF018495063F8E6AED18290FA5">Yamamori, Luna</mods:namePart>
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<mods:namePart id="06072D453E5A437310C31793BFC49FEC">Imada, Yume</mods:namePart>
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<treatment id="22B3ABC733915CC7A3062A1197061F58" ID-GBIF-Taxon="205344559" LSID="urn:lsid:zoobank.org:act:3599CBDF-77C3-4B23-B42B-18F6FD336819" httpUri="http://treatment.plazi.org/id/22B3ABC733915CC7A3062A1197061F58" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="E41E1A88AB0C1D3AD374DFD245A9C3C8" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="8222E2FBFCE85497BC3DCA47016E5D98" pageId="0" pageNumber="1">
12. 
<taxonomicName id="9F8B2691369BBDD7557755BEE91E10A3" ID-CoL="BDJLC" LSID="https://zoobank.org/3599CBDF-77C3-4B23-B42B-18F6FD336819" authority="Kato" authorityName="Kato" authorityYear="2022" class="Insecta" family="Agromyzidae" genus="Phytoliriomyza" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Phytoliriomyza cometiformis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cometiformis" status="sp. nov.">Phytoliriomyza cometiformis Kato</taxonomicName>
<taxonomicNameLabel id="58398267264696EF663725003B5EB05C" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="A95FD75824E7AF9D5CB8FD7A411C583C" pageId="0" pageNumber="1" type="description">
<paragraph id="81E8D53709588D536558849E86C5C1B3" pageId="0" pageNumber="1">
<figureCitation id="C8F0457D282D75A88D085F9EE20680E7" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">Figs 22</figureCitation>
<figureCitation id="0D50CDF5C31A1C29DA47216E9F97A87F" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female morphology and larval ecology of Phytoliriomyza cometiformis sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C habitat at type locality D-G mined thalli of Reboulia hemisphaerica orientalis (D at type locality E at Ukawa F, G at Sagiura). An arrow in F indicates an internal puparium." figureDoi="10.3897/zookeys.1133.94530.figure23" httpUri="https://binary.pensoft.net/fig/775396" pageId="0" pageNumber="1">, 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="426F0406B571EACAA397DFF694C49650" pageId="0" pageNumber="1" type="material examined">
<paragraph id="1A8C58745643506BE0F777CAF8146696" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="9189C730D5EBB2240FDD143CE87B9860" pageId="0" pageNumber="1">
<emphasis id="6C98953405EC75BB9D254E3C70012CD3" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
<emphasis id="F94455239362B1570EC8277BDDB35C40" bold="true" pageId="0" pageNumber="1">:</emphasis>
Japan: 1♂ (MK-AG-789), Ashizuri-misaki, Tosashimizu, Kochi Pref. (
<geoCoordinate id="193104F6960F838C40F676BF88E8F10C" degrees="32.7298" direction="north" orientation="latitude" precision="5" value="32.7298">32.7298°N</geoCoordinate>
, 
<geoCoordinate id="025FC33A48556B455A151A8B5167D0E6" degrees="132.9971" direction="east" orientation="longitude" precision="5" value="132.9971">132.9971°E</geoCoordinate>
, 75 m asl), 26-II-2011 (as larva), emerged on 1-IV-2011 NSMT-I-Dip 31954. 
<emphasis id="53BE1E3C4541F24B405320DE5B9E89BF" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
<emphasis id="07EA01C3157842B6473823F25137C0DB" bold="true" pageId="0" pageNumber="1">:</emphasis>
Japan: 1♂1♀ (MK-AG-a427, a428), same data as holotype, emerged on 30-III-2011, NSMT-I-Dip 31955, 31955; 1♀ (MK-AG-a461), Hachijo Is., Tokyo Pref., 24-IV-2001 (as larva), emerged on 3-V-2001, NSMT-I-Dip 31957; 1♂ (MK-AG-411), Sagiura, Taisha, Izumo, Shimane Pref., 31-III-2015 (as larva), emerged on 30-IV-2015, NSMT-I-Dip 31958.
</paragraph>
</subSubSection>
<subSubSection id="835D79BC4D2120DEC5C816BF385A07F2" pageId="0" pageNumber="1" type="other material">
<paragraph id="CB19BF48A6C43D7C9AF3ED14C6974D30" pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph id="2BD885191BD9F1E0FF74523865CD82F1" pageId="0" pageNumber="1">Japan: 6♂7♀, Izu-oshima Is. Tokyo Pref., 22-III-2009 (as larva), emerged on 15-20-IV-2009; 2♀, Yugashima, Izu, Shizuoka Pref., 7-III-2012 (as larva), emerged on 12-14-IV-2012; 4♂8♀, Sinjo, Mihama, Mikata, Fukui Pref., 11-III-2012 (as larva), emerged on 18-23-IV-2012; 3♂2♀, Shimaji-gawa, Ujitachi, Ise, Mie Pref., 3-IV-2010 (as larva), emerged on 22-IV-2-V-2010; 2♂1♀, Urashima-jinja, Honjo-hama, Ine, Kyoto Pref., 31-VII-2011 (as larva), emerged on 22-III-2011; 2♂5♀, Ukawa, Tango, Kyotango, Kyoto Pref., 5-III-2021 (as larva), emerged on 2-8-IV-2021; 1♀, Sannoko, Kawakami, Higashi-yoshino, Nara Pref., 26-II-2016 (as larva), emerged on 11-IV-2016; 1♀, Tazukawa-keikoku, Katsuura, Tokushima Pref., 11-X-2016 (as larva), emerged on 19-IV-2011.</paragraph>
</subSubSection>
<subSubSection id="7F0EED6EC69489548FAF301517C8FB81" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="CA8E59587B007BFC46A8B04F929E4A07" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="4459217680727A3F017E63D85CC7036D" pageId="0" pageNumber="1">A large yellow species (wing length 2.0-2.2 mm) having a pruinose dark gray scutum with an oval yellow pattern extending from the mid-posterior margin to the scutellum, a black 1st flagellomere, yellow maxillary palpus, yellow halteres, and yellow legs. Male epandrium inner-laterally with a hand-like comb comprising</paragraph>
<paragraph id="C490E15093FD5528DD1D7938B2408FBF" pageId="0" pageNumber="1">
Four or five basally fused, long, tubercle-like setae. Larva mines the thallus of 
<taxonomicName id="FEF1ADA5A6872C7E70498406646188F2" class="Marchantiopsida" family="Aytoniaceae" genus="Reboulia" kingdom="Plantae" lsidName="Reboulia hemisphaerica subsp. orientalis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="hemisphaerica" subSpecies="orientalis">
<emphasis id="19064E8D07CB3A5C7D26FDFD1B430AC8" italics="true" pageId="0" pageNumber="1">Reboulia hemisphaerica orientalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D3506C3316DEA47D3CB5F7E3082D7B07" pageId="0" pageNumber="1" type="description">
<paragraph id="0BA0DB9F5ED493F82E4A6137AD5F4538" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="389C65803F4D5FC8E4E2E8112B486E14" pageId="0" pageNumber="1">
<emphasis id="C046B1BFC1FBF8696F3FB0A96BB685FC" bold="true" pageId="0" pageNumber="1">Adult male</emphasis>
(Fig. 
<figureCitation id="C821004F831697924D15D6130E111D05" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22A-D</figureCitation>
).
</paragraph>
<paragraph id="8BAE96F75C1263B724B6D6A8F6870EA4" pageId="0" pageNumber="1">
<emphasis id="DBD192081CBAD49A012C8CFF4B3CEB67" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="2F5BCAD72A9A9BF91128B7DE30AA0353" bold="true" pageId="0" pageNumber="1">:</emphasis>
Head entirely yellow, with ocellar tubercle brown, and back of head dark brown (Fig. 
<figureCitation id="26E2B94E1CE3F85CDBDC01B4B3F8CCA1" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22C</figureCitation>
). Antenna porrect; first flagellomere black, pedicel yellow and scape light yellow. Arista subbasal, brown, pubescent. Face, gena, parafacial and postgena yellow. Proboscis normal, yellow; palpus yellow, cylindrical (Fig. 
<figureCitation id="F7AECC198496DF4AAA36684A934A06F6" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22C</figureCitation>
). 
<emphasis id="97B05D6F666837E78273881E54A8E3F9" bold="true" italics="true" pageId="0" pageNumber="1">Chaetotaxy</emphasis>
<emphasis id="BD319A4E52CFEB0D2CCAA8634B55C2D2" bold="true" pageId="0" pageNumber="1">:</emphasis>
Front orbitals three pairs; one ori directed inward; two ors directed upward (Fig. 
<figureCitation id="2E96F0C5310447D71D98619CE0C48BB2" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22B</figureCitation>
). Orbital setulae minute and erect, in a single row.
</paragraph>
<caption id="FE910866505774A7CC399FAAE46CCB4E" doi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1" start="Figure 22" startId="F22">
<paragraph id="8B0927A063B924D58C9CC2EF92C03964" pageId="0" pageNumber="1">
<emphasis id="B78281F0EEAD15846B417623ECACC542" bold="true" pageId="0" pageNumber="1">Figure 22.</emphasis>
<taxonomicName id="042B451B1A581A262B6A58336E073667" authorityName="Kato" authorityYear="2022" class="Insecta" family="Dendrocerotaceae" genus="Phytoliriomyza" kingdom="Animalia" lsidName="Phytoliriomyza cometiformis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cometiformis">
<emphasis id="E5F28C9C7C663D1950147B7B49074EB6" italics="true" pageId="0" pageNumber="1">Phytoliriomyza cometiformis</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="138726FD1F486D61F68E41B78BFFF011" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
holotype male 
<emphasis id="72AEF3675D1B6040BF383FDE73699227" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus 
<emphasis id="8C3CC3540CE3072766E0E6C41DEA5629" bold="true" pageId="0" pageNumber="1">B</emphasis>
lateral 
<emphasis id="16318A0BE5132AFA10AA4DB5E25EA90B" bold="true" pageId="0" pageNumber="1">C</emphasis>
frontal 
<emphasis id="BFD19E6B2B48181B940558E093B12483" bold="true" pageId="0" pageNumber="1">D</emphasis>
dorsal 
<emphasis id="18C05FE9BB33D2F6C5C1A2A89ED71F55" bold="true" pageId="0" pageNumber="1">E</emphasis>
paratype female, dorsal (MK-AG-a428) 
<emphasis id="4579AF406AE3493639CE230D77582626" bold="true" pageId="0" pageNumber="1">F-J</emphasis>
male genitalia (
<emphasis id="2712B4B256C4AAD667249A21D98DD805" bold="true" pageId="0" pageNumber="1">F-H</emphasis>
at type locality 
<emphasis id="BD5F15DAC30598720C92AEB680CB083D" bold="true" pageId="0" pageNumber="1">I, J</emphasis>
at Sagiura) 
<emphasis id="93C12D768B4E5E6A1261E6BCAF13F52D" bold="true" pageId="0" pageNumber="1">F</emphasis>
whole genitalia, ventral 
<emphasis id="FA510FA48CF506C5EA111AFF4AFBDA24" bold="true" pageId="0" pageNumber="1">G</emphasis>
phallic complex, lateral 
<emphasis id="A632257D130BA5FA9AADCC3767F42082" bold="true" pageId="0" pageNumber="1">H, J</emphasis>
epandrium, ventral 
<emphasis id="152BF3070F02E8F03D5ECB9BC428FAA9" bold="true" pageId="0" pageNumber="1">I</emphasis>
ejaculatory apodeme, lateral.
</paragraph>
</caption>
<paragraph id="B9D92AE7D3C694ECAB49FD6C2676417A" pageId="0" pageNumber="1">
<emphasis id="87268B4E1C785C3D82EA98278E68035F" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
<emphasis id="A7F36DC6EF96A01F199BC505C55A8327" bold="true" pageId="0" pageNumber="1">:</emphasis>
Thorax subshiny. Scutum yellow with a black medial stripe on anterior 2/3, one pair of black suborbicular presutural patches confluent with the medial stripe, and a pair of wide black bands on anterior 7/8, which is confluent with the presutural patches. Scutellum yellow with lateral margins brown (Fig. 
<figureCitation id="4C96BAA78FC7DDDFCBBC71E478BE4397" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22D</figureCitation>
). Subscutellum yellow. Mediotergite brown, anatergite yellow, katatergite brown (Fig. 
<figureCitation id="2CAC212BA27355B5CDFD2EC0DE1398CD" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22B</figureCitation>
). Pleuron largely yellow; propleuron with small brown patch on anterior-dorsal corner; anepisternum with a small spot on dorsal corner and a large brown patch near ventral margin; anepimeron with a narrow brown patch on anterior corner; katepisternum and meron with brown patches on venter (Fig. 
<figureCitation id="E89C59BDF2DCBDB37986805111A02302" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22B</figureCitation>
). Haltere yellow. Calypter margin and hairs gray. Leg segments yellow; tibia and tarsus darker. 
<emphasis id="84983053EBD4E630CC8250F0963ABED4" bold="true" italics="true" pageId="0" pageNumber="1">Chaetotaxy</emphasis>
<emphasis id="30C4811E04FE911DE7D511A5275A595F" bold="true" pageId="0" pageNumber="1">:</emphasis>
Scutum with 1+3 dorsocentrals, shortened anteriorly (Fig. 
<figureCitation id="5222DABDD088992FAD01DDEFC4ADD821" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22D</figureCitation>
). Acrostichal setulae eight or nine pairs in two rows. 
<emphasis id="737288AADD74E39660497A52336AD744" bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
<emphasis id="F45535CD01AC99579953C7421F897740" bold="true" pageId="0" pageNumber="1">:</emphasis>
Wing length 2.0-2.2&amp;nbsp;mm, costa reaching M1 (Fig. 
<figureCitation id="CF93D01A0B624F86065519CAEE1856FF" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22A</figureCitation>
). Length of ultimate section of vein M4 divided by penultimate section 1.8-1.9.
</paragraph>
<paragraph id="36ACAAD1D9BC28B06A20768AD1E6024D" pageId="0" pageNumber="1">
<emphasis id="DDA530B679AF41DD01BCC5F6E8C420FB" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
<emphasis id="CB07CED76DD81EFD913B02AF57C2200F" bold="true" pageId="0" pageNumber="1">:</emphasis>
Abdomen dorsally subshiny yellow (Fig. 
<figureCitation id="FFA28432D76E8E613B2A1BD3B98A8AC4" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22A</figureCitation>
). 
<emphasis id="19544A86C0F164C71C426082226C1C74" bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
<emphasis id="518F07FBD54BFB470A617FC37FDCB240" bold="true" pageId="0" pageNumber="1">:</emphasis>
(Fig. 
<figureCitation id="99339FE491B00FBD167678B62C9B623E" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22F-J</figureCitation>
) Epandrium dark brown, rounded apically; inner-lateral margin with a row of 5-7 short tubercle-like setae; inner-anterior surface with a comb comprising three or four fused long tubercle-like setae; inner-lateral surface with an enlarged protrusion bearing a strong tubercle-like seta (Fig. 
<figureCitation id="FD52176E9ECD5A0A6E95DCB62F209941" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22H, J</figureCitation>
). Surstylus rounded and bilobed; anterior lobe sparsely setose apically; posterior lobe with one stout tubercle-like seta and a long normal seta. Cercus narrow, setose. Subepandrial sclerite with a pair of plate-like arms, each with a dorsal lobe bearing a seta (Fig. 
<figureCitation id="B129D689531F8423110A71F91D96BF8A" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22H</figureCitation>
). Hypandrium slightly sclerotized along outer margin. Postgonite bare and goose barnacle-shaped (Fig. 
<figureCitation id="8950EDA5A6B6F5E15D2734B76E30D77B" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22G</figureCitation>
). Phallophorus with deep incision below, articulated with phallapodeme, fused to epiphallus (Fig. 
<figureCitation id="C7249C0727F31CA2EBFB0C7DDC56BBD5" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22F, G</figureCitation>
). Basiphallus with long narrow right sclerites and a short basal sclerite. Hypophallus broad and membranous; lateral margins lightly sclerotized, medially with a pair of fused narrow sclerites (Fig. 
<figureCitation id="0E29A650CFDCD7FEB921CD6CC484CF49" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22G</figureCitation>
). Mesophallus dark, cylindrical, widest basally, as long as distiphallus (Fig. 
<figureCitation id="ECF6CFC40A249D80C4EA2F18A3B9AB3C" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22G</figureCitation>
). Distiphallus comprising one pair of stout tubules; basal 1/3 composed of ventral dark subtriangular sclerite and weaker medial region; distal 2/3 cylindrical, dorsally and laterally pigmented, with truncated, unpigmented apex (Fig. 
<figureCitation id="3C99C91FE0597F7B865526FAF66F5A25" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22G</figureCitation>
). Ejaculatory apodeme pale brown, fan-shaped with long stalk; sclerite of sperm pump with lateral extension; sperm pump clear (Fig. 
<figureCitation id="5A72B00EFAE2E560188F20A56A752FEB" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22I</figureCitation>
).
</paragraph>
<paragraph id="E9F8CAA115B16E3F0CFAE236B6256111" pageId="0" pageNumber="1">
<emphasis id="5CF1823D99D628402FF687A36ED60FB8" bold="true" pageId="0" pageNumber="1">Female</emphasis>
(Fig. 
<figureCitation id="FA05C1EC486502525BDAE3EAE778A3A6" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure22" httpUri="https://binary.pensoft.net/fig/775395" pageId="0" pageNumber="1">22E</figureCitation>
). Similar to male, but larger. Wing length 2.3 mm. 
<emphasis id="7AB2C416464A1F58AAE4C34F77BBE8E6" bold="true" italics="true" pageId="0" pageNumber="1">Postabdomen</emphasis>
<emphasis id="670ABF732F5A2ED9943B07467FBAD704" bold="true" pageId="0" pageNumber="1">:</emphasis>
(Fig. 
<figureCitation id="498BD5DD38A2559AB981A809A7A0C3E3" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female morphology and larval ecology of Phytoliriomyza cometiformis sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C habitat at type locality D-G mined thalli of Reboulia hemisphaerica orientalis (D at type locality E at Ukawa F, G at Sagiura). An arrow in F indicates an internal puparium." figureDoi="10.3897/zookeys.1133.94530.figure23" httpUri="https://binary.pensoft.net/fig/775396" pageId="0" pageNumber="1">23A, B</figureCitation>
) Oviscape dark brown, setigerous (Fig. 
<figureCitation id="A4689933F802C29DAD81E2AFBF8058F9" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female morphology and larval ecology of Phytoliriomyza cometiformis sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C habitat at type locality D-G mined thalli of Reboulia hemisphaerica orientalis (D at type locality E at Ukawa F, G at Sagiura). An arrow in F indicates an internal puparium." figureDoi="10.3897/zookeys.1133.94530.figure23" httpUri="https://binary.pensoft.net/fig/775396" pageId="0" pageNumber="1">23A</figureCitation>
). Tergite 10 trifurcate, laterally uniting narrow pleural sclerites (Fig. 
<figureCitation id="6FA5C6A9EAD7429B4A878F04C1A47221" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female morphology and larval ecology of Phytoliriomyza cometiformis sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C habitat at type locality D-G mined thalli of Reboulia hemisphaerica orientalis (D at type locality E at Ukawa F, G at Sagiura). An arrow in F indicates an internal puparium." figureDoi="10.3897/zookeys.1133.94530.figure23" httpUri="https://binary.pensoft.net/fig/775396" pageId="0" pageNumber="1">23B</figureCitation>
). Each cercus with two stout, apical, trichoid sensilla, 1/3 length of cercus (Fig. 
<figureCitation id="F12E3467D43662A6DE761BDE4C66092F" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female morphology and larval ecology of Phytoliriomyza cometiformis sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C habitat at type locality D-G mined thalli of Reboulia hemisphaerica orientalis (D at type locality E at Ukawa F, G at Sagiura). An arrow in F indicates an internal puparium." figureDoi="10.3897/zookeys.1133.94530.figure23" httpUri="https://binary.pensoft.net/fig/775396" pageId="0" pageNumber="1">23B</figureCitation>
). Spermathecae semi-orbicular, with truncate proximal ends (Fig. 
<figureCitation id="76A14225871C89D992B0072ABE0261C8" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female morphology and larval ecology of Phytoliriomyza cometiformis sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C habitat at type locality D-G mined thalli of Reboulia hemisphaerica orientalis (D at type locality E at Ukawa F, G at Sagiura). An arrow in F indicates an internal puparium." figureDoi="10.3897/zookeys.1133.94530.figure23" httpUri="https://binary.pensoft.net/fig/775396" pageId="0" pageNumber="1">23A</figureCitation>
).
</paragraph>
<caption id="2E33FD477DAA891C1EB2A6FC3D46739D" doi="10.3897/zookeys.1133.94530.figure23" httpUri="https://binary.pensoft.net/fig/775396" pageId="0" pageNumber="1" start="Figure 23" startId="F23">
<paragraph id="DC6082C843D84099F5BC0987C036F8E1" pageId="0" pageNumber="1">
<emphasis id="1F211B70D0B93F082794D2699F22BF95" bold="true" pageId="0" pageNumber="1">Figure 23.</emphasis>
Female morphology and larval ecology of 
<taxonomicName id="980C30D67479B3AEF7B33C8E70F88FAB" authorityName="Kato" authorityYear="2022" class="Insecta" family="Dendrocerotaceae" genus="Phytoliriomyza" kingdom="Animalia" lsidName="Phytoliriomyza cometiformis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cometiformis">
<emphasis id="6F50DD670BCC8E4E691C95A7C8C314C5" italics="true" pageId="0" pageNumber="1">Phytoliriomyza cometiformis</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="3851E2C743442F50A8891292605FA751" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
female postabdomen 
<emphasis id="DADA0B4B795B87D81F4778EA84118371" bold="true" pageId="0" pageNumber="1">A</emphasis>
oviscape and spermatheca 
<emphasis id="341CE8823C9C8AD58BD8C1A06D9068E3" bold="true" pageId="0" pageNumber="1">B</emphasis>
tergite 10 
<emphasis id="D0D1CDB26569B2FF62D809A17A642791" bold="true" pageId="0" pageNumber="1">C</emphasis>
habitat at type locality 
<emphasis id="7624AE028E55EDB3678092E667F3C26C" bold="true" pageId="0" pageNumber="1">D-G</emphasis>
mined thalli of 
<taxonomicName id="D201EDDCF788D1C92153874F4DC3F1CF" class="Marchantiopsida" family="Aytoniaceae" genus="Reboulia" kingdom="Plantae" lsidName="Reboulia hemisphaerica subsp. orientalis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="hemisphaerica" subSpecies="orientalis">
<emphasis id="088F9E59B1AA1C4D2E16539FB688B321" italics="true" pageId="0" pageNumber="1">Reboulia hemisphaerica orientalis</emphasis>
</taxonomicName>
(
<emphasis id="EB2B8AC03564316D1B00D26AE4E61C1F" bold="true" pageId="0" pageNumber="1">D</emphasis>
at type locality 
<emphasis id="685787BDF67CEC5A33E4EE62875046AC" bold="true" pageId="0" pageNumber="1">E</emphasis>
at Ukawa 
<emphasis id="CAD5A75120CB47995B6EA975A5527311" bold="true" pageId="0" pageNumber="1">F, G</emphasis>
at Sagiura). An arrow in 
<emphasis id="769DD543D4365BE90B6AD328DB8A779C" bold="true" pageId="0" pageNumber="1">F</emphasis>
indicates an internal puparium.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8C367BBFBD7CEF252354BD41CA3ABF95" pageId="0" pageNumber="1" type="etymology">
<paragraph id="D5C1E44FEAAA96915486BB42BA2DDE35" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="418DFA697213DB9B5FE1A9FADF1378DD" pageId="0" pageNumber="1">
The specific name (
<emphasis id="AB6003AD2384D25B9DA5025BFFA24C2C" italics="true" pageId="0" pageNumber="1">cometiformis</emphasis>
= comet-shaped) refers to the oblong trail-leaving yellow pattern against the black background on the scutum, which resembles a comet.
</paragraph>
</subSubSection>
<subSubSection id="FEBE921E06905B84E5786F194B043DA3" pageId="0" pageNumber="1" type="japanese name">
<paragraph id="EDDA6CE81C311DA1F1D398B3CA5421E4" pageId="0" pageNumber="1">Japanese name.</paragraph>
<paragraph id="92026C8BB5101BBFC74BDFE02C4A1DC2" pageId="0" pageNumber="1">Suisei-jingasagoke-hamoguribae.</paragraph>
</subSubSection>
<subSubSection id="4B974F75B137B1F07D7C0A8DA0474EB7" pageId="0" pageNumber="1" type="host plant">
<paragraph id="21005C672901AA3604817BE61555B56D" pageId="0" pageNumber="1">Host plant.</paragraph>
<paragraph id="AD1E402952840ED327406C1378F03366" pageId="0" pageNumber="1">
<taxonomicName id="AF04A5E645B09D8649E252C7203B37A2" class="Marchantiopsida" family="Aytoniaceae" genus="Reboulia" kingdom="Plantae" lsidName="Reboulia hemisphaerica subsp. orientalis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="hemisphaerica" subSpecies="orientalis">
<emphasis id="6728F46BF1CE58FF17FBB8A1808ECB0E" italics="true" pageId="0" pageNumber="1">Reboulia hemisphaerica orientalis</emphasis>
</taxonomicName>
(
<taxonomicName id="54DB0776DB83620594654B9CDB388FB0" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="family">Aytoniaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="91244B76314A0C4F52B68583ED1BF7B2" pageId="0" pageNumber="1" type="mine">
<paragraph id="3472F95753A399F86ACE613EEAA2F17C" pageId="0" pageNumber="1">Mine.</paragraph>
<paragraph id="A9216B1E155D66FE6224F59BD07948C2" pageId="0" pageNumber="1">
Larvae construct linear-blotch mines in the thallus, and pupate in the mines (Fig. 
<figureCitation id="8563CB531DE6A892C840DF1A94CF0C2F" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female morphology and larval ecology of Phytoliriomyza cometiformis sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C habitat at type locality D-G mined thalli of Reboulia hemisphaerica orientalis (D at type locality E at Ukawa F, G at Sagiura). An arrow in F indicates an internal puparium." figureDoi="10.3897/zookeys.1133.94530.figure23" httpUri="https://binary.pensoft.net/fig/775396" pageId="0" pageNumber="1">23D-G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2843C8701D100229A7DE48FF501F4B3" pageId="0" pageNumber="1" type="biological notes">
<paragraph id="D4E0271CA0AFF70D4623FC431494532E" pageId="0" pageNumber="1">Biological notes.</paragraph>
<paragraph id="7F00B2A8F0ED184BB1CC88C35A44DF9F" pageId="0" pageNumber="1">
The habitats of this species are rocky cliffs in warm temperate evergreen forests (Fig. 
<figureCitation id="43508432AD963EA6FCECE9DD13A58881" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female morphology and larval ecology of Phytoliriomyza cometiformis sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C habitat at type locality D-G mined thalli of Reboulia hemisphaerica orientalis (D at type locality E at Ukawa F, G at Sagiura). An arrow in F indicates an internal puparium." figureDoi="10.3897/zookeys.1133.94530.figure23" httpUri="https://binary.pensoft.net/fig/775396" pageId="0" pageNumber="1">23C</figureCitation>
). Our rearing records suggest that this species is univoltine, and that adults emerge from overwintered pupae in spring.
