327 lines
51 KiB
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327 lines
51 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang & Liang, Hongbin" docDate="2013" docId="224D43D1FFA2C2CB848A4803DECBC180" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Allocota Motschulsky 1859" docType="treatment" docVersion="5" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="48" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="47" updateTime="1668155586682" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Shi, Hongliang</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zhou, Hongzhang</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-04-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>284</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>129</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-284-1</mods:identifier>
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<mods:identifier type="ZooBank">33B15A74746D4A82A865EA1E7E55A9BB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">192EFF970E2BFFDF3161FFE53C4BFFAA</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578323</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152042271" LSID="urn:lsid:plazi:treatment:224D43D1FFA2C2CB848A4803DECBC180" httpUri="http://treatment.plazi.org/id/224D43D1FFA2C2CB848A4803DECBC180" lastPageId="48" lastPageNumber="48" pageId="46" pageNumber="47">
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<subSubSection pageId="46" pageNumber="47" type="nomenclature">
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<paragraph pageId="46" pageNumber="47">
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<pageBreakToken pageId="46" pageNumber="47" start="start">Genus</pageBreakToken>
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<taxonomicName LSID="224D43D1-FFA2-C2CB-848A-4803DECBC180" authority="Motschulsky, 1859" authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Allocota" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Allocota Motschulsky, 1859</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="46" pageNumber="47" type="reference_group">
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<paragraph pageId="46" pageNumber="47">
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<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Allocota</taxonomicName>
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<bibRefCitation author="Motschulsky, V" journalOrPublisher="Etudes Entomologiques" pageId="88" pageNumber="89" pagination="25 - 118" refId="B73" refString="Motschulsky, V, 1859. Entomologie Speciale. Insectes des Indes Orientales, et de Contrees Analogues. 2: de serie. Etudes Entomologiques 8: 25 - 118" title="Entomologie Speciale. Insectes des Indes Orientales, et de Contrees Analogues. 2: de serie." volume="8" year="1859">Motschulsky 1859</bibRefCitation>
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: 29;
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<bibRefCitation author="Chaudoir, BM" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou" pageId="86" pageNumber="87" pagination="188 - 268" refId="B35" refString="Chaudoir, BM, 1877. Genres nouveaux et especes inedites de la famille des Carabiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 52 (1): 188 - 268" title="Genres nouveaux et especes inedites de la famille des Carabiques." volume="52" year="1877">Chaudoir 1877</bibRefCitation>
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: 203;
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<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="87" pageNumber="88" pagination="269 - 579" refId="B58" refString="Jedlicka, A, 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae)." volume="28" year="1963">
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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1963
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</bibRefCitation>
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: 304 (in part);
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<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="87" pageNumber="88" pagination="757 - 799" refId="B62" refString="Kirschenhofer, E, 1996. Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil. Linzer Biologische Beitraege 28 (2): 757 - 799" title="Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil." volume="28" year="1996">Kirschenhofer 1996</bibRefCitation>
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: 763 (in part).
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</paragraph>
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<paragraph pageId="46" pageNumber="47">
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Shi & Zhou & Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Taicona" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Taicona</taxonomicName>
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<bibRefCitation author="Bates, HW" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="85" pageNumber="86" pagination="229 - 322" refId="B23" refString="Bates, HW, 1873. On the Geodephagous Coleoptera of Japan. Fam. Carabidae. The Transactions of the Entomological Society of London 1873 (2): 229 - 322" title="On the Geodephagous Coleoptera of Japan. Fam. Carabidae." volume="1873" year="1873">Bates 1873</bibRefCitation>
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: 314;
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<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="87" pageNumber="88" pagination="269 - 579" refId="B58" refString="Jedlicka, A, 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae)." volume="28" year="1963">
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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1963
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</bibRefCitation>
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: 448;
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<bibRefCitation author="Habu, A" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="87" pageNumber="88" refId="B48" refString="Habu, A, 1967. Fauna Japonica, Carabidae Truncatipennes Group (Insecta: Coleoptera). Biogeographical Society of Japan. Japan, 338 pp." title="Fauna Japonica, Carabidae Truncatipennes Group (Insecta: Coleoptera). Biogeographical Society of Japan. Japan, 338 pp." year="1967">Habu 1967</bibRefCitation>
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: 140.
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<emphasis bold="true" pageId="46" pageNumber="47">Syn. n.</emphasis>
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[Synonym]
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</paragraph>
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</subSubSection>
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<subSubSection pageId="46" pageNumber="47" type="type-species">
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<paragraph pageId="46" pageNumber="47">Type-species:</paragraph>
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<paragraph pageId="46" pageNumber="47">
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<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="viridipennis">
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<emphasis italics="true" pageId="46" pageNumber="47">Allocota viridipennis</emphasis>
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</taxonomicName>
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Motschulsky, 1859, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="46" pageNumber="47" type="diagnosis">
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<paragraph pageId="46" pageNumber="47">Diagnosis.</paragraph>
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<paragraph pageId="46" pageNumber="47">Dorsal side glabrous; mandibles strongly widened; posterior supraorbital setae near eyes, insertions not tumid; pronotum narrow, ratio PW/PL less than 1.4; lateral setae of pronotum absent; elytral 3rd and 5th intervals with four or more setigerous pores, subequally placed; 7th intervals sometimes with setigerous pores; protibiae with cleaning spur reduced or absent; males with two pairs of setigerous pores on terminal sternum.</paragraph>
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<paragraph pageId="46" pageNumber="47">
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This genus is closest to
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Shi & Zhou & Liang" baseAuthorityYear="1829" class="Insecta" family="Carabidae" genus="Physodera" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="46" pageNumber="47">Physodera</emphasis>
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</taxonomicName>
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; their differences are presented in the diagnosis of
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Shi & Zhou & Liang" baseAuthorityYear="1829" class="Insecta" family="Carabidae" genus="Physodera" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="46" pageNumber="47">Physodera</emphasis>
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</taxonomicName>
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. In general appearance,
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<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="46" pageNumber="47">Allocota</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Diamella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="46" pageNumber="47">Diamella</emphasis>
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</taxonomicName>
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, but can be distinguished from the latter by the glabrous surface and vertex not tumid.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="generic characters">
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<paragraph pageId="46" pageNumber="47">Generic characters.</paragraph>
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<paragraph lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
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Body length 6.0 to 8.7 mm, slender, pronotum narrow. Elytra strongly metallic, sometimes disc more or less reddish. Head and pronotum glabrous, elytra glabrous except primary setigerous pores; elytra usually with faint isodiametric microsculpture.
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<emphasis bold="true" pageId="46" pageNumber="47">Head</emphasis>
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glabrous; eyes hemispherical, strongly prominent; tempora slightly longer than half length of eyes, gradually narrowed behind eyes, not expanded; vertex not tumid. Antennae extended to about elytral basal one-third; 1st antennomere gradually narrowed to base, 3rd longer than 4th. Labrum smooth, without secondary setae, slightly widened to apex, apical margin more or less emarginate; mandibles strongly widened, outer margin rounded (
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<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="46" pageNumber="47">Fig. 150</figureCitation>
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), surface glabrous; terminal maxillary palpomeres fusiform in both sexes; terminal labial palpomeres more or less widened in males, fusiform, truncate or slightly securiform; less widened in females; ligular apex truncate, with four long setae; paraglossae membranous, as long as ligula, adnate; mentum tooth simple, apex rounded or sharp, with two setae near base; submentum with two long setae; genae glabrous beneath eyes.
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<emphasis bold="true" pageId="46" pageNumber="47">Pronotum</emphasis>
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slightly wider than long, nearly as wide as head; disc glabrous, front angles with a few setae, lateral margins glabrous; mid-lateral setae absent; pronotal base briefly but distinctly lobed; lateral margins slightly expanded in middle; hind angles more or less distinct; basal foveae shallow.
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<emphasis bold="true" pageId="46" pageNumber="47">Elytra</emphasis>
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narrow, slightly convex, slightly widened to apex; apex truncate or slightly curved, sutural angles not projected, outer angles rounded; disc without distinct depression; intervals flat, without additional pubescence; basal margination only reaching 3rd interval; basal pores present; 3rd and 5th intervals with four or more setigerous pores, sometimes 7th interval also with setigerous pores; 7th and 8th intervals strongly tumid near apex.
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<emphasis bold="true" pageId="46" pageNumber="47">Ventral side</emphasis>
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nearly glabrous; males with terminal sternum moderately emarginate apically, with two pairs of setae (
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<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="46" pageNumber="47">Fig. 145</figureCitation>
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); females with terminal sternum straight or slightly curved apically, with two pairs of setae, rarely one additional setae present on one side (the right side in
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<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="46" pageNumber="47">Fig. 146</figureCitation>
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).
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<emphasis bold="true" pageId="46" pageNumber="47">Legs</emphasis>
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short; protibiae with cleaning spur absent (
|
||
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="46" pageNumber="47">Fig. 140</figureCitation>
|
||
), or very fine and short (
|
||
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="46" pageNumber="47">Fig. 141</figureCitation>
|
||
),
|
||
<pageBreakToken pageId="47" pageNumber="48" start="start">distant</pageBreakToken>
|
||
from inner margin; tarsi widened; 4th tarsomere bifid, claws pectinate; males with adhesive hairs on first three protarsomeres well developed (two whole rows), those on first three mesotarsomeres rudimentary (two rows, weakly present near apex) or well developed (two whole rows) on some mesotarsomeres.
|
||
<emphasis bold="true" pageId="47" pageNumber="48">Male genitalia</emphasis>
|
||
with median lobe of aedeagus not twisted, slightly bent to right side in dorsal view; apical orifice opened apically; apical lamella short; dorsal surface with some fine setae subapically; internal sac with main flagellum moderately thick, apex not reaching apical orifice, trumpet-form expansion small; apical bursa absent; secondary flagellum distinct.
|
||
<emphasis bold="true" pageId="47" pageNumber="48">Female genitalia.</emphasis>
|
||
Spermatheca tubular, with more or less ring-sculpture, inserted at the joining of the bursa copulatrix and common oviduct; spermathecal gland with basal part very fine, inserted at middle of spermatheca; spermatheca not distinctly bent. Apical segment of ovipositor very short and wide, apex rounded, with membranous extension slightly widened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="47" pageNumber="48" type="distribution">
|
||
<paragraph pageId="47" pageNumber="48">Distribution</paragraph>
|
||
<paragraph pageId="47" pageNumber="48">
|
||
(
|
||
<figureCitation captionStart="Map 6" captionStartId="F50" captionText="Map 6. Known distribution of genus Allocota: ● Allocota aurata (Bates) ★ Allocota cyanipennis Heller ▲ Allocota viridipennis Motschulsky ◆ Allocota bicolor sp. n." httpUri="https://binary.pensoft.net/fig/29728" pageId="47" pageNumber="48">Map 6</figureCitation>
|
||
). East Asia and Southeast Asia: Japan, China, Indo-China Peninsula, Malay Peninsula, Philippine Islands, Borneo, Sumatra, Java, Sulawesi. Not discovered in South Asia or the eastern part of the Malay Archipelago.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="47" pageNumber="48" type="monophyly and relationships">
|
||
<paragraph pageId="47" pageNumber="48">Monophyly and relationships.</paragraph>
|
||
<paragraph pageId="47" pageNumber="48">
|
||
Monophyly of
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota</emphasis>
|
||
</taxonomicName>
|
||
is suggested by the following apomorphic character states: (1) cleaning spur on protibiae reduced or absent; (2) setigerous pores on 5th interval not restricted to basal half; (3) apical segment of ovipositor only slightly longer than basal width.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="taxonomic comments">
|
||
<paragraph pageId="47" pageNumber="48">Taxonomic comments.</paragraph>
|
||
<paragraph pageId="47" pageNumber="48">
|
||
We studied the type series and other material of
|
||
<taxonomicName authorityName="Bates" authorityYear="1873" class="Insecta" family="Carabidae" genus="Taicona" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="aurata">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Taicona aurata</emphasis>
|
||
</taxonomicName>
|
||
Bates (type-species of
|
||
<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Shi & Zhou & Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Taicona" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Taicona</emphasis>
|
||
</taxonomicName>
|
||
), and a male of
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="viridipennis">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota viridipennis</emphasis>
|
||
</taxonomicName>
|
||
Motschulsky (type-species of
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota</emphasis>
|
||
</taxonomicName>
|
||
) from Java (type locality) that had been compared with the type by Andrewes. Except for color and body size, the only significant difference between them is that setigerous pores are present on the 7th interval in
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota</emphasis>
|
||
</taxonomicName>
|
||
. Even the male genitalia show no obvious differences at the species level (see the detailed discussion under
|
||
<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="aurata">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota aurata</emphasis>
|
||
</taxonomicName>
|
||
). So it can be inferred that these two species are extremely closely related. As the pore distribution on the 7th interval is not constant even at the species-level (see discussion under
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="viridipennis">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota viridipennis</emphasis>
|
||
</taxonomicName>
|
||
), we herein synonymize
|
||
<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Shi & Zhou & Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Taicona" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Taicona</emphasis>
|
||
</taxonomicName>
|
||
Bates with
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota</emphasis>
|
||
</taxonomicName>
|
||
Motschulsky.
|
||
</paragraph>
|
||
<paragraph pageId="47" pageNumber="48">
|
||
A total of seven species was included in
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota</emphasis>
|
||
</taxonomicName>
|
||
before the present study, but four of them should be transferred to other genera. In the present paper, we combine species previously included in
|
||
<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Shi & Zhou & Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Taicona" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Taicona</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota</emphasis>
|
||
</taxonomicName>
|
||
, propose two new synonyms, and describe a new species. Therefore, in the present concept of
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota</emphasis>
|
||
</taxonomicName>
|
||
, a total of four species is included. Three of these are very closely allied with each other, and strictly allopatric; while the fourth one,
|
||
<taxonomicName authorityName="Shi & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota bicolor</emphasis>
|
||
</taxonomicName>
|
||
sp. n., is quite different from the others and sympatric with
|
||
<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="aurata">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota aurata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="47" pageNumber="48">
|
||
<emphasis bold="true" pageId="47" pageNumber="48">
|
||
Key to species of
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota</emphasis>
|
||
</taxonomicName>
|
||
Motschulsky
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
|
||
<table inLine="true" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
|
||
<tr pageId="47" pageNumber="48">
|
||
<td colspan="1" pageId="47" pageNumber="48" rowspan="1">1</td>
|
||
<td colspan="1" pageId="47" pageNumber="48" rowspan="1">
|
||
Protibiae with cleaning spur present, but very fine (
|
||
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="47" pageNumber="48">Fig. 141</figureCitation>
|
||
); males with terminal labial palpomeres moderately widened, apex truncate; males with adhesive hairs rudimentary on first three mesotarsomeres; 7th interval of elytra with setigerous pores; internal sac of aedeagus without setose area
|
||
</td>
|
||
<td colspan="1" pageId="47" pageNumber="48" rowspan="1">
|
||
<taxonomicName authorityName="Shi & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis italics="true" pageId="47" pageNumber="48">Allocota bicolor</emphasis>
|
||
</taxonomicName>
|
||
sp. n.
|
||
</td>
|
||
</tr>
|
||
<tr pageId="48" pageNumber="49">
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">
|
||
<pageBreakToken pageId="48" pageNumber="49" start="start">-</pageBreakToken>
|
||
</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">
|
||
Protibiae with cleaning spur absent (
|
||
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="48" pageNumber="49">Fig. 140</figureCitation>
|
||
); males with terminal labial palpomeres slightly widened in middle, not truncate; males with adhesive hairs well developed on first two mesotarsomeres, rudimentary on 3rd mesotarsomeres; 7th interval of elytra usually without setigerous pores (except some specimens from Java and Borneo); internal sac of aedeagus with setose area near middle
|
||
</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">2</td>
|
||
</tr>
|
||
<tr pageId="48" pageNumber="49">
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">2</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">Elytra greenish to cupreous; metasternum and abdomen not distinctly darker than prosternum; species from Japan, Taiwan, and Asian mainland</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">
|
||
<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="aurata">
|
||
<emphasis italics="true" pageId="48" pageNumber="49">Allocota aurata</emphasis>
|
||
</taxonomicName>
|
||
(Bates)
|
||
</td>
|
||
</tr>
|
||
<tr pageId="48" pageNumber="49">
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">-</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">Elytra more or less bluish; metasternum and abdomen much darker than prosternum; species from Malay Archipelago and Malay Peninsula</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">3</td>
|
||
</tr>
|
||
<tr pageId="48" pageNumber="49">
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">3</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">
|
||
Setae on front angles of pronotum very short (
|
||
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="48" pageNumber="49">Fig. 151</figureCitation>
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); internal sac of aedeagus with setose area divided into two parts, apex of secondary flagellum simple; species from Malaya, Borneo, Sumatra, Java
|
||
</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="viridipennis">
|
||
<emphasis italics="true" pageId="48" pageNumber="49">Allocota viridipennis</emphasis>
|
||
</taxonomicName>
|
||
Motschulsky
|
||
</td>
|
||
</tr>
|
||
<tr pageId="48" pageNumber="49">
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">-</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">
|
||
Setae on front angles of pronotum much longer (
|
||
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="48" pageNumber="49">Fig. 152</figureCitation>
|
||
); internal sac of aedeagus with setose area divided into three parts, apex of secondary flagellum forming a large triangular sclerite; species from the Philippines and Sulawesi
|
||
</td>
|
||
<td colspan="1" pageId="48" pageNumber="49" rowspan="1">
|
||
<taxonomicName authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="cyanipennis">
|
||
<emphasis italics="true" pageId="48" pageNumber="49">Allocota cyanipennis</emphasis>
|
||
</taxonomicName>
|
||
Heller
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |