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<document id="69499FEFDEC159249BD4DFBEE2F20464" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFE86FEDDFDDEFE33A851E854" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia erythrothrix Tebbitt &amp; Moonlight" docType="treatment" docVersion="7" lastPageNumber="293" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="292" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="9606990AE8EF60B9D96FA1A8991C6470" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3D66FC53B0A95E5A2A88451583CBD3C7">
<mods:title id="368D29C8E28FA2776F58240631FE8B83">The genus Begonia (Begoniaceae) in Peru</mods:title>
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<mods:namePart id="F6964691B44751493690E9253BFC2389">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="8352E5625268DFF563113D6ADFF1399C">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
<mods:nameIdentifier id="08351A913377E56A35079AE2119FFACD" type="email">moonligp@tcd.ie</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="F9F0F8F0CAC9E010D0D39687F7D017B7">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="CA4B1169565E66A1D4E29D056CAFF592">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="8A7E9A0988A6C12E1778D890CC08D015" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="A2C22BA28E2B4E920A1F833C57E58C79">Purvis, David A.</mods:namePart>
<mods:affiliation id="B0F14E1E9F1EDB2DF14B9BD37A2EFF12">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="9B6A464F02B7D1468244A92EE82D0FD0" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
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<mods:role id="A0767E369DF7599BDF2C7A3DD7B6D7C9">
<mods:roleTerm id="9F5D64DA7E38061189055587F2C33159">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="551F7630E38F8D99D203895DD50AEF7B">Delves, Jay</mods:namePart>
<mods:affiliation id="2F3D3DAFB16DAE65180584ED5DC2F305">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="AC217D2E5288E9FC0F19711CE4AC0BA4" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
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<mods:namePart id="1CD6A1386FF6373BBCD07A57FCB28ADC">Allen, Josh P.</mods:namePart>
<mods:affiliation id="EEFB49F8DE49BFF8FCCD804509FEEACF">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="0A5B8549AD9869E9449FDD802997EFE7" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4DEF74C8356FD45B73079022C1184E20">Reynel, Carlos</mods:namePart>
<mods:affiliation id="9EECA78CA2320B578DDA15786C510487">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="54E06D90DC219BEF8EB4DC636C4B74CD" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
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<mods:title id="B246F7147C0D7C9CDC3880148D6BA4DB">European Journal of Taxonomy</mods:title>
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<mods:part id="D387FF9C84B509811E446EAA8B267B58">
<mods:date id="0E351A0E0BFBF8AA9316C354E03E7250">2023</mods:date>
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<mods:number id="D0FD3F664DF8460D8897447A9BB8E7A5">2023-07-18</mods:number>
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<mods:number id="B74F127431A21FC5A83E9648F8F69A0D">881</mods:number>
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<mods:url id="A8536E82D2348238951DF1FAC8CD2BA1">http://dx.doi.org/10.5852/ejt.2023.881.2175</mods:url>
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<mods:classification id="4EFED9C12B3E079F19E1808F3456C736">journal article</mods:classification>
<mods:identifier id="955B0DDD52226BA1B34D8BF1A53ABA4F" type="DOI">https://doi.org/10.5852/ejt.2023.881.2175</mods:identifier>
<mods:identifier id="651AFB790DCB56025DA34B7A7871B83B" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="B0BE165B15214B7D8EECE2BB6136F32F" type="Zenodo-Dep">8178280</mods:identifier>
</mods:mods>
<treatment id="03B26B4BFE86FEDDFDDEFE33A851E854" LSID="urn:lsid:plazi:treatment:03B26B4BFE86FEDDFDDEFE33A851E854" httpUri="http://treatment.plazi.org/id/03B26B4BFE86FEDDFDDEFE33A851E854" lastPageId="292" lastPageNumber="293" pageId="291" pageNumber="292">
<subSubSection id="C30189D6FE86FEDAFDDEFE33A90AEBDF" box="[519,1069,395,422]" pageId="291" pageNumber="292" type="nomenclature">
<paragraph id="8BA4DA5DFE86FEDAFDDEFE33A90AEBDF" blockId="291.[519,1069,395,461]" box="[519,1069,395,422]" pageId="291" pageNumber="292">
72.
<taxonomicName id="4C1BA1DEFE86FEDAFDEAFE33A90AEBDF" ID-CoL="7SFJY" authority="Tebbitt &amp; Moonlight" authorityName="Tebbitt &amp; Moonlight" box="[563,1069,395,422]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="291" pageNumber="292" phylum="Tracheophyta" rank="species" species="erythrothrix">
<emphasis id="B96F064FFE86FEDAFDEAFE33AE15EBDF" bold="true" box="[563,818,395,422]" italics="true" pageId="291" pageNumber="292">Begonia erythrothrix</emphasis>
Tebbitt &amp; Moonlight
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE86FEDAFD3FFE0CA910E86B" pageId="291" pageNumber="292" type="description">
<paragraph id="8BA4DA5DFE86FEDAFD3FFE0CAE69EBB4" blockId="291.[519,1069,395,461]" box="[742,846,435,461]" pageId="291" pageNumber="292">
<figureCitation id="1320C6D8FE86FEDAFD3FFE0CAE69EBB4" box="[742,846,435,461]" captionStart="Fig" captionStartId="288.[189,232,1857,1883]" captionTargetBox="[194,1390,1193,1812]" captionTargetId="figure-313@288.[189,1398,1183,1818]" captionTargetPageId="288" captionText="Fig. 90. Distribution of the Begonia sect. Ignota in Peru and surrounding countries. A. B. bifurcata L.B.Sm. &amp; B.G.Schub. (red), B. erythrothrix Tebbitt &amp; Moonlight (red), and B. occultata J.P.Allen &amp; Moonlight sp. nov. (green). B. B. speculum Moonlight &amp; Tebbitt (red), B. thyrsoidea Irmsch., and B. urubambensis Tebbitt (green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178495" httpUri="https://zenodo.org/record/8178495/files/figure.png" pageId="291" pageNumber="292">Fig. 90A</figureCitation>
</paragraph>
<paragraph id="8BA4DA5DFE86FEDAFF64FE47A910E86B" blockId="291.[189,1399,504,687]" box="[189,1079,504,530]" pageId="291" pageNumber="292">
<emphasis id="B96F064FFE86FEDAFF64FE47AF0CE86B" box="[189,555,504,530]" italics="true" pageId="291" pageNumber="292">Edinburgh Journal of Botany</emphasis>
74 (2): 116 (
<bibRefCitation id="EF8AA7ACFE86FEDAFD05FE47A90CE86B" author="Moonlight &amp; Tebbitt" box="[732,1067,504,530]" firstAuthor="Moonlight" pageId="291" pageNumber="292" pagination="111 - 122" refId="ref229847" refString="Moonlight P. W. &amp; Tebbitt M. C. 2016. Two new Peruvian species of Begonia (Begoniaceae) and an amended description of Begonia thyrsoidea. Edinburgh Journal of Botany 74 (2): 111 - 122. https: // doi. org / 10.1017 / S 0960428616000299" type="journal article" year="2016">Moonlight &amp; Tebbitt 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="8BA4DA5DFE86FEDAFB9CFE46AC15E8D6" blockId="291.[189,1399,504,687]" pageId="291" pageNumber="292">
–
<emphasis id="B96F064FFE86FEDAFBB8FE47A9B8E86B" bold="true" box="[1121,1183,504,530]" pageId="291" pageNumber="292">Type</emphasis>
:
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<collectingCountry id="F30C9ACDFE86FEDAFB6DFE46A9D8E86A" box="[1204,1279,505,531]" name="Peru" pageId="291" pageNumber="292">PERU</collectingCountry>
–
<emphasis id="B96F064FFE86FEDAFAF0FE47AF79E840" bold="true" pageId="291" pageNumber="292">
<collectingRegion id="49DF14BFFE86FEDAFAF0FE47AC60E840" country="Peru" name="Cusco" pageId="291" pageNumber="292">Cusco Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FE86FEDAFE8DFDA0AF79E840" box="[340,606,542,569]" pageId="291" pageNumber="292">Prov. La Convención</collectingCounty>
</emphasis>
•
<collectingMunicipality id="6BC04027FE86FEDAFDAFFDA0AE07E840" box="[630,800,543,569]" pageId="291" pageNumber="292">Dist. Echarate</collectingMunicipality>
,
<location id="8EC48C86FE86FEDAFCF2FD9FAEC5E840" LSID="urn:lsid:plazi:treatment:03B26B4BFE86FEDDFDDEFE33A851E854:8EC48C86FE86FEDAFCF2FD9FAEC5E840" box="[811,994,543,570]" country="Peru" county="Prov. La Convencion" latitude="-12.85" longLatPrecision="1292" longitude="-73.5" municipality="Dist. Echarate" name="E rio Apurimac" pageId="291" pageNumber="292" stateProvince="Cusco Region">E río Apurimac</location>
,
<locationDeviation id="9A16C37FFE86FEDAFC37FD9FA99FE840" box="[1006,1208,543,570]" pageId="291" pageNumber="292">NE Pueblo Libre</locationDeviation>
,
<locationDeviation id="9A16C37FFE86FEDAFB1AFD9FAF2AE819" pageId="291" pageNumber="292">up mountain of Anchihuay &amp; Bellavista</locationDeviation>
,
<location id="8EC48C86FE86FEDAFDC2FDF9AE54E819" LSID="urn:lsid:plazi:treatment:03B26B4BFE86FEDDFDDEFE33A851E854:8EC48C86FE86FEDAFDC2FDF9AE54E819" box="[539,883,582,608]" country="Peru" county="Prov. La Convencion" latitude="-12.85" longLatPrecision="1292" longitude="-73.5" municipality="Dist. Echarate" name="south Cordillera Vilcabamba" pageId="291" pageNumber="292" stateProvince="Cusco Region">south Cordillera Vilcabamba</location>
;
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,
<geoCoordinate id="EE2FBC9AFE86FEDAFC21FDFAA94BE818" box="[1016,1132,581,609]" degrees="73" direction="west" minutes="30" orientation="longitude" pageId="291" pageNumber="292" precision="925" value="-73.5">73°30′ W</geoCoordinate>
;
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<quantity id="4CE377B8FE86FEDAFBA3FDF9A9F2E818" box="[1146,1237,582,609]" metricMagnitude="3" metricUnit="m" metricValue="2.445" pageId="291" pageNumber="292" unit="m" value="2445.0">2445 m</quantity>
a.s.l.
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;
<date id="FFA5FC9DFE86FEDAFAFFFDF8AC0FE8F1" pageId="291" pageNumber="292" value="1998-08-03">
<collectingDate id="EFE10575FE86FEDAFAFFFDF8AC0FE8F1" pageId="291" pageNumber="292" value="1998-08-03">3 Aug. 1998</collectingDate>
</date>
;
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,
<collectorName id="26EEBF8BFE86FEDAFE73FDD1AF7BE8FE" box="[426,604,621,647]" pageId="291" pageNumber="292">
R. de la
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</collectorName>
&amp;
<collectorName id="26EEBF8BFE86FEDAFD58FDD1AE22E8FE" box="[641,773,621,648]" pageId="291" pageNumber="292">S. Udvardy</collectorName>
23400
</emphasis>
;
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:
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;
</materialsCitation>
<materialsCitation id="3B73D000FE86FEDAFBFCFDD2AC15E8D6" collectingDate="1998-08-03" collectionCode="US, USM" collectorName="P. Nunez &amp; R. de la Colina &amp; S. Udvardy" country="Peru" county="Prov. La Convencion" elevation="2445" latitude="-12.85" location="E rio Apurimac" longLatPrecision="1292" longitude="-73.5" municipality="Dist. Echarate" pageId="291" pageNumber="292" specimenCode="US00625233" specimenCount="1" stateProvince="Cusco Region" typeStatus="isotypes">
<typeStatus id="54A064FFFE86FEDAFBFCFDD2A9ADE8FE" box="[1061,1162,621,647]" pageId="291" pageNumber="292">isotypes</typeStatus>
:
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[
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],
<collectionCode id="ED0A4298FE86FEDAFF33FD2AAC0AE8D6" box="[234,301,661,687]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="291" pageNumber="292">USM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE86FEDAFF64FD65A81BE95E" box="[189,1340,730,807]" pageId="291" pageNumber="292" type="etymology">
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<heading id="D0EC6D31FE86FEDAFF64FD65AC62E88D" bold="true" box="[189,325,730,756]" fontSize="11" level="3" pageId="291" pageNumber="292" reason="3">
<emphasis id="B96F064FFE86FEDAFF64FD65AC62E88D" bold="true" box="[189,325,730,756]" pageId="291" pageNumber="292">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE86FEDAFF64FCB3A81BE95E" blockId="291.[189,1340,780,807]" box="[189,1340,780,807]" pageId="291" pageNumber="292">
The epithet derives from the Greek words ‘
<emphasis id="B96F064FFE86FEDAFD61FCB3AE29E95F" box="[696,782,780,806]" italics="true" pageId="291" pageNumber="292">erythro</emphasis>
’ and ‘
<emphasis id="B96F064FFE86FEDAFC82FCB3AEB4E95F" box="[859,915,780,806]" italics="true" pageId="291" pageNumber="292">thrix</emphasis>
’, meaning the ‘red haired’
<taxonomicName id="4C1BA1DEFE86FEDAFB0BFCB2A812E95F" baseAuthorityName="Moonlight &amp; Fuentes" baseAuthorityYear="2022" box="[1234,1333,781,806]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="291" pageNumber="292" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFE86FEDAFB0BFCB2A812E95F" box="[1234,1333,781,806]" italics="true" pageId="291" pageNumber="292">Begonia</emphasis>
</taxonomicName>
.
</paragraph>
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<subSubSection id="C30189D6FE86FEDAFF64FCEEAE96E9BC" pageId="291" pageNumber="292" type="materials_examined">
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<emphasis id="B96F064FFE86FEDAFF64FCEEAC9FE912" bold="true" box="[189,440,849,875]" pageId="291" pageNumber="292">Specimen examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE86FEDAFF64FC3BAE96E9BC" blockId="291.[189,1399,898,966]" pageId="291" pageNumber="292">
<materialsCitation id="3B73D000FE86FEDAFF64FC3BAE96E9BC" collectingDate="1968-06-29" collectionCode="US" collectorName="T. R. Dudley" country="Peru" county="Prov. La Convencion" latitude="-12.633333" location="Knox's cascade" longLatPrecision="1293" longitude="-73.05" pageId="291" pageNumber="292" specimenCode="US01925935" specimenCount="1" stateProvince="Cusco Region">
<collectingCountry id="F30C9ACDFE86FEDAFF64FC3BAC2FE9E7" box="[189,264,900,926]" name="Peru" pageId="291" pageNumber="292">PERU</collectingCountry>
–
<emphasis id="B96F064FFE86FEDAFEFEFC3BAFD6E9E7" bold="true" box="[295,753,899,926]" pageId="291" pageNumber="292">
<collectingRegion id="49DF14BFFE86FEDAFEFEFC3BACFEE9E7" box="[295,473,900,926]" country="Peru" name="Cusco" pageId="291" pageNumber="292">Cusco Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FE86FEDAFE3FFC3BAFD6E9E7" box="[486,753,899,926]" pageId="291" pageNumber="292">Prov. La Convención</collectingCounty>
</emphasis>
•
<location id="8EC48C86FE86FEDAFCD3FC3BAEE3E9E7" LSID="urn:lsid:plazi:treatment:03B26B4BFE86FEDDFDDEFE33A851E854:8EC48C86FE86FEDAFCD3FC3BAEE3E9E7" box="[778,964,900,926]" country="Peru" county="Prov. La Convencion" latitude="-12.633333" longLatPrecision="1293" longitude="-73.05" name="Knox's cascade" pageId="291" pageNumber="292" stateProvince="Cusco Region">Knox’s cascade</location>
,
<locationDeviation id="9A16C37FFE86FEDAFC08FC3BA9D9E9E7" box="[977,1278,898,926]" pageId="291" pageNumber="292">
ca
<quantity id="4CE377B8FE86FEDAFC2AFC3BA909E9E7" box="[1011,1070,900,926]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="291" pageNumber="292" unit="km" value="2.0">2 km</quantity>
NW of camp 2½
</locationDeviation>
;
<geoCoordinate id="EE2FBC9AFE86FEDAFAD3FC3DA856E9E7" box="[1290,1393,898,926]" degrees="12" direction="south" minutes="38" orientation="latitude" pageId="291" pageNumber="292" precision="925" value="-12.633333">12°38′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFE86FEDAFF64FC14AC08E9BC" box="[189,303,937,965]" degrees="73" direction="west" minutes="03" orientation="longitude" pageId="291" pageNumber="292" precision="925" value="-73.05">73°03′ W</geoCoordinate>
;
<date id="FFA5FC9DFE86FEDAFEE2FC14ACF2E9BC" box="[315,469,939,965]" pageId="291" pageNumber="292" value="1968-06-29">
<collectingDate id="EFE10575FE86FEDAFEE2FC14ACF2E9BC" box="[315,469,939,965]" pageId="291" pageNumber="292" value="1968-06-29">29 Jun. 1968</collectingDate>
</date>
;
<emphasis id="B96F064FFE86FEDAFE38FC13AF98E9BC" box="[481,703,939,965]" italics="true" pageId="291" pageNumber="292">
<collectorName id="26EEBF8BFE86FEDAFE38FC13AF4AE9BC" box="[481,621,939,965]" pageId="291" pageNumber="292">T.R. Dudley</collectorName>
10618
</emphasis>
;
<collectionCode id="ED0A4298FE86FEDAFD17FC14AFD4E9BC" box="[718,755,939,965]" pageId="291" pageNumber="292">US</collectionCode>
[
<specimenCode id="DBBD7226FE86FEDAFCDCFC14AE87E9BF" box="[773,928,939,966]" pageId="291" pageNumber="292">US01925935</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE86FEDAFF64FC4FA850ED08" pageId="291" pageNumber="292" type="description">
<paragraph id="8BA4DA5DFE86FEDAFF64FC4FAC68EE73" blockId="291.[189,335,1008,1034]" box="[189,335,1008,1034]" pageId="291" pageNumber="292">
<heading id="D0EC6D31FE86FEDAFF64FC4FAC68EE73" bold="true" box="[189,335,1008,1034]" fontSize="11" level="3" pageId="291" pageNumber="292" reason="3">
<emphasis id="B96F064FFE86FEDAFF64FC4FAC68EE73" bold="true" box="[189,335,1008,1034]" pageId="291" pageNumber="292">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE86FEDAFF64FB9DA850ED08" blockId="291.[187,1400,1058,1905]" pageId="291" pageNumber="292">
Acaulescent, rhizomatous herb, to
<quantity id="4CE377B8FE86FEDAFDBFFB9DAF93EE44" box="[614,692,1058,1085]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="291" pageNumber="292" unit="cm" value="15.0">15 cm</quantity>
high.
<emphasis id="B96F064FFE86FEDAFCDEFB9DAE4AEE45" box="[775,877,1058,1084]" italics="true" pageId="291" pageNumber="292">Rhizome</emphasis>
ellipsoid, 0.1–0.8 ×
<quantity id="4CE377B8FE86FEDAFBA8FB9CA994EE44" box="[1137,1203,1059,1085]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="291" pageNumber="292" unit="cm" value="4.0">4 cm</quantity>
, with numerous growing points.
<emphasis id="B96F064FFE86FEDAFE58FBF5ACF8EE1D" box="[385,479,1098,1124]" italics="true" pageId="291" pageNumber="292">Stipules</emphasis>
persistent, ovate, 0.3–1 ×
<quantity id="4CE377B8FE86FEDAFCC5FBF5AE89EE1D" box="[796,942,1098,1124]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="291" pageNumber="292" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 mm</quantity>
, apex acute, opaque, brown, glabrous, margin entire, aciliate.
<emphasis id="B96F064FFE86FEDAFE01FBCEAF0DEEF3" box="[472,554,1137,1162]" italics="true" pageId="291" pageNumber="292">Leaves</emphasis>
1–4, alternate, basifixed; petiole
<quantity id="4CE377B8FE86FEDAFC1DFBCFA97DEEF2" box="[964,1114,1136,1163]" metricMagnitude="-2" metricUnit="m" metricValue="5.85" metricValueMax="10.5" metricValueMin="1.2" pageId="291" pageNumber="292" unit="cm" value="5.85" valueMax="10.5" valueMin="1.2">1.2–10.5 cm</quantity>
long, colour unknown, densely hispid, hairs red; blade asymmetrical, ovate, to 11 ×
<quantity id="4CE377B8FE86FEDAFC7FFB27AEC1EECB" box="[934,998,1176,1202]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="291" pageNumber="292" unit="cm" value="6.0">6 cm</quantity>
, succulent, apex acuminate, base cordate, basal lobes not overlapping, sinus to
<quantity id="4CE377B8FE86FEDAFD16FB00AE06EEA0" box="[719,801,1215,1241]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="291" pageNumber="292" unit="mm" value="10.0">10 mm</quantity>
deep, margin serrate, ciliate, upper surface yellow-green, pink to red on the veins, sparsely pubescent, lower surface red-purple maculate, sparsely to moderately pubescent, veins palmate, ca 6 veined from the base.
<emphasis id="B96F064FFE86FEDAFC65FAB2A943EF5E" box="[956,1124,1293,1319]" italics="true" pageId="291" pageNumber="292">Inflorescences</emphasis>
1–3, bisexual, axillary, erect, an asymmetrical cyme, with 4 branches, bearing up to 4 staminate flowers and 4 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FE86FEDAFE23FAE4AF79EF0F" box="[506,606,1371,1398]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="291" pageNumber="292" unit="cm" value="11.5">11.5 cm</quantity>
long, colour unknown, moderately glandular hispid, bracts late deciduous, ovate, ca 3 ×
<quantity id="4CE377B8FE86FEDAFE36FA3DAF70EFE4" box="[495,599,1410,1437]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="291" pageNumber="292" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
, opaque, colour unknown, glabrous, apex acute, apiculate, margin entire, aciliate.
<emphasis id="B96F064FFE86FEDAFEB7FA16AF66EFBA" box="[366,577,1449,1475]" italics="true" pageId="291" pageNumber="292">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FE86FEDAFD0DFA16AE02EFBD" box="[724,805,1449,1476]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="291" pageNumber="292" unit="mm" value="18.0">18 mm</quantity>
long, glabrous; tepals 4, spreading, outer 2 elliptic, 0.6–2 ×
<quantity id="4CE377B8FE86FEDAFEFDFA6FAC84EF92" box="[292,419,1488,1515]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="291" pageNumber="292" unit="mm" value="0.65" valueMax="1.0" valueMin="0.3">0.3–1 mm</quantity>
, apex obtuse, white tinged pink or red, glabrous, margin entire, aciliate, inner 2 obovate, 0.6–2.2 ×
<quantity id="4CE377B8FE86FEDAFE7CFA48AF10EC6B" box="[421,567,1527,1554]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="291" pageNumber="292" unit="mm" value="0.75" valueMax="1.2" valueMin="0.3">0.3–1.2 mm</quantity>
, apex obtuse, white, glabrous, margin entire, aciliate; stamens 15–30, projecting, yellow, filaments
<quantity id="4CE377B8FE86FEDAFDC7F9A1AFA7EC40" box="[542,640,1566,1593]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="291" pageNumber="292" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, fused into a
<quantity id="4CE377B8FE86FEDAFCB2F9A1AEE8EC40" box="[875,975,1566,1593]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="291" pageNumber="292" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long column, anthers sub-globose, 0.5–0.75 ×
<quantity id="4CE377B8FE86FEDAFE9BF9FAACBCEC19" box="[322,411,1605,1632]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="291" pageNumber="292" unit="mm" value="0.5">0.5 mm</quantity>
long, dehiscing via lateral slits, connectives not projecting, symmetrically basifixed.
<emphasis id="B96F064FFE86FEDAFF64F9D3ACA1ECFF" box="[189,390,1644,1670]" italics="true" pageId="291" pageNumber="292">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FE86FEDAFDF9F9D3AF5CECFE" box="[544,635,1644,1671]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="291" pageNumber="292" unit="mm" value="1.8">1.8 mm</quantity>
long; bracteoles lacking; tepals 5, subequal, deciduouss in fruit, spreading, the largest ovate to elliptic, 6.5–13 ×
<quantity id="4CE377B8FE86FEDAFD29F92BAE73ECD7" box="[752,852,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="291" pageNumber="292" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
, apex subacute, white, glabrous, margin entire, aciliate, the smallest elliptic, 6.5–1.3 ×
<quantity id="4CE377B8FE86FEDAFD58F905AE37ECAC" box="[641,784,1722,1749]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="9.5" metricValueMin="0.6" pageId="291" pageNumber="292" unit="mm" value="5.05" valueMax="9.5" valueMin="0.6">0.6–9.5 mm</quantity>
, apex obtuse, white, glabrous, margin entire, aciliate; ovary body broadly ellipsoid, 3–10 ×
<quantity id="4CE377B8FE86FEDAFDA4F95EAFC5EC85" box="[637,738,1761,1788]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="291" pageNumber="292" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
, colour unknown, glabrous, unequally 3-winged, wings subtriangular, largest 3–10 ×
<quantity id="4CE377B8FE86FEDAFDCAF8B6AF51ED5A" box="[531,630,1801,1827]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="291" pageNumber="292" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">2–7 mm</quantity>
, smallest 3–10 ×
<quantity id="4CE377B8FE86FEDAFC98F8B6AE83ED5A" box="[833,932,1801,1827]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="291" pageNumber="292" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
; 3-locular, placentae unknown; styles 3, yellow, fused at the base,
<quantity id="4CE377B8FE86FEDAFE34F890AF42ED33" box="[493,613,1839,1866]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="291" pageNumber="292" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
long, 2-lobed, stigmatic papillae in a spiral band.
<emphasis id="B96F064FFE86FEDAFB6FF88FA850ED33" box="[1206,1399,1840,1866]" italics="true" pageId="291" pageNumber="292">Fruiting pedicel</emphasis>
to
<quantity id="4CE377B8FE86FEDAFF02F8E8AC0AED08" box="[219,301,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="291" pageNumber="292" unit="mm" value="22.0">22 mm</quantity>
long.
<emphasis id="B96F064FFE86FEDAFEACF8E8ACD5ED08" box="[373,498,1879,1905]" italics="true" pageId="291" pageNumber="292">Fruit body</emphasis>
ovoid, to 10 ×
<quantity id="4CE377B8FE86FEDAFD7CF8E8AFDDED08" box="[677,762,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="291" pageNumber="292" unit="mm" value="10.0">10 mm</quantity>
, drying brown, wings same shape and size as in ovary.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE86FEDAFF64F823AE9BED91" pageId="291" pageNumber="292" type="conservation">
<paragraph id="8BA4DA5DFE86FEDAFF64F823AF4AEDCF" blockId="291.[189,621,1948,1974]" box="[189,621,1948,1974]" pageId="291" pageNumber="292">
<heading id="D0EC6D31FE86FEDAFF64F823AF4AEDCF" bold="true" box="[189,621,1948,1974]" fontSize="11" level="3" pageId="291" pageNumber="292" reason="3">
<emphasis id="B96F064FFE86FEDAFF64F823AF4AEDCF" bold="true" box="[189,621,1948,1974]" pageId="291" pageNumber="292">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE86FEDAFF64F871AE9BED91" blockId="291.[189,956,1998,2025]" box="[189,956,1998,2025]" pageId="291" pageNumber="292">
Assessed by
<bibRefCitation id="EF8AA7ACFE86FEDAFE8CF871AFBAED91" author="Moonlight &amp; Tebbitt" box="[341,669,1998,2024]" firstAuthor="Moonlight" pageId="291" pageNumber="292" pagination="111 - 122" refId="ref229847" refString="Moonlight P. W. &amp; Tebbitt M. C. 2016. Two new Peruvian species of Begonia (Begoniaceae) and an amended description of Begonia thyrsoidea. Edinburgh Journal of Botany 74 (2): 111 - 122. https: // doi. org / 10.1017 / S 0960428616000299" type="journal article" year="2016">Moonlight &amp; Tebbitt (2016)</bibRefCitation>
as Data Deficient (DD).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE81FEDDFF64FEB7A927EB0C" pageId="292" pageNumber="293" type="discussion">
<paragraph id="8BA4DA5DFE81FEDDFF64FEB7AC97EB5A" blockId="292.[189,432,264,291]" box="[189,432,264,291]" pageId="292" pageNumber="293">
<heading id="D0EC6D31FE81FEDDFF64FEB7AC97EB5A" bold="true" box="[189,432,264,291]" fontSize="11" level="3" pageId="292" pageNumber="293" reason="3">
<emphasis id="B96F064FFE81FEDDFF64FEB7AC97EB5A" bold="true" box="[189,432,264,291]" pageId="292" pageNumber="293">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE81FEDDFF64FE87A927EB0C" blockId="292.[189,1399,311,374]" pageId="292" pageNumber="293">
<taxonomicName id="4C1BA1DEFE81FEDDFF64FE87AC90EB28" box="[189,439,311,337]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="292" pageNumber="293" phylum="Tracheophyta" rank="species" species="erythrothrix">
<emphasis id="B96F064FFE81FEDDFF64FE87AC90EB28" box="[189,439,311,337]" italics="true" pageId="292" pageNumber="293">Begonia erythrothrix</emphasis>
</taxonomicName>
is the only
<taxonomicName id="4C1BA1DEFE81FEDDFD9FFE87AE22EB28" box="[582,773,311,337]" pageId="292" pageNumber="293">
<emphasis id="B96F064FFE81FEDDFD9FFE87AF8EEB28" box="[582,681,312,337]" italics="true" pageId="292" pageNumber="293">Begonia</emphasis>
species
</taxonomicName>
in
<collectingCountry id="F30C9ACDFE81FEDDFCF4FE87AE44EB2B" box="[813,867,312,338]" name="Peru" pageId="292" pageNumber="293">Peru</collectingCountry>
with a creeping, aboveground rhizome; four tepals on the staminate flower; and an indumentum of red, hispid hairs.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE81FEDDFF64FE22A851E854" pageId="292" pageNumber="293" type="distribution">
<paragraph id="8BA4DA5DFE81FEDDFF64FE22ACD2EBCE" blockId="292.[189,501,413,439]" box="[189,501,413,439]" pageId="292" pageNumber="293">
<heading id="D0EC6D31FE81FEDDFF64FE22ACD2EBCE" bold="true" box="[189,501,413,439]" fontSize="11" level="3" pageId="292" pageNumber="293" reason="3">
<emphasis id="B96F064FFE81FEDDFF64FE22ACD2EBCE" bold="true" box="[189,501,413,439]" pageId="292" pageNumber="293">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE81FEDDFF64FE73A851E854" blockId="292.[188,1398,460,558]" pageId="292" pageNumber="293">
Endemic to
<collectingCountry id="F30C9ACDFE81FEDDFE92FE72ACA6EB9E" box="[331,385,461,487]" name="Peru" pageId="292" pageNumber="293">Peru</collectingCountry>
and
<collectingRegion id="49DF14BFFE81FEDDFE60FE73AF78EB9F" box="[441,607,460,486]" country="Peru" name="Cusco" pageId="292" pageNumber="293">Cusco Region</collectingRegion>
(
<figureCitation id="1320C6D8FE81FEDDFDB6FE73AFFEEB9F" box="[623,729,460,486]" captionStart="Fig" captionStartId="288.[189,232,1857,1883]" captionTargetBox="[194,1390,1193,1812]" captionTargetId="figure-313@288.[189,1398,1183,1818]" captionTargetPageId="288" captionText="Fig. 90. Distribution of the Begonia sect. Ignota in Peru and surrounding countries. A. B. bifurcata L.B.Sm. &amp; B.G.Schub. (red), B. erythrothrix Tebbitt &amp; Moonlight (red), and B. occultata J.P.Allen &amp; Moonlight sp. nov. (green). B. B. speculum Moonlight &amp; Tebbitt (red), B. thyrsoidea Irmsch., and B. urubambensis Tebbitt (green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178495" httpUri="https://zenodo.org/record/8178495/files/figure.png" pageId="292" pageNumber="293">Fig. 90A</figureCitation>
) where it is found in middle and possibly lower montane Forest at an elevation of ca
<quantity id="4CE377B8FE81FEDDFE22FE50AFBBE873" box="[507,668,495,522]" metricMagnitude="3" metricUnit="m" metricValue="1.7975" metricValueMax="2.445" metricValueMin="1.15" pageId="292" pageNumber="293" unit="m" value="1797.5" valueMax="2445.0" valueMin="1150.0">1150–2445 m</quantity>
a.s.l.
<taxonomicName id="4C1BA1DEFE81FEDDFD07FE4FAEF2E873" box="[734,981,496,522]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="292" pageNumber="293" phylum="Tracheophyta" rank="species" species="erythrothrix">
<emphasis id="B96F064FFE81FEDDFD07FE4FAEF2E873" box="[734,981,496,522]" italics="true" pageId="292" pageNumber="293">Begonia erythrothrix</emphasis>
</taxonomicName>
is rhizomatous and may die back to its rhizome at some point of the year. The species has been collected in flower and fruit in June and July.
</paragraph>
</subSubSection>
</treatment>
</document>
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