</paragraph>
</subSubSection>
<subSubSection id="08FBF2ED2D1E206EC9D4DA652F22B776" pageId="0" pageNumber="1" type="distribution">
<paragraph id="41CBC539A3BB9BBB7396B7D9CB655E68" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="C023300336D2C8B78691BCF2727486AD" pageId="0" pageNumber="1">Japan (Honshu, Shikoku).</paragraph>
</subSubSection>
<subSubSection id="F57542CD927F61712F966CA6960609FB" pageId="0" pageNumber="1" type="remarks">
<paragraph id="52C822F82550A13E1AE0E74E49E69311" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="5E25791EC1AD57B26964A7DE485D0275" pageId="0" pageNumber="1">
This species resembles 
<taxonomicName id="BD66914FE427C49E32979903E9695E19" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. igniculus" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="igniculus">
<emphasis id="D1133B9227818D9BF82733580403D9BD" italics="true" pageId="0" pageNumber="1">P. igniculus</emphasis>
</taxonomicName>
and 
<taxonomicName id="0289AF2B26D9A21D9ECAC709D395F818" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. luna" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="luna">
<emphasis id="2A8E5B82E5329381C7BD80E9C9939AC3" italics="true" pageId="0" pageNumber="1">P. luna</emphasis>
</taxonomicName>
in having a pair of black lateral band on scutum and wholly yellow scutellum; it is distinguished from them by the number of tubercle-like setae in a comb of male epandrium (3-4 in 
<taxonomicName id="71B4C1A625CE5A01F2C6DD0925716CF7" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. cometiformis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="cometiformis">
<emphasis id="A7DF7C181AC76BC9F9F759D138FAF7A0" italics="true" pageId="0" pageNumber="1">P. cometiformis</emphasis>
</taxonomicName>
; 5-6 in 
<taxonomicName id="D1C2B5722A56AB62D715783FFAC91878" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. igniculus" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="igniculus">
<emphasis id="0A3EB9075FBAD346F5E85DE3FBCCD8E0" italics="true" pageId="0" pageNumber="1">P. igniculus</emphasis>
</taxonomicName>
; 7-8 in 
<taxonomicName id="2D3351391004064107745688A71E6B26" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. luna" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="luna">
<emphasis id="312437C0D90271E5A90B060C74E1B97D" italics="true" pageId="0" pageNumber="1">P. luna</emphasis>
</taxonomicName>
). The morphology of male epandrium of this species closely resembles that of &quot; 
<taxonomicName id="D0C3F931B5D8766961058110CF696153" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. dorsata" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="dorsata">
<emphasis id="87AA92C19702134B303F32C5D2E387A1" italics="true" pageId="0" pageNumber="1">P. dorsata</emphasis>
</taxonomicName>
&quot; in 
<bibRefCitation id="26DE55EE3070D87885C0DB882FDA0E67" DOI="https://doi.org/10.18655/Agromyzidae.Vol.3" author="Papp, L" journalOrPublisher="Pars Ltd., Hungary. Nagykovacsi" pageId="0" pageNumber="1" refId="B28" refString="Papp, L, Cerny, M, 2017. Agromyzidae (Diptera) of Hungary (Vol. 3). Phytomyzinae II. Pars Ltd., Hungary. Nagykovacsi, DOI: https://doi.org/10.18655/Agromyzidae.Vol.3" title="Agromyzidae (Diptera) of Hungary (Vol. 3). Phytomyzinae II." url="https://doi.org/10.18655/Agromyzidae.Vol.3" year="2017">
Papp and 
<normalizedToken id="25B880A3908A2C69D449C440E069F6A7" originalValue="Černý">Cerny</normalizedToken>
(2017
</bibRefCitation>
: fig. 101E), suggesting that this species is distributed also in Europe.
</paragraph>
</subSubSection>
</treatment>
</document>