treatments-xml/data/03/F2/87/03F287C0FF85FFADFF61FB82069AFA09.xml

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<mods:title id="6FD171E0962B885CF2A890A78BED4935">A new species of the pontoniine shrimp genus Hamopontonia Bruce, 1970 associated with caryophyllid coral Euphyllia glabrescens (Chamisso &amp; Eysenhardt 1821) in Nhatrang Bay, Vietnam</mods:title>
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<treatment id="03F287C0FF85FFADFF61FB82069AFA09" ID-DOI="http://doi.org/10.5281/zenodo.3510364" ID-GBIF-Taxon="119431944" ID-Zenodo-Dep="3510364" LSID="urn:lsid:plazi:treatment:03F287C0FF85FFADFF61FB82069AFA09" httpUri="http://treatment.plazi.org/id/03F287C0FF85FFADFF61FB82069AFA09" lastPageId="8" lastPageNumber="139" pageId="1" pageNumber="132">
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<paragraph id="8BE436D6FF85FFA4FF61FB82072AFBA7" blockId="1.[151,594,1081,1142]" box="[151,594,1081,1106]" pageId="1" pageNumber="132">
<heading id="D0AC81BAFF85FFA4FF61FB82072AFBA7" bold="true" box="[151,594,1081,1106]" fontSize="10" level="2" pageId="1" pageNumber="132" reason="2">
<emphasis id="B92FEAC4FF85FFA4FF61FB82072AFBA7" bold="true" box="[151,594,1081,1106]" pageId="1" pageNumber="132">
<taxonomicName id="4C5B4D55FF85FFA4FF61FB820489FBA7" ID-CoL="eff5cab1-d6ef-42b5-9d31-e0effa68d85b" box="[151,497,1081,1106]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF85FFA4FF61FB820489FBA7" bold="true" box="[151,497,1081,1106]" italics="true" pageId="1" pageNumber="132">Hamopontonia nhatrangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57BFFF85FFA4FE0EFB82072AFBA7" box="[504,594,1081,1106]" pageId="1" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BE436D6FF85FFA4FF61FBE60429FB83" blockId="1.[151,594,1081,1142]" box="[151,337,1117,1142]" pageId="1" pageNumber="132">
(
<figureCitation id="13602A53FF85FFA4FF69FBE60472FB83" box="[159,266,1117,1142]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1802,1824]" captionStartId-1="3.[151,250,1954,1976]" captionStartId-2="4.[151,250,1953,1975]" captionStartId-3="5.[151,250,1924,1946]" captionTargetBox-0="[168,1416,851,1758]" captionTargetBox-1="[196,1368,198,1933]" captionTargetBox-2="[192,1406,210,1919]" captionTargetBox-3="[196,1385,212,1872]" captionTargetId-0="figure@2.[151,1436,844,1778]" captionTargetId-1="figure@3.[196,1392,193,1933]" captionTargetId-2="figure@4.[173,1413,193,1931]" captionTargetId-3="figure@5.[196,1386,193,1873]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Hamopontonia nhatrangensis sp. nov.: a — ovigerous female; b — mature male." captionText-1="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." captionText-2="FIGURE 3. Hamopontonia nhatrangensis sp. nov., ovigerous female: a — pereiopod I; b — major pereiopod II; c — same, chela; d — minor pereiopod II; e — same, chela; f — pereiopod III; g — same, distal part of propodus and dactylus." captionText-3="FIGURE 4. Coloration of Hamopontonia nhatrangensis sp. nov.: a — holotype, ovigerous female, dorsal view; b — ovigerous female, dorsal view; c—same specimen, lateral view; d — mature male, lateral view; e — same specimen, dorsal view; h — same specimens among tentacles of its host, f — different ovigerous females, dorsal and lateral views (g, h represent the same specimen)." httpUri-0="https://zenodo.org/record/230074/files/figure.png" httpUri-1="https://zenodo.org/record/230075/files/figure.png" httpUri-2="https://zenodo.org/record/230076/files/figure.png" httpUri-3="https://zenodo.org/record/230077/files/figure.png" pageId="1" pageNumber="132">Figs. 1–4</figureCitation>
, 
<figureCitation id="13602A53FF85FFA4FEE0FBE6045CFB83" box="[278,292,1117,1142]" captionStart="FIGURE 5" captionStartId="6.[151,250,1914,1936]" captionTargetBox="[151,1436,193,1891]" captionTargetId="figure@6.[151,1436,193,1892]" captionTargetPageId="6" captionText="FIGURE 5. Coral host Euphyllia glabrescens and Hamopontonia nhatrangensis sp. nov.: a, c — whole colony of host coral Euphyllia glabrescens; b — part of colony with ovigerous female H. nhatrangensis sp. nov. (arrow) in situ; d – f — ovigerous female of Hamopontonia nhatrangensis sp. nov. in natural habitat—among tentacles of coral host E. glabrescens." httpUri="https://zenodo.org/record/230078/files/figure.png" pageId="1" pageNumber="132">5</figureCitation>
<emphasis id="B92FEAC4FF85FFA4FED2FBE60430FB83" box="[292,328,1117,1142]" italics="true" pageId="1" pageNumber="132">d–f</emphasis>
)
</paragraph>
<paragraph id="8BE436D6FF85FFA4FF61FB1D0796FB4E" blockId="1.[151,750,1189,1211]" box="[151,750,1189,1211]" pageId="1" pageNumber="132">
<treatmentCitationGroup id="AB4B11F8FF85FFA4FF61FB1D0796FB4E" box="[151,750,1189,1211]" pageId="1" pageNumber="132">
<taxonomicName id="4C5B4D55FF85FFA4FF61FB1D04ECFB4E" ID-CoL="10cbb7e2-108f-441d-90e2-be4b941496ba" box="[151,404,1189,1211]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="fungicola">
<emphasis id="B92FEAC4FF85FFA4FF61FB1D04ECFB4E" box="[151,404,1189,1211]" italics="true" pageId="1" pageNumber="132">Hamopontonia fungicola</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0AFA10C7FF85FFA4FE45FB1E070AFB4E" ID-DOI="http://doi.org/10.5281/zenodo.3507700" ID-Zenodo-Dep="3507700" author="Marin" box="[435,626,1189,1211]" httpUri="http://treatment.plazi.org/id/E24987F87948F92EFF6D2F1FFC85DA1D" page="348" pageId="1" pageNumber="132" year="2012">
<bibRefCitation id="EFCA4B27FF85FFA4FE45FB1E0738FB4E" author="Marin" box="[435,576,1189,1211]" pageId="1" pageNumber="132" refString="Marin, I. (2012 a) The &quot; Hamopontonia corallicola &quot; Bruce, 1970 species complex (Crustacea, Decapoda, Palaemonidae): new records and new species from the Great Barrier Reef, Australia. Zootaxa, 3272, 35 - 56." type="journal article" year="2012" yearSuffix="a">Marin, 2012a</bibRefCitation>
: 348
</treatmentCitation>
, Pl. 53C, D.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<paragraph id="8BE436D6FF85FFA4FF61FB5C01A7F9B1" blockId="1.[151,1437,1255,2037]" pageId="1" pageNumber="132">
<emphasis id="B92FEAC4FF85FFA4FF61FB5C04FDFAF5" bold="true" box="[151,389,1255,1280]" pageId="1" pageNumber="132">Material examined.</emphasis>
<typeStatus id="54E08874FF85FFA4FE67FB5C0484FAF5" box="[401,508,1255,1280]" pageId="1" pageNumber="132" type="holotype">Holotype</typeStatus>
, ovigerous female—Pacific Ocean, South 
<collectingCountry id="F34C7646FF85FFA4FC04FB5C014DFAF5" box="[1010,1077,1255,1280]" name="China" pageId="1" pageNumber="132">China</collectingCountry>
Sea, 
<emphasis id="B92FEAC4FF85FFA4FB8AFB5C0198FAF5" bold="true" box="[1148,1248,1255,1280]" pageId="1" pageNumber="132">
<collectingCountry id="F34C7646FF85FFA4FB8AFB5C0198FAF5" box="[1148,1248,1255,1280]" name="Vietnam" pageId="1" pageNumber="132">Vietnam</collectingCountry>
</emphasis>
, Nhatrang Bay, south-east part of Tre 
<collectingCountry id="F34C7646FF85FFA4FE6BFAB7049CFAD0" box="[413,484,1292,1317]" name="Iceland" pageId="1" pageNumber="132">Island</collectingCountry>
, 
<geoCoordinate id="EE6F5011FF85FFA4FE02FAB707D8FAD0" box="[500,672,1292,1317]" direction="north" orientation="latitude" pageId="1" pageNumber="132" precision="1" value="12.190598">
12°11’26.15” 
<geoCoordinate id="EE6F5011FF85FFA4FD7FFAB707D8FAD0" box="[649,672,1292,1317]" direction="north" orientation="latitude" pageId="1" pageNumber="132" precision="925" value="109.316666">N</geoCoordinate>
</geoCoordinate>
109°19′51.99″E, on coral 
<taxonomicName id="4C5B4D55FF85FFA4FC2FFAB701ADFAD0" box="[985,1237,1292,1317]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="1" pageNumber="132" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF85FFA4FC2FFAB701ADFAD0" box="[985,1237,1292,1317]" italics="true" pageId="1" pageNumber="132">Euphyllia glabrescens</emphasis>
</taxonomicName>
, depth 7 meters, collected by I. Marin, 
<collectionCode id="ED4AAE13FF85FFA4FE61FA8B048FFABD" box="[407,503,1328,1352]" pageId="1" pageNumber="132">SCUBA</collectionCode>
, 
<date id="FFE51016FF85FFA4FDF2FA8B07CBFABD" box="[516,691,1327,1352]" pageId="1" pageNumber="132" value="2014-03-22">22 March 2014</date>
(
<collectionCode id="ED4AAE13FF85FFA4FD32FA8B065AFABD" box="[708,802,1328,1352]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="132">ZMMU</collectionCode>
). 
<typeStatus id="54E08874FF85FFA4FCC0FA8B06DFFABD" box="[822,935,1328,1352]" pageId="1" pageNumber="132" type="paratype">Paratypes</typeStatus>
: Pacific Ocean, South 
<collectingCountry id="F34C7646FF85FFA4FB5DFA940196FABD" box="[1195,1262,1327,1352]" name="China" pageId="1" pageNumber="132">China</collectingCountry>
Sea, 
<emphasis id="B92FEAC4FF85FFA4FAD9FA9400EBFABD" bold="true" box="[1327,1427,1327,1352]" pageId="1" pageNumber="132">
<collectingCountry id="F34C7646FF85FFA4FAD9FA9400EBFABD" box="[1327,1427,1327,1352]" name="Vietnam" pageId="1" pageNumber="132">Vietnam</collectingCountry>
</emphasis>
, Nhatrang Bay—1 ovigerous female, 1 male—Mun 
<collectingCountry id="F34C7646FF85FFA4FD23FAEF0664FA98" box="[725,796,1364,1389]" name="Iceland" pageId="1" pageNumber="132">Island</collectingCountry>
, south-eastern bay, on coral 
<taxonomicName id="4C5B4D55FF85FFA4FBAEFAEF0028FA98" box="[1112,1360,1364,1389]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="1" pageNumber="132" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF85FFA4FBAEFAEF0028FA98" box="[1112,1360,1364,1389]" italics="true" pageId="1" pageNumber="132">Euphyllia glabrescens</emphasis>
</taxonomicName>
, depth 10 meters, collected by I. Marin, 
<collectionCode id="ED4AAE13FF85FFA4FDD1FAC307FFFA65" box="[551,647,1400,1424]" pageId="1" pageNumber="132">SCUBA</collectionCode>
, 
<date id="FFE51016FF85FFA4FD6EFACC0601FA65" box="[664,889,1399,1424]" pageId="1" pageNumber="132" value="2010-11-15">15 November 2010</date>
(
<collectionCode id="ED4AAE13FF85FFA4FC7BFAC30689FA65" box="[909,1009,1400,1424]" pageId="1" pageNumber="132">LEMMI</collectionCode>
); 1 ovigerous female, 1 male—Mun 
<collectingCountry id="F34C7646FF85FFA4FF61FA2705A6FA40" box="[151,222,1436,1461]" name="Iceland" pageId="1" pageNumber="132">Island</collectingCountry>
, north side, on coral 
<taxonomicName id="4C5B4D55FF85FFA4FE30FA2707C7FA40" box="[454,703,1436,1461]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="1" pageNumber="132" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF85FFA4FE30FA2707C7FA40" box="[454,703,1436,1461]" italics="true" pageId="1" pageNumber="132">Euphyllia glabrescens</emphasis>
</taxonomicName>
, depth 5 meters, collected by I. Marin, 
<collectionCode id="ED4AAE13FF85FFA4FB80FA2701AEFA41" box="[1142,1238,1436,1460]" pageId="1" pageNumber="132">SCUBA</collectionCode>
, 
<date id="FFE51016FF85FFA4FB17FA2700E4FA41" box="[1249,1436,1436,1461]" pageId="1" pageNumber="132" value="2012-10-15">15 October 2012</date>
(
<collectionCode id="ED4AAE13FF85FFA4FF69FA7B047BFA2D" box="[159,259,1472,1496]" pageId="1" pageNumber="132">LEMMI</collectionCode>
); 1 ovigerous female—same locality, 
<date id="FFE51016FF85FFA4FD37FA7B06FFFA2D" box="[705,903,1471,1496]" pageId="1" pageNumber="132" value="2012-10-21">21 October 2012</date>
(
<collectionCode id="ED4AAE13FF85FFA4FC6DFA7B0687FA2D" box="[923,1023,1472,1496]" pageId="1" pageNumber="132">LEMMI</collectionCode>
); 1 ovigerous female, 1 male—Tre 
<collectingCountry id="F34C7646FF85FFA4FF61FA5F05A6FA08" box="[151,222,1508,1533]" name="Iceland" pageId="1" pageNumber="132">Island</collectingCountry>
, south-east side, Light house, on coral 
<taxonomicName id="4C5B4D55FF85FFA4FD41FA5F06CEFA08" box="[695,950,1508,1533]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="1" pageNumber="132" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF85FFA4FD41FA5F06CEFA08" box="[695,950,1508,1533]" italics="true" pageId="1" pageNumber="132">Euphyllia glabrescens</emphasis>
</taxonomicName>
, depth 12 meters, collected by I. Marin, 
<collectionCode id="ED4AAE13FF85FFA4FF61F9B3058FF9D5" box="[151,247,1544,1568]" pageId="1" pageNumber="132">SCUBA</collectionCode>
, 0 
<date id="FFE51016FF85FFA4FEE7F9B304E6F9D5" box="[273,414,1543,1568]" pageId="1" pageNumber="132" value="2013-04-03">3 April 2013</date>
(
<collectionCode id="ED4AAE13FF85FFA4FE5AF9B30768F9D5" box="[428,528,1544,1568]" pageId="1" pageNumber="132">LEMMI</collectionCode>
); 1 ovigerous female, 1 male—Tre 
<collectingCountry id="F34C7646FF85FFA4FC6FF9BC0698F9D5" box="[921,992,1543,1568]" name="Iceland" pageId="1" pageNumber="132">Island</collectingCountry>
, south-east part near Dam Bay, on coral 
<taxonomicName id="4C5B4D55FF85FFA4FF61F99704E8F9B0" box="[151,400,1580,1605]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="1" pageNumber="132" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF85FFA4FF61F99704E8F9B0" box="[151,400,1580,1605]" italics="true" pageId="1" pageNumber="132">Euphyllia glabrescens</emphasis>
</taxonomicName>
, depth 8 meters, collected by I. Marin, 
<collectionCode id="ED4AAE13FF85FFA4FCBFF99706D1F9B1" box="[841,937,1580,1604]" pageId="1" pageNumber="132">SCUBA</collectionCode>
, 
<date id="FFE51016FF85FFA4FC43F9970118F9B1" box="[949,1120,1580,1605]" pageId="1" pageNumber="132" value="2014-03-25">25 March 2014</date>
(
<collectionCode id="ED4AAE13FF85FFA4FB98F99701AAF9B1" box="[1134,1234,1580,1604]" pageId="1" pageNumber="132">LEMMI</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C341655DFF85FFA2FF31F9F400F0F9FD" lastPageId="7" lastPageNumber="138" pageId="1" pageNumber="132" type="description">
<paragraph id="8BE436D6FF85FFA4FF31F9F4066BF8B5" blockId="1.[151,1437,1255,2037]" pageId="1" pageNumber="132">
<emphasis id="B92FEAC4FF85FFA4FF31F9F40421F99D" bold="true" box="[199,345,1615,1640]" pageId="1" pageNumber="132">Description.</emphasis>
Medium-sized pontoniine shrimp with cylindrical slightly depressed body. Carapace swollen in medial part, smooth, with large antennal tooth, without hepatic or supraorbital teeth (
<figureCitation id="13602A53FF85FFA4FB56F9CF018BF979" box="[1184,1267,1652,1677]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="1" pageNumber="132">Figs. 2</figureCitation>
<emphasis id="B92FEAC4FF85FFA4FB05F9CE0065F978" box="[1267,1309,1652,1677]" italics="true" pageId="1" pageNumber="132">
a 
<emphasis id="B92FEAC4FF85FFA4FAF7F9CF0077F97E" bold="true" box="[1281,1295,1652,1675]" italics="true" pageId="1" pageNumber="132">–</emphasis>
d
</emphasis>
). Rostrum compressed, pointing distally, straight, reaching to distal margin of basal antennular segment; rostral formula 1 
<emphasis id="B92FEAC4FF85FFA4FF53F90705CBF926" bold="true" box="[165,179,1724,1747]" italics="true" pageId="1" pageNumber="132">–</emphasis>
3+4 
<emphasis id="B92FEAC4FF85FFA4FF28F9070594F926" bold="true" box="[222,236,1724,1747]" italics="true" pageId="1" pageNumber="132">–</emphasis>
5/0 (
<figureCitation id="13602A53FF85FFA4FED7F907041CF921" box="[289,356,1724,1749]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="1" pageNumber="132">Fig. 2</figureCitation>
<emphasis id="B92FEAC4FF85FFA4FE92F90604F4F921" box="[356,396,1724,1748]" italics="true" pageId="1" pageNumber="132">
a 
<emphasis id="B92FEAC4FF85FFA4FE84F90704F8F926" bold="true" box="[370,384,1724,1747]" italics="true" pageId="1" pageNumber="132">–</emphasis>
c
</emphasis>
); dorsal rostral carina well developed, reaching to 1/3 of carapace length, with well marked small dorsal teeth; ventral rostral carina poorly developed, unarmed; proximal lateral rostral lamina feebly developed (
<figureCitation id="13602A53FF85FFA4FEEAF8BF0419F8E9" box="[284,353,1796,1821]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="1" pageNumber="132">Fig. 2</figureCitation>
<emphasis id="B92FEAC4FF85FFA4FE97F8BE04D4F8E8" box="[353,428,1796,1821]" italics="true" pageId="1" pageNumber="132">a, b, d</emphasis>
). Orbit well developed, deep, unarmed; inferior orbital angle bluntly produced distally. Pterygostomial angle produced, bluntly rounded distally.
</paragraph>
<paragraph id="8BE436D6FF85FFA4FF31F8F70582F801" blockId="1.[151,1437,1255,2037]" pageId="1" pageNumber="132">
Abdominal somites smooth; pleura of abdominal somites I–V rounded (
<figureCitation id="13602A53FF85FFA4FC0FF8F70145F891" box="[1017,1085,1868,1893]" captionStart="FIGURE 1" captionStartId="2.[151,250,1802,1824]" captionTargetBox="[168,1416,851,1758]" captionTargetId="figure@2.[151,1436,844,1778]" captionTargetPageId="2" captionText="FIGURE 1. Hamopontonia nhatrangensis sp. nov.: a — ovigerous female; b — mature male." httpUri="https://zenodo.org/record/230074/files/figure.png" pageId="1" pageNumber="132">Fig. 1</figureCitation>
). Telson (
<figureCitation id="13602A53FF85FFA4FB47F8F7018DF891" box="[1201,1269,1868,1893]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="1" pageNumber="132">
Fig. 
<quantity id="4CA39B33FF85FFA4FB11F8F7018DF891" box="[1255,1269,1868,1892]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="1" pageNumber="132" unit="g" value="2.0">2</quantity>
</figureCitation>
<emphasis id="B92FEAC4FF85FFA4FB03F8F60058F890" box="[1269,1312,1868,1893]" italics="true" pageId="1" pageNumber="132">g, h</emphasis>
) relatively stout, about 3 times as long as proximal width, narrow distally, distal margin of telson armed with two hook-like projections (teeth) turned downward (
<figureCitation id="13602A53FF85FFA4FDB7F82F07FCF859" box="[577,644,1940,1965]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="1" pageNumber="132">
Fig. 
<date id="FFE51016FF85FFA4FD80F82F07FCF859" box="[630,644,1940,1964]" pageId="1" pageNumber="132">2</date>
</figureCitation>
<emphasis id="B92FEAC4FF85FFA4FD72F82E07F4F859" box="[644,652,1941,1964]" italics="true" pageId="1" pageNumber="132">i</emphasis>
) and distinct notch medially (
<figureCitation id="13602A53FF85FFA4FC29F82F015AF859" box="[991,1058,1940,1965]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="1" pageNumber="132">
Fig. 
<quantity id="4CA39B33FF85FFA4FBE2F82F015AF859" box="[1044,1058,1940,1964]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="1" pageNumber="132" unit="g" value="2.0">2</quantity>
</figureCitation>
<emphasis id="B92FEAC4FF85FFA4FBD4F82E0148F859" box="[1058,1072,1941,1964]" italics="true" pageId="1" pageNumber="132">g</emphasis>
), with 3 pairs of medium dorsal submarginal spines at 0.6, 0.8 and 0.95 of telson length, with paired most distal tiny dorsal spines in some specimen (
<figureCitation id="13602A53FF85FFA4FF69F867059AF801" box="[159,226,2012,2037]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="1" pageNumber="132">
Fig. 
<date id="FFE51016FF85FFA4FF22F867059AF801" box="[212,226,2012,2036]" pageId="1" pageNumber="132">2</date>
</figureCitation>
<emphasis id="B92FEAC4FF85FFA4FF17F8660591F801" box="[225,233,2013,2036]" italics="true" pageId="1" pageNumber="132">i</emphasis>
).
</paragraph>
<paragraph id="8BE436D6FF86FFA7FF31FF230709FF21" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="133">
Eyes well developed, large; eyestalk smooth, cylindrical, about 1.5 times as long as wide; cornea well developed, medium, subovate (
<figureCitation id="13602A53FF86FFA7FE03FF07074FFF21" box="[501,567,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="2" pageNumber="133">Fig. 2</figureCitation>
<emphasis id="B92FEAC4FF86FFA7FDC1FF060719FF20" box="[567,609,188,213]" italics="true" pageId="2" pageNumber="133">
a 
<emphasis id="B92FEAC4FF86FFA7FDB3FF07072BFF26" bold="true" box="[581,595,188,211]" italics="true" pageId="2" pageNumber="133">–</emphasis>
d
</emphasis>
).
</paragraph>
<paragraph id="8BE436D6FF86FFA7FF31FF640199FE72" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="133">
Antennula (
<figureCitation id="13602A53FF86FFA7FEA4FF6404E3FF0D" box="[338,411,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="2" pageNumber="133">Fig. 2</figureCitation>
<emphasis id="B92FEAC4FF86FFA7FE6DFF5A04DFFF0D" box="[411,423,225,248]" italics="true" pageId="2" pageNumber="133">e</emphasis>
) well developed; basal segment stout, about 1.8 times longer than wide, with well developed slender acute stylocerite, with distolateral angle bearing large acute triangular tooth and small rounded medial projection; ventromesial tooth small; intermediate segment stout, about as long as wide; distal segment stout, as long as wide, equal to previous segment; proximal part of upper antennular flagellum with 8–9 separate segments, shorter ramus with more than 15 segments; about 9 or 10 groups of aesthetascs present.
</paragraph>
<paragraph id="8BE436D6FF86FFA7FF31FE2F0056FE00" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="133">
Antenna (
<figureCitation id="13602A53FF86FFA7FEC1FE2F0404FE59" box="[311,380,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="2" pageNumber="133">Fig. 2</figureCitation>
<emphasis id="B92FEAC4FF86FFA7FE8AFE2F04FCFE59" box="[380,388,404,428]" italics="true" pageId="2" pageNumber="133">f</emphasis>
) well developed, basicerite about 1.5 times longer than wide, smooth, without distoventral tooth; scaphocerite wide, about 1.5 times longer than maximal width, overreaching the distal margin of antennular peduncle (
<figureCitation id="13602A53FF86FFA7FEFDFE670435FE01" box="[267,333,476,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="2" pageNumber="133">Fig. 2</figureCitation>
<emphasis id="B92FEAC4FF86FFA7FEBBFE670423FE00" box="[333,347,476,501]" italics="true" pageId="2" pageNumber="133">d</emphasis>
), distolateral tooth well developed, acute, small, not reaching to distal margin of blade.
</paragraph>
<paragraph id="8BE436D6FF86FFA7FF31FE44017CFD95" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="133">Mouthparts characteristic for the genus and previously described species. Maxilliped III with slender unarmed segments; ischiomeral segment about 5 times longer than wide; antepenultimate segment about 3 times longer than wide; penultimate segment tapering distally, about 2.5 times longer than wide.</paragraph>
<paragraph id="8BE436D6FF86FFA7FF31FDD7078AFCE0" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="133">
Pereiopod I (
<figureCitation id="13602A53FF86FFA7FEABFDD704DAFD71" box="[349,418,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[192,1406,210,1919]" captionTargetId="figure@4.[173,1413,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Hamopontonia nhatrangensis sp. nov., ovigerous female: a — pereiopod I; b — major pereiopod II; c — same, chela; d — minor pereiopod II; e — same, chela; f — pereiopod III; g — same, distal part of propodus and dactylus." httpUri="https://zenodo.org/record/230076/files/figure.png" pageId="2" pageNumber="133">Fig. 3</figureCitation>
<emphasis id="B92FEAC4FF86FFA7FE57FDD604D7FD71" box="[417,431,621,644]" italics="true" pageId="2" pageNumber="133">a</emphasis>
) with relatively slender unarmed segments, without specific features; coxal segment as long as wide, with distoventral lobe; basis as long as wide; ischium about twice longer than wide; merus slender, about 4.5 times as long as wide; carpus about 6 times as long as wide, subequal to merus; propodus about 3 times longer than wide, cylindrical, slightly flaring distally, fingers simple and slender, about 4 times as long as wide, with straight, smooth cutting margins and simple tips.
</paragraph>
<caption id="DF24665EFF86FFA7FF61F8B10140F8D5" box="[151,1080,1802,1824]" httpUri="https://zenodo.org/record/230074/files/figure.png" pageId="2" pageNumber="133" targetBox="[168,1416,851,1758]" targetPageId="2">
<paragraph id="8BE436D6FF86FFA7FF61F8B10140F8D5" blockId="2.[151,1080,1802,1824]" box="[151,1080,1802,1824]" pageId="2" pageNumber="133">
<emphasis id="B92FEAC4FF86FFA7FF61F8B1046BF8EA" bold="true" box="[151,275,1802,1824]" pageId="2" pageNumber="133">FIGURE 1.</emphasis>
<taxonomicName id="4C5B4D55FF86FFA7FEEFF8B0073EF8D5" box="[281,582,1802,1824]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF86FFA7FEEFF8B0073EF8D5" box="[281,582,1802,1824]" italics="true" pageId="2" pageNumber="133">Hamopontonia nhatrangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57BFFF86FFA7FDBBF8B107E5F8EA" box="[589,669,1802,1823]" pageId="2" pageNumber="133" rank="species">
<emphasis id="B92FEAC4FF86FFA7FDBBF8B107E5F8EA" bold="true" box="[589,669,1802,1823]" pageId="2" pageNumber="133">sp. nov.</emphasis>
</taxonomicNameLabel>
: 
<emphasis id="B92FEAC4FF86FFA7FD5DF8B007C0F8D5" box="[683,696,1803,1824]" italics="true" pageId="2" pageNumber="133">a</emphasis>
—ovigerous female; 
<emphasis id="B92FEAC4FF86FFA7FC7BF8B106E1F8D5" box="[909,921,1802,1824]" italics="true" pageId="2" pageNumber="133">b</emphasis>
—mature male.
</paragraph>
</caption>
<caption id="DF24665EFF87FFA6FF61F8190119F803" httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="3" pageNumber="134" targetBox="[196,1368,198,1933]" targetPageId="3">
<paragraph id="8BE436D6FF87FFA6FF61F8190119F803" blockId="3.[151,1436,1954,2038]" pageId="3" pageNumber="134">
<emphasis id="B92FEAC4FF87FFA6FF61F8190460F842" bold="true" box="[151,280,1954,1976]" pageId="3" pageNumber="134">FIGURE 2.</emphasis>
<taxonomicName id="4C5B4D55FF87FFA6FED5F818072DF84D" box="[291,597,1954,1976]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF87FFA6FED5F818072DF84D" box="[291,597,1954,1976]" italics="true" pageId="3" pageNumber="134">Hamopontonia nhatrangensis</emphasis>
</taxonomicName>
<emphasis id="B92FEAC4FF87FFA6FD97F81807CEF84D" bold="true" box="[609,694,1955,1976]" pageId="3" pageNumber="134">
<taxonomicNameLabel id="A21C57BFFF87FFA6FD97F81807CEF84D" box="[609,694,1955,1976]" pageId="3" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ovigerous female (
<emphasis id="B92FEAC4FF87FFA6FC66F8180695F84D" box="[912,1005,1954,1976]" italics="true" pageId="3" pageNumber="134">a, b, d–j</emphasis>
), male (
<emphasis id="B92FEAC4FF87FFA6FBBAF818012FF84D" box="[1100,1111,1955,1976]" italics="true" pageId="3" pageNumber="134">c</emphasis>
): 
<emphasis id="B92FEAC4FF87FFA6FB84F8180107F84D" box="[1138,1151,1955,1976]" italics="true" pageId="3" pageNumber="134">a</emphasis>
–c –carapace, lateral view; 
<emphasis id="B92FEAC4FF87FFA6FF61F87A05DCF822" box="[151,164,1985,2007]" italics="true" pageId="3" pageNumber="134">d</emphasis>
—carapace, dorsal view; 
<emphasis id="B92FEAC4FF87FFA6FE5CF87904CDF822" box="[426,437,1986,2007]" italics="true" pageId="3" pageNumber="134">e</emphasis>
—antennula; 
<emphasis id="B92FEAC4FF87FFA6FDB7F87A0730F822" box="[577,584,1985,2007]" italics="true" pageId="3" pageNumber="134">f</emphasis>
—antenna; g—sixth abdominal somite, telson and uropods, dorsal view; 
<emphasis id="B92FEAC4FF87FFA6FACAF87A0031F822" box="[1340,1353,1985,2007]" italics="true" pageId="3" pageNumber="134">h</emphasis>
—same, lateral view; 
<emphasis id="B92FEAC4FF87FFA6FEEDF85A045AF803" box="[283,290,2017,2038]" italics="true" pageId="3" pageNumber="134">i</emphasis>
—distal margin of telson, lateral view; 
<emphasis id="B92FEAC4FF87FFA6FD59F85A07CEF803" box="[687,694,2017,2038]" italics="true" pageId="3" pageNumber="134">j</emphasis>
—distolateral margin of uropodal exopod.
</paragraph>
</caption>
<caption id="DF24665EFF80FFA1FF61F81A01C3F820" httpUri="https://zenodo.org/record/230076/files/figure.png" pageId="4" pageNumber="135" targetBox="[192,1406,210,1919]" targetPageId="4">
<paragraph id="8BE436D6FF80FFA1FF61F81A01C3F820" blockId="4.[151,1436,1953,2005]" pageId="4" pageNumber="135">
<emphasis id="B92FEAC4FF80FFA1FF61F81A046AF843" bold="true" box="[151,274,1953,1975]" pageId="4" pageNumber="135">FIGURE 3.</emphasis>
<taxonomicName id="4C5B4D55FF80FFA1FEEEF819073DF842" box="[280,581,1953,1975]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF80FFA1FEEEF819073DF842" box="[280,581,1953,1975]" italics="true" pageId="4" pageNumber="135">Hamopontonia nhatrangensis</emphasis>
</taxonomicName>
<emphasis id="B92FEAC4FF80FFA1FDBDF81A07E3F843" bold="true" box="[587,667,1953,1974]" pageId="4" pageNumber="135">
<taxonomicNameLabel id="A21C57BFFF80FFA1FDBDF81A07E3F843" box="[587,667,1953,1974]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ovigerous female: 
<emphasis id="B92FEAC4FF80FFA1FC92F8190609F842" box="[868,881,1954,1975]" italics="true" pageId="4" pageNumber="135">a</emphasis>
—pereiopod I; 
<emphasis id="B92FEAC4FF80FFA1FBFDF81A016FF842" box="[1035,1047,1953,1975]" italics="true" pageId="4" pageNumber="135">b</emphasis>
—major pereiopod II; 
<emphasis id="B92FEAC4FF80FFA1FB0DF819007EF842" box="[1275,1286,1954,1975]" italics="true" pageId="4" pageNumber="135">c</emphasis>
—same, chela; 
<emphasis id="B92FEAC4FF80FFA1FF61F80405DCF820" box="[151,164,1983,2005]" italics="true" pageId="4" pageNumber="135">d</emphasis>
—minor pereiopod II; 
<emphasis id="B92FEAC4FF80FFA1FE7FF87B04ECF820" box="[393,404,1984,2005]" italics="true" pageId="4" pageNumber="135">e</emphasis>
—same, chela; 
<emphasis id="B92FEAC4FF80FFA1FDC7F8040740F820" box="[561,568,1983,2005]" italics="true" pageId="4" pageNumber="135">f</emphasis>
—pereiopod III; 
<emphasis id="B92FEAC4FF80FFA1FD12F87B0788F820" box="[740,752,1984,2005]" italics="true" pageId="4" pageNumber="135">g</emphasis>
—same, distal part of propodus and dactylus.
</paragraph>
</caption>
<caption id="DF24665EFF81FFA0FF61F83F047EF800" httpUri="https://zenodo.org/record/230077/files/figure.png" pageId="5" pageNumber="136" targetBox="[196,1385,212,1872]" targetPageId="5">
<paragraph id="8BE436D6FF81FFA0FF61F83F047EF800" blockId="5.[151,1436,1924,2037]" pageId="5" pageNumber="136">
<emphasis id="B92FEAC4FF81FFA0FF61F83F046CF86C" bold="true" box="[151,276,1924,1946]" pageId="5" pageNumber="136">FIGURE 4.</emphasis>
Coloration of 
<taxonomicName id="4C5B4D55FF81FFA0FE5AF83E07A3F86F" box="[428,731,1924,1946]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF81FFA0FE5AF83E07A3F86F" box="[428,731,1924,1946]" italics="true" pageId="5" pageNumber="136">Hamopontonia nhatrangensis</emphasis>
</taxonomicName>
<emphasis id="B92FEAC4FF81FFA0FD14F83F064BF86C" bold="true" box="[738,819,1924,1945]" pageId="5" pageNumber="136">
<taxonomicNameLabel id="A21C57BFFF81FFA0FD14F83F064BF86C" box="[738,819,1924,1945]" pageId="5" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 
<emphasis id="B92FEAC4FF81FFA0FCB5F83E0637F86F" box="[835,847,1925,1946]" italics="true" pageId="5" pageNumber="136">a</emphasis>
—holotype, ovigerous female, dorsal view; 
<emphasis id="B92FEAC4FF81FFA0FAE4F83F0067F86F" box="[1298,1311,1924,1946]" italics="true" pageId="5" pageNumber="136">b</emphasis>
—ovigerous female, dorsal view; c—same specimen, lateral view; 
<emphasis id="B92FEAC4FF81FFA0FD4BF81907B1F84D" box="[701,713,1954,1976]" italics="true" pageId="5" pageNumber="136">d</emphasis>
—mature male, lateral view; 
<emphasis id="B92FEAC4FF81FFA0FC05F8180686F84D" box="[1011,1022,1955,1976]" italics="true" pageId="5" pageNumber="136">e</emphasis>
—same specimen, dorsal view; 
<emphasis id="B92FEAC4FF81FFA0FAB4F8190036F84D" box="[1346,1358,1954,1976]" italics="true" pageId="5" pageNumber="136">h</emphasis>
—same specimens among tentacles of its host, 
<emphasis id="B92FEAC4FF81FFA0FDC8F87A073DF822" box="[574,581,1985,2007]" italics="true" pageId="5" pageNumber="136">f</emphasis>
—different ovigerous females, dorsal and lateral views (
<emphasis id="B92FEAC4FF81FFA0FB56F87901B2F822" box="[1184,1226,1985,2007]" italics="true" pageId="5" pageNumber="136">g, h</emphasis>
represent the same specimen).
</paragraph>
</caption>
<caption id="DF24665EFF82FFA3FF61F8C1006EF838" httpUri="https://zenodo.org/record/230078/files/figure.png" pageId="6" pageNumber="137" targetBox="[151,1436,193,1891]" targetPageId="6">
<paragraph id="8BE436D6FF82FFA3FF61F8C1006EF838" blockId="6.[151,1436,1914,1997]" pageId="6" pageNumber="137">
<emphasis id="B92FEAC4FF82FFA3FF61F8C1046DF87A" bold="true" box="[151,277,1914,1936]" pageId="6" pageNumber="137">FIGURE 5.</emphasis>
Coral host 
<taxonomicName id="4C5B4D55FF82FFA3FE67F8C1070CF865" box="[401,628,1914,1936]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="6" pageNumber="137" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF82FFA3FE67F8C1070CF865" box="[401,628,1914,1936]" italics="true" pageId="6" pageNumber="137">Euphyllia glabrescens</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5B4D55FF82FFA3FD5CF8C006A2F865" box="[682,986,1914,1936]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF82FFA3FD5CF8C006A2F865" box="[682,986,1914,1936]" italics="true" pageId="6" pageNumber="137">Hamopontonia nhatrangensis</emphasis>
</taxonomicName>
<emphasis id="B92FEAC4FF82FFA3FC14F8C1014DF87A" bold="true" box="[994,1077,1914,1935]" pageId="6" pageNumber="137">
<taxonomicNameLabel id="A21C57BFFF82FFA3FC14F8C1014DF87A" box="[994,1077,1914,1935]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 
<emphasis id="B92FEAC4FF82FFA3FBB3F8C00113F865" box="[1093,1131,1915,1936]" italics="true" pageId="6" pageNumber="137">a, c</emphasis>
—whole colony of host coral 
<taxonomicName id="4C5B4D55FF82FFA3FF61F8230403F85B" box="[151,379,1944,1966]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="6" pageNumber="137" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF82FFA3FF61F8230403F85B" box="[151,379,1944,1966]" italics="true" pageId="6" pageNumber="137">Euphyllia glabrescens</emphasis>
</taxonomicName>
; 
<emphasis id="B92FEAC4FF82FFA3FE7DF82304EFF85B" box="[395,407,1944,1966]" italics="true" pageId="6" pageNumber="137">b</emphasis>
—part of colony with ovigerous female 
<taxonomicName id="4C5B4D55FF82FFA3FCC9F8210688F85B" box="[831,1008,1944,1967]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF82FFA3FCC9F8210688F85B" box="[831,1008,1944,1967]" italics="true" pageId="6" pageNumber="137">H. nhatrangensis</emphasis>
</taxonomicName>
<emphasis id="B92FEAC4FF82FFA3FC0CF8220136F85B" bold="true" box="[1018,1102,1945,1966]" pageId="6" pageNumber="137">
<taxonomicNameLabel id="A21C57BFFF82FFA3FC0CF8220136F85B" box="[1018,1102,1945,1966]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(arrow) 
<emphasis id="B92FEAC4FF82FFA3FB5AF8220196F85B" box="[1196,1262,1945,1966]" italics="true" pageId="6" pageNumber="137">in situ</emphasis>
; 
<emphasis id="B92FEAC4FF82FFA3FB08F8230067F85B" box="[1278,1311,1944,1966]" italics="true" pageId="6" pageNumber="137">d–f</emphasis>
—ovigerous female of 
<taxonomicName id="4C5B4D55FF82FFA3FF0BF8030754F838" box="[253,556,1975,1997]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF82FFA3FF0BF8030754F838" box="[253,556,1975,1997]" italics="true" pageId="6" pageNumber="137">Hamopontonia nhatrangensis</emphasis>
</taxonomicName>
<emphasis id="B92FEAC4FF82FFA3FDC7F80C07FAF839" bold="true" box="[561,642,1975,1996]" pageId="6" pageNumber="137">
<taxonomicNameLabel id="A21C57BFFF82FFA3FDC7F80C07FAF839" box="[561,642,1975,1996]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in natural habitat—among tentacles of coral host 
<taxonomicName id="4C5B4D55FF82FFA3FB8AF8030068F838" box="[1148,1296,1975,1997]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="6" pageNumber="137" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF82FFA3FB8AF8030068F838" box="[1148,1296,1975,1997]" italics="true" pageId="6" pageNumber="137">E. glabrescens</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE436D6FF83FFA2FF31FF2C0683FE25" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
Pereiopods II similar in shape and dissimilar in size (
<figureCitation id="13602A53FF83FFA2FCD2FF2C060AFF45" box="[804,882,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1802,1824]" captionTargetBox="[168,1416,851,1758]" captionTargetId="figure@2.[151,1436,844,1778]" captionTargetPageId="2" captionText="FIGURE 1. Hamopontonia nhatrangensis sp. nov.: a — ovigerous female; b — mature male." httpUri="https://zenodo.org/record/230074/files/figure.png" pageId="7" pageNumber="138">Figs. 1</figureCitation>
, 
<figureCitation id="13602A53FF83FFA2FC77FF2306F7FF45" box="[897,911,152,176]" captionStart="FIGURE 3" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[192,1406,210,1919]" captionTargetId="figure@4.[173,1413,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Hamopontonia nhatrangensis sp. nov., ovigerous female: a — pereiopod I; b — major pereiopod II; c — same, chela; d — minor pereiopod II; e — same, chela; f — pereiopod III; g — same, distal part of propodus and dactylus." httpUri="https://zenodo.org/record/230076/files/figure.png" pageId="7" pageNumber="138">3</figureCitation>
<emphasis id="B92FEAC4FF83FFA2FC79FF2C06C2FF45" box="[911,954,151,176]" italics="true" pageId="7" pageNumber="138">b, d</emphasis>
); with robust and smooth segments; coxal segment as long as wide, with small distoventral lobe; basis as long as wide; ischium about 3 
<emphasis id="B92FEAC4FF83FFA2FB42FF0701BAFF20" bold="true" box="[1204,1218,188,213]" pageId="7" pageNumber="138">–</emphasis>
4 times longer than wide; merus robust, about 3.5 
<emphasis id="B92FEAC4FF83FFA2FE1FFF64048FFF0D" bold="true" box="[489,503,223,248]" pageId="7" pageNumber="138">–</emphasis>
4 times longer than wide, with straight lateral margins; carpus triangular in shape, stout, flaring distally, slightly overlapping carpo-propodal articulation, with smooth distal margin (
<figureCitation id="13602A53FF83FFA2FB15FEBF005EFEE9" box="[1251,1318,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[192,1406,210,1919]" captionTargetId="figure@4.[173,1413,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Hamopontonia nhatrangensis sp. nov., ovigerous female: a — pereiopod I; b — major pereiopod II; c — same, chela; d — minor pereiopod II; e — same, chela; f — pereiopod III; g — same, distal part of propodus and dactylus." httpUri="https://zenodo.org/record/230076/files/figure.png" pageId="7" pageNumber="138">Fig. 3</figureCitation>
<emphasis id="B92FEAC4FF83FFA2FAD3FEBE0034FEE9" box="[1317,1356,261,284]" italics="true" pageId="7" pageNumber="138">c, e</emphasis>
); palm (
<figureCitation id="13602A53FF83FFA2FF69FE9C059AFEB5" box="[159,226,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[192,1406,210,1919]" captionTargetId="figure@4.[173,1413,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Hamopontonia nhatrangensis sp. nov., ovigerous female: a — pereiopod I; b — major pereiopod II; c — same, chela; d — minor pereiopod II; e — same, chela; f — pereiopod III; g — same, distal part of propodus and dactylus." httpUri="https://zenodo.org/record/230076/files/figure.png" pageId="7" pageNumber="138">Fig. 3</figureCitation>
<emphasis id="B92FEAC4FF83FFA2FF15FE920472FEB5" box="[227,266,297,320]" italics="true" pageId="7" pageNumber="138">c, e</emphasis>
) smooth, cylindrical, about 3.5 
<emphasis id="B92FEAC4FF83FFA2FD9BFE930703FECA" bold="true" box="[621,635,296,319]" italics="true" pageId="7" pageNumber="138">–</emphasis>
4 times as long as wide, slightly narrowing medially; fingers relatively slender, about twice shorter than palm, about 3 times as long as wide; fixed finger (pollex) compressed, about 3 times longer than wide, with two medial teeth situated in proximal 1/3 of length, with simple sharp curved tip, with small medial depression opposite to tooth on movable finger; movable finger (dactylus) slender, compressed, with large tooth situated in proximal 1/3 of cutting margin, with simple curved tip.
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF3BFE670774FD5D" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
Pereiopod III (
<figureCitation id="13602A53FF83FFA2FE87FE6704CBFE01" box="[369,435,476,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[192,1406,210,1919]" captionTargetId="figure@4.[173,1413,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Hamopontonia nhatrangensis sp. nov., ovigerous female: a — pereiopod I; b — major pereiopod II; c — same, chela; d — minor pereiopod II; e — same, chela; f — pereiopod III; g — same, distal part of propodus and dactylus." httpUri="https://zenodo.org/record/230076/files/figure.png" pageId="7" pageNumber="138">Fig. 3</figureCitation>
<emphasis id="B92FEAC4FF83FFA2FE45FE6704C3FE01" box="[435,443,476,500]" italics="true" pageId="7" pageNumber="138">f</emphasis>
) with smooth slender unarmed segments; coxal segment as long as wide, unarmed; basis as long as wide; ischium robust, about twice longer than wide; merus slender, about 4 times as long as wide; carpus about twice longer than wide, about half shorter than propodus and merus; propodus relatively stout, about 4.5–5 times as long as proximal its width, with smooth unarmed parallel margins, without ventral or distoventral teeth (
<figureCitation id="13602A53FF83FFA2FF69FDD70599FD71" box="[159,225,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[192,1406,210,1919]" captionTargetId="figure@4.[173,1413,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Hamopontonia nhatrangensis sp. nov., ovigerous female: a — pereiopod I; b — major pereiopod II; c — same, chela; d — minor pereiopod II; e — same, chela; f — pereiopod III; g — same, distal part of propodus and dactylus." httpUri="https://zenodo.org/record/230076/files/figure.png" pageId="7" pageNumber="138">
Fig. 
<quantity id="4CA39B33FF83FFA2FF25FDD70599FD71" box="[211,225,620,644]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="7" pageNumber="138" unit="g" value="3.0">3</quantity>
</figureCitation>
<emphasis id="B92FEAC4FF83FFA2FF17FDD60597FD71" box="[225,239,621,644]" italics="true" pageId="7" pageNumber="138">g</emphasis>
); dactylus (
<figureCitation id="13602A53FF83FFA2FE85FDD704CDFD71" box="[371,437,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[192,1406,210,1919]" captionTargetId="figure@4.[173,1413,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Hamopontonia nhatrangensis sp. nov., ovigerous female: a — pereiopod I; b — major pereiopod II; c — same, chela; d — minor pereiopod II; e — same, chela; f — pereiopod III; g — same, distal part of propodus and dactylus." httpUri="https://zenodo.org/record/230076/files/figure.png" pageId="7" pageNumber="138">
Fig. 
<quantity id="4CA39B33FF83FFA2FE51FDD704CDFD71" box="[423,437,620,644]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="7" pageNumber="138" unit="g" value="3.0">3</quantity>
</figureCitation>
<emphasis id="B92FEAC4FF83FFA2FE43FDD604BBFD71" box="[437,451,621,644]" italics="true" pageId="7" pageNumber="138">g</emphasis>
) simple, with basal part about 1.5 times as long as wide, with simple slender and curved unguis. Pereiopods III–V similar.
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF31FD0F0146FD05" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
Pleopods normal, without specific features. Uropods slender, not exceeding telson; distolateral margin of uropodal exopod (
<figureCitation id="13602A53FF83FFA2FE95FD6C04DDFD05" box="[355,421,727,752]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="7" pageNumber="138">Fig. 2</figureCitation>
<emphasis id="B92FEAC4FF83FFA2FE53FD6204D5FD05" box="[421,429,729,752]" italics="true" pageId="7" pageNumber="138">j</emphasis>
) with small movable distolateral spine, fixed tooth absent.
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF25FD4704CEFCCD" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
Males morphologically similar to females, but smaller in size, with more developed strongly dissimilar pereiopods II (
<figureCitation id="13602A53FF83FFA2FECDFCA40405FCCD" box="[315,381,799,824]" captionStart="FIGURE 1" captionStartId="2.[151,250,1802,1824]" captionTargetBox="[168,1416,851,1758]" captionTargetId="figure@2.[151,1436,844,1778]" captionTargetPageId="2" captionText="FIGURE 1. Hamopontonia nhatrangensis sp. nov.: a — ovigerous female; b — mature male." httpUri="https://zenodo.org/record/230074/files/figure.png" pageId="7" pageNumber="138">Fig. 1</figureCitation>
, 
<figureCitation id="13602A53FF83FFA2FE7DFC9B04E1FCCD" box="[395,409,800,824]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[196,1368,198,1933]" captionTargetId="figure@3.[196,1392,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. Hamopontonia nhatrangensis sp. nov., ovigerous female (a, b, d – j), male (c): a – c – carapace, lateral view; d — carapace, dorsal view; e — antennula; f — antenna; g—sixth abdominal somite, telson and uropods, dorsal view; h — same, lateral view; i — distal margin of telson, lateral view; j — distolateral margin of uropodal exopod." httpUri="https://zenodo.org/record/230075/files/figure.png" pageId="7" pageNumber="138">2</figureCitation>
<emphasis id="B92FEAC4FF83FFA2FE6FFC9A04DDFCCD" box="[409,421,801,824]" italics="true" pageId="7" pageNumber="138">c</emphasis>
).
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF31FCFF0676FC8A" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
<emphasis id="B92FEAC4FF83FFA2FF31FCFF0405FCA8" bold="true" box="[199,381,836,861]" pageId="7" pageNumber="138">Measurements.</emphasis>
Medium-sized pontoniine shrimp. The largest studied ovigerous female has pcl. 
<quantity id="4CA39B33FF83FFA2FAF0FCFF002FFCA9" box="[1286,1367,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="7" pageNumber="138" unit="mm" value="21.0">21 mm</quantity>
, tl. 
<quantity id="4CA39B33FF83FFA2FA76FCFF05BEFC8A" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="138" unit="mm" value="35.0">35 mm</quantity>
; the largest studied male has pcl. 
<quantity id="4CA39B33FF83FFA2FDCBFCD307F6FC8A" box="[573,654,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="138" unit="mm" value="18.0">18 mm</quantity>
, tl. 
<quantity id="4CA39B33FF83FFA2FD41FCD30671FC8A" box="[695,777,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="138" unit="mm" value="28.0">28 mm</quantity>
.
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF31FC370185FC3D" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
<emphasis id="B92FEAC4FF83FFA2FF31FC370437FC50" bold="true" box="[199,335,908,933]" pageId="7" pageNumber="138">Coloration.</emphasis>
Coloration of the new species clearly imitates the coloration of tentacles of host coral 
<taxonomicName id="4C5B4D55FF83FFA2FAD9FC3707F3FC3D" authority="Chamisso &amp; Eysenhardt, 1821" authorityName="Chamisso &amp; Eysenhardt" authorityYear="1821" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="7" pageNumber="138" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF83FFA2FAD9FC370464FC3D" italics="true" pageId="7" pageNumber="138">Euphyllia glabrescens</emphasis>
(Chamisso &amp; Eysenhardt, 1821)
</taxonomicName>
(Hexacorallia, 
<taxonomicName id="4C5B4D55FF83FFA2FCCFFC1406B8FC3D" box="[825,960,943,968]" class="Anthozoa" kingdom="Animalia" order="Scleractinia" pageId="7" pageNumber="138" phylum="Cnidaria" rank="order">Scleractinia</taxonomicName>
, 
<taxonomicName id="4C5B4D55FF83FFA2FC3DFC140100FC3D" box="[971,1144,943,968]" class="Anthozoa" family="Caryophyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="7" pageNumber="138" phylum="Cnidaria" rank="family">Caryophyllidae</taxonomicName>
(
<figureCitation id="13602A53FF83FFA2FB71FC1401B1FC3D" box="[1159,1225,943,968]" captionStart="FIGURE 5" captionStartId="6.[151,250,1914,1936]" captionTargetBox="[151,1436,193,1891]" captionTargetId="figure@6.[151,1436,193,1892]" captionTargetPageId="6" captionText="FIGURE 5. Coral host Euphyllia glabrescens and Hamopontonia nhatrangensis sp. nov.: a, c — whole colony of host coral Euphyllia glabrescens; b — part of colony with ovigerous female H. nhatrangensis sp. nov. (arrow) in situ; d – f — ovigerous female of Hamopontonia nhatrangensis sp. nov. in natural habitat—among tentacles of coral host E. glabrescens." httpUri="https://zenodo.org/record/230078/files/figure.png" pageId="7" pageNumber="138">Fig. 5</figureCitation>
<emphasis id="B92FEAC4FF83FFA2FB3FFC140195FC3D" box="[1225,1261,943,968]" italics="true" pageId="7" pageNumber="138">d–f</emphasis>
).
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF31FC6F01EEFB55" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
Females. Body and appendages generally transparent, covered with tiny red dots; cornea black, eyestalk transparent, covered with tiny dots; almost all dorsal surface of carapace and dorsal half of its lateral margins covered with uniformly white or white-pinky patch; dorsal surface and dorsal half of pleurae of abdominal somite II with large semicircular white-pinky patch, dorsal surface and dorsal half of pleurae of abdominal somite III with large circular white-pinky patch, dorsal surface of abdominal somite V mostly covered with small semicircular or oval white-pinky spot, sometimes this spot absent, especially in smaller specimens (
<figureCitation id="13602A53FF83FFA2FBB2FB3C01F1FB55" box="[1092,1161,1159,1184]" captionStart="FIGURE 4" captionStartId="5.[151,250,1924,1946]" captionTargetBox="[196,1385,212,1872]" captionTargetId="figure@5.[196,1386,193,1873]" captionTargetPageId="5" captionText="FIGURE 4. Coloration of Hamopontonia nhatrangensis sp. nov.: a — holotype, ovigerous female, dorsal view; b — ovigerous female, dorsal view; c—same specimen, lateral view; d — mature male, lateral view; e — same specimen, dorsal view; h — same specimens among tentacles of its host, f — different ovigerous females, dorsal and lateral views (g, h represent the same specimen)." httpUri="https://zenodo.org/record/230077/files/figure.png" pageId="7" pageNumber="138">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF31FB1707EEFAC5" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
Males. General coloration similar to females, with less number of dorsal patches. Body and appendages transparent, covered with tiny red dots; cornea black, eyestalk transparent, covered with tiny dots; dorsal surface of carapace covered with medium uniformly white or white-pinky patch; with uniformly white or white-pinky oval dorsal patch on abdominal somite III (
<figureCitation id="13602A53FF83FFA2FDB5FAAC07F0FAC5" box="[579,648,1303,1328]" captionStart="FIGURE 4" captionStartId="5.[151,250,1924,1946]" captionTargetBox="[196,1385,212,1872]" captionTargetId="figure@5.[196,1386,193,1873]" captionTargetPageId="5" captionText="FIGURE 4. Coloration of Hamopontonia nhatrangensis sp. nov.: a — holotype, ovigerous female, dorsal view; b — ovigerous female, dorsal view; c—same specimen, lateral view; d — mature male, lateral view; e — same specimen, dorsal view; h — same specimens among tentacles of its host, f — different ovigerous females, dorsal and lateral views (g, h represent the same specimen)." httpUri="https://zenodo.org/record/230077/files/figure.png" pageId="7" pageNumber="138">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF31FA8700F0F9FD" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
<emphasis id="B92FEAC4FF83FFA2FF31FA87047EFAA0" bold="true" box="[199,262,1340,1365]" pageId="7" pageNumber="138">Host.</emphasis>
The species was found in association with the caryophyllid coral 
<taxonomicName id="4C5B4D55FF83FFA2FBF7FA870416FA8D" authority="Chamisso &amp; Eysenhardt, 1821" authorityName="Chamisso &amp; Eysenhardt" authorityYear="1821" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="7" pageNumber="138" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF83FFA2FBF7FA870184FAA0" box="[1025,1276,1340,1365]" italics="true" pageId="7" pageNumber="138">Euphyllia glabrescens</emphasis>
(Chamisso &amp; Eysenhardt, 1821)
</taxonomicName>
. The coral-associated pontoniine shrimp 
<taxonomicName id="4C5B4D55FF83FFA2FCACFADA0068FA8D" authority="Marin &amp; Anker, 2005" authorityName="Marin &amp; Anker" authorityYear="2005" box="[858,1296,1375,1400]" class="Malacostraca" family="Palaemonidae" genus="Vir" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="euphyllius">
<emphasis id="B92FEAC4FF83FFA2FCACFADA0684FA8D" box="[858,1020,1375,1400]" italics="true" pageId="7" pageNumber="138">Vir euphyllius</emphasis>
Marin &amp; Anker, 2005
</taxonomicName>
(Crustacea, 
<taxonomicName id="4C5B4D55FF83FFA2FF61FA3F0472FA68" box="[151,266,1412,1437]" class="Malacostraca" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="order">Decapoda</taxonomicName>
, 
<taxonomicName id="4C5B4D55FF83FFA2FEEFFA3F04C1FA68" box="[281,441,1412,1437]" class="Malacostraca" family="Palaemonidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="family">Palaemonidae</taxonomicName>
, 
<taxonomicName id="4C5B4D55FF83FFA2FE3FFA3F0729FA68" box="[457,593,1412,1437]" class="Malacostraca" family="Palaemonidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="subFamily" subFamily="Pontoniinae">Pontoniinae</taxonomicName>
) is the other pontoniine species living on colonies of 
<taxonomicName id="4C5B4D55FF83FFA2FB25FA3E0003FA68" box="[1235,1403,1412,1437]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="7" pageNumber="138" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF83FFA2FB25FA3E0003FA68" box="[1235,1403,1412,1437]" italics="true" pageId="7" pageNumber="138">E. glabrescens</emphasis>
</taxonomicName>
in Nhatrang Bay, 
<collectingCountry id="F34C7646FF83FFA2FEB2FA1C04DBFA35" box="[324,419,1447,1472]" name="Vietnam" pageId="7" pageNumber="138">Vietnam</collectingCountry>
(Marin &amp; Anker, 2005; 
<bibRefCitation id="EFCA4B27FF83FFA2FD34FA1C0618FA35" author="Marin" box="[706,864,1447,1472]" pageId="7" pageNumber="138" refString="Marin, I. N. (2007 a) Notes on taxonomy and biology of the symbiotic shrimp Vir euphyllius Marin &amp; Anker, 2005 (Decapoda, Palaemonidae, Pontoniinae), associated with hammer corals Euphyllia spp (Cnidaria, Caryophyllidae). Invertebrate Zoology, 4 (1), 15 - 23." type="journal article" year="2007" yearSuffix="a">Marin, 2007a</bibRefCitation>
). At the same time, both species have been never observed on the same coral colony, possibly showing a competitive pair of pontoniine shrimp species (see remarks of similar competitive pair 
<taxonomicName id="4C5B4D55FF83FFA2FE3EFA54067DF9FD" authority="Marin, 2012" authorityName="Marin" authorityYear="2012" box="[456,773,1519,1544]" class="Malacostraca" family="Palaemonidae" genus="Philarius" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="condi">
<emphasis id="B92FEAC4FF83FFA2FE3EFA54070DF9FD" box="[456,629,1519,1544]" italics="true" pageId="7" pageNumber="138">Philarius condi</emphasis>
Marin, 2012
</taxonomicName>
— 
<taxonomicName id="4C5B4D55FF83FFA2FCD4FA5401BDF9FD" authority="De Man, 1902" authorityName="De Man" authorityYear="1902" box="[802,1221,1519,1544]" class="Malacostraca" family="Palaemonidae" genus="Harpilius" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B92FEAC4FF83FFA2FCD4FA540166F9FD" box="[802,1054,1519,1544]" italics="true" pageId="7" pageNumber="138">Harpilius consobrinus</emphasis>
De Man, 1902
</taxonomicName>
<bibRefCitation id="EFCA4B27FF83FFA2FB3AFA5400FCF9FD" author="Marin" box="[1228,1412,1519,1544]" pageId="7" pageNumber="138" refString="Marin, I. (2012 b) A new species of the genus Philarius Holthuis, 1952 (Crustacea: Decapoda: Palaemonidae) from the Great Barrier Reef of Australia. Zootaxa, 3182, 43 - 50." type="journal article" year="2012" yearSuffix="b">(Marin, 2012b))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C341655DFF83FFADFF31F9AF01DBFDED" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138" type="distribution">
<paragraph id="8BE436D6FF83FFA2FF31F9AF0029F9D8" blockId="7.[151,1437,151,2013]" box="[199,1361,1556,1581]" pageId="7" pageNumber="138">
<emphasis id="B92FEAC4FF83FFA2FF31F9AF0418F9D8" bold="true" box="[199,352,1556,1581]" pageId="7" pageNumber="138">Distribution.</emphasis>
The species is presently known exclusively from its 
<typeStatus id="54E08874FF83FFA2FC5CF9AE06A2F9D8" box="[938,986,1557,1581]" pageId="7" pageNumber="138">type</typeStatus>
locality, Nhatrang Bay, 
<collectingCountry id="F34C7646FF83FFA2FB1CF9AF0035F9D8" box="[1258,1357,1556,1581]" name="Vietnam" pageId="7" pageNumber="138">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF31F98C078DF981" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="138">
<emphasis id="B92FEAC4FF83FFA2FF31F98C04B5F9A5" bold="true" box="[199,461,1591,1616]" pageId="7" pageNumber="138">Differential diagnosis.</emphasis>
The new species can be easily differentiated from congeners by specific coloration and occupied host (following 
<bibRefCitation id="EFCA4B27FF83FFA2FE40F9E6073AF980" author="Bruce" box="[438,578,1628,1653]" pageId="7" pageNumber="138" refString="Bruce, A. J. (1986) Three new species of commensal shrimps from Port Essington, Arnhem Land, northern Australia (Crustacea: Decapoda: Palaemonidae). The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 3, 143 - 166." type="journal article" year="1986">Bruce, 1986</bibRefCitation>
; 
<bibRefCitation id="EFCA4B27FF83FFA2FDBBF9E70791F981" author="Marin" box="[589,745,1628,1653]" pageId="7" pageNumber="138" refString="Marin, I. (2012 a) The &quot; Hamopontonia corallicola &quot; Bruce, 1970 species complex (Crustacea, Decapoda, Palaemonidae): new records and new species from the Great Barrier Reef, Australia. Zootaxa, 3272, 35 - 56." type="journal article" year="2012" yearSuffix="a">Marin, 2012a</bibRefCitation>
).
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF31F93A06C0F96D" blockId="7.[151,1437,151,2013]" box="[199,952,1663,1688]" pageId="7" pageNumber="138">
Moreover, 
<taxonomicName id="4C5B4D55FF83FFA2FEB4F93A07E8F96D" box="[322,656,1663,1688]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF83FFA2FEB4F93A07E8F96D" box="[322,656,1663,1688]" italics="true" pageId="7" pageNumber="138">Hamopontonia nhatrangensis</emphasis>
</taxonomicName>
<emphasis id="B92FEAC4FF83FFA2FD61F9C40788F96D" bold="true" box="[663,752,1663,1688]" pageId="7" pageNumber="138">
<taxonomicNameLabel id="A21C57BFFF83FFA2FD61F9C40788F96D" box="[663,752,1663,1688]" pageId="7" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated:
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF61F91E00E4F949" blockId="7.[151,1437,151,2013]" box="[151,1436,1700,1725]" pageId="7" pageNumber="138">
- from 
<taxonomicName id="4C5B4D55FF83FFA2FEF0F91E04E4F949" box="[262,412,1701,1724]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="essingtoni">
<emphasis id="B92FEAC4FF83FFA2FEF0F91E04E4F949" box="[262,412,1701,1724]" italics="true" pageId="7" pageNumber="138">H. essingtoni</emphasis>
</taxonomicName>
by absence of specific blunt median process between hook-like distolateral processes on
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF33F97C0477F915" blockId="7.[151,1437,151,2013]" box="[197,271,1735,1760]" pageId="7" pageNumber="138">telson.</paragraph>
<paragraph id="8BE436D6FF83FFA2FF61F95600E4F8F0" blockId="7.[151,1437,151,2013]" box="[151,1436,1772,1797]" pageId="7" pageNumber="138">
<emphasis id="B92FEAC4FF83FFA2FF61F95605D8F8F1" box="[151,160,1773,1796]" italics="true" pageId="7" pageNumber="138">-</emphasis>
from 
<taxonomicName id="4C5B4D55FF83FFA2FEF2F95604D9F8F0" box="[260,417,1772,1797]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="corallicola">
<emphasis id="B92FEAC4FF83FFA2FEF2F95604D9F8F0" box="[260,417,1772,1797]" italics="true" pageId="7" pageNumber="138">H. corallicola</emphasis>
</taxonomicName>
by swollen carapace, more slender and shorter rostrum, shorter dorsal spines on telson and
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF33F8AA06E4F8DD" blockId="7.[151,1437,151,2013]" box="[197,924,1807,1833]" pageId="7" pageNumber="138">stouter segments of ambulatory pereiopods, especially propodus.</paragraph>
<paragraph id="8BE436D6FF83FFA2FF61F88E00E4F8B8" blockId="7.[151,1437,151,2013]" box="[151,1436,1844,1869]" pageId="7" pageNumber="138">
- from 
<taxonomicName id="4C5B4D55FF83FFA2FEFFF88E04DCF8B8" box="[265,420,1844,1869]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="physogyra">
<emphasis id="B92FEAC4FF83FFA2FEFFF88E04DCF8B8" box="[265,420,1844,1869]" italics="true" pageId="7" pageNumber="138">H. physogyra</emphasis>
</taxonomicName>
by slender rostrum, shorter dorsal spines on telson and stouter ambulatory pereiopods
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF33F8EC0482F885" blockId="7.[151,1437,151,2013]" box="[197,506,1879,1904]" pageId="7" pageNumber="138">without distoventral spines.</paragraph>
<paragraph id="8BE436D6FF83FFA2FF61F8C600E4F860" blockId="7.[151,1437,151,2013]" box="[151,1436,1916,1941]" pageId="7" pageNumber="138">
- rom 
<taxonomicName id="4C5B4D55FF83FFA2FF0DF8C604F0F860" box="[251,392,1916,1941]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="fungicola">
<emphasis id="B92FEAC4FF83FFA2FF0DF8C604F0F860" box="[251,392,1916,1941]" italics="true" pageId="7" pageNumber="138">H. fungicola</emphasis>
</taxonomicName>
by shorter rostrum, shorter dorsal spines on telson, smaller and dissimilar in size pereiopods
</paragraph>
<paragraph id="8BE436D6FF83FFA2FF33F81A0712F84C" blockId="7.[151,1437,151,2013]" box="[197,618,1952,1977]" pageId="7" pageNumber="138">II and stouter ambulatory pereiopods.</paragraph>
<paragraph id="8BE436D6FF83FFADFF31F87F016AFE59" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1436,151,429]" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138">
The new species is morphologically most similar to 
<taxonomicName id="4C5B4D55FF83FFA2FCCFF87E06ADF828" box="[825,981,1988,2013]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="physogyra">
<emphasis id="B92FEAC4FF83FFA2FCCFF87E06ADF828" box="[825,981,1988,2013]" italics="true" pageId="7" pageNumber="138">H. physogyra</emphasis>
</taxonomicName>
and 
<emphasis id="B92FEAC4FF83FFA2FBE3F87E01D5F828" box="[1045,1197,1988,2013]" italics="true" pageId="7" pageNumber="138">
<taxonomicName id="4C5B4D55FF83FFA2FBE3F87E01D1F828" box="[1045,1193,1988,2013]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="fungicola">H. fungicola</taxonomicName>
,
</emphasis>
resembling them in coloration (see 
<bibRefCitation id="EFCA4B27FF8CFFADFEB2FF2C0499FF45" author="Marin" box="[324,481,151,176]" pageId="8" pageNumber="139" refString="Marin, I. (2012 a) The &quot; Hamopontonia corallicola &quot; Bruce, 1970 species complex (Crustacea, Decapoda, Palaemonidae): new records and new species from the Great Barrier Reef, Australia. Zootaxa, 3272, 35 - 56." type="journal article" year="2012" yearSuffix="a">Marin, 2012a</bibRefCitation>
). At the same time, the coloration of the new species clearly differs from 
<taxonomicName id="4C5B4D55FF8CFFADFAEBFF2200E4FF45" box="[1309,1436,151,176]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="physogy">
<emphasis id="B92FEAC4FF8CFFADFAEBFF2200E4FF45" box="[1309,1436,151,176]" italics="true" pageId="8" pageNumber="139">H. physogy</emphasis>
</taxonomicName>
by being generally translucent with tiny small dots on the body and appendages as well as having uniformly white or pinky-white colored dorsal patches on carapace and abdomen (vs. body and appendages covered with numerous white and blue dots as well as dorsal patches consisting of white spots in the centre of the patch fringed with broad white or creamy band in 
<taxonomicName id="4C5B4D55FF8CFFADFE32FE920727FEB5" box="[452,607,295,320]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="physogyra">
<emphasis id="B92FEAC4FF8CFFADFE32FE920727FEB5" box="[452,607,295,320]" italics="true" pageId="8" pageNumber="139">H. physogyra</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFCA4B27FF8CFFADFD55FE9C063BFEB5" author="Marin" box="[675,835,295,320]" pageId="8" pageNumber="139" refString="Marin, I. (2012 a) The &quot; Hamopontonia corallicola &quot; Bruce, 1970 species complex (Crustacea, Decapoda, Palaemonidae): new records and new species from the Great Barrier Reef, Australia. Zootaxa, 3272, 35 - 56." type="journal article" year="2012" yearSuffix="a">Marin, 2012a</bibRefCitation>
: 55, fig. 13)). The new species also can be clearly separated by coloration from 
<taxonomicName id="4C5B4D55FF8CFFADFE12FEF60709FE90" box="[484,625,332,357]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="fungicola">
<emphasis id="B92FEAC4FF8CFFADFE12FEF60709FE90" box="[484,625,332,357]" italics="true" pageId="8" pageNumber="139">H. fungicola</emphasis>
</taxonomicName>
by having smaller dorsal spots on the carapace, not covering almost all dorsal surface, as well as a different pattern and smaller dorsal spots on abdominal somites II and III not almost covering the abdominal pleurae (see 
<bibRefCitation id="EFCA4B27FF8CFFADFDC2FE2F07B7FE59" author="Marin" box="[564,719,404,429]" pageId="8" pageNumber="139" refString="Marin, I. (2012 a) The &quot; Hamopontonia corallicola &quot; Bruce, 1970 species complex (Crustacea, Decapoda, Palaemonidae): new records and new species from the Great Barrier Reef, Australia. Zootaxa, 3272, 35 - 56." type="journal article" year="2012" yearSuffix="a">Marin, 2012a</bibRefCitation>
: fig. 7; present paper, fig. 4).
</paragraph>
<paragraph id="8BE436D6FF8CFFADFF61FE4401DBFDED" blockId="8.[151,1187,511,536]" box="[151,1187,511,536]" pageId="8" pageNumber="139">
<emphasis id="B92FEAC4FF8CFFADFF61FE4406C0FDED" bold="true" box="[151,952,511,536]" pageId="8" pageNumber="139">
Renewed key to the species of the genus 
<taxonomicName id="4C5B4D55FF8CFFADFD80FDBB06C0FDED" authority="Bruce, 1970" authorityName="Bruce" authorityYear="1970" box="[630,952,511,536]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEAC4FF8CFFADFD80FDBB065CFDED" bold="true" box="[630,804,512,536]" italics="true" pageId="8" pageNumber="139">Hamopontonia</emphasis>
<bibRefCitation id="EFCA4B27FF8CFFADFCDDFE4406C0FDED" author="Bruce" box="[811,952,511,536]" pageId="8" pageNumber="139" refString="Bruce, A. J. (1970) Notes on some Indo-Pacific Pontoniinae. XV. Hamopontonia corallicola gen. nov., sp. nov., a new pontoniid shrimp from Hong Kong. Crustaceana, 18, 37 - 48. http: // dx. doi. org / 10.1163 / 156854070 x 00040" type="journal article" year="1970">Bruce, 1970</bibRefCitation>
</taxonomicName>
</emphasis>
(after 
<bibRefCitation id="EFCA4B27FF8CFFADFBF5FE4401E3FDED" author="Marin" box="[1027,1179,511,536]" pageId="8" pageNumber="139" refString="Marin, I. (2012 a) The &quot; Hamopontonia corallicola &quot; Bruce, 1970 species complex (Crustacea, Decapoda, Palaemonidae): new records and new species from the Great Barrier Reef, Australia. Zootaxa, 3272, 35 - 56." type="journal article" year="2012" yearSuffix="a">Marin, 2012a</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C341655DFF8CFFADFF61FDFC00E4FBBA" pageId="8" pageNumber="139" type="key">
<key id="FFCA61E7FF8CFFADFF61FDFC00E4FBBA" pageId="8" pageNumber="139">
<keyStep id="30AF2E73FF8CFFADFF61FDFC00E3FD27" pageId="8" pageNumber="139">
<paragraph id="8BE436D6FF8CFFADFF61FDFC00E3FD62" blockId="8.[151,1436,583,1103]" pageId="8" pageNumber="139">
<keyLead id="30AA95E3FF8CFFADFF61FDFC00E3FD62" pageId="8" pageNumber="139">
1. Distal margin of telson with small blunt median process between hook-like distolateral processes. Small species; with translucent body and appendages covered with numerous tiny red dots. Without large bright colored spots or patches on dorsal surface of carapace and abdominal somites. Associated with 
<taxonomicName id="4C5B4D55FF8CFFADFD2BFD3A013AFD62" box="[733,1090,641,663]" class="Insecta" family="Pocilloporidae" genus="Stylophora" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="pistillata">
<emphasis id="B92FEAC4FF8CFFADFD2BFD3A06D8FD62" box="[733,928,641,663]" italics="true" pageId="8" pageNumber="139">Stylophora pistillata</emphasis>
(Pocilloporidae)
</taxonomicName>
....... 
<taxonomicName id="4C5B4D55FF8CFFADFB54FD3900E3FD62" ID-CoL="ba896df8-6c42-48a6-9a85-a138a92ca22c" authority="Bruce, 1986" authorityName="Bruce" authorityYear="1986" box="[1186,1435,641,663]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="essingtoni">
<emphasis id="B92FEAC4FF8CFFADFB54FD390058FD62" box="[1186,1312,642,663]" italics="true" pageId="8" pageNumber="139">H. essingtoni</emphasis>
<bibRefCitation id="EFCA4B27FF8CFFADFAD0FD3900E3FD62" author="Bruce" box="[1318,1435,641,663]" pageId="8" pageNumber="139" refString="Bruce, A. J. (1986) Three new species of commensal shrimps from Port Essington, Arnhem Land, northern Australia (Crustacea: Decapoda: Palaemonidae). The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 3, 143 - 166." type="journal article" year="1986">Bruce, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE436D6FF8CFFADFF61FD2400E3FD27" blockId="8.[151,1436,583,1103]" pageId="8" pageNumber="139">
<keyLead id="30AA95E3FF8CFFADFF61FD2400E3FD27" pageId="8" pageNumber="139">- Distal margin of telson without blunt median process between hook-like distolateral processes. Large bright colored spots or patches present on dorsal surface of carapace and abdomen.................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E73FF8CFFADFF61FD6200E3FC90" pageId="8" pageNumber="139">
<paragraph id="8BE436D6FF8CFFADFF61FD6200E4FCDF" blockId="8.[151,1436,583,1103]" pageId="8" pageNumber="139">
<keyLead id="30AA95E3FF8CFFADFF61FD6200E4FCDF" pageId="8" pageNumber="139">
2. Pereiopods II large, longer than carapace length, equal in size. Almost all dorsal and lateral surface of carapace and pleura of abdominal somites II–III covered with large white or creamy colored patches. Associated with fungiid corals, mainly with 
<taxonomicName id="4C5B4D55FF8CFFADFF17FCAF0744FCDF" ID-CoL="3KB7Y" box="[225,572,788,810]" class="Anthozoa" family="Fungiidae" genus="Heliofungia" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="species" species="actiniformis">
<emphasis id="B92FEAC4FF8CFFADFF17FCAF04B4FCDF" box="[225,460,788,810]" italics="true" pageId="8" pageNumber="139">Heliofungia actiniformis</emphasis>
(Fungidae)
</taxonomicName>
.................................................. 
<taxonomicName id="4C5B4D55FF8CFFADFB50FCAE00E4FCDF" ID-CoL="10cbb7e2-108f-441d-90e2-be4b941496ba" authority="Marin, 2012" authorityName="Marin" authorityYear="2012" box="[1190,1436,788,810]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="fungicola">
<emphasis id="B92FEAC4FF8CFFADFB50FCAE0066FCDF" box="[1190,1310,788,810]" italics="true" pageId="8" pageNumber="139">H. fungicola</emphasis>
Marin, 2012
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE436D6FF8CFFADFF61FC8900E3FC90" blockId="8.[151,1436,583,1103]" pageId="8" pageNumber="139">
<keyLead id="30AA95E3FF8CFFADFF61FC8900E3FC90" pageId="8" pageNumber="139">- Pereiopods II equal or smaller than carapace length, unequal in size. Carapace and abdominal somites with only dorsal and upper half of lateral margin covered with brightly colored patches............................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E73FF8CFFADFF61FCD700E3FC2F" pageId="8" pageNumber="139">
<paragraph id="8BE436D6FF8CFFADFF61FCD700E4FC55" blockId="8.[151,1436,583,1103]" pageId="8" pageNumber="139">
<keyLead id="30AA95E3FF8CFFADFF61FCD700E4FC55" pageId="8" pageNumber="139">
3. Rostrum relatively short, reaching to distal margin of basal antennular segment. Associated with 
<taxonomicName id="4C5B4D55FF8CFFADFB34FCD7041CFC6A" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF8CFFADFB34FCD700E4FC77" box="[1218,1436,876,898]" italics="true" pageId="8" pageNumber="139">Euphyllia glabrescens</emphasis>
(Euphyllidae)
</taxonomicName>
.................................................................... 
<taxonomicName id="4C5B4D55FF8CFFADFB55FC310031FC6A" ID-CoL="eff5cab1-d6ef-42b5-9d31-e0effa68d85b" box="[1187,1353,905,927]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="nhatrangensis" status="sp. nov.">
<emphasis id="B92FEAC4FF8CFFADFB55FC310031FC6A" box="[1187,1353,905,927]" italics="true" pageId="8" pageNumber="139">H. nhatrangensis</emphasis>
</taxonomicName>
<emphasis id="B92FEAC4FF8CFFADFAB9FC3100E4FC55" bold="true" box="[1359,1436,906,928]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A21C57BFFF8CFFADFAB9FC3100E4FC55" box="[1359,1436,906,928]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE436D6FF8CFFADFF61FC1C00E3FC2F" blockId="8.[151,1436,583,1103]" pageId="8" pageNumber="139">
<keyLead id="30AA95E3FF8CFFADFF61FC1C00E3FC2F" pageId="8" pageNumber="139">- Rostrum long, significantly overreaching the distal margin of basal antennular segment reaching to distal margin of antennula peduncle........................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E73FF8CFFADFF61FC5A00E4FBBA" pageId="8" pageNumber="139">
<paragraph id="8BE436D6FF8CFFADFF61FC5A00E3FBE0" blockId="8.[151,1436,583,1103]" pageId="8" pageNumber="139">
<keyLead id="30AA95E3FF8CFFADFF61FC5A00E3FBE0" pageId="8" pageNumber="139">
4. Rostrum with well developed deep dorsal carina. Single large dorsal circle patch presents on dorsal surface of abdominal somite III. Associated with caryophyllid coral 
<emphasis id="B92FEAC4FF8CFFADFD69FC4406F9FBE0" box="[671,897,1023,1045]" italics="true" pageId="8" pageNumber="139">
<taxonomicName id="4C5B4D55FF8CFFADFD69FC44067EFBE0" ID-CoL="a7050964-7795-489f-8ca4-2d3e7bb0665c" box="[671,774,1023,1045]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="physogyra">Physogyra</taxonomicName>
lichtensteini
</emphasis>
(
<taxonomicName id="4C5B4D55FF8CFFADFC7BFC44015CFBE0" box="[909,1060,1023,1045]" class="Anthozoa" family="Caryophyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="family">Caryophyllidae</taxonomicName>
)................... 
<taxonomicName id="4C5B4D55FF8CFFADFAECFBBB00E3FBE0" ID-CoL="a7050964-7795-489f-8ca4-2d3e7bb0665c" box="[1306,1435,1023,1045]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="physogyra">
<emphasis id="B92FEAC4FF8CFFADFAECFBBB00E3FBE0" box="[1306,1435,1023,1045]" italics="true" pageId="8" pageNumber="139">H. physogyra</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE436D6FF8CFFADFF61FBA600E4FBBA" blockId="8.[151,1436,583,1103]" pageId="8" pageNumber="139">
<keyLead id="30AA95E3FF8CFFADFF61FBA600E4FBBA" pageId="8" pageNumber="139">
- Rostrum with shallow dorsal carina. Dorsal surface of abdominal somites I–IV covered with brightly colored patches. Associated with poritid coral 
<taxonomicName id="4C5B4D55FF8CFFADFE4DFB82075CFBBA" ID-CoL="87G7S" box="[443,548,1081,1103]" class="Anthozoa" family="Poritidae" genus="Goniopora" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="genus">
<emphasis id="B92FEAC4FF8CFFADFE4DFB82075CFBBA" box="[443,548,1081,1103]" italics="true" pageId="8" pageNumber="139">Goniopora</emphasis>
</taxonomicName>
spp (
<taxonomicName id="4C5B4D55FF8CFFADFDAEFB8207CAFBBA" ID-CoL="84KPB" box="[600,690,1081,1103]" class="Anthozoa" family="Poritidae" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="family">Poritidae</taxonomicName>
)................................................. 
<taxonomicName id="4C5B4D55FF8CFFADFAE3FB8100E4FBBA" ID-CoL="5d0d23d1-bcbc-479a-9bc4-45b92724caae" box="[1301,1436,1081,1103]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="corallicola">
<emphasis id="B92FEAC4FF8CFFADFAE3FB8100E4FBBA" box="[1301,1436,1081,1103]" italics="true" pageId="8" pageNumber="139">H. corallicola</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="C341655DFF8CFFADFF31FBC7069AFA09" pageId="8" pageNumber="139" type="discussion">
<paragraph id="8BE436D6FF8CFFADFF31FBC707B6FA65" blockId="8.[151,1437,1148,1533]" pageId="8" pageNumber="139">
<emphasis id="B92FEAC4FF8CFFADFF31FBC70443FB60" bold="true" box="[199,315,1148,1173]" pageId="8" pageNumber="139">Remarks.</emphasis>
The Torch coral 
<taxonomicName id="4C5B4D55FF8CFFADFE0FFBC7078AFB60" box="[505,754,1148,1173]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF8CFFADFE0FFBC7078AFB60" box="[505,754,1148,1173]" italics="true" pageId="8" pageNumber="139">Euphyllia glabrescens</emphasis>
</taxonomicName>
belongs to the family 
<taxonomicName id="4C5B4D55FF8CFFADFC1CFBC7010CFB60" box="[1002,1140,1148,1173]" class="Anthozoa" family="Euphyllidae" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="family">Euphyllidae</taxonomicName>
, not the 
<taxonomicName id="4C5B4D55FF8CFFADFB24FBC7003DFB60" box="[1234,1349,1148,1173]" class="Anthozoa" family="Fungiidae" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="family">Fungiidae</taxonomicName>
, but the colonies of 
<taxonomicName id="4C5B4D55FF8CFFADFED2FB1A04B5FB4D" box="[292,461,1183,1208]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF8CFFADFED2FB1A04B5FB4D" box="[292,461,1183,1208]" italics="true" pageId="8" pageNumber="139">E. glabrescens</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5B4D55FF8CFFADFDFBFB240126FB4D" authority="Quoy &amp; Gaimard, 1833" authorityName="Quoy &amp; Gaimard" authorityYear="1833" box="[525,1118,1183,1208]" class="Anthozoa" family="Fungiidae" genus="Heliofungia" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="species" species="actiniformis">
<emphasis id="B92FEAC4FF8CFFADFDFBFB24065EFB4D" box="[525,806,1183,1208]" italics="true" pageId="8" pageNumber="139">Heliofungia actiniformis</emphasis>
(Quoy &amp; Gaimard, 1833)
</taxonomicName>
(Hexacorallia, 
<taxonomicName id="4C5B4D55FF8CFFADFAE7FB2400E0FB4D" box="[1297,1432,1183,1208]" class="Anthozoa" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="order">Scleractinia</taxonomicName>
, Fungidae) are greatly similar (see 
<figureCitation id="13602A53FF8CFFADFDE3FB7F072DFB28" box="[533,597,1220,1245]" captionStart="FIGURE 5" captionStartId="6.[151,250,1914,1936]" captionTargetBox="[151,1436,193,1891]" captionTargetId="figure@6.[151,1436,193,1892]" captionTargetPageId="6" captionText="FIGURE 5. Coral host Euphyllia glabrescens and Hamopontonia nhatrangensis sp. nov.: a, c — whole colony of host coral Euphyllia glabrescens; b — part of colony with ovigerous female H. nhatrangensis sp. nov. (arrow) in situ; d – f — ovigerous female of Hamopontonia nhatrangensis sp. nov. in natural habitat—among tentacles of coral host E. glabrescens." httpUri="https://zenodo.org/record/230078/files/figure.png" pageId="8" pageNumber="139">Fig. 5</figureCitation>
<emphasis id="B92FEAC4FF8CFFADFDA3FB7E0706FB29" box="[597,638,1221,1244]" italics="true" pageId="8" pageNumber="139">a–c</emphasis>
; 
<bibRefCitation id="EFCA4B27FF8CFFADFD7AFB7E0662FB29" author="Veron" box="[652,794,1220,1244]" pageId="8" pageNumber="139" refString="Veron J. E. N. (2000) Corals of the World. Vol. 2. Townsville. Australian Institute of Marine Science, 435 pp." type="book" year="2000">Veron, 2000</bibRefCitation>
). The coloration of the tentacles of 
<taxonomicName id="4C5B4D55FF8CFFADFB52FB7E003FFB28" box="[1188,1351,1220,1245]" class="Anthozoa" family="Euphyllidae" genus="Euphyllia" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="species" species="glabrescens">
<emphasis id="B92FEAC4FF8CFFADFB52FB7E003FFB28" box="[1188,1351,1220,1245]" italics="true" pageId="8" pageNumber="139">E. glabrescens</emphasis>
</taxonomicName>
bears a striking resemblance to the coloration of tentacles of 
<taxonomicName id="4C5B4D55FF8CFFADFCE2FB5206BBFAF5" box="[788,963,1256,1280]" class="Anthozoa" family="Fungiidae" genus="Heliofungia" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="species" species="actiniformis">
<emphasis id="B92FEAC4FF8CFFADFCE2FB5206BBFAF5" box="[788,963,1256,1280]" italics="true" pageId="8" pageNumber="139">H. actiniformis</emphasis>
</taxonomicName>
. At the same time, a multi-year survey (
<date id="FFE51016FF8CFFADFF68FAB70466FAD0" box="[158,286,1292,1317]" pageId="8" pageNumber="139" value="2005" valueMax="2014">2005–2014</date>
years) of coral reef communities in Nhatrang Bay revealed the absence of 
<taxonomicName id="4C5B4D55FF8CFFADFB9CFAB70006FAD1" box="[1130,1406,1292,1317]" class="Anthozoa" family="Fungiidae" genus="Heliofungia" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="species" species="actiniformis">
<emphasis id="B92FEAC4FF8CFFADFB9CFAB70006FAD1" box="[1130,1406,1292,1317]" italics="true" pageId="8" pageNumber="139">Heliofungia actiniformis</emphasis>
</taxonomicName>
in the Bay and adjacent localitites (
<bibRefCitation id="EFCA4B27FF8CFFADFDF9FA8B0119FABD" author="Latypov" box="[527,1121,1327,1352]" pageId="8" pageNumber="139" refString="Latypov Yu. Ya. (2006) The common corals of Vietnam: field handbook. Vladivostok: Far Eastern National University Press. 148 pp. [in Russian]" type="book" year="2006">Latypov, 2006; Yu. Latypov, I. Marin, pers. observ.</bibRefCitation>
) as well as two specialized associated pontoniine shrimp species, 
<taxonomicName id="4C5B4D55FF8CFFADFD92FAEE0781FA98" box="[612,761,1364,1389]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="fungicola">
<emphasis id="B92FEAC4FF8CFFADFD92FAEE0781FA98" box="[612,761,1364,1389]" italics="true" pageId="8" pageNumber="139">H. fungicola</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5B4D55FF8CFFADFCB6FAEE0199FA99" authority="Bruce, 1977" authorityName="Bruce" authorityYear="1977" box="[832,1249,1364,1389]" class="Malacostraca" family="Palaemonidae" genus="Cuapetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="kororensis">
<emphasis id="B92FEAC4FF8CFFADFCB6FAEE0154FA98" box="[832,1068,1364,1389]" italics="true" pageId="8" pageNumber="139">Cuapetes kororensis</emphasis>
(Bruce, 1977)
</taxonomicName>
, specific to 
<taxonomicName id="4C5B4D55FF8CFFADFA77FAEE0466FA65" class="Anthozoa" family="Fungiidae" genus="Heliofungia" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="139" phylum="Cnidaria" rank="species" species="actiniformis">
<emphasis id="B92FEAC4FF8CFFADFA77FAEE0466FA65" italics="true" pageId="8" pageNumber="139">H. actiniformis</emphasis>
</taxonomicName>
, have never been observed in the Bay.
</paragraph>
<paragraph id="8BE436D6FF8CFFADFF31FA27069AFA09" blockId="8.[151,1437,1148,1533]" pageId="8" pageNumber="139">
This record increase the number of scleractinian coral-associated pontoniine shrimp species known from Nhatrang Bay, 
<collectingCountry id="F34C7646FF8CFFADFEB0FA0404DDFA2D" box="[326,421,1471,1496]" name="Vietnam" pageId="8" pageNumber="139">Vietnam</collectingCountry>
up to 31, while 
<taxonomicName id="4C5B4D55FF8CFFADFD9BFA7A0786FA2D" box="[621,766,1471,1496]" class="Malacostraca" family="Palaemonidae" genus="Hamopontonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="fungicola">
<emphasis id="B92FEAC4FF8CFFADFD9BFA7A0786FA2D" box="[621,766,1471,1496]" italics="true" pageId="8" pageNumber="139">H. fungicola</emphasis>
</taxonomicName>
should be excluded from the list (Marin &amp; Anker, 2005; 
<bibRefCitation id="EFCA4B27FF8CFFADFF61FA5F044AFA08" author="Marin" box="[151,306,1508,1533]" pageId="8" pageNumber="139" refString="Marin, I. (2007 b) The coral-associated shrimp genus Pontonides (Caridea, Palaemonidae, Pontoniinae) in Nhatrang Bay, Vietnam, with description of two new species. Zootaxa, 1653, 1 - 21." type="journal article" year="2007" yearSuffix="b">Marin, 2007b</bibRefCitation>
, 
<bibRefCitation id="EFCA4B27FF8CFFADFECBFA5F0401FA09" author="Marin" box="[317,377,1508,1532]" pageId="8" pageNumber="139" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Vir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, Leiden, 82 (35), 375 - 390." type="journal article" year="2008">2008</bibRefCitation>
; reviews in 
<bibRefCitation id="EFCA4B27FF8CFFADFE09FA5F0669FA09" author="Marin" box="[511,785,1508,1533]" pageId="8" pageNumber="139" refString="Marin, I. N. &amp; Savinkin, O. V. (2007) Further records and preliminary list of pontoniine (Caridea: Palaemonidae: Pontoniinae) and hymenocerid (Caridea: Hymenoceridae) shrimps from Nhatrang Bay. In: Britayev, T. A., Pavlov, D. S. (Eds.), Benthic fauna of the Bay of Nhatrang, Southern Vietnam. Vol. 2. KMK-Press, Moscow, pp. 175 - 208." type="book" year="2007">Marin &amp; Savinkin, 2007</bibRefCitation>
and 
<bibRefCitation id="EFCA4B27FF8CFFADFCB1FA5F06AEFA09" author="Marin" box="[839,982,1508,1533]" pageId="8" pageNumber="139" refString="Marin, I. N. (2013) New records and associations of pontoniine shrimps (Crustacea: Decapoda: Caridea: Palaemonidae: Pontoniinae) from the Nhatrang Bay, Vietnam; with taxonomic remarks on some species from the Indo-West Pacific region. In: Britayev, T. A., Pavlov, D. S. (Eds.), Benthic fauna of the Bay of Nhatrang, Southern Vietnam. Vol. 2. KMK- Press, Moscow, pp. 339 - 397." type="book" year="2013">Marin, 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>