125 lines
18 KiB
XML
125 lines
18 KiB
XML
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<mods:title id="8B9723634ADB3E297998136B8059E298">The weevil genera Nyphaeba Pascoe and Pantiala Pascoe and the problems of an unstable nomenclature in orphaned taxa</mods:title>
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<mods:namePart id="8BEFC99525C5C14C3D2744B7634ACB1F">Riedel, Alexander</mods:namePart>
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<mods:title id="838D520D50DF0C51173BAEEB58F2FB6B">Zootaxa</mods:title>
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<mods:date id="3A7F383F4EE744E82FE5EA7DB377ECB6">2017</mods:date>
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<mods:identifier id="6C61832CAB68A45DCAAAF2F55BE6A578" type="DOI">10.11646/zootaxa.4244.3.6</mods:identifier>
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<treatment id="03F287B6067CFF9954F3CB53FAAB2D71" ID-DOI="http://doi.org/10.5281/zenodo.6041767" ID-GBIF-Taxon="127848131" ID-Zenodo-Dep="6041767" LSID="urn:lsid:plazi:treatment:03F287B6067CFF9954F3CB53FAAB2D71" httpUri="http://treatment.plazi.org/id/03F287B6067CFF9954F3CB53FAAB2D71" lastPageId="2" lastPageNumber="379" pageId="1" pageNumber="378">
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<subSubSection id="C341652B067CFF9A54F3CB53FEC02EEB" box="[151,364,1661,1687]" pageId="1" pageNumber="378" type="nomenclature">
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<paragraph id="8BE436A0067CFF9A54F3CB53FEC02EEB" blockId="1.[151,364,1661,1687]" box="[151,364,1661,1687]" pageId="1" pageNumber="378">
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<heading id="D0AC81CC067CFF9A54F3CB53FEC02EEB" bold="true" box="[151,364,1661,1687]" fontSize="11" level="1" pageId="1" pageNumber="378" reason="1">
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<taxonomicName id="4C5B4D23067CFF9A54F3CB53FEC02EEB" ID-CoL="8HN8M" authority="Pascoe" authorityName="Pascoe" box="[151,364,1661,1687]" class="Insecta" family="Curculionidae" genus="Nyphaeba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067CFF9A54F3CB53FEC02EEB" bold="true" box="[151,364,1661,1687]" pageId="1" pageNumber="378">
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<emphasis id="B92FEAB2067CFF9A54F3CB53FEBD2EEB" bold="true" box="[151,273,1661,1687]" italics="true" pageId="1" pageNumber="378">Nyphaeba</emphasis>
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Pascoe
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C341652B067CFF9A54F3CBEAFE632EA6" box="[151,463,1732,1754]" pageId="1" pageNumber="378" type="reference_group">
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<paragraph id="8BE436A0067CFF9A54F3CBEAFE632EA6" blockId="1.[151,1103,1732,1784]" box="[151,463,1732,1754]" pageId="1" pageNumber="378">
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<treatmentCitationGroup id="AB4B118E067CFF9A54F3CBEAFE632EA6" box="[151,463,1732,1754]" pageId="1" pageNumber="378">
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<taxonomicName id="4C5B4D23067CFF9A54F3CBEAFE672EA6" ID-CoL="8HN8M" authority="Pascoe, 1871a: 203" authorityName="Pascoe" authorityPageNumber="203" authorityYear="1871" box="[151,459,1732,1754]" class="Insecta" family="Curculionidae" genus="Nyphaeba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067CFF9A54F3CBEAFF522EA6" box="[151,254,1732,1754]" italics="true" pageId="1" pageNumber="378">Nyphaeba</emphasis>
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<treatmentCitation id="0AFA10B1067CFF9A5560CBEBFE672EA6" author="Pascoe" box="[260,459,1732,1754]" page="203" pageId="1" pageNumber="378" year="1871">
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<bibRefCitation id="EFCA4B51067CFF9A5560CBEBFE342EA6" author="Pascoe" box="[260,408,1732,1754]" pageId="1" pageNumber="378" refString="Pascoe, F. P. (1871 a) Catalogue of Zygopinae, a subfamily of Curculionidae, found by Mr Wallace in the Eastern Archipelago. Annals and Magazine of Natural History, Series 4, 7 (39), 198 - 222." type="journal article" year="1871" yearSuffix="a">Pascoe, 1871a</bibRefCitation>
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: 203
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C341652B067CFF9A54F3CBCDFBE32E84" box="[151,1103,1762,1784]" pageId="1" pageNumber="378" type="materials_examined">
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<paragraph id="8BE436A0067CFF9A54F3CBCDFBE32E84" blockId="1.[151,1103,1732,1784]" box="[151,1103,1762,1784]" pageId="1" pageNumber="378">
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Type species:
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<taxonomicName id="4C5B4D23067CFF9A5542CBCCFDD82E84" authority="Pascoe" authorityName="Pascoe" box="[294,628,1762,1784]" class="Insecta" family="Curculionidae" genus="Nyphaeba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="monommoides">
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<emphasis id="B92FEAB2067CFF9A5542CBCCFD892E84" box="[294,549,1762,1784]" italics="true" pageId="1" pageNumber="378">Nyphaeba monommoides</emphasis>
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Pascoe
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</taxonomicName>
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, by subsequent monotypy:
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<bibRefCitation id="EFCA4B51067CFF9A57ECCBCDFBB42E84" author="Pascoe" box="[904,1048,1762,1784]" pageId="1" pageNumber="378" refString="Pascoe, F. P. (1871 b) Catalogue of Zygopinae, a subfamily of Curculionidae, found by Mr Wallace in the Eastern Archipelago. Annals and Magazine of Natural History, Series 4, 7 (40), 258 - 266, pls. XV - XVI." type="journal article" year="1871" yearSuffix="b">Pascoe 1871b</bibRefCitation>
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: 265.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C341652B067CFF9954F3CA0BFAAB2D71" lastPageId="2" lastPageNumber="379" pageId="1" pageNumber="378" type="description">
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<paragraph id="8BE436A0067CFF9954F3CA0BFE6C2CDC" blockId="1.[151,1436,1829,2034]" lastBlockId="2.[151,1437,151,1293]" lastPageId="2" lastPageNumber="379" pageId="1" pageNumber="378">
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<emphasis id="B92FEAB2067CFF9A54F3CA0BFE912F42" bold="true" box="[151,317,1829,1854]" pageId="1" pageNumber="378">Redescription</emphasis>
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. Length
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<quantity id="4CA39B45067CFF9A55CCCA0BFDFD2F41" box="[424,593,1829,1854]" metricMagnitude="-3" metricUnit="m" metricValue="4.49" metricValueMax="5.3" metricValueMin="3.68" pageId="1" pageNumber="378" unit="mm" value="4.49" valueMax="5.3" valueMin="3.68">3.68–5.30 mm</quantity>
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. Habitus subovate, sides subparallel; in profile convex, with constriction between pronotum and elytron. Integument largely subglabrous, with small patches of scales. Rostrum ca. 0.75 X length of pronotum; parallel-sided or weakly converging to apex; in profile weakly curved; dorsoventrally compressed; in apical half slightly more slender in female; in both sexes dorsal surface without carinae, subglabrous and sparsely punctate in apical half, coarsely punctate in basal half. Antennae inserted slightly in front of middle. Scrobe parallel to rostral contour, meeting anterior margin of eye. Scape subequal to funicle, subglabrous; funicle robust, subglabrous, with sparse thin setae. Club subovate. Eyes large, with subtriangular outline, in dorsolateral position, medially approximate, separated by 0.4 X basal width of rostrum. Forehead coarsely punctate. Pronotum 1.5-1.6 X wider than long; widest at base, sides subparallel in basal half, markedly converging in apical half; with shallow subapical constriction; dorsally densely punctate, laterally coarsely punctate or foveate. Postocular lobe subangular. Scutellar shield distinct, subglabrous. Elytron with 10 distinct striae; stria 10 abbreviated at level of metacoxa, separated by lateral carina originating from elytral base below humerus; sutural interval behind scutellum with swelling and /or distinct sculpture; base sinuate or almost straight; subapically on ventral side without stridulatory file. Wings fully developed. Thoracic venter (
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<figureCitation id="13602A25067FFF995156CCBAFA2329D1" box="[1330,1423,404,429]" captionStart="FIGURE 5" captionStartId="9.[151,250,1927,1949]" captionTargetBox="[195,1349,303,1889]" captionTargetId="figure@9.[195,1392,303,1902]" captionTargetPageId="9" captionText="FIGURE 5 a, c – d. Nyphaeba mimica sp. n., holotype; (a) venter (c) detail of ventrite 1 with row of short ridges marked by arrows (d) posterior surface of metafemur with patch of ridges, presumably acting as stridulatory device together with structures shown in (c). Fig. 5 b. Pantiala illusa Pascoe, venter." httpUri="https://zenodo.org/record/428754/files/figure.png" pageId="2" pageNumber="379">Fig. 5a</figureCitation>
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). Mesothoracic receptacle broadly U-shaped, wider than long, cavernous, internally glabrous, with ventral rim swollen, punctate-rugose, sparsely squamose. Metaventrite with coarse punctures; anteromedially and posteromedially with deep fovea, anterior fovea transverse, posterior fovea elongate. Metanepisternum distinct, subglabrous except one longitudinal row of coarse punctures; anapleural suture complete, without sclerolepidia. Legs. Femora ventrally sulcate; profemur in basal 1/3 with distinct tooth (with indistinct angulation in
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<taxonomicName id="4C5B4D23067FFF995144CF67FF5F2AF8" authority="Faust" authorityName="Faust" class="Insecta" family="Curculionidae" genus="Pantiala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="germari">
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<emphasis id="B92FEAB2067FFF995144CF67FA372A1C" box="[1312,1435,585,608]" italics="true" pageId="2" pageNumber="379">N. germari</emphasis>
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(Faust))
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</taxonomicName>
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; meso- and metafemur edentate. Metafemur with posterior face flattened, at middle with patch of fine parallel ridges (
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<figureCitation id="13602A25067FFF99552DCFA1FE212AD4" box="[329,397,655,680]" captionStart="FIGURE 5" captionStartId="9.[151,250,1927,1949]" captionTargetBox="[195,1349,303,1889]" captionTargetId="figure@9.[195,1392,303,1902]" captionTargetPageId="9" captionText="FIGURE 5 a, c – d. Nyphaeba mimica sp. n., holotype; (a) venter (c) detail of ventrite 1 with row of short ridges marked by arrows (d) posterior surface of metafemur with patch of ridges, presumably acting as stridulatory device together with structures shown in (c). Fig. 5 b. Pantiala illusa Pascoe, venter." httpUri="https://zenodo.org/record/428754/files/figure.png" pageId="2" pageNumber="379">Fig. 5</figureCitation>
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d). Tibiae with dorsal contour convex, with fine carina; apically with strong uncus, without premucro; mesotibia subbasally with dorsal angulation. Tarsi elongate, slender, third tarsomere 1.5 X as wide as second tarsomere. Abdominal venter with ventrites 1-2 convex, punctate. Ventrite 1 with anterior margin bordered by dense row of coarse punctures; with row of short ridges curving along posterolateral angle (
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<figureCitation id="13602A25067FFF9950D0CFD2FAA92B69" box="[1204,1285,764,789]" captionStart="FIGURE 5" captionStartId="9.[151,250,1927,1949]" captionTargetBox="[195,1349,303,1889]" captionTargetId="figure@9.[195,1392,303,1902]" captionTargetPageId="9" captionText="FIGURE 5 a, c – d. Nyphaeba mimica sp. n., holotype; (a) venter (c) detail of ventrite 1 with row of short ridges marked by arrows (d) posterior surface of metafemur with patch of ridges, presumably acting as stridulatory device together with structures shown in (c). Fig. 5 b. Pantiala illusa Pascoe, venter." httpUri="https://zenodo.org/record/428754/files/figure.png" pageId="2" pageNumber="379">Fig. 5c</figureCitation>
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), presumably acting together with metafemur as stridulatory device; intercoxal process subangulate, wider than long. Ventrites 3- 5 flat. Ventrite 5 short, ca.
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<date id="FFE51060067FFF9955A5CE6AFE462B20" box="[449,490,836,860]" pageId="2" pageNumber="379">3 X</date>
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as wide as long.
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<emphasis id="B92FEAB2067FFF9956C8CE6AFCD72B21" box="[684,891,836,861]" italics="true" pageId="2" pageNumber="379">Female terminalia</emphasis>
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. Tergite VII wider than long; microsetae absent; basal margin straight; apical margin convex. Tergite VIII longer than wide; apical margin subtruncate, crenulate. Sternite VIII elongate, narrowly Y-shaped, with apical arms separate and parallel for half its length; membrane ventrally free to apex of apodeme, forming a large elongate pocket. Bursa without sclerites. Spermathecal duct arising at junction of oviduct and bursa.
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<emphasis id="B92FEAB2067FFF99563BCEFAFCB92B91" box="[607,789,980,1005]" italics="true" pageId="2" pageNumber="379">Male terminalia</emphasis>
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. Tergite VII wider than long; without microsetae. Tergite VIII apically subtruncate; dorsally with plumose scales. Sternum VIII with hemisternites small, nude. Spiculum gastrale slightly shorter than penis, straight. Tegmen with parameres and apodeme slightly shorter than width of tegminal ring. Body of penis shorter than apodemes, flattened; apex subtruncate or rounded, without setae; laterally well-sclerotized, medially weakly sclerotized, containing large flat endophallic sclerite shaped like an arrow-head; transfer apparatus distinct.
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</paragraph>
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<paragraph id="8BE436A0067FFF9954A3C982FAAB2D71" blockId="2.[151,1437,151,1293]" pageId="2" pageNumber="379">
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<emphasis id="B92FEAB2067FFF9954A3C982FEAA2CB9" bold="true" box="[199,262,1196,1221]" pageId="2" pageNumber="379">Note.</emphasis>
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<taxonomicName id="4C5B4D23067FFF995575C982FE2F2CB9" box="[273,387,1196,1221]" class="Insecta" family="Curculionidae" genus="Nyphaeba" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067FFF995575C982FE2F2CB9" box="[273,387,1196,1221]" italics="true" pageId="2" pageNumber="379">Nyphaeba</emphasis>
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</taxonomicName>
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has well-developed wings and metanepisterna and therefore it needs to be removed from
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<taxonomicName id="4C5B4D23067FFF9954F3C9E1FED12C94" authority="Lacordaire" authorityName="Lacordaire" box="[151,381,1231,1256]" class="Gastropoda" family="Tylodinidae" genus="Tylodina" kingdom="Animalia" order="Umbraculida" pageId="2" pageNumber="379" phylum="Mollusca" rank="genus">Tylodina Lacordaire</taxonomicName>
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of
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<taxonomicName id="4C5B4D23067FFF9955CDC9E1FC882C94" authority="Schoenherr. In" authorityName="Schoenherr. In" box="[425,804,1231,1256]" class="Insecta" family="Curculionidae" genus="Cryptorhynchinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="genus">Cryptorhynchinae Schoenherr. In</taxonomicName>
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the phylogeny of
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<bibRefCitation id="EFCA4B51067FFF99579AC9E1FB4D2C94" box="[1022,1249,1231,1256]" pageId="2" pageNumber="379" refString="Riedel, A., Tanzler, R., Pons, J., Suhardjono, Y. R. & Balke, M. (2016) Large-scale molecular phylogeny of Cryptorhynchinae (Coleoptera, Curculionidae) from multiple genes suggests American origin and later Australian radiation. Systematic Entomology, 41 (2), 492 - 503." type="journal article">
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Riedel
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<emphasis id="B92FEAB2067FFF995035C9FFFB202C94" box="[1105,1164,1231,1256]" italics="true" pageId="2" pageNumber="379">et al.</emphasis>
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(2016)
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</bibRefCitation>
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it is part of the “
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<taxonomicName id="4C5B4D23067FFF9954C7C9DAFE862D71" box="[163,298,1268,1293]" class="Insecta" family="Curculionidae" genus="Rhynchodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067FFF9954C7C9DAFE862D71" box="[163,298,1268,1293]" italics="true" pageId="2" pageNumber="379">Rhynchodes</emphasis>
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</taxonomicName>
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-group” which is supported by the structure of the female sternite VIII (new placement).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |