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<document id="A51011EDBC510D866FDBCEC088ED7541" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFF4CFF12FDD9FC0FA954E948" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia scorpiocaulis Moonlight &amp; Tebbitt" docType="treatment" docVersion="7" lastPageNumber="236" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="234" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="A238D753F0F0EA10FADC7BF9708308D4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="590205A55268DA7E3558EBC29BEF6DDA">
<mods:title id="6A8059B475218A63192466DA659CA70A">The genus Begonia (Begoniaceae) in Peru</mods:title>
</mods:titleInfo>
<mods:name id="792AF5220280E4DB5ECCE9B6ED93F757" type="personal">
<mods:role id="D7A954A95AA6E9C3856006A0AD67E990">
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<mods:namePart id="8E3731E0A813EB3C7C284C1A1E50FBA1">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="A151AA701AF8962E3CF3D2B1BE0FAACD">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
<mods:nameIdentifier id="EF2B20B2B5BC6CC0F9B3FC861A3A0A6E" type="email">moonligp@tcd.ie</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="2183F4A841FAF0F6E780B4D433C3F19C">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="4EE0AED192D5801B9CDD89989E5AD24C">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="DC6E1BC632E0CC80B12FA160A1ACAE12" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="8647CA11D93AC5803A1672E68E0B9FAC" type="personal">
<mods:role id="D04A977F923775BF16AE670357BFEF3F">
<mods:roleTerm id="66BC6B362F0A2F4E69EAC3EA0EFEB5DA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA11C18718B3D580ADBEBD9FFC20657D">Purvis, David A.</mods:namePart>
<mods:affiliation id="DA2B3C566FDB4A6F8683E34201F26144">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="CDB51E53357D0B5F3A50C7EF9E26EA8D" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="62527265127E748AB430153D1E09A930" type="personal">
<mods:role id="DAC8A3F03698FA04476D2CAA6428FE2A">
<mods:roleTerm id="306F77E8B136E0ACFB671C7AE9842150">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F570A9D1992BA77B049BF3D67B67C3E7">Delves, Jay</mods:namePart>
<mods:affiliation id="C5B7EB1D52585E8833D7713295229333">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="6C98A68F42B49E9F90CC9D4B1B1E7F69" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="D587B4925249B14A096F2437AB211C84" type="personal">
<mods:role id="966A20A1A416BE941BD3256DABB3CBF2">
<mods:roleTerm id="62633B3B0FE68300A47BC2AAC2E69F1D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC1077010795255E8753ED2063150C3C">Allen, Josh P.</mods:namePart>
<mods:affiliation id="4EB1FAEEBC4B4196BDB5CF62AA449FF4">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="2DAB66CF8AAC2282703FB9E3E735CFB3" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="F22092355C16A5EA731E01EBDF5D39A8">Reynel, Carlos</mods:namePart>
<mods:affiliation id="13C1B1A9656A90A9A6A99686DDC21447">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="23EECA9908AEFFB56FF63756EFA45BBF" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A49C8F421F5B2E178CB9E7A6C37D39DD">text</mods:typeOfResource>
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<mods:title id="911F6181C2B99C8D8298870477ED9357">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="E4F29604D6E2A3327D191BF43A07440B">
<mods:date id="7DE40E9D9F7A91B23E9E326498255722">2023</mods:date>
<mods:detail id="0B757D1BD415E21A8940AA7E3704515C" type="pubDate">
<mods:number id="2130EA924F73A8FC9551D6935722F689">2023-07-18</mods:number>
</mods:detail>
<mods:detail id="8CA85B62F70BEF3EA95B26A9B2030697" type="volume">
<mods:number id="AD003217899C28B8AB060D367F3842E9">881</mods:number>
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<mods:start id="7F45C26FBCEEFD1100D9FFEC4A6C79D8">1</mods:start>
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<mods:url id="611DA94B9FE9228C46A1A6882EE8C2C1">http://dx.doi.org/10.5852/ejt.2023.881.2175</mods:url>
</mods:location>
<mods:classification id="67FF064388705F313E816E0ACB3CD013">journal article</mods:classification>
<mods:identifier id="BA73A5748400288E2FEFFC632C95A23A" type="DOI">https://doi.org/10.5852/ejt.2023.881.2175</mods:identifier>
<mods:identifier id="0A64CB254FE724D415B69C03B81A6BA8" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="9C93E2602734E7F07F046D7EEFCED17B" type="Zenodo-Dep">8178280</mods:identifier>
</mods:mods>
<treatment id="03B26B4BFF4CFF12FDD9FC0FA954E948" LSID="urn:lsid:plazi:treatment:03B26B4BFF4CFF12FDD9FC0FA954E948" httpUri="http://treatment.plazi.org/id/03B26B4BFF4CFF12FDD9FC0FA954E948" lastPageId="235" lastPageNumber="236" pageId="233" pageNumber="234">
<subSubSection id="C30189D6FF4CFF10FDD9FC0FA914E9B3" box="[512,1075,943,971]" pageId="233" pageNumber="234" type="nomenclature">
<paragraph id="8BA4DA5DFF4CFF10FDD9FC0FA914E9B3" blockId="233.[512,1075,943,1007]" box="[512,1075,943,971]" pageId="233" pageNumber="234">
59.
<taxonomicName id="4C1BA1DEFF4CFF10FDF5FC0FA914E9B3" ID-CoL="7SHG4" authority="Moonlight &amp; Tebbitt" authorityName="Moonlight &amp; Tebbitt" box="[556,1075,943,970]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="233" pageNumber="234" phylum="Tracheophyta" rank="species" species="scorpiocaulis">
<emphasis id="B96F064FFF4CFF10FDF5FC0FAE1EE9B3" bold="true" box="[556,825,943,970]" italics="true" pageId="233" pageNumber="234">Begonia scorpiocaulis</emphasis>
Moonlight &amp; Tebbitt
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF4CFF10FD17FC6BAE41E996" box="[718,870,980,1007]" pageId="233" pageNumber="234" type="description">
<paragraph id="8BA4DA5DFF4CFF10FD17FC6BAE41E996" blockId="233.[512,1075,943,1007]" box="[718,870,980,1007]" pageId="233" pageNumber="234">
<figureCitation id="1320C6D8FF4CFF10FD17FC6BAE1BE997" box="[718,828,980,1006]" captionStart="Fig" captionStartId="224.[189,232,1857,1883]" captionTargetBox="[194,1392,1193,1812]" captionTargetId="figure-357@224.[189,1398,1183,1818]" captionTargetPageId="224" captionText="Fig. 71. Distribution of the maynensis group of Begonia sect. Knesebeckia (Klotzsch) A.DC. in Peru and surrounding countries. A. B. albomaculata C.DC. (red), B. chemillenensis Moonlight (blue) and B. scorpiocaulisMoonlight &amp; Tebbitt (green). B. B. maynensis A.DC. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178452" httpUri="https://zenodo.org/record/8178452/files/figure.png" pageId="233" pageNumber="234">Figs 71A</figureCitation>
,
<figureCitation id="1320C6D8FF4CFF10FC90FC6BAE41E996" box="[841,870,980,1007]" captionStart="Fig" captionStartId="234.[189,232,1892,1918]" captionTargetBox="[302,1285,285,1815]" captionTargetId="figure-14@234.[287,1300,265,1855]" captionTargetPageId="234" captionText="Fig. 75. Begonia scorpiocaulis Moonlight &amp; Tebbitt. A. Habit. B. Pistillate flower, side view. C. Bracteole. D. Bract. E. Pistil, front view. F. Pistil, back view. Illustration by Claire Banks from A.H. Gentry 27367 (MO). Reproduced from Moonlight et al. (2017b), with the permission of Edinburgh Journal of Botany." figureDoi="http://doi.org/10.5281/zenodo.8178462" httpUri="https://zenodo.org/record/8178462/files/figure.png" pageId="233" pageNumber="234">75</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF4CFF10FF64FBA9AECDEE49" box="[189,1002,1046,1073]" pageId="233" pageNumber="234" type="reference_group">
<paragraph id="8BA4DA5DFF4CFF10FF64FBA9AECDEE49" blockId="233.[189,1399,1046,1182]" box="[189,1002,1046,1073]" pageId="233" pageNumber="234">
<emphasis id="B96F064FFF4CFF10FF64FBA9AF38EE49" box="[189,543,1046,1073]" italics="true" pageId="233" pageNumber="234">Edinburgh Journal of Botany</emphasis>
74 (2): 185 (
<bibRefCitation id="EF8AA7ACFF4CFF10FD18FBA9AEF9EE49" author="Moonlight" box="[705,990,1046,1072]" etAl="et al." firstAuthor="Moonlight" pageId="233" pageNumber="234" pagination="179 - 198" refId="ref229967" refString="Moonlight P. W., Hughes M. &amp; Tebbitt M. C. 2017 b. Taxonomy of Begonia albomaculata and description of two new species endemic to Peru. Edinburgh Journal of Botany 74 (2): 179 - 198. https: // doi. org / 10.1017 / S 0960428617000075" type="journal article" year="2017" yearSuffix="b">
Moonlight
<emphasis id="B96F064FFF4CFF10FC93FBA8AEAFEE49" box="[842,904,1046,1072]" italics="true" pageId="233" pageNumber="234">et al.</emphasis>
2017b
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF4CFF10FC2AFBA8ACFBEEE7" pageId="233" pageNumber="234" type="materials_examined">
<paragraph id="8BA4DA5DFF4CFF10FC2AFBA8ACFBEEE7" blockId="233.[189,1399,1046,1182]" pageId="233" pageNumber="234">
–
<emphasis id="B96F064FFF4CFF10FBD7FBA8A96DEE48" bold="true" box="[1038,1098,1047,1073]" pageId="233" pageNumber="234">Type</emphasis>
:
<collectingCountry id="F30C9ACDFF4CFF10FB82FBA8A981EE48" box="[1115,1190,1047,1073]" name="Peru" pageId="233" pageNumber="234">PERU</collectingCountry>
•
<materialsCitation id="3B73D000FF4CFF10FB1DFBA9ACF2EEE7" collectingDate="1979-10-29" collectionCode="MO" collectorName="A. H. Gentry &amp; J. Terborgh &amp; J. Aronson &amp; R. Ramirez" elevation="660" latitude="-12.65" location="serrania across rio alto Madre de Dios from Shintuya" longLatPrecision="1293" longitude="-71.28333" municipality="Pantiacolla" pageId="233" pageNumber="234" specimenCode="MO-2154624" specimenCount="1" stateProvince="Madre de Dios Region" typeStatus="holotype">
<collectingRegion id="49DF14BFFF4CFF10FB1DFBA9AC61EE2C" country="Peru" name="Madre de Dios" pageId="233" pageNumber="234">Madre de Dios Region</collectingRegion>
:
<collectingMunicipality id="6BC04027FF4CFF10FE8AFB84ACFFEE2C" box="[339,472,1083,1109]" pageId="233" pageNumber="234">Pantiacolla</collectingMunicipality>
,
<location id="8EC48C86FF4CFF10FE3FFB84A940EE2C" LSID="urn:lsid:plazi:treatment:03B26B4BFF4CFF12FDD9FC0FA954E948:8EC48C86FF4CFF10FE3FFB84A940EE2C" box="[486,1127,1083,1109]" latitude="-12.65" longLatPrecision="1293" longitude="-71.28333" municipality="Pantiacolla" name="serrania across rio alto Madre de Dios from Shintuya" pageId="233" pageNumber="234" stateProvince="Madre de Dios Region">serranía across río alto Madre de Dios from Shintuya</location>
; [
<geoCoordinate id="EE2FBC9AFF4CFF10FBA5FB86A9C3EE2C" box="[1148,1252,1081,1109]" degrees="12" direction="south" minutes="39" orientation="latitude" pageId="233" pageNumber="234" precision="925" value="-12.65">12°39′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF4CFF10FB2AFB86A84DEE2C" box="[1267,1386,1081,1109]" degrees="71" direction="west" minutes="17" orientation="longitude" pageId="233" pageNumber="234" precision="925" value="-71.28333">71°17′ W</geoCoordinate>
];
<elevation id="00363D6EFF4CFF10FF33FBE0AC8BEE00" box="[234,428,1119,1145]" metricMagnitude="2" metricUnit="m" metricValue="6.6" metricValueMax="8.4" metricValueMin="4.8" pageId="233" pageNumber="234" unit="m" value="660.0" valueMax="840.0" valueMin="480.0">
<quantity id="4CE377B8FF4CFF10FF33FBE0AC49EE00" box="[234,366,1119,1145]" metricMagnitude="2" metricUnit="m" metricValue="6.6" metricValueMax="8.4" metricValueMin="4.8" pageId="233" pageNumber="234" unit="m" value="660.0" valueMax="840.0" valueMin="480.0">480–840 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF4CFF10FE61FBE0AF77EE00" box="[440,592,1119,1145]" pageId="233" pageNumber="234" value="1979-10-29">
<collectingDate id="EFE10575FF4CFF10FE61FBE0AF77EE00" box="[440,592,1119,1145]" pageId="233" pageNumber="234" value="1979-10-29">29 Oct. 1979</collectingDate>
</date>
;
<emphasis id="B96F064FFF4CFF10FD80FBE0A9DFEE00" box="[601,1272,1119,1145]" italics="true" pageId="233" pageNumber="234">
<collectorName id="26EEBF8BFF4CFF10FD80FBE0AFCEEE00" box="[601,745,1119,1145]" pageId="233" pageNumber="234">A.H. Gentry</collectorName>
,
<collectorName id="26EEBF8BFF4CFF10FD2AFBDFAE5AEE00" box="[755,893,1119,1145]" pageId="233" pageNumber="234">J. Terborgh</collectorName>
,
<collectorName id="26EEBF8BFF4CFF10FC5EFBDFA923EE00" box="[903,1028,1119,1145]" pageId="233" pageNumber="234">J. Aronson</collectorName>
&amp;
<collectorName id="26EEBF8BFF4CFF10FBFEFBDFA98EEE00" box="[1063,1193,1120,1145]" pageId="233" pageNumber="234">R. Ramirez</collectorName>
27367
</emphasis>
;
<typeStatus id="54A064FFFF4CFF10FAD1FBE0A854EE00" box="[1288,1395,1119,1145]" pageId="233" pageNumber="234">holotype</typeStatus>
:
<collectionCode id="ED0A4298FF4CFF10FF30FB3CAC3FEEE4" box="[233,280,1155,1181]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="233" pageNumber="234" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FF4CFF10FEF2FB3CACEDEEE7" box="[299,458,1155,1182]" pageId="233" pageNumber="234">MO-2154624</specimenCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF4CFF10FF64FB79A9C2EF76" box="[189,1253,1222,1296]" pageId="233" pageNumber="234" type="etymology">
<paragraph id="8BA4DA5DFF4CFF10FF64FB79AC62EE99" blockId="233.[189,325,1222,1248]" box="[189,325,1222,1248]" pageId="233" pageNumber="234">
<heading id="D0EC6D31FF4CFF10FF64FB79AC62EE99" bold="true" box="[189,325,1222,1248]" fontSize="11" level="3" pageId="233" pageNumber="234" reason="3">
<emphasis id="B96F064FFF4CFF10FF64FB79AC62EE99" bold="true" box="[189,325,1222,1248]" pageId="233" pageNumber="234">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF4CFF10FF64FB4AA9C2EF76" blockId="233.[189,1253,1269,1296]" box="[189,1253,1269,1296]" pageId="233" pageNumber="234">Named for the contorted stem of the species, which resembles the coiled tail of a scorpion.</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF4CFF12FF64FA88AC15EB3E" lastPageId="235" lastPageNumber="236" pageId="233" pageNumber="234" type="description">
<paragraph id="8BA4DA5DFF4CFF10FF64FA88AC68EF28" blockId="233.[189,335,1335,1361]" box="[189,335,1335,1361]" pageId="233" pageNumber="234">
<heading id="D0EC6D31FF4CFF10FF64FA88AC68EF28" bold="true" box="[189,335,1335,1361]" fontSize="11" level="3" pageId="233" pageNumber="234" reason="3">
<emphasis id="B96F064FFF4CFF10FF64FA88AC68EF28" bold="true" box="[189,335,1335,1361]" pageId="233" pageNumber="234">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF4CFF12FF64FAD8AC15EB3E" blockId="233.[189,1401,1383,2025]" lastBlockId="235.[189,1399,265,327]" lastPageId="235" lastPageNumber="236" pageId="233" pageNumber="234">
Caulescent herb, to
<quantity id="4CE377B8FF4CFF10FE6CFAD8AF25EFFB" box="[437,514,1383,1410]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="233" pageNumber="234" unit="cm" value="20.0">20 cm</quantity>
high.
<emphasis id="B96F064FFF4CFF10FD8FFAD8AFA8EFF8" box="[598,655,1383,1409]" italics="true" pageId="233" pageNumber="234">Stem</emphasis>
erect, unbranched; internodes to
<quantity id="4CE377B8FF4CFF10FBF6FAD8A949EFFB" box="[1071,1134,1383,1410]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="233" pageNumber="234" unit="cm" value="2.0">2 cm</quantity>
long at the base and progressively shorter towards the apex, to
<quantity id="4CE377B8FF4CFF10FD11FA34AE37EFDF" box="[712,784,1419,1446]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="233" pageNumber="234" unit="mm" value="5.0">5 mm</quantity>
thick, succulent, brown, glabrous.
<emphasis id="B96F064FFF4CFF10FB19FA34A839EFDC" box="[1216,1310,1419,1445]" italics="true" pageId="233" pageNumber="234">Stipules</emphasis>
tardily deciduous, lanceolate, 4–10 ×
<quantity id="4CE377B8FF4CFF10FDF9FA0FAFA3EFB3" box="[544,644,1456,1482]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="233" pageNumber="234" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
, apex acuminate to mucronate, opaque, brown, glabrous, margin entire, aciliate.
<emphasis id="B96F064FFF4CFF10FEACFA6BACE0EF94" box="[373,455,1492,1517]" italics="true" pageId="233" pageNumber="234">Leaves</emphasis>
&gt; 3, alternate, basifixed; petiole
<quantity id="4CE377B8FF4CFF10FC94FA6BAE82EF97" box="[845,933,1492,1518]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="233" pageNumber="234" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 cm</quantity>
long, colour unknown, glabrous; blade asymmetrical, elliptic to oblanceolate, to 12.5 ×
<quantity id="4CE377B8FF4CFF10FCDDFA47AE7DEC6B" box="[772,858,1528,1555]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="233" pageNumber="234" unit="cm" value="3.5">3.5 cm</quantity>
, succulent, apex acuminate, base rounded on the broad side of the lamina, cuneate on the narrow side, margin irregularly dentate, ciliate, upper colour unknown, glabrous, lower surface colour unknown, glabrous, veins palmate-pinnate, with 6–8 veins from the base, with 3–4 secondary veins on the broad side of the lamina, 1–3 on the narrow side.
<emphasis id="B96F064FFF4CFF10FF64F937AC42ECDB" box="[189,357,1672,1698]" italics="true" pageId="233" pageNumber="234">Inflorescences</emphasis>
at least one per stem, presumed bisexual, axillary, erect, cymose, with up to 4 branches, presumed to bear up to 8 staminate flowers and 8 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FF4CFF10FAE0F912A850ECBE" box="[1337,1399,1709,1735]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="233" pageNumber="234" unit="cm" value="6.0">6 cm</quantity>
long, colour unknown, glabrous, bracts late deciduous, narrowly-lanceolate, ca 3 ×
<quantity id="4CE377B8FF4CFF10FB43F96EA9C5EC95" box="[1178,1250,1745,1772]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="233" pageNumber="234" unit="mm" value="1.0">1 mm</quantity>
, translucent, colour unknown, glabrous, acute, margin denticulate, ciliate.
<emphasis id="B96F064FFF4CFF10FC7BF949A95EED76" box="[930,1145,1781,1808]" italics="true" pageId="233" pageNumber="234">Staminate flowers</emphasis>
: unknown.
<emphasis id="B96F064FFF4CFF10FAD2F94AAC35ED4A" italics="true" pageId="233" pageNumber="234">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF4CFF10FE6AF8A6AF2DED4D" box="[435,522,1817,1844]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="233" pageNumber="234" unit="mm" value="15.0">15 mm</quantity>
long; bracteoles 3, directly beneath the ovary, lanceolate to ovate, ca 5 ×
<quantity id="4CE377B8FF4CFF10FF01F881AC05ED21" box="[216,290,1854,1880]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="233" pageNumber="234" unit="mm" value="3.0">3 mm</quantity>
, translucent, colour known, glabrous, apex acute to obtuse, margin serrate, ciliate; tepals 5, subequal, persistent in fruit, spreading, lanceolate to ovate, 4–7 ×
<quantity id="4CE377B8FF4CFF10FC39F8DDA960ED05" box="[992,1095,1890,1916]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="233" pageNumber="234" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, apex obtuse to rounded, orange, glabrous, margin entire, aciliate; ovary body ellipsoid, ca 6 ×
<quantity id="4CE377B8FF4CFF10FC32F839A915EDD9" box="[1003,1074,1926,1952]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="233" pageNumber="234" unit="mm" value="4.0">4 mm</quantity>
, colour unknown, glabrous, unequally 3-winged, wings triangular, the largest 7–9 ×
<quantity id="4CE377B8FF4CFF10FC87F815AEC5EDBC" box="[862,994,1962,1989]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="233" pageNumber="234" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
, the smallest two 6–8 ×
<quantity id="4CE377B8FF4CFF10FAD2F815A856EDBC" box="[1291,1393,1962,1989]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="233" pageNumber="234" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free, ca
<quantity id="4CE377B8FF4CFF10FAEDF871A85EED90" box="[1332,1401,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="233" pageNumber="234" unit="mm" value="5.0">5 mm</quantity>
long, once-divided, stigmatic papillae in a spirally twisted band.
<emphasis id="B96F064FFF4EFF12FC3FFEB5A98BEB5A" box="[998,1196,265,291]" italics="true" pageId="235" pageNumber="236">Fruiting pedicel</emphasis>
unknown.
<emphasis id="B96F064FFF4EFF12FAE3FEB5A850EB5A" box="[1338,1399,266,291]" italics="true" pageId="235" pageNumber="236">Fruit</emphasis>
unknown.
</paragraph>
</subSubSection>
<caption id="DF648AD5FF4FFF13FF64F8DBACBDED90" ID-DOI="http://doi.org/10.5281/zenodo.8178462" ID-Zenodo-Dep="8178462" httpUri="https://zenodo.org/record/8178462/files/figure.png" pageId="234" pageNumber="235" startId="234.[189,232,1892,1918]" targetBox="[302,1285,285,1815]" targetPageId="234">
<paragraph id="8BA4DA5DFF4FFF13FF64F8DBACBDED90" blockId="234.[189,1399,1892,2025]" pageId="234" pageNumber="235">
<emphasis id="B96F064FFF4FFF13FF64F8DBAC04ED06" bold="true" box="[189,291,1892,1919]" pageId="234" pageNumber="235">Fig. 75.</emphasis>
<taxonomicName id="4C1BA1DEFF4FFF13FEEAF8D9A9AFED06" authority="Moonlight &amp; Tebbitt. A. Habit. B. Pistillate" box="[307,1160,1892,1919]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="234" pageNumber="235" phylum="Tracheophyta" rank="species" species="scorpiocaulis">
<emphasis id="B96F064FFF4FFF13FEEAF8D9AF61ED06" box="[307,582,1893,1919]" italics="true" pageId="234" pageNumber="235">Begonia scorpiocaulis</emphasis>
Moonlight &amp; Tebbitt.
<emphasis id="B96F064FFF4FFF13FCA8F8DBAEA1ED07" bold="true" box="[881,902,1892,1918]" pageId="234" pageNumber="235">A</emphasis>
. Habit.
<emphasis id="B96F064FFF4FFF13FC2FF8DAA92DED06" bold="true" box="[1014,1034,1893,1919]" pageId="234" pageNumber="235">B</emphasis>
. Pistillate
</taxonomicName>
flower, side view.
<emphasis id="B96F064FFF4FFF13FF64F837ADF5EDDB" bold="true" box="[189,210,1928,1954]" pageId="234" pageNumber="235">C</emphasis>
. Bracteole.
<emphasis id="B96F064FFF4FFF13FEB9F837AC52EDDB" bold="true" box="[352,373,1928,1954]" pageId="234" pageNumber="235">D</emphasis>
. Bract.
<emphasis id="B96F064FFF4FFF13FE08F837ACC2EDDB" bold="true" box="[465,485,1928,1954]" pageId="234" pageNumber="235">E</emphasis>
. Pistil, front view.
<emphasis id="B96F064FFF4FFF13FD1DF837AFF1EDDB" bold="true" box="[708,726,1928,1954]" pageId="234" pageNumber="235">F</emphasis>
. Pistil, back view. Illustration by Claire Banks from
<emphasis id="B96F064FFF4FFF13FA99F837AC43EDBC" italics="true" pageId="234" pageNumber="235">A.H. Gentry 27367</emphasis>
(MO). Reproduced from
<bibRefCitation id="EF8AA7ACFF4FFF13FD46F814AEE9EDBC" author="Moonlight" box="[671,974,1963,1989]" etAl="et al." firstAuthor="Moonlight" pageId="234" pageNumber="235" pagination="179 - 198" refId="ref229967" refString="Moonlight P. W., Hughes M. &amp; Tebbitt M. C. 2017 b. Taxonomy of Begonia albomaculata and description of two new species endemic to Peru. Edinburgh Journal of Botany 74 (2): 179 - 198. https: // doi. org / 10.1017 / S 0960428617000075" type="journal article" year="2017" yearSuffix="b">
Moonlight
<emphasis id="B96F064FFF4FFF13FCFFF813AE43EDBC" box="[806,868,1963,1989]" italics="true" pageId="234" pageNumber="235">et al.</emphasis>
(2017b)
</bibRefCitation>
, with the permission of
<emphasis id="B96F064FFF4FFF13FB20F814ACB4ED91" italics="true" pageId="234" pageNumber="235">Edinburgh Journal of Botany</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C30189D6FF4EFF12FF64FECFAEB2EBC3" pageId="235" pageNumber="236" type="conservation">
<paragraph id="8BA4DA5DFF4EFF12FF64FECFAF4AEBF3" blockId="235.[189,621,368,394]" box="[189,621,368,394]" pageId="235" pageNumber="236">
<heading id="D0EC6D31FF4EFF12FF64FECFAF4AEBF3" bold="true" box="[189,621,368,394]" fontSize="11" level="3" pageId="235" pageNumber="236" reason="3">
<emphasis id="B96F064FFF4EFF12FF64FECFAF4AEBF3" bold="true" box="[189,621,368,394]" pageId="235" pageNumber="236">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF4EFF12FF64FE1FAEB2EBC3" blockId="235.[189,917,416,443]" box="[189,917,416,443]" pageId="235" pageNumber="236">
Assessed by
<bibRefCitation id="EF8AA7ACFF4EFF12FE8CFE1FAF50EBC3" author="Moonlight" box="[341,631,416,442]" etAl="et al." firstAuthor="Moonlight" pageId="235" pageNumber="236" pagination="179 - 198" refId="ref229967" refString="Moonlight P. W., Hughes M. &amp; Tebbitt M. C. 2017 b. Taxonomy of Begonia albomaculata and description of two new species endemic to Peru. Edinburgh Journal of Botany 74 (2): 179 - 198. https: // doi. org / 10.1017 / S 0960428617000075" type="journal article" year="2017" yearSuffix="b">
Moonlight
<emphasis id="B96F064FFF4EFF12FE01FE1EAF34EBC3" box="[472,531,416,442]" italics="true" pageId="235" pageNumber="236">et al.</emphasis>
(2017b)
</bibRefCitation>
as Data Deficient (DD).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF4EFF12FF64FE5CAC76E828" pageId="235" pageNumber="236" type="discussion">
<paragraph id="8BA4DA5DFF4EFF12FF64FE5CAC97EB84" blockId="235.[189,432,483,509]" box="[189,432,483,509]" pageId="235" pageNumber="236">
<heading id="D0EC6D31FF4EFF12FF64FE5CAC97EB84" bold="true" box="[189,432,483,509]" fontSize="11" level="3" pageId="235" pageNumber="236" reason="3">
<emphasis id="B96F064FFF4EFF12FF64FE5CAC97EB84" bold="true" box="[189,432,483,509]" pageId="235" pageNumber="236">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF4EFF12FF64FDABAC76E828" blockId="235.[189,1398,531,594]" pageId="235" pageNumber="236">
<taxonomicName id="4C1BA1DEFF4EFF12FF64FDABACE0E854" box="[189,455,531,557]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="235" pageNumber="236" phylum="Tracheophyta" rank="species" species="scorpiocaulis">
<emphasis id="B96F064FFF4EFF12FF64FDABACE0E854" box="[189,455,531,557]" italics="true" pageId="235" pageNumber="236">Begonia scorpiocaulis</emphasis>
</taxonomicName>
is the only
<taxonomicName id="4C1BA1DEFF4EFF12FD94FDABAE2BE854" box="[589,780,531,557]" pageId="235" pageNumber="236">
<emphasis id="B96F064FFF4EFF12FD94FDABAF97E854" box="[589,688,532,557]" italics="true" pageId="235" pageNumber="236">Begonia</emphasis>
species
</taxonomicName>
from Andean South America that has yellow flowers but no tuber.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF4EFF12FF64FDC5A954E948" pageId="235" pageNumber="236" type="distribution">
<paragraph id="8BA4DA5DFF4EFF12FF64FDC5ACD2E8ED" blockId="235.[189,501,634,660]" box="[189,501,634,660]" pageId="235" pageNumber="236">
<heading id="D0EC6D31FF4EFF12FF64FDC5ACD2E8ED" bold="true" box="[189,501,634,660]" fontSize="11" level="3" pageId="235" pageNumber="236" reason="3">
<emphasis id="B96F064FFF4EFF12FF64FDC5ACD2E8ED" bold="true" box="[189,501,634,660]" pageId="235" pageNumber="236">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF4EFF12FF64FD15A954E948" blockId="235.[187,1399,682,818]" pageId="235" pageNumber="236">
Endemic to
<collectingCountry id="F30C9ACDFF4EFF12FE91FD15AC59E8BD" box="[328,382,682,708]" name="Peru" pageId="235" pageNumber="236">Peru</collectingCountry>
and known only from the
<typeStatus id="54A064FFFF4EFF12FD72FD15AFF9E8BD" box="[683,734,682,708]" pageId="235" pageNumber="236">type</typeStatus>
collection collected in
<collectingRegion id="49DF14BFFF4EFF12FC30FD15A9C8E8BD" box="[1001,1263,682,708]" country="Peru" name="Madre de Dios" pageId="235" pageNumber="236">Madre de Dios Region</collectingRegion>
(
<figureCitation id="1320C6D8FF4EFF12FB24FD15A84EE8BD" box="[1277,1385,682,708]" captionStart="Fig" captionStartId="224.[189,232,1857,1883]" captionTargetBox="[194,1392,1193,1812]" captionTargetId="figure-357@224.[189,1398,1183,1818]" captionTargetPageId="224" captionText="Fig. 71. Distribution of the maynensis group of Begonia sect. Knesebeckia (Klotzsch) A.DC. in Peru and surrounding countries. A. B. albomaculata C.DC. (red), B. chemillenensis Moonlight (blue) and B. scorpiocaulisMoonlight &amp; Tebbitt (green). B. B. maynensis A.DC. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178452" httpUri="https://zenodo.org/record/8178452/files/figure.png" pageId="235" pageNumber="236">Fig. 71A</figureCitation>
). Found at an altitude of
<quantity id="4CE377B8FF4EFF12FE03FD70AF44E890" box="[474,611,719,745]" metricMagnitude="2" metricUnit="m" metricValue="6.6" metricValueMax="8.4" metricValueMin="4.8" pageId="235" pageNumber="236" unit="m" value="660.0" valueMax="840.0" valueMin="480.0">480–840 m</quantity>
a.s.l. in an area of Amazonian or lower montane forest.
<taxonomicName id="4C1BA1DEFF4EFF12FACDFD70AC7AE974" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="235" pageNumber="236" phylum="Tracheophyta" rank="species" species="scorpiocaulis">
<emphasis id="B96F064FFF4EFF12FACDFD70AC7AE974" italics="true" pageId="235" pageNumber="236">Begonia scorpiocaulis</emphasis>
</taxonomicName>
was collected as an epiphyte, which is surprising as its closest relatives
<taxonomicName id="4C1BA1DEFF4EFF12FB72FD4BA848E974" box="[1195,1391,755,781]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="235" pageNumber="236" phylum="Tracheophyta" rank="species" species="albomaculata">
<emphasis id="B96F064FFF4EFF12FB72FD4BA848E974" box="[1195,1391,755,781]" italics="true" pageId="235" pageNumber="236">B. albomaculata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1BA1DEFF4EFF12FF64FCA7ACAAE948" box="[189,397,791,817]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="235" pageNumber="236" phylum="Tracheophyta" rank="species" species="chemillenensis">
<emphasis id="B96F064FFF4EFF12FF64FCA7ACAAE948" box="[189,397,791,817]" italics="true" pageId="235" pageNumber="236">B. chemillenensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1BA1DEFF4EFF12FE14FCA7AF4EE948" box="[461,617,792,817]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="235" pageNumber="236" phylum="Tracheophyta" rank="species" species="maynensis">
<emphasis id="B96F064FFF4EFF12FE14FCA7AF4EE948" box="[461,617,792,817]" italics="true" pageId="235" pageNumber="236">B. maynensis</emphasis>
</taxonomicName>
are principally collected as terrestrial herbs.
</paragraph>
</subSubSection>
</treatment>
</document>
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