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<mods:title id="3C9992405F0D0F5E5FCFBC5358CFE638">Geometridae (Lepidoptera) of the Baikal region: identification keys and annotated catalogue with notes to DNA barcoding. Part 2. Archiearinae, Geometrinae, Sterrhinae</mods:title>
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6. 
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<emphasis id="B92FEA9D690CFFB8FF1641E1FE1E08F2" bold="true" box="[181,480,1227,1254]" italics="true" pageId="29" pageNumber="30">Hemistola zimmermanni</emphasis>
(Hedemann, 1879)
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<paragraph id="8BE4368F690CFFB8FF344020FAE00936" blockId="29.[151,1437,1290,1498]" box="[151,1310,1290,1314]" pageId="29" pageNumber="30">
(adults 
<figureCitation id="13602A0A690CFFB8FF414020FE7E0936" box="[226,384,1290,1314]" captionStart="FIGURE 14" captionStartId="37.[151,250,1206,1230]" captionTargetBox="[152,1434,182,1181]" captionTargetId="figure-15@37.[151,1435,181,1182]" captionTargetPageId="37" captionText="FIGURE 14. Lectotypes of (A) Hemistola zimmermanni Hedemann and (B) H. intermedia Djakonov." figureDoi="http://doi.org/10.5281/zenodo.7965108" httpUri="https://zenodo.org/record/7965108/files/figure.png" pageId="29" pageNumber="30">Figs. 14A, 14B</figureCitation>
, 
<figureCitation id="13602A0A690CFFB8FE294020FE460936" box="[394,440,1290,1314]" captionStart="FIGURE 16" captionStartId="61.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@61.[151,1435,181,1836]" captionTargetPageId="61" captionText="FIGURE 16. Adults of Siberian Geometrinae. A—Thetidia chlorophyllaria; B—T. smaragdaria; C—Comibaena amoenaria; D—Hemistola intermedia Lectotype; E—Geometra zimmermanni Lectotype." figureDoi="http://doi.org/10.5281/zenodo.7965116" httpUri="https://zenodo.org/record/7965116/files/figure.png" pageId="29" pageNumber="30">16D</figureCitation>
, 
<figureCitation id="13602A0A690CFFB8FE614020FE0E0936" box="[450,496,1290,1314]" captionStart="FIGURE 17" captionStartId="62.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@62.[151,1435,181,1836]" captionTargetPageId="62" captionText="FIGURE 17. Adults of Siberian Geometrinae. A—Hemistola zimmermanni; B—H. chrysoprasaria; C—Thalera fimbrialis; D—T. chlorosaria." figureDoi="http://doi.org/10.5281/zenodo.7965120" httpUri="https://zenodo.org/record/7965120/files/figure.png" pageId="29" pageNumber="30">17A</figureCitation>
, 
<figureCitation id="13602A0A690CFFB8FE584020FD9E0936" box="[507,608,1290,1314]" captionStart="FIGURE 57" captionStartId="102.[151,250,1867,1891]" captionTargetBox="[151,1436,181,1844]" captionTargetId="figure-25@102.[151,1436,181,1844]" captionTargetPageId="102" captionText="FIGURE 57. Adults of some Siberian Geometrinae in live. A—Thetidia chlorophyllaria (male); B–D—H. zimmermanni (B— male, C, D—females); E—Chlorissa viridata (male); F—Thalera chlorosaria (male). Photos by Marya Dementyeva (Irkutsk)." figureDoi="http://doi.org/10.5281/zenodo.7965236" httpUri="https://zenodo.org/record/7965236/files/figure.png" pageId="29" pageNumber="30">57B–57D</figureCitation>
; male gen. 
<figureCitation id="13602A0A690CFFB8FD704020FCBE0936" box="[723,832,1290,1314]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="31.[151,250,719,743]" captionStartId-1="32.[151,250,1866,1890]" captionStartId-2="33.[151,250,1867,1891]" captionTargetBox-0="[209,1378,187,687]" captionTargetBox-1="[162,1425,189,1838]" captionTargetBox-2="[162,1423,191,1844]" captionTargetId-0="figure-15@31.[200,1387,181,695]" captionTargetId-1="figure-25@32.[151,1435,181,1842]" captionTargetId-2="figure-15@33.[153,1434,181,1844]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionText-0="FIGURE 8. Aedeagi of Siberian and Far Eastern Hemistola. A–D—aedeagi of “Hemistola intermedia” from Krasnoyarsky Kray and Khakassia; E–H—aedeagi of H. zimmermanni from Zabaikalsky Kray, Khabarovsky Kray and Primorsky Kray." captionText-1="FIGURE 9. Genital segments and sternites A8 of Siberian “Hemistola intermedia” (Krasnoyarsky Kray and Khakassia). A, C, E, G—genital segments; B, D, F—sternites A8." captionText-2="FIGURE 10. Genital segments and sternites A8 of Far Eastern Hemistola zimmermanni (Zabaikalsky Kray, Khabarovsky Kray and Primorsky Kray). A, C, E, G—genital segments; B, D, F, H—sternites A8." figureDoi-0="http://doi.org/10.5281/zenodo.7965091" figureDoi-1="http://doi.org/10.5281/zenodo.7965093" figureDoi-2="http://doi.org/10.5281/zenodo.7965096" httpUri-0="https://zenodo.org/record/7965091/files/figure.png" httpUri-1="https://zenodo.org/record/7965093/files/figure.png" httpUri-2="https://zenodo.org/record/7965096/files/figure.png" pageId="29" pageNumber="30">Figs. 8–10</figureCitation>
, 
<figureCitation id="13602A0A690CFFB8FCE84020FC500936" box="[843,942,1290,1314]" captionStart="FIGURE 34" captionStartId="79.[151,250,1866,1890]" captionTargetBox="[163,1428,200,1836]" captionTargetId="figure-15@79.[155,1432,181,1842]" captionTargetPageId="79" captionText="FIGURE 34. Male genitalia of Siberian Geometrinae. A–C—Hemistola chrysoprasaria; D–F—H. zimmermanni; G, H— Hemithea aestivaria; I–K—Thalera fimbrialis; L–N—T. chlorosaria. A, E, G, I, M—genital segment; B, F, H, J, N—phallos; B, F—vesica everted; K—valva, dorsal view; L—dorso-lateral view of genital segment. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965180" httpUri="https://zenodo.org/record/7965180/files/figure.png" pageId="29" pageNumber="30">34D–34F</figureCitation>
; female gen. 
<figureCitation id="13602A0A690CFFB8FB964020FB790936" box="[1077,1159,1290,1314]" captionStart="FIGURE 11" captionStartId="34.[151,250,1867,1891]" captionTargetBox="[162,1427,186,1838]" captionTargetId="figure-25@34.[154,1433,181,1844]" captionTargetPageId="34" captionText="FIGURE 11. Female genitalia of Siberian Hemistola zimmermanni. A–D—samples from Irkutskaya Oblast; E–H—samples from Buryatia." figureDoi="http://doi.org/10.5281/zenodo.7965100" httpUri="https://zenodo.org/record/7965100/files/figure.png" pageId="29" pageNumber="30">Figs. 11</figureCitation>
, 
<figureCitation id="13602A0A690CFFB8FB314020FB530936" box="[1170,1197,1290,1314]" captionStart="FIGURE 12" captionStartId="35.[151,250,1071,1095]" captionTargetBox="[165,1420,187,1041]" captionTargetId="figure-15@35.[158,1429,181,1048]" captionTargetPageId="35" captionText="FIGURE 12. Female genitalia of Siberian and Far Eastern Hemistola zimmermanni. A, B—samples from Buryatia; C—sample from Zabaikalsky Kray; D—sample from Primorsky Kray." figureDoi="http://doi.org/10.5281/zenodo.7965102" httpUri="https://zenodo.org/record/7965102/files/figure.png" pageId="29" pageNumber="30">12</figureCitation>
, 
<figureCitation id="13602A0A690CFFB8FB1B4020FAE90936" box="[1208,1303,1290,1314]" captionStart="FIGURE 37" captionStartId="82.[151,250,1862,1886]" captionTargetBox="[153,1423,192,1832]" captionTargetId="figure-25@82.[152,1435,181,1838]" captionTargetPageId="82" captionText="FIGURE 37. Female genitalia of Siberian Geometrinae. A—Geometra papilionaria; B—Thetidia chlorophyllaria; C—T. smaragdaria; D—Comibaena amoenaria; E—Hemistola chrysoprasaria; F, G—H. zimmermanni; H—Hemithea aestivaria. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965188" httpUri="https://zenodo.org/record/7965188/files/figure.png" pageId="29" pageNumber="30">37F, 37G</figureCitation>
)
</paragraph>
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<treatmentCitation id="0AFA109E690CFFB8FF344002FD8E0954" author="Viidalepp, J." box="[151,624,1320,1344]" page="846" pageId="29" pageNumber="30" year="1976">
<taxonomicName id="4C5B4D0C690CFFB8FF344002FD8E0954" ID-CoL="6M329" authority=": Viidalepp, 1976: 846" authorityName="Viidalepp" authorityPageNumber="846" authorityYear="1976" box="[151,624,1320,1344]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D690CFFB8FF344002FE710954" box="[151,399,1320,1344]" italics="true" pageId="29" pageNumber="30">Hemistola zimmermanni</emphasis>
: 
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; 
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<bibRefCitation id="EFCA4B7E690CFFB8FDDA4002FD2A0954" author="Viidalepp, J." box="[633,724,1320,1344]" pageId="29" pageNumber="30" refId="ref74481" refString="Viidalepp, J. (1996) Checklist of the Geometridae (Lepidoptera) of the former U. S. S. R. Apollo Books, Stenstrup, 111 pp." type="book" year="1996">1996: 61</bibRefCitation>
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; 
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; 
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; 
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Mironov 
<emphasis id="B92FEA9D690CFFB8FB504003FAE10954" box="[1267,1311,1320,1344]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
., 2008: 205
</bibRefCitation>
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; 
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. (Locus typicus: “Chingan” [
<collectingCountry id="F34C761F690CFFB8FC87406DFC93094B" box="[804,877,1351,1375]" name="Russia" pageId="29" pageNumber="30">Russia</collectingCountry>
, Far East, Khingan Mountains]. 
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<typeStatus id="54E0882D690CFFB8FB17406DFAE1094B" box="[1204,1311,1351,1375]" pageId="29" pageNumber="30" type="lectotype">Lectotype</typeStatus>
</emphasis>
♁, herewith designated to stabilize nomenclature (ZIN)).
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<taxonomicName id="4C5B4D0C690CFFB8FF3440AEFDE50988" ID-CoL="3KSZB" authority="Djakonov, 1926" authorityName="Djakonov" authorityYear="1926" box="[151,539,1412,1436]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D690CFFB8FF3440AEFE8C0988" box="[151,370,1412,1436]" italics="true" pageId="29" pageNumber="30">Hemistola intermedia</emphasis>
<bibRefCitation id="EFCA4B7E690CFFB8FED440AEFDE50988" author="Djakonov, A. M." box="[375,539,1412,1436]" pageId="29" pageNumber="30" pagination="1 - 78" refId="ref70583" refString="Djakonov, A. M. (1926) Zur Kenntnis der Geometriden fauna des Minussinsk Bezirks (Sibirien, Jenissej Gouv.). Jahrbuch des Martjanov'schen Staatsmuseums in Minussinsk (Sibirien), 4 (1), 1 - 78. [in Russian and German]" type="journal article" year="1926">Djakonov, 1926</bibRefCitation>
</taxonomicName>
: 
<treatmentCitation id="0AFA109E690CFFB8FD8040AEFCA70988" author="Beljaev, E. A. &amp; Mironov, V. G." box="[547,857,1412,1437]" page="254" pageId="29" pageNumber="30" year="2019">
<bibRefCitation id="EFCA4B7E690CFFB8FD8040AEFCA70988" author="Beljaev, E. A. &amp; Mironov, V. G." box="[547,857,1412,1437]" pageId="29" pageNumber="30" pagination="235 - 281" refId="ref68795" refString="Beljaev, E. A. &amp; Mironov, V. G. (2019) Geometridae. In: Sinev, S. Yu. (Ed.), Catalogue of the Lepidoptera of Russia. 2 nd Edition. Zoological Institute RAS, St. Petersburg, pp. 235 - 281. [in Russian]" type="book chapter" year="2019">Beljaev &amp; Mironov, 2019: 254</bibRefCitation>
</treatmentCitation>
. (Locus typicus: Minusinsk [
<collectingCountry id="F34C761F690CFFB8FBDB40AEFB3F0988" box="[1144,1217,1412,1436]" name="Russia" pageId="29" pageNumber="30">Russia</collectingCountry>
, Krasnoyarsky Kray]. 
<emphasis id="B92FEA9D690CFFB8FF644089FECC09AF" bold="true" box="[199,306,1443,1467]" pageId="29" pageNumber="30">
<typeStatus id="54E0882D690CFFB8FF644089FECC09AF" box="[199,306,1443,1467]" pageId="29" pageNumber="30" type="lectotype">Lectotype</typeStatus>
</emphasis>
♁, herewith designated to stabilize nomenclature (ZIN)).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE4368F690CFFB8FF3440E8FCFA09CE" blockId="29.[151,1437,1290,1498]" box="[151,772,1474,1498]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="AB4B11A1690CFFB8FF3440E8FCFA09CE" box="[151,772,1474,1498]" pageId="29" pageNumber="30">
<emphasis id="B92FEA9D690CFFB8FF3440E8FF5A09CD" box="[151,164,1474,1497]" italics="true" pageId="29" pageNumber="30">?</emphasis>
<treatmentCitation id="0AFA109E690CFFB8FF0940E8FCFE09CE" author="Viidalepp, J." box="[170,768,1474,1498]" page="61" pageId="29" pageNumber="30" year="1996">
<taxonomicName id="4C5B4D0C690CFFB8FF0940E8FCFE09CE" ID-CoL="3KSZB" authority=": Viidalepp, 1996: 61" authorityName="Viidalepp" authorityPageNumber="61" authorityYear="1996" box="[170,768,1474,1498]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="chrysoprasaria" subSpecies="intermedia">
<emphasis id="B92FEA9D690CFFB8FF0940E8FDD609CE" box="[170,552,1474,1498]" italics="true" pageId="29" pageNumber="30">Hemistola chrysoprasaria intermedia</emphasis>
: 
<bibRefCitation id="EFCA4B7E690CFFB8FD9640E8FCFE09CE" author="Viidalepp, J." box="[565,768,1474,1498]" pageId="29" pageNumber="30" refId="ref74481" refString="Viidalepp, J. (1996) Checklist of the Geometridae (Lepidoptera) of the former U. S. S. R. Apollo Books, Stenstrup, 111 pp." type="book" year="1996">Viidalepp, 1996: 61</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3416504690CFFBAFF34432BFAEB0B4F" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BE4368F690CFFB8FF34432BFBF80ADB" blockId="29.[151,1437,1537,1926]" pageId="29" pageNumber="30">
<emphasis id="B92FEA9D690CFFB8FF34432BFDB80A0F" bold="true" box="[151,582,1537,1563]" pageId="29" pageNumber="30">
Type material examined. 
<typeStatus id="54E0882D690CFFB8FE73432BFDB80A0F" box="[464,582,1537,1563]" pageId="29" pageNumber="30" type="lectotype">Lectotype</typeStatus>
</emphasis>
♁ of 
<taxonomicName id="4C5B4D0C690CFFB8FD324328FB8F0A0F" authority="Hedemann, 1879" authorityName="Hedemann" authorityYear="1879" box="[657,1137,1537,1564]" class="Insecta" family="Geometridae" genus="Geometra" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D690CFFB8FD324328FC5A0A0F" box="[657,932,1538,1564]" italics="true" pageId="29" pageNumber="30">Geometra zimmermanni</emphasis>
Hedemann, 1879
</taxonomicName>
(hereby designated) (
<figureCitation id="13602A0A690CFFB8FACC432BFF330A2B" captionStart="FIGURE 14" captionStartId="37.[151,250,1206,1230]" captionTargetBox="[152,1434,182,1181]" captionTargetId="figure-15@37.[151,1435,181,1182]" captionTargetPageId="37" captionText="FIGURE 14. Lectotypes of (A) Hemistola zimmermanni Hedemann and (B) H. intermedia Djakonov." figureDoi="http://doi.org/10.5281/zenodo.7965108" httpUri="https://zenodo.org/record/7965108/files/figure.png" pageId="29" pageNumber="30">Fig. 14A</figureCitation>
): [
<emphasis id="B92FEA9D690CFFB8FF48430FFEBD0A2B" bold="true" box="[235,323,1573,1599]" pageId="29" pageNumber="30">
<collectingCountry id="F34C761F690CFFB8FF48430FFEC10A2B" box="[235,319,1573,1599]" name="Russia" pageId="29" pageNumber="30">Russia</collectingCountry>
:
</emphasis>
] &lt;small brownish quadrat with handwritten text&gt; ‘Chingan | Geb. | 28/7. [18]77’, &lt;small whitish quadrat with handwritten text&gt; ’61.’, &lt;yellowish rectangular label with printed text&gt; ‘Колл. Вел. КнЯЗЯ | НиколаЯ | Михайловича.[collection of Grand Duke Nikolai Mikhailovich]’, &lt;large rectangular label with red handwritten text&gt; ‘ 
<taxonomicName id="4C5B4D0C690CFFB8FF3843BBFEF10ABF" authorityName="Linnaeus" authorityYear="1758" box="[155,271,1681,1707]" class="Insecta" family="Geometridae" genus="Geometra" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Geometra</taxonomicName>
| 
<taxonomicName id="4C5B4D0C690CFFB8FE8743BBFDDB0ABF" authority="Hedem." authorityName="Hedemann" authorityYear="1879" box="[292,549,1681,1707]" class="Insecta" family="Geometridae" genus="Geometra" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="zimmermanni">Zimmermanni Hedem.</taxonomicName>
| Type.’, &lt;red rectangle with handwritten text&gt; ‘&lt;underlined&gt; 
<typeStatus id="54E0882D690CFFB8FB4943BBFA620ABF" box="[1258,1436,1681,1707]" pageId="29" pageNumber="30" type="lectotype">LECTOTYPUS</typeStatus>
♁ | 
<taxonomicName id="4C5B4D0C690CFFB8FF63439FFED00ADB" authorityName="Linnaeus" authorityYear="1758" box="[192,302,1717,1743]" class="Insecta" family="Geometridae" genus="Geometra" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Geometra</taxonomicName>
| zimmermanni | Hedemann, 1879 | Design. I. A. Makhov 2023’.
</paragraph>
<paragraph id="8BE4368F690CFFB8FF6443F3FB5E0B97" blockId="29.[151,1437,1537,1926]" pageId="29" pageNumber="30">
<emphasis id="B92FEA9D690CFFB8FF6443F3FEC30AE7" bold="true" box="[199,317,1753,1779]" pageId="29" pageNumber="30">
<typeStatus id="54E0882D690CFFB8FF6443F3FEC30AE7" box="[199,317,1753,1779]" pageId="29" pageNumber="30" type="lectotype">Lectotype</typeStatus>
</emphasis>
♁ of 
<taxonomicName id="4C5B4D0C690CFFB8FED843F3FCDA0AE7" authority="Djakonov, 1926" authorityName="Djakonov" authorityYear="1926" box="[379,804,1753,1779]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D690CFFB8FED843F3FD930AE7" box="[379,621,1753,1779]" italics="true" pageId="29" pageNumber="30">Hemistola intermedia</emphasis>
<bibRefCitation id="EFCA4B7E690CFFB8FDD043F3FCDA0AE7" author="Djakonov, A. M." box="[627,804,1753,1779]" pageId="29" pageNumber="30" pagination="1 - 78" refId="ref70583" refString="Djakonov, A. M. (1926) Zur Kenntnis der Geometriden fauna des Minussinsk Bezirks (Sibirien, Jenissej Gouv.). Jahrbuch des Martjanov'schen Staatsmuseums in Minussinsk (Sibirien), 4 (1), 1 - 78. [in Russian and German]" type="journal article" year="1926">Djakonov, 1926</bibRefCitation>
</taxonomicName>
(hereby designated) (
<figureCitation id="13602A0A690CFFB8FBB643F3FB840AE7" box="[1045,1146,1753,1779]" captionStart="FIGURE 14" captionStartId="37.[151,250,1206,1230]" captionTargetBox="[152,1434,182,1181]" captionTargetId="figure-15@37.[151,1435,181,1182]" captionTargetPageId="37" captionText="FIGURE 14. Lectotypes of (A) Hemistola zimmermanni Hedemann and (B) H. intermedia Djakonov." figureDoi="http://doi.org/10.5281/zenodo.7965108" httpUri="https://zenodo.org/record/7965108/files/figure.png" pageId="29" pageNumber="30">Fig. 14B</figureCitation>
): [
<emphasis id="B92FEA9D690CFFB8FB3443F3FF210B03" bold="true" pageId="29" pageNumber="30">
<collectingCountry id="F34C761F690CFFB8FB3443F3FB160AE7" box="[1175,1256,1753,1779]" name="Russia" pageId="29" pageNumber="30">Russia</collectingCountry>
, Krasnoyarsky Kray:
</emphasis>
] &lt;small rectangular label with printed text&gt; ‘Минусинск [Minusinsk] | &lt;handwritten&gt; 6.VII.[19]24 | &lt;printed text&gt; Филипьев [Filipjev]’ [on the back of the label:] &lt;handwritten text&gt; ‘ГрЯДы’ [Gryady]; &lt;handwritten text&gt; ‘ 
<taxonomicName id="4C5B4D0C690CFFB8FF02426FFE2B0B4B" authority="Djak." authorityName="Djak." box="[161,469,1861,1887]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D690CFFB8FF02426FFE6C0B4B" box="[161,402,1861,1887]" italics="true" pageId="29" pageNumber="30">Hemistola intermedia</emphasis>
Djak.
</taxonomicName>
| &lt;printed text&gt;A. Djakonov det.’, &lt;red rectangle with handwritten text&gt; ‘&lt;underlined&gt; 
<typeStatus id="54E0882D690CFFB8FF344243FEB70B97" box="[151,329,1897,1923]" pageId="29" pageNumber="30" type="lectotype">LECTOTYPUS</typeStatus>
♁ | 
<taxonomicName id="4C5B4D0C690CFFB8FEDA4243FE120B97" authorityName="Warren" authorityYear="1893" box="[377,492,1897,1923]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Hemistola</taxonomicName>
| intermedia | 
<bibRefCitation id="EFCA4B7E690CFFB8FD284243FCC00B97" author="Djakonov, A. M." box="[651,830,1897,1923]" pageId="29" pageNumber="30" pagination="1 - 78" refId="ref70583" refString="Djakonov, A. M. (1926) Zur Kenntnis der Geometriden fauna des Minussinsk Bezirks (Sibirien, Jenissej Gouv.). Jahrbuch des Martjanov'schen Staatsmuseums in Minussinsk (Sibirien), 4 (1), 1 - 78. [in Russian and German]" type="journal article" year="1926">Djakonov, 1926</bibRefCitation>
| Design. I. A. Makhov | 2023’.
</paragraph>
<caption id="DF246607690FFFBBFF34426AFD700B68" ID-DOI="http://doi.org/10.5281/zenodo.7965087" ID-Zenodo-Dep="7965087" httpUri="https://zenodo.org/record/7965087/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1856,1880]" targetBox="[151,1436,187,1826]" targetPageId="30" targetType="figure">
<paragraph id="8BE4368F690FFFBBFF34426AFD700B68" blockId="30.[151,1436,1856,1916]" pageId="30" pageNumber="31">
<emphasis id="B92FEA9D690FFFBBFF34426AFEED0B4C" bold="true" box="[151,275,1856,1880]" pageId="30" pageNumber="31">FIGURE 7.</emphasis>
Male antennae of 
<taxonomicName id="4C5B4D0C690FFFBBFE73426AFDC60B4C" authorityName="Warren" authorityYear="1893" box="[464,568,1856,1880]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEA9D690FFFBBFE73426AFDC60B4C" box="[464,568,1856,1880]" italics="true" pageId="30" pageNumber="31">Hemistola</emphasis>
</taxonomicName>
. A, C, E, G, I, K, M, O—antenna; B, D, F, H, J, L, N, P—10 
<superScript id="7C2E9BC7690FFFBBFB00426AFB500B5A" attach="left" box="[1187,1198,1856,1870]" fontSize="6" pageId="30" pageNumber="31">th</superScript>
flagellomere. M, N, O, P—antennae aligned in length. Scale bar—1 mm.
</paragraph>
</caption>
<caption id="DF246607690EFFBAFF3447E5FA960F18" ID-DOI="http://doi.org/10.5281/zenodo.7965091" ID-Zenodo-Dep="7965091" httpUri="https://zenodo.org/record/7965091/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,719,743]" targetBox="[209,1378,187,687]" targetPageId="31" targetType="figure">
<paragraph id="8BE4368F690EFFBAFF3447E5FA960F18" blockId="31.[151,1437,719,780]" pageId="31" pageNumber="32">
<emphasis id="B92FEA9D690EFFBAFF3447E5FEEB0EF3" bold="true" box="[151,277,719,743]" pageId="31" pageNumber="32">FIGURE 8.</emphasis>
Aedeagi of Siberian and Far Eastern 
<taxonomicName id="4C5B4D0C690EFFBAFD0047FAFCF50EFC" authorityName="Warren" authorityYear="1893" box="[675,779,720,744]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEA9D690EFFBAFD0047FAFCF50EFC" box="[675,779,720,744]" italics="true" pageId="31" pageNumber="32">Hemistola</emphasis>
</taxonomicName>
. A–D—aedeagi of “ 
<taxonomicName id="4C5B4D0C690EFFBAFC4047FAFB3C0EFC" box="[995,1218,720,744]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D690EFFBAFC4047FAFB3C0EFC" box="[995,1218,720,744]" italics="true" pageId="31" pageNumber="32">Hemistola intermedia</emphasis>
</taxonomicName>
” from Krasnoyarsky Kray and Khakassia; E–H—aedeagi of 
<taxonomicName id="4C5B4D0C690EFFBAFD8447DEFD2E0F1F" authorityName="Hedemann" authorityYear="1879" box="[551,720,756,779]" class="Insecta" family="Geometridae" genus="Geometra" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D690EFFBAFD8447DEFD2E0F1F" box="[551,720,756,779]" italics="true" pageId="31" pageNumber="32">H. zimmermanni</emphasis>
</taxonomicName>
from Zabaikalsky Kray, Khabarovsky Kray and Primorsky Kray.
</paragraph>
</caption>
<paragraph id="8BE4368F690EFFBAFF644674FAEB0B4F" blockId="31.[151,1437,861,1885]" pageId="31" pageNumber="32">
<emphasis id="B92FEA9D690EFFBAFF644674FD630F6C" bold="true" box="[199,669,861,888]" pageId="31" pageNumber="32">
Further material. 
<emphasis id="B92FEA9D690EFFBAFE0C4677FD6A0F6C" bold="true" box="[431,660,861,888]" italics="true" pageId="31" pageNumber="32">Krasnoyarsky Kray</emphasis>
:
</emphasis>
1 ♁: “Окр. Минусинска 
<date id="FFE5104F690EFFBAFC4A4674FB9A0F6C" box="[1001,1124,862,888]" pageId="31" pageNumber="32" value="1928-07-09">9.VII.1928</date>
Кожанчиков [Vicinity of Minusinsk, 
<date id="FFE5104F690EFFBAFE8346A8FE630F88" box="[288,413,898,924]" pageId="31" pageNumber="32" value="1928-07-09">9.VII.1928</date>
, Kozhantchikov]” [on the back of the label:] “Тагарск. остр. [Tagarsky Island]”; 1 ♁: “МинусинскЕнис. г. 
<date id="FFE5104F690EFFBAFE30468CFDE00FD5" box="[403,542,934,961]" pageId="31" pageNumber="32" value="1925-07-21">21.VII.1925</date>
. Кравченко и Цыганков [Minusinsk, Yeniseskaya Governorate, 
<date id="FFE5104F690EFFBAFAAF468CFA660FD5" box="[1292,1432,934,961]" pageId="31" pageNumber="32" value="1925-07-21">21.VII.1925</date>
, Kravchenko &amp; Tsygankov]” [on the back of the label:] “Тагар. остр. [Tagarsky Island]”; 1 ♁: “МинусинскЕнис. г. 
<date id="FFE5104F690EFFBAFF0D46C7FEC4081C" box="[174,314,1005,1032]" pageId="31" pageNumber="32" value="1925-07-16">16.VII.1925</date>
. Кравченко и Цыганков” [on the back of the label:] “Тагар. остр. [Tagarsky Island]” | “ 
<taxonomicName id="4C5B4D0C690EFFBAFA8A46C7FE92083F" authority="Hed." authorityName="Hedemann" authorityYear="1879" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D690EFFBAFA8A46C7FED1083F" italics="true" pageId="31" pageNumber="32">Hemistola zimmermanni</emphasis>
Hed.
</taxonomicName>
det. W. Koshantschikov”; 1 ♁: “МинусинскЕнис. г. 
<date id="FFE5104F690EFFBAFC63413BFBB20838" box="[960,1100,1041,1068]" pageId="31" pageNumber="32" value="1925-07-16">16.VII.1925</date>
. Кравченко и Цыганков” [on the back of the label:] “Тагар. остр.”; 1 ♁: “Окр. Минусинска 
<date id="FFE5104F690EFFBAFCF0411EFC22085A" box="[851,988,1076,1102]" pageId="31" pageNumber="32" value="1936-07-23">23.VII.1936</date>
Кожанчиков” [on the back of the label:] “ГрЯДы”; 1 ♁: “Окр. Минусинска 
<date id="FFE5104F690EFFBAFD964172FD430866" box="[565,701,1112,1138]" pageId="31" pageNumber="32" value="1928-07-19">19.VII.1928</date>
Кожанчиков” [on the back of the label:] “Таг. остр. [Tagarsky Island]”; 1 ♁: “Окр. Минусинска 
<date id="FFE5104F690EFFBAFD8C4151FD460881" box="[559,696,1147,1173]" pageId="31" pageNumber="32" value="1937-07-12">12.VII.1937</date>
Кожанчиков” [on the back of the label:] “ГрЯДы”; 1 ♁: “Окр. Минусинска 
<date id="FFE5104F690EFFBAFE9141B4FE4208AD" box="[306,444,1182,1209]" pageId="31" pageNumber="32" value="1937-07-15">15.VII.1937</date>
Кожанчиков” [on the back of the label:] “ГрЯДы”; 1 ♁: “Окр. Минусинска 
<date id="FFE5104F690EFFBAFAB041B5FA6208AD" box="[1299,1436,1183,1209]" pageId="31" pageNumber="32" value="1937-07-31">31.VII.1937</date>
Кожанчиков” [on the back of the label:] “ГрЯДы”; 1 ♁: “Окр. Минусинска 
<date id="FFE5104F690EFFBAFC4041E8FB9208C8" box="[995,1132,1218,1244]" pageId="31" pageNumber="32" value="1936-07-23">23.VII.1936</date>
Кожанчиков” [on the back of the label:] “ГрЯДы”; 1 ♁: “Окр. Минусинска 
<date id="FFE5104F690EFFBAFD0C41CCFCC60914" box="[687,824,1254,1280]" pageId="31" pageNumber="32" value="1937-07-12">12.VII.1937</date>
Кожанчиков” [on the back of the label:] “ГрЯДы”; 1 ♁: “МинусинскЕнис. г. 
<date id="FFE5104F690EFFBAFE284023FDE90930" box="[395,535,1289,1316]" pageId="31" pageNumber="32" value="1925-07-14">14.VII.1925</date>
. Кравченко и Цыганков” [on the back of the label:] “Тагар. остр.”; Minusinsky Distr., close to Bystraya vill., Grjady, steppe hills (S-exposition slope: 
<geoCoordinate id="EE6F5048690EFFBAFC1D4006FBB40953" box="[958,1098,1324,1351]" degrees="53" direction="north" minutes="44" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="21" value="53.739166">53°44’21”N</geoCoordinate>
, 
<geoCoordinate id="EE6F5048690EFFBAFBF64007FB200953" box="[1109,1246,1325,1351]" degrees="91" direction="east" minutes="33" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="38" value="91.56056">91°33’38”E</geoCoordinate>
, 
<specimenCount id="9D5DFD06690EFFBAFB484006FAC10952" box="[1259,1343,1324,1351]" pageId="31" pageNumber="32" type="generic">
<quantity id="4CA39B6A690EFFBAFB484006FAC50952" box="[1259,1339,1324,1351]" metricMagnitude="2" metricUnit="m" metricValue="3.5799999999999996" pageId="31" pageNumber="32" unit="m" value="358.0">358 m</quantity>
.
</specimenCount>
a.s.l.), at light, 
<date id="FFE5104F690EFFBAFF79407AFEA2097E" box="[218,348,1360,1386]" pageId="31" pageNumber="32" value="2020-06-19">
19. 
<collectionCode id="ED4AAE4A690EFFBAFF5A407AFEE7097E" box="[249,281,1360,1386]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="31" pageNumber="32">VI</collectionCode>
.2020
</date>
, 1 ♁ (GenBank ID: 
<collectionCode id="ED4AAE4A690EFFBAFDE5407AFD91097E" box="[582,623,1360,1386]" pageId="31" pageNumber="32">OQ</collectionCode>
362117), 
<collectionCode id="ED4AAE4A690EFFBAFD78407AFD0F097E" box="[731,753,1360,1386]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="32">R</collectionCode>
. Maksimov [
<collectionCode id="ED4AAE4A690EFFBAFC27407AFC45097E" box="[900,955,1360,1386]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="31" pageNumber="32" type="Museum">ZIN</collectionCode>
]. 
<emphasis id="B92FEA9D690EFFBAFC6D4065FB2B097E" bold="true" box="[974,1237,1359,1386]" italics="true" pageId="31" pageNumber="32">
Republic of 
<collectingRegion id="499FF86D690EFFBAFBFE4065FB2B097E" box="[1117,1237,1359,1386]" country="Russia" name="Khakasiya" pageId="31" pageNumber="32">Khakassia</collectingRegion>
</emphasis>
: Ust’-Abakansky Distr., Khakassky state National Reserve, Oglahty area, motley grass steppe, rocky hills (
<geoCoordinate id="EE6F5048690EFFBAFBDB4059FAFA099A" box="[1144,1284,1395,1422]" degrees="53" direction="north" minutes="59" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="17" value="53.988056">53°59’17”N</geoCoordinate>
, 
<geoCoordinate id="EE6F5048690EFFBAFAAC4059FA690999" box="[1295,1431,1395,1421]" degrees="91" direction="east" minutes="29" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="46" value="91.49611">91°29’46”E</geoCoordinate>
, 
<specimenCount id="9D5DFD06690EFFBAFF3440BDFF1209A5" box="[151,236,1431,1457]" pageId="31" pageNumber="32" type="generic">
<quantity id="4CA39B6A690EFFBAFF3440BDFF1609A5" box="[151,232,1431,1457]" metricMagnitude="2" metricUnit="m" metricValue="3.09" pageId="31" pageNumber="32" unit="m" value="309.0">309 m</quantity>
.
</specimenCount>
a.s.l.), at light, 
<date id="FFE5104F690EFFBAFE3B40BDFDEB09A5" box="[408,533,1431,1457]" pageId="31" pageNumber="32" value="2018-07-03">3.VII.2018</date>
, 2 ♁ (GenBank ID: 
<collectionCode id="ED4AAE4A690EFFBAFCA440BDFCCE09A6" box="[775,816,1431,1458]" pageId="31" pageNumber="32">OQ</collectionCode>
362115, OQ362116), 
<collectionCode id="ED4AAE4A690EFFBAFB8B40BDFBC009A5" box="[1064,1086,1431,1457]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="32">R</collectionCode>
. Maksimov [
<collectionCode id="ED4AAE4A690EFFBAFB7640BDFAF209A5" box="[1237,1292,1431,1457]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="31" pageNumber="32" type="Museum">ZIN</collectionCode>
]. 
<emphasis id="B92FEA9D690EFFBAFA8340BCFF1C09C1" bold="true" italics="true" pageId="31" pageNumber="32">Irkutskaya Oblast</emphasis>
: Sarma, 
<date id="FFE5104F690EFFBAFEEB4090FE2D09C0" box="[328,467,1466,1492]" pageId="31" pageNumber="32" value="2019-07-22">22.VII.2019</date>
, 1 ♁, 
<specimenCount id="9D5DFD06690EFFBAFDBB4090FDBA09C1" box="[536,580,1466,1493]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
; Tazheranskaya steppe, 
<date id="FFE5104F690EFFBAFCF54090FC1F09C0" box="[854,993,1466,1492]" pageId="31" pageNumber="32" value="2019-07-23">23.VII.2019</date>
, 9 ♁, 
<specimenCount id="9D5DFD06690EFFBAFB844090FBAC09C1" box="[1063,1106,1466,1493]" pageId="31" pageNumber="32" type="female">5 ♀</specimenCount>
; same loc., 
<date id="FFE5104F690EFFBAFB794090FA9809C0" box="[1242,1382,1466,1492]" pageId="31" pageNumber="32" value="2020-07-28">28.VII.2020</date>
, 
<specimenCount id="9D5DFD06690EFFBAFAD24090FA6209C1" box="[1393,1436,1466,1493]" pageId="31" pageNumber="32" type="female">4 ♀</specimenCount>
(GenBank ID: 
<collectionCode id="ED4AAE4A690EFFBAFE9D40F4FE9909EC" box="[318,359,1502,1528]" pageId="31" pageNumber="32">OK</collectionCode>
073174), I. Makhov [
<collectionCode id="ED4AAE4A690EFFBAFDF740F4FD7509EC" box="[596,651,1502,1528]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="31" pageNumber="32" type="Museum">ZIN</collectionCode>
]. 
<emphasis id="B92FEA9D690EFFBAFD3E40F4FCFC09EC" bold="true" box="[669,770,1502,1528]" italics="true" pageId="31" pageNumber="32">
<collectingRegion id="499FF86D690EFFBAFD3E40F4FCFC09EC" box="[669,770,1502,1528]" country="Russia" name="Buryatiya" pageId="31" pageNumber="32">Buryatia</collectingRegion>
</emphasis>
: Ulan-Ude, 
<date id="FFE5104F690EFFBAFC2E40F4FBE609ED" box="[909,1048,1502,1529]" pageId="31" pageNumber="32" value="1956-07-24">24.VII.1956</date>
, 1 ♁; same loc., 
<date id="FFE5104F690EFFBAFB7640F4FAA109ED" box="[1237,1375,1502,1529]" pageId="31" pageNumber="32" value="1956-07-11">11.VII.1956</date>
, 1 ♁, unknown collector; Kurba, 
<date id="FFE5104F690EFFBAFE71432BFD790A0F" box="[466,647,1537,1563]" pageId="31" pageNumber="32" value="1973-07-16" valueMax="1973-07-22" valueMin="1973-07-16">16–22.VII.1973</date>
, 2 ♁; Petropavlovka, 
<date id="FFE5104F690EFFBAFC27432BFBF10A0F" box="[900,1039,1537,1563]" pageId="31" pageNumber="32" value="1974-07-27">27.VII.1974</date>
, 1 ♁, A. Tarmaeva [
<collectionCode id="ED4AAE4A690EFFBAFB59432BFABE0A0F" box="[1274,1344,1537,1563]" pageId="31" pageNumber="32">ISEA</collectionCode>
]; Dobo-Yenkhor, 
<date id="FFE5104F690EFFBAFEA6430FFE6F0A54" box="[261,401,1573,1600]" pageId="31" pageNumber="32" value="2008-07-15">15.VII.2008</date>
, 1 ♁, 
<collectionCode id="ED4AAE4A690EFFBAFE78430FFE130A2B" box="[475,493,1573,1599]" name="Tavera, Department of Geology and Geophysics" pageId="31" pageNumber="32">T</collectionCode>
. Gordeeva [
<collectionCode id="ED4AAE4A690EFFBAFDD4430FFD3E0A2B" box="[631,704,1573,1599]" pageId="31" pageNumber="32">IGEB</collectionCode>
]; Tatarsky Klyuch, 
<date id="FFE5104F690EFFBAFC02430FFBE00A2B" box="[929,1054,1573,1599]" pageId="31" pageNumber="32" value="2020-07-02">2.VII.2020</date>
, 1 ♁; Gusinoye, 
<date id="FFE5104F690EFFBAFB46430FFA9C0A2B" box="[1253,1378,1573,1599]" pageId="31" pageNumber="32" value="2016-07-01">1.VII.2016</date>
, 1 ♁ (GenBank ID: 
<collectionCode id="ED4AAE4A690EFFBAFEE34362FE930A76" box="[320,365,1608,1634]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="31" pageNumber="32">MZ</collectionCode>
148333), 
<specimenCount id="9D5DFD06690EFFBAFE754362FDFC0A77" box="[470,514,1608,1635]" pageId="31" pageNumber="32" type="female">2 ♀</specimenCount>
(GenBank ID: 
<collectionCode id="ED4AAE4A690EFFBAFD114362FD200A76" box="[690,734,1608,1634]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="31" pageNumber="32">MZ</collectionCode>
148334, 
<collectionCode id="ED4AAE4A690EFFBAFC9C4362FC950A77" box="[831,875,1608,1635]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="31" pageNumber="32">MZ</collectionCode>
148335), 
<specimenCount id="9D5DFD06690EFFBAFC754362FBFF0A77" box="[982,1025,1608,1635]" pageId="31" pageNumber="32" type="female">3 ♀</specimenCount>
, 8 spms.; Beloozersk, 
<date id="FFE5104F690EFFBAFAA74362FA7C0A76" box="[1284,1410,1608,1634]" pageId="31" pageNumber="32" value="2016-07-03">3.VII.2016</date>
, 2 spms.; 
<collectingRegion id="499FF86D690EFFBAFF454346FECE0A92" box="[230,304,1644,1670]" country="Germany" name="Bayern" pageId="31" pageNumber="32">Bayan</collectingRegion>
, 
<date id="FFE5104F690EFFBAFE9F4341FE280A92" box="[316,470,1643,1670]" pageId="31" pageNumber="32" value="2016-07-05" valueMax="2016-07-06" valueMin="2016-07-05">5–6.VII.2016</date>
, 2 spms.; M. Tasarkhay, 
<date id="FFE5104F690EFFBAFD494346FC7D0A92" box="[746,899,1644,1670]" pageId="31" pageNumber="32" value="2016-07-07" valueMax="2016-07-08" valueMin="2016-07-07">7–8.VII.2016</date>
, 2 ♁, 
<specimenCount id="9D5DFD06690EFFBAFC654346FC0E0A92" box="[966,1008,1644,1670]" pageId="31" pageNumber="32" type="female">2 ♀</specimenCount>
, 6 spms.; Tarbagatay, 
<date id="FFE5104F690EFFBAFB4A4346FA8A0A92" box="[1257,1396,1644,1670]" pageId="31" pageNumber="32" value="2018-07-13">13.VII.2018</date>
, 13 ♁ (GenBank ID: 
<collectionCode id="ED4AAE4A690EFFBAFEFB43A5FE7A0ABD" box="[344,388,1679,1705]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="31" pageNumber="32">MZ</collectionCode>
148328, 
<collectionCode id="ED4AAE4A690EFFBAFE4043A5FDF10ABD" box="[483,527,1679,1705]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="31" pageNumber="32">MZ</collectionCode>
148329); same loc., 
<date id="FFE5104F690EFFBAFD4C43A5FC840ABD" box="[751,890,1679,1705]" pageId="31" pageNumber="32" value="2021-07-23">23.VII.2021</date>
, 40 ♁, 
<specimenCount id="9D5DFD06690EFFBAFC6943A5FC0A0ABE" box="[970,1012,1679,1706]" pageId="31" pageNumber="32" type="female">2 ♀</specimenCount>
, I. Makhov [
<collectionCode id="ED4AAE4A690EFFBAFB2143BAFB470ABE" box="[1154,1209,1680,1706]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="31" pageNumber="32" type="Museum">ZIN</collectionCode>
]. 
<emphasis id="B92FEA9D690EFFBAFB6943A5FA6A0ABD" bold="true" box="[1226,1428,1679,1706]" italics="true" pageId="31" pageNumber="32">Zabaikalsky Kray</emphasis>
: Borzinsky Distr., Daursky Nature Reserve, Adon-Chelon, 
<date id="FFE5104F690EFFBAFC8C4399FC480AD9" box="[815,950,1715,1741]" pageId="31" pageNumber="32" value="2022-08-01">1.VIII.2022</date>
, 
<specimenCount id="9D5DFD06690EFFBAFC674399FC0E0AD9" box="[964,1008,1715,1741]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
, A. Stekolshchikov; Sretensky Distr., Shilka river valley, 4.5 km NE of Molodovsk vill. (
<geoCoordinate id="EE6F5048690EFFBAFD4D43FCFC860AE5" box="[750,888,1750,1777]" degrees="52" direction="north" minutes="16" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="58" value="52.282776">52°16’58”N</geoCoordinate>
<geoCoordinate id="EE6F5048690EFFBAFC2143FCFBE60AE5" box="[898,1048,1750,1777]" degrees="117" direction="east" minutes="55" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="30" value="117.924995">117°55’30”E</geoCoordinate>
), 
<date id="FFE5104F690EFFBAFB9343FCFB440AE4" box="[1072,1210,1750,1776]" pageId="31" pageNumber="32" value="2020-07-11">11.VII.2020</date>
, 2 ♁, 
<specimenCount id="9D5DFD06690EFFBAFAA543FCFACA0AE5" box="[1286,1332,1750,1777]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
; Kuenga river valley, 
<quantity id="4CA39B6A690EFFBAFE8443D0FE9C0B07" box="[295,354,1785,1812]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="km" value="3.0">3 km</quantity>
N of Dunayevo vill. (
<geoCoordinate id="EE6F5048690EFFBAFDC243D3FD150B00" box="[609,747,1785,1812]" degrees="52" direction="north" minutes="05" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="53" value="52.098053">52°05’53”N</geoCoordinate>
<geoCoordinate id="EE6F5048690EFFBAFD5743D0FC740B00" box="[756,906,1786,1812]" degrees="117" direction="east" minutes="02" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="24" value="117.04">117°02’24”E</geoCoordinate>
), 
<date id="FFE5104F690EFFBAFC0343D0FBE00B00" box="[928,1054,1786,1812]" pageId="31" pageNumber="32" value="2020-07-08">8.VII.2020</date>
, 10 ♁ (GenBank ID: 
<collectionCode id="ED4AAE4A690EFFBAFAB443D0FABD0B00" box="[1303,1347,1786,1812]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="31" pageNumber="32">MZ</collectionCode>
148330, 
<collectionCode id="ED4AAE4A690EFFBAFF344237FF3D0B23" box="[151,195,1821,1847]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="31" pageNumber="32">MZ</collectionCode>
148331, 
<collectionCode id="ED4AAE4A690EFFBAFE874237FEAE0B23" box="[292,336,1821,1847]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="31" pageNumber="32">MZ</collectionCode>
148332); same loc., 
<date id="FFE5104F690EFFBAFD904237FD4E0B23" box="[563,688,1821,1847]" pageId="31" pageNumber="32" value="2020-07-09">9.VII.2020</date>
, 3 ♁, 
<specimenCount id="9D5DFD06690EFFBAFD564237FCDE0B2C" box="[757,800,1821,1848]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
; same loc., 
<date id="FFE5104F690EFFBAFC044237FBCC0B23" box="[935,1074,1821,1847]" pageId="31" pageNumber="32" value="2020-07-10">10.VII.2020</date>
, 3 ♁, Nerchinsky Distr., Nercha river valley, vicinity of Nerchinsk vill. (
<geoCoordinate id="EE6F5048690EFFBAFDF6426AFD200B4F" box="[597,734,1856,1883]" degrees="51" direction="north" minutes="59" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="01" value="51.983612">51°59’01”N</geoCoordinate>
<geoCoordinate id="EE6F5048690EFFBAFD46426AFC850B4F" box="[741,891,1856,1883]" degrees="116" direction="east" minutes="31" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="15" value="116.520836">116°31’15”E</geoCoordinate>
), 
<date id="FFE5104F690EFFBAFC33426BFBF30B4F" box="[912,1037,1857,1883]" pageId="31" pageNumber="32" value="2020-07-08">8.VII.2020</date>
, 2 ♁, I. Makhov [
<collectionCode id="ED4AAE4A690EFFBAFB70426BFAF40B4F" box="[1235,1290,1857,1883]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="31" pageNumber="32" type="Museum">ZIN</collectionCode>
].
</paragraph>
</subSubSection>
<caption id="DF2466076931FF85FF344260FD870B93" ID-DOI="http://doi.org/10.5281/zenodo.7965093" ID-Zenodo-Dep="7965093" httpUri="https://zenodo.org/record/7965093/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1866,1890]" targetBox="[162,1425,189,1838]" targetPageId="32" targetType="figure">
<paragraph id="8BE4368F6931FF85FF344260FD870B93" blockId="32.[151,1436,1866,1927]" pageId="32" pageNumber="33">
<emphasis id="B92FEA9D6931FF85FF344260FEED0B76" bold="true" box="[151,275,1866,1890]" pageId="32" pageNumber="33">FIGURE 9.</emphasis>
Genital segments and sternites A8 of Siberian “ 
<taxonomicName id="4C5B4D0C6931FF85FD5F4260FC260B76" box="[764,984,1866,1890]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6931FF85FD5F4260FC260B76" box="[764,984,1866,1890]" italics="true" pageId="32" pageNumber="33">Hemistola intermedia</emphasis>
</taxonomicName>
” (Krasnoyarsky Kray and Khakassia). A, C, E, G—genital segments; B, D, F—sternites A8.
</paragraph>
</caption>
<caption id="DF2466076930FF84FF344261FC4A0B9C" ID-DOI="http://doi.org/10.5281/zenodo.7965096" ID-Zenodo-Dep="7965096" httpUri="https://zenodo.org/record/7965096/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,1867,1891]" targetBox="[162,1423,191,1844]" targetPageId="33" targetType="figure">
<paragraph id="8BE4368F6930FF84FF344261FC4A0B9C" blockId="33.[151,1437,1867,1928]" pageId="33" pageNumber="34">
<emphasis id="B92FEA9D6930FF84FF344261FEE10B70" bold="true" box="[151,287,1867,1892]" pageId="33" pageNumber="34">FIGURE 10.</emphasis>
Genital segments and sternites A8 of Far Eastern 
<taxonomicName id="4C5B4D0C6930FF84FCB44266FBF10B77" authorityName="Hedemann" authorityYear="1879" box="[791,1039,1868,1892]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6930FF84FCB44266FBF10B77" box="[791,1039,1868,1892]" italics="true" pageId="33" pageNumber="34">Hemistola zimmermanni</emphasis>
</taxonomicName>
(Zabaikalsky Kray, Khabarovsky Kray and Primorsky Kray). A, C, E, G—genital segments; B, D, F, H—sternites A8.
</paragraph>
</caption>
<caption id="DF2466076933FF87FF344261FED30B9C" ID-DOI="http://doi.org/10.5281/zenodo.7965100" ID-Zenodo-Dep="7965100" httpUri="https://zenodo.org/record/7965100/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,250,1867,1891]" targetBox="[162,1427,186,1838]" targetPageId="34" targetType="figure">
<paragraph id="8BE4368F6933FF87FF344261FED30B9C" blockId="34.[151,1436,1867,1928]" pageId="34" pageNumber="35">
<emphasis id="B92FEA9D6933FF87FF344261FEDF0B70" bold="true" box="[151,289,1867,1892]" pageId="34" pageNumber="35">FIGURE 11.</emphasis>
Female genitalia of Siberian 
<taxonomicName id="4C5B4D0C6933FF87FDF44266FCAC0B77" authorityName="Hedemann" authorityYear="1879" box="[599,850,1868,1892]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6933FF87FDF44266FCAC0B77" box="[599,850,1868,1892]" italics="true" pageId="34" pageNumber="35">Hemistola zimmermanni</emphasis>
</taxonomicName>
. A–D—samples from Irkutskaya Oblast; E–H—samples from Buryatia.
</paragraph>
</caption>
<caption id="DF2466076932FF86FF344105FD140878" ID-DOI="http://doi.org/10.5281/zenodo.7965102" ID-Zenodo-Dep="7965102" httpUri="https://zenodo.org/record/7965102/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,1071,1095]" targetBox="[165,1420,187,1041]" targetPageId="35" targetType="figure">
<paragraph id="8BE4368F6932FF86FF344105FD140878" blockId="35.[151,1436,1071,1132]" pageId="35" pageNumber="36">
<emphasis id="B92FEA9D6932FF86FF344105FEE10853" bold="true" box="[151,287,1071,1095]" pageId="35" pageNumber="36">FIGURE 12.</emphasis>
Female genitalia of Siberian and Far Eastern 
<taxonomicName id="4C5B4D0C6932FF86FD4F411AFC1A0853" authorityName="Hedemann" authorityYear="1879" box="[748,996,1072,1096]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6932FF86FD4F411AFC1A0853" box="[748,996,1072,1096]" italics="true" pageId="35" pageNumber="36">Hemistola zimmermanni</emphasis>
</taxonomicName>
. A, B—samples from Buryatia; C—sample from Zabaikalsky Kray; D—sample from Primorsky Kray.
</paragraph>
</caption>
<subSubSection id="C34165046932FF86FF644197FE37090B" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BE4368F6932FF86FF644197FBF908EF" blockId="35.[151,1437,1213,2022]" pageId="35" pageNumber="36">
<emphasis id="B92FEA9D6932FF86FF644197FE9F08C3" bold="true" box="[199,353,1213,1239]" pageId="35" pageNumber="36">Distribution.</emphasis>
Siberian–Far Eastern, subboreal. From southern part of Krasnoyarsky Kray and 
<collectingRegion id="499FF86D6932FF86FAAA4197FA8208C3" box="[1289,1404,1213,1239]" country="Russia" name="Khakasiya" pageId="35" pageNumber="36">Khakassia</collectingRegion>
to Primorye. Outside 
<collectingCountry id="F34C761F6932FF86FECF41CBFE4908EF" box="[364,439,1249,1275]" name="Russia" pageId="35" pageNumber="36">Russia</collectingCountry>
in Mongolia, N and NE 
<collectingCountry id="F34C761F6932FF86FD6F41CBFCEC08EF" box="[716,786,1249,1275]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
, N and central 
<collectingCountry id="F34C761F6932FF86FC1841CBFBFD08EF" box="[955,1027,1249,1275]" name="South Korea" pageId="35" pageNumber="36">Korea</collectingCountry>
.
</paragraph>
<paragraph id="8BE4368F6932FF86FF64402FFE37090B" blockId="35.[151,1437,1213,2022]" box="[199,457,1285,1311]" pageId="35" pageNumber="36">
<emphasis id="B92FEA9D6932FF86FF64402FFEAC090B" bold="true" box="[199,338,1285,1311]" pageId="35" pageNumber="36">Hostplants:</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34165046932FF83FF644002FC600F2D" lastPageId="38" lastPageNumber="39" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8BE4368F6932FF86FF644002FF050A03" blockId="35.[151,1437,1213,2022]" pageId="35" pageNumber="36">
<emphasis id="B92FEA9D6932FF86FF644002FEC50956" bold="true" box="[199,315,1320,1346]" pageId="35" pageNumber="36">Remarks.</emphasis>
The species 
<taxonomicName id="4C5B4D0C6932FF86FE6A4003FD9D0956" authority="Djakonov, 1926" box="[457,611,1320,1346]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6932FF86FE6A4003FD9D0956" box="[457,611,1320,1346]" italics="true" pageId="35" pageNumber="36">H. intermedia</emphasis>
</taxonomicName>
was described from the environs of Minusinsk (the south of Krasnoyarsky Kray) by 
<bibRefCitation id="EFCA4B7E6932FF86FEA94066FE2F0972" author="Djakonov, A. M." box="[266,465,1356,1382]" pageId="35" pageNumber="36" pagination="1 - 78" refId="ref70583" refString="Djakonov, A. M. (1926) Zur Kenntnis der Geometriden fauna des Minussinsk Bezirks (Sibirien, Jenissej Gouv.). Jahrbuch des Martjanov'schen Staatsmuseums in Minussinsk (Sibirien), 4 (1), 1 - 78. [in Russian and German]" type="journal article" year="1926">Djakonov (1926)</bibRefCitation>
. He noted that 
<taxonomicName id="4C5B4D0C6932FF86FD294066FCD40972" authority="Djakonov, 1926" box="[650,810,1356,1382]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6932FF86FD294066FCD40972" box="[650,810,1356,1382]" italics="true" pageId="35" pageNumber="36">H. intermedia</emphasis>
</taxonomicName>
on the average smaller than 
<taxonomicName id="4C5B4D0C6932FF86FB274066FABC0971" authorityName="Hedemann" authorityYear="1879" box="[1156,1346,1356,1381]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6932FF86FB274066FABC0971" box="[1156,1346,1356,1381]" italics="true" pageId="35" pageNumber="36">H. zimmermanni</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5B4D0C6932FF86FA224066FEBC099D" authorityName=": Berlov &amp; Berlov" authorityYear="2006" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="chrysoprasaria">
<emphasis id="B92FEA9D6932FF86FA224066FEBC099D" italics="true" pageId="35" pageNumber="36">H. chrysoprasaria</emphasis>
</taxonomicName>
, however, the main differences of this taxon from the closer 
<taxonomicName id="4C5B4D0C6932FF86FBA3405AFB42099D" authorityName="Hedemann" authorityYear="1879" box="[1024,1212,1392,1417]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6932FF86FBA3405AFB42099D" box="[1024,1212,1392,1417]" italics="true" pageId="35" pageNumber="36">H. zimmermanni</emphasis>
</taxonomicName>
were the shortened antennal pectinations and relatively longer coecum of aedeagus: “the strongly elongated coecum in the new species is considerably longer and thinner [than in 
<taxonomicName id="4C5B4D0C6932FF86FDDC409DFCC409C4" authorityName="Hedemann" authorityYear="1879" box="[639,826,1463,1488]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6932FF86FDDC409DFCC409C4" box="[639,826,1463,1488]" italics="true" pageId="35" pageNumber="36">H. zimmermanni</emphasis>
</taxonomicName>
]: it is 3.5 times shorter than the length of the phallos, while in 
<taxonomicName id="4C5B4D0C6932FF86FF5B40F0FE4C09E7" authorityName="Hedemann" authorityYear="1879" box="[248,434,1498,1523]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6932FF86FF5B40F0FE4C09E7" box="[248,434,1498,1523]" italics="true" pageId="35" pageNumber="36">H. zimmermanni</emphasis>
</taxonomicName>
(and in 
<taxonomicName id="4C5B4D0C6932FF86FDAE40F0FD9309E7" baseAuthorityName="Hubner" baseAuthorityYear="1789" box="[525,621,1498,1523]" class="Insecta" family="Geometridae" genus="Geometra" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="vernaria">
<emphasis id="B92FEA9D6932FF86FDAE40F0FD9309E7" box="[525,621,1498,1523]" italics="true" pageId="35" pageNumber="36">vernaria</emphasis>
</taxonomicName>
[= 
<taxonomicName id="4C5B4D0C6932FF86FD3040F0FC9E09E7" authorityName=": Berlov &amp; Berlov" authorityYear="2006" box="[659,864,1497,1523]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="chrysoprasaria">
<emphasis id="B92FEA9D6932FF86FD3040F0FC9E09E7" box="[659,864,1497,1523]" italics="true" pageId="35" pageNumber="36">H. chrysoprasaria</emphasis>
</taxonomicName>
]) coecum is 4.5 times shorter than the length of the phallos.”
</paragraph>
<paragraph id="8BE4368F6932FF86FF64430AFC8A0B1B" blockId="35.[151,1437,1213,2022]" pageId="35" pageNumber="36">
The status of the taxon 
<taxonomicName id="4C5B4D0C6932FF86FE77430BFD8E0A2E" authority="Djakonov, 1926" box="[468,624,1568,1594]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6932FF86FE77430BFD8E0A2E" box="[468,624,1568,1594]" italics="true" pageId="35" pageNumber="36">H. intermedia</emphasis>
</taxonomicName>
remained unclear. 
<bibRefCitation id="EFCA4B7E6932FF86FCEA430AFC160A2F" author="Wehrli, E." box="[841,1000,1568,1595]" pageId="35" pageNumber="36" pagination="8 - 30" refId="ref74946" refString="Wehrli, E. (1929) Beitrag zur Geometriden-Fauna von Minussinsk Sibirien, Gouv. Jenissej. Ezhegodnik Gosudarstvannogo Muzeya imeni N. M. Martyanova, 6 (1), 8 - 30. [in German]" type="journal article" year="1929">Wehrli (1929)</bibRefCitation>
supposed that 
<taxonomicName id="4C5B4D0C6932FF86FB30430BFAD10A2E" authority="Djakonov, 1926" box="[1171,1327,1568,1594]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6932FF86FB30430BFAD10A2E" box="[1171,1327,1568,1594]" italics="true" pageId="35" pageNumber="36">H. intermedia</emphasis>
</taxonomicName>
might be either a mere form of 
<taxonomicName id="4C5B4D0C6932FF86FE3F436FFDA70A4A" authorityName="Hedemann" authorityYear="1879" box="[412,601,1605,1630]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6932FF86FE3F436FFDA70A4A" box="[412,601,1605,1630]" italics="true" pageId="35" pageNumber="36">H. zimmermanni</emphasis>
</taxonomicName>
or a young species incompletely differentiated from 
<taxonomicName id="4C5B4D0C6932FF86FB66436FFA6B0A4A" authorityName=": Berlov &amp; Berlov" authorityYear="2006" box="[1221,1429,1604,1630]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="chrysoprasaria">
<emphasis id="B92FEA9D6932FF86FB66436FFA6B0A4A" box="[1221,1429,1604,1630]" italics="true" pageId="35" pageNumber="36">H. chrysoprasaria</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EFCA4B7E6932FF86FF34434DFEAA0A95" author="Viidalepp, J." box="[151,340,1639,1665]" pageId="35" pageNumber="36" refId="ref74481" refString="Viidalepp, J. (1996) Checklist of the Geometridae (Lepidoptera) of the former U. S. S. R. Apollo Books, Stenstrup, 111 pp." type="book" year="1996">Viidalepp (1996)</bibRefCitation>
interpreted 
<taxonomicName id="4C5B4D0C6932FF86FE79434DFDAD0A95" box="[474,595,1639,1665]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6932FF86FE79434DFDAD0A95" box="[474,595,1639,1665]" italics="true" pageId="35" pageNumber="36">intermedia</emphasis>
</taxonomicName>
as a subspecies of 
<taxonomicName id="4C5B4D0C6932FF86FC804342FC110A95" authorityName=": Berlov &amp; Berlov" authorityYear="2006" box="[803,1007,1639,1665]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="chrysoprasaria">
<emphasis id="B92FEA9D6932FF86FC804342FC110A95" box="[803,1007,1639,1665]" italics="true" pageId="35" pageNumber="36">H. chrysoprasaria</emphasis>
</taxonomicName>
. In the first edition of the Catalogue of the 
<taxonomicName id="4C5B4D0C6932FF86FF6343A1FEB90AB1" authorityName="Linnaeus" authorityYear="1758" box="[192,327,1675,1701]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
of 
<collectingCountry id="F34C761F6932FF86FEC843A1FE480AB1" box="[363,438,1675,1701]" name="Russia" pageId="35" pageNumber="36">Russia</collectingCountry>
(
<bibRefCitation id="EFCA4B7E6932FF86FE6043A1FD5B0AB1" author="Mironov, V. G. &amp; Beljaev, E. A. &amp; Vasilenko, S. V." box="[451,677,1675,1701]" pageId="35" pageNumber="36" pagination="190 - 227" refId="ref73212" refString="Mironov, V. G., Beljaev, E. A. &amp; Vasilenko, S. V. (2008) Sem. Geometridae. In: Sinev, S. Yu. (Ed.), Catalogue of the Lepidoptera of Russia. KMK Scientific Press Ltd., St. Petersburg-Moscow, pp. 190 - 227. [in Russian]" type="book chapter" year="2008">
Mironov 
<emphasis id="B92FEA9D6932FF86FD8D43A6FDA00AB1" box="[558,606,1675,1701]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. 2008
</bibRefCitation>
) 
<taxonomicName id="4C5B4D0C6932FF86FD1043A6FCB00AB1" authority="Djakonov, 1926" box="[691,846,1675,1701]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6932FF86FD1043A6FCB00AB1" box="[691,846,1675,1701]" italics="true" pageId="35" pageNumber="36">H. intermedia</emphasis>
</taxonomicName>
was not mentioned, however, later, E.A. Beljaev and V.G. Mironov considered 
<taxonomicName id="4C5B4D0C6932FF86FE1B4385FDAC0ADC" authority="Djakonov, 1926" box="[440,594,1710,1736]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6932FF86FE1B4385FDAC0ADC" box="[440,594,1710,1736]" italics="true" pageId="35" pageNumber="36">H. intermedia</emphasis>
</taxonomicName>
a good species (
<bibRefCitation id="EFCA4B7E6932FF86FCAB4384FC5C0ADC" author="Beljaev, E. A." box="[776,930,1710,1736]" pageId="35" pageNumber="36" refId="ref68981" refString="Beljaev, E. A. (2016) Fam. Geometridae - Pyadenitzi. In: Lelei, A. S. (Ed.), Annotated catalogue of the insects of Russian Far East. Vol. 2. Lepidoptera. Dal'nauka, Vladivostok, pp. 518 - 666. [in Russian]" type="book" year="2016">Beljaev 2016</bibRefCitation>
; 
<bibRefCitation id="EFCA4B7E6932FF86FC0F4384FB370ADC" author="Beljaev, E. A. &amp; Mironov, V. G." box="[940,1225,1710,1736]" pageId="35" pageNumber="36" pagination="235 - 281" refId="ref68795" refString="Beljaev, E. A. &amp; Mironov, V. G. (2019) Geometridae. In: Sinev, S. Yu. (Ed.), Catalogue of the Lepidoptera of Russia. 2 nd Edition. Zoological Institute RAS, St. Petersburg, pp. 235 - 281. [in Russian]" type="book chapter" year="2019">Beljaev &amp; Mironov 2019</bibRefCitation>
). The moths of this group from nearby regions (Republic of 
<collectingRegion id="499FF86D6932FF86FDC343F8FD650AF8" box="[608,667,1746,1772]" country="Russia" name="Tyva" pageId="35" pageNumber="36">Tuva</collectingRegion>
, Altaisky Kray, Irkutskaya Oblast) have traditionally been identified as 
<taxonomicName id="4C5B4D0C6932FF86FF1543DCFE8E0B1B" authorityName="Hedemann" authorityYear="1879" box="[182,368,1782,1807]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6932FF86FF1543DCFE8E0B1B" box="[182,368,1782,1807]" italics="true" pageId="35" pageNumber="36">H. zimmermanni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCA4B7E6932FF86FEDC43DFFDD10B1B" author="Viidalepp, J." box="[383,559,1781,1807]" pageId="35" pageNumber="36" pagination="842 - 852" refId="ref74401" refString="Viidalepp, J. (1976) List of geometrid moths (Lepidoptera, Geometridae) of the fauna of USSR, I. Entomological review, 55 (4), 842 - 852. [in Russian]" type="journal article" year="1976">Viidalepp 1976</bibRefCitation>
, 
<bibRefCitation id="EFCA4B7E6932FF86FD9943DFFD8B0B1B" author="Viidalepp, J." box="[570,629,1781,1807]" pageId="35" pageNumber="36" refId="ref74481" refString="Viidalepp, J. (1996) Checklist of the Geometridae (Lepidoptera) of the former U. S. S. R. Apollo Books, Stenstrup, 111 pp." type="book" year="1996">1996</bibRefCitation>
; 
<bibRefCitation id="EFCA4B7E6932FF86FD2343DFFC970B1B" author="Volynkin, A. V. &amp; Trilikauskas, L. A. &amp; Baghirov, R. T-O. &amp; Burmistrov, M. V. &amp; Byvaltsev, A. M. &amp; Vasilenko, S. V. &amp; Vishnevskaya, M. S. &amp; Danilov, Yu. N. &amp; Dudko, A. Yu. &amp; Dudko, R. Yu. &amp; Knyshov, A. A. &amp; Kosova, O. V. &amp; Kostrov, D. V. &amp; Krugova, T. M. &amp; Kuznetsova, R. O. &amp; Kuzmenkin, D. V. &amp; Legalov, A. A. &amp; Lvovsky, A. L. &amp; Namyatova, A. A. &amp; Nedoshivina, S. V. &amp; Perunov, Yu. E. &amp; Reschikov, A. V. &amp; Sinev, S. Yu. &amp; Solovarov, V. V. &amp; Tyumaseva, Z. I. &amp; Udalov, I. A. &amp; Ustyuzhanin, P. Ya. &amp; Filimonov, R. V. &amp; Tshernyshev, S. E. &amp; Tshesnokova, S. V. &amp; Sheikin, S. D. &amp; Shcherbakov, M. V. &amp; Yanygina, L. V." box="[640,873,1781,1808]" pageId="35" pageNumber="36" refId="ref74637" refString="Volynkin, A. V., Trilikauskas, L. A., Baghirov, R. T-O., Burmistrov, M. V., Byvaltsev, A. M., Vasilenko, S. V., Vishnevskaya, M. S., Danilov, Yu. N., Dudko, A. Yu., Dudko, R. Yu., Knyshov, A. A., Kosova, O. V., Kostrov, D. V., Krugova, T. M., Kuznetsova, R. O., Kuzmenkin, D. V., Legalov, A. A., Lvovsky, A. L., Namyatova, A. A., Nedoshivina, S. V., Perunov, Yu. E., Reschikov, A. V., Sinev, S. Yu., Solovarov, V. V., Tyumaseva, Z. I., Udalov, I. A., Ustyuzhanin, P. Ya., Filimonov, R. V., Tshernyshev, S. E., Tshesnokova, S. V., Sheikin, S. D., Shcherbakov, M. V. &amp; Yanygina, L. V. (2011) Invertebrates of the Tigirek Strict Nature Reserve (an annotated check-list). In: Davydov, E. A. (Ed.), Biota of Tigirek Nature Reserve. Proceedings of the Tigirek State Natural Reserve. Vol. 4. Tigirek State Reserve, Barnaul, pp. 165 - 226. [in Russian] https: // doi. org / 10.53005 / 20767390 _ 2011 _ 4 _ 165" type="book" year="2011">
Volynkin 
<emphasis id="B92FEA9D6932FF86FD4D43DCFCE10B1B" box="[750,799,1781,1807]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. 2011
</bibRefCitation>
).
</paragraph>
<paragraph id="8BE4368F6932FF80FF644233FF0E0AB7" blockId="35.[151,1437,1213,2022]" lastBlockId="37.[151,1437,1312,2023]" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36">
When comparing “ 
<taxonomicName id="4C5B4D0C6932FF86FE0E4230FDAC0B27" authority="Djakonov, 1926" box="[429,594,1817,1843]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6932FF86FE0E4230FDAC0B27" box="[429,594,1817,1843]" italics="true" pageId="35" pageNumber="36">H. intermedia</emphasis>
</taxonomicName>
” from the 
<typeStatus id="54E0882D6932FF86FD414233FCED0B27" box="[738,787,1817,1843]" pageId="35" pageNumber="36">type</typeStatus>
locality and nearby regions with the Far Eastern 
<taxonomicName id="4C5B4D0C6932FF86FA224230FECF0B42" authorityName="Hedemann" authorityYear="1879" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6932FF86FA224230FECF0B42" italics="true" pageId="35" pageNumber="36">H. zimmermanni</emphasis>
</taxonomicName>
, indeed, the difference in the size of the individuals and in the length of the antennal pectinations is obvious. The moths from the eastern part of the range are generally much larger than those from the west; however, the serial material from different regions (Krasnoyarsky Kray, Irkutskaya Oblast, Buryatia, Zabaikalsky Kray, Amurskaya Oblast, Khabarovsky Kray, Primorsky Kray, 
<collectingCountry id="F34C761F6932FF86FCBD428DFC700BD5" box="[798,910,1959,1985]" name="Mongolia" pageId="35" pageNumber="36">Mongolia</collectingCountry>
) shows that individuals vary markedly in size. In one location, along with moths of the usual size, very small individuals are sometimes found. In my opinion, the main diagnostic feature of 
<taxonomicName id="4C5B4D0C6934FF80FE77400BFD8B092E" authority="Djakonov, 1926" box="[468,629,1312,1338]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6934FF80FE77400BFD8B092E" box="[468,629,1312,1338]" italics="true" pageId="37" pageNumber="38">H. intermedia</emphasis>
</taxonomicName>
indicated by Djakonov (shortened pectinations of antennae) fits into variability spectrum of 
<taxonomicName id="4C5B4D0C6934FF80FE3D406FFDA4094A" authorityName="Hedemann" authorityYear="1879" box="[414,602,1349,1374]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6934FF80FE3D406FFDA4094A" box="[414,602,1349,1374]" italics="true" pageId="37" pageNumber="38">H. zimmermanni</emphasis>
</taxonomicName>
. The moths of 
<taxonomicName id="4C5B4D0C6934FF80FCA7406FFC3E094A" authorityName="Hedemann" authorityYear="1879" box="[772,960,1349,1374]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6934FF80FCA7406FFC3E094A" box="[772,960,1349,1374]" italics="true" pageId="37" pageNumber="38">H. zimmermanni</emphasis>
</taxonomicName>
considerably vary in size throughout their range, and as a result, different parts and appendages (including the processes of flagellomeres) vary proportionally. For clarity, we can compare the antennae in 
<figureCitation id="13602A0A6934FF80FD3B40A6FD2109B3" box="[664,735,1420,1447]" captionStart="FIGURE 7" captionStartId="30.[151,250,1856,1880]" captionTargetBox="[151,1436,187,1826]" captionTargetId="figure-25@30.[151,1436,181,1832]" captionTargetPageId="30" captionText="FIGURE 7. Male antennae of Hemistola. A, C, E, G, I, K, M, O—antenna; B, D, F, H, J, L, N, P—10th flagellomere. M, N, O, P—antennae aligned in length. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965087" httpUri="https://zenodo.org/record/7965087/files/figure.png" pageId="37" pageNumber="38">Fig. 7</figureCitation>
. It shows the antennae of males from the western part of the range to the eastern one (A, C, E, G, I, K). On the right, the 10 
<superScript id="7C2E9BC76934FF80FCC7409AFC8F09AA" attach="left" box="[868,881,1456,1470]" fontSize="6" pageId="37" pageNumber="38">th</superScript>
flagellomeres of the corresponding antennae are presented separately. When comparing the antenna of the male from the 
<typeStatus id="54E0882D6934FF80FC7340FFFBFF09FB" box="[976,1025,1493,1519]" pageId="37" pageNumber="38">type</typeStatus>
locality of 
<taxonomicName id="4C5B4D0C6934FF80FB2740FFFADD09FA" authority="Djakonov, 1926" box="[1156,1315,1492,1518]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6934FF80FB2740FFFADD09FA" box="[1156,1315,1492,1518]" italics="true" pageId="37" pageNumber="38">H. intermedia</emphasis>
</taxonomicName>
(
<figureCitation id="13602A0A6934FF80FA9140FEFF470A07" captionStart="FIGURE 7" captionStartId="30.[151,250,1856,1880]" captionTargetBox="[151,1436,187,1826]" captionTargetId="figure-25@30.[151,1436,181,1832]" captionTargetPageId="30" captionText="FIGURE 7. Male antennae of Hemistola. A, C, E, G, I, K, M, O—antenna; B, D, F, H, J, L, N, P—10th flagellomere. M, N, O, P—antennae aligned in length. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965087" httpUri="https://zenodo.org/record/7965087/files/figure.png" pageId="37" pageNumber="38">Figs. 7A, 7B</figureCitation>
) and the antenna of the male from Khabarovsky Kray (
<figureCitation id="13602A0A6934FF80FC8C40D2FC4F0A07" box="[815,945,1528,1555]" captionStart="FIGURE 7" captionStartId="30.[151,250,1856,1880]" captionTargetBox="[151,1436,187,1826]" captionTargetId="figure-25@30.[151,1436,181,1832]" captionTargetPageId="30" captionText="FIGURE 7. Male antennae of Hemistola. A, C, E, G, I, K, M, O—antenna; B, D, F, H, J, L, N, P—10th flagellomere. M, N, O, P—antennae aligned in length. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965087" httpUri="https://zenodo.org/record/7965087/files/figure.png" pageId="37" pageNumber="38">Figs. 7I, 7J</figureCitation>
), it can be seen that flagellomere processes are indeed different. However, the difference in the total length of the antenna must also be taken into account. At the bottom of the figure these two antennae are aligned in length. It is obvious here that the difference in length of the processes of flagellomeres is hardly noticeable, although these antennae belong to tentatively different species.
</paragraph>
<caption id="DF2466076935FF81FF34424EFC6E0BD0" ID-DOI="http://doi.org/10.5281/zenodo.7965104" ID-Zenodo-Dep="7965104" httpUri="https://zenodo.org/record/7965104/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1892,1916]" targetBox="[155,1432,185,1863]" targetPageId="36" targetType="figure">
<paragraph id="8BE4368F6935FF81FF34424EFC6E0BD0" blockId="36.[151,1436,1892,1988]" pageId="36" pageNumber="37">
<emphasis id="B92FEA9D6935FF81FF34424EFEDD0B68" bold="true" box="[151,291,1892,1916]" pageId="36" pageNumber="37">FIGURE 13.</emphasis>
Bayesian tree of 
<taxonomicName id="4C5B4D0C6935FF81FE41424EFDB40B68" authorityName="Warren" authorityYear="1893" box="[482,586,1892,1916]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEA9D6935FF81FE41424EFDB40B68" box="[482,586,1892,1916]" italics="true" pageId="36" pageNumber="37">Hemistola</emphasis>
</taxonomicName>
based on analysis of 
<emphasis id="B92FEA9D6935FF81FC97424EFC9E0B6F" box="[820,864,1892,1915]" italics="true" pageId="36" pageNumber="37">COI</emphasis>
gene sequences. The numbers above the branches are Bayesian posterior probabilities. Support values less than 0.7 are not shown. The sequences of specimens from type locality of 
<taxonomicName id="4C5B4D0C6935FF81FF344286FE8A0BD0" box="[151,372,1964,1988]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6935FF81FF344286FE8A0BD0" box="[151,372,1964,1988]" italics="true" pageId="36" pageNumber="37">Hemistola intermedia</emphasis>
</taxonomicName>
and adjacent area (Khakassia) are highlighted in red.
</paragraph>
</caption>
<caption id="DF2466076934FF80FF34419CFB5D08DA" ID-DOI="http://doi.org/10.5281/zenodo.7965108" ID-Zenodo-Dep="7965108" box="[151,1187,1206,1231]" httpUri="https://zenodo.org/record/7965108/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,1206,1230]" targetBox="[152,1434,182,1181]" targetPageId="37" targetType="figure">
<paragraph id="8BE4368F6934FF80FF34419CFB5D08DA" blockId="37.[151,1187,1206,1231]" box="[151,1187,1206,1231]" pageId="37" pageNumber="38">
<emphasis id="B92FEA9D6934FF80FF34419CFEDE08DA" bold="true" box="[151,288,1206,1230]" pageId="37" pageNumber="38">FIGURE 14.</emphasis>
Lectotypes of (A) 
<taxonomicName id="4C5B4D0C6934FF80FE43419CFCB308DA" authority="Hedemann" authorityName="Hedemann" authorityYear="1879" box="[480,845,1206,1230]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6934FF80FE43419CFD2708DA" box="[480,729,1206,1230]" italics="true" pageId="37" pageNumber="38">Hemistola zimmermanni</emphasis>
Hedemann
</taxonomicName>
and (B) 
<taxonomicName id="4C5B4D0C6934FF80FC05419DFB5D08DA" authority="Djakonov." authorityName="Djakonov." box="[934,1187,1206,1230]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6934FF80FC05419DFBCD08DA" box="[934,1075,1206,1230]" italics="true" pageId="37" pageNumber="38">H. intermedia</emphasis>
Djakonov.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BE4368F6934FF80FF644386FAE60B6F" blockId="37.[151,1437,1312,2023]" pageId="37" pageNumber="38">
The second features used by Djakonov as diagnostic for 
<taxonomicName id="4C5B4D0C6934FF80FCF14387FC110AD2" authority="Djakonov, 1926" box="[850,1007,1708,1734]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6934FF80FCF14387FC110AD2" box="[850,1007,1708,1734]" italics="true" pageId="37" pageNumber="38">H. intermedia</emphasis>
</taxonomicName>
is the length of coecum of aedeagus. I have studied at least 15 samples from different locations and found no difference in the coecum length between “ 
<taxonomicName id="4C5B4D0C6934FF80FF0743DFFEBE0B1A" authority="Djakonov, 1926" box="[164,320,1780,1806]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6934FF80FF0743DFFEBE0B1A" box="[164,320,1780,1806]" italics="true" pageId="37" pageNumber="38">H. intermedia</emphasis>
</taxonomicName>
” and 
<taxonomicName id="4C5B4D0C6934FF80FE2543DFFDBC0B1A" authorityName="Hedemann" authorityYear="1879" box="[390,578,1781,1806]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6934FF80FE2543DFFDBC0B1A" box="[390,578,1781,1806]" italics="true" pageId="37" pageNumber="38">H. zimmermanni</emphasis>
</taxonomicName>
. Several phalloses from the males collected in different regions (from west to east) are presented in 
<figureCitation id="13602A0A6934FF80FE0E4232FE0F0B27" box="[429,497,1816,1843]" captionStart="FIGURE 8" captionStartId="31.[151,250,719,743]" captionTargetBox="[209,1378,187,687]" captionTargetId="figure-15@31.[200,1387,181,695]" captionTargetPageId="31" captionText="FIGURE 8. Aedeagi of Siberian and Far Eastern Hemistola. A–D—aedeagi of “Hemistola intermedia” from Krasnoyarsky Kray and Khakassia; E–H—aedeagi of H. zimmermanni from Zabaikalsky Kray, Khabarovsky Kray and Primorsky Kray." figureDoi="http://doi.org/10.5281/zenodo.7965091" httpUri="https://zenodo.org/record/7965091/files/figure.png" pageId="37" pageNumber="38">Fig. 8</figureCitation>
. Thus, the diagnostic characters the lengths of the coecum, as well as the length of antennal pectinations seem to be unreasonable. Regarding to the male genital segment and sternite A8, there are also no evident differences between “ 
<taxonomicName id="4C5B4D0C6934FF80FDA4424BFD5C0B6E" authority="Djakonov, 1926" box="[519,674,1888,1914]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6934FF80FDA4424BFD5C0B6E" box="[519,674,1888,1914]" italics="true" pageId="37" pageNumber="38">H. intermedia</emphasis>
</taxonomicName>
” (
<figureCitation id="13602A0A6934FF80FD1D424AFCAC0B6F" box="[702,850,1888,1915]" captionStart="FIGURE 9" captionStartId="32.[151,250,1866,1890]" captionTargetBox="[162,1425,189,1838]" captionTargetId="figure-25@32.[151,1435,181,1842]" captionTargetPageId="32" captionText="FIGURE 9. Genital segments and sternites A8 of Siberian “Hemistola intermedia” (Krasnoyarsky Kray and Khakassia). A, C, E, G—genital segments; B, D, F—sternites A8." figureDoi="http://doi.org/10.5281/zenodo.7965093" httpUri="https://zenodo.org/record/7965093/files/figure.png" pageId="37" pageNumber="38">Figs. 9A–9G</figureCitation>
) and 
<taxonomicName id="4C5B4D0C6934FF80FC33424BFBB40B6E" authorityName="Hedemann" authorityYear="1879" box="[912,1098,1889,1914]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6934FF80FC33424BFBB40B6E" box="[912,1098,1889,1914]" italics="true" pageId="37" pageNumber="38">H. zimmermanni</emphasis>
</taxonomicName>
(
<figureCitation id="13602A0A6934FF80FBF9424AFAF20B6F" box="[1114,1292,1888,1915]" captionStart="FIGURE 10" captionStartId="33.[151,250,1867,1891]" captionTargetBox="[162,1423,191,1844]" captionTargetId="figure-15@33.[153,1434,181,1844]" captionTargetPageId="33" captionText="FIGURE 10. Genital segments and sternites A8 of Far Eastern Hemistola zimmermanni (Zabaikalsky Kray, Khabarovsky Kray and Primorsky Kray). A, C, E, G—genital segments; B, D, F, H—sternites A8." figureDoi="http://doi.org/10.5281/zenodo.7965096" httpUri="https://zenodo.org/record/7965096/files/figure.png" pageId="37" pageNumber="38">Figs. 10A–10H</figureCitation>
).
</paragraph>
<paragraph id="8BE4368F6934FF83FF6442AFFCC90D09" blockId="37.[151,1437,1312,2023]" lastBlockId="38.[151,1437,151,825]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
It is noteworthy that the shape and size of lamella postvaginalis in female genitalia are also somewhat variable (
<figureCitation id="13602A0A6934FF80FF3C4282FF050BD7" box="[159,251,1960,1987]" captionStart="FIGURE 11" captionStartId="34.[151,250,1867,1891]" captionTargetBox="[162,1427,186,1838]" captionTargetId="figure-25@34.[154,1433,181,1844]" captionTargetPageId="34" captionText="FIGURE 11. Female genitalia of Siberian Hemistola zimmermanni. A–D—samples from Irkutskaya Oblast; E–H—samples from Buryatia." figureDoi="http://doi.org/10.5281/zenodo.7965100" httpUri="https://zenodo.org/record/7965100/files/figure.png" pageId="37" pageNumber="38">Figs. 11</figureCitation>
, 
<figureCitation id="13602A0A6934FF80FEA44283FED80BD7" box="[263,294,1961,1987]" captionStart="FIGURE 12" captionStartId="35.[151,250,1071,1095]" captionTargetBox="[165,1420,187,1041]" captionTargetId="figure-15@35.[158,1429,181,1048]" captionTargetPageId="35" captionText="FIGURE 12. Female genitalia of Siberian and Far Eastern Hemistola zimmermanni. A, B—samples from Buryatia; C—sample from Zabaikalsky Kray; D—sample from Primorsky Kray." figureDoi="http://doi.org/10.5281/zenodo.7965102" httpUri="https://zenodo.org/record/7965102/files/figure.png" pageId="37" pageNumber="38">12</figureCitation>
). Occasionally, the appearance of the lamella postvaginalis is clearly different in moths collected from the same location on the same date (compare 
<figureCitation id="13602A0A6934FF80FD3742E6FC990BF3" box="[660,871,1996,2023]" captionStart="FIGURE 11" captionStartId="34.[151,250,1867,1891]" captionTargetBox="[162,1427,186,1838]" captionTargetId="figure-25@34.[154,1433,181,1844]" captionTargetPageId="34" captionText="FIGURE 11. Female genitalia of Siberian Hemistola zimmermanni. A–D—samples from Irkutskaya Oblast; E–H—samples from Buryatia." figureDoi="http://doi.org/10.5281/zenodo.7965100" httpUri="https://zenodo.org/record/7965100/files/figure.png" pageId="37" pageNumber="38">Figs. 11A and 11B</figureCitation>
; 
<figureCitation id="13602A0A6934FF80FCD142E6FBB80BF3" box="[882,1094,1996,2023]" captionStart="FIGURE 11" captionStartId="34.[151,250,1867,1891]" captionTargetBox="[162,1427,186,1838]" captionTargetId="figure-25@34.[154,1433,181,1844]" captionTargetPageId="34" captionText="FIGURE 11. Female genitalia of Siberian Hemistola zimmermanni. A–D—samples from Irkutskaya Oblast; E–H—samples from Buryatia." figureDoi="http://doi.org/10.5281/zenodo.7965100" httpUri="https://zenodo.org/record/7965100/files/figure.png" pageId="37" pageNumber="38">Figs. 11C and 11D</figureCitation>
; 
<figureCitation id="13602A0A6934FF80FBF142E6FA690BF3" box="[1106,1431,1996,2023]" captionStart="FIGURE 11" captionStartId="34.[151,250,1867,1891]" captionTargetBox="[162,1427,186,1838]" captionTargetId="figure-25@34.[154,1433,181,1844]" captionTargetPageId="34" captionText="FIGURE 11. Female genitalia of Siberian Hemistola zimmermanni. A–D—samples from Irkutskaya Oblast; E–H—samples from Buryatia." figureDoi="http://doi.org/10.5281/zenodo.7965100" httpUri="https://zenodo.org/record/7965100/files/figure.png" pageId="37" pageNumber="38">Figs. 11E, 11F, 11G and 11H</figureCitation>
; 
<figureCitation id="13602A0A6937FF83FF3445BDFE930CA5" box="[151,365,151,177]" captionStart="FIGURE 12" captionStartId="35.[151,250,1071,1095]" captionTargetBox="[165,1420,187,1041]" captionTargetId="figure-15@35.[158,1429,181,1048]" captionTargetPageId="35" captionText="FIGURE 12. Female genitalia of Siberian and Far Eastern Hemistola zimmermanni. A, B—samples from Buryatia; C—sample from Zabaikalsky Kray; D—sample from Primorsky Kray." figureDoi="http://doi.org/10.5281/zenodo.7965102" httpUri="https://zenodo.org/record/7965102/files/figure.png" pageId="38" pageNumber="39">Figs. 12A and 12B</figureCitation>
). Nevertheless, when studying the serial material, these differences form a series with a gradual transition of shape of lamella postvaginalis, therefore, in my opinion, they are not diagnostic. At the same time, the female genitalia of 
<taxonomicName id="4C5B4D0C6937FF83FE0745F5FD8A0CED" authorityName=": Berlov &amp; Berlov" authorityYear="2006" box="[420,628,223,249]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="chrysoprasaria">
<emphasis id="B92FEA9D6937FF83FE0745F5FD8A0CED" box="[420,628,223,249]" italics="true" pageId="38" pageNumber="39">H. chrysoprasaria</emphasis>
</taxonomicName>
reliably differ from those presented in 
<figureCitation id="13602A0A6937FF83FB9D45F5FB620CED" box="[1086,1180,223,249]" captionStart="FIGURE 11" captionStartId="34.[151,250,1867,1891]" captionTargetBox="[162,1427,186,1838]" captionTargetId="figure-25@34.[154,1433,181,1844]" captionTargetPageId="34" captionText="FIGURE 11. Female genitalia of Siberian Hemistola zimmermanni. A–D—samples from Irkutskaya Oblast; E–H—samples from Buryatia." figureDoi="http://doi.org/10.5281/zenodo.7965100" httpUri="https://zenodo.org/record/7965100/files/figure.png" pageId="38" pageNumber="39">Figs. 11</figureCitation>
, 
<figureCitation id="13602A0A6937FF83FB0845F5FB390CED" box="[1195,1223,223,249]" captionStart="FIGURE 12" captionStartId="35.[151,250,1071,1095]" captionTargetBox="[165,1420,187,1041]" captionTargetId="figure-15@35.[158,1429,181,1048]" captionTargetPageId="35" captionText="FIGURE 12. Female genitalia of Siberian and Far Eastern Hemistola zimmermanni. A, B—samples from Buryatia; C—sample from Zabaikalsky Kray; D—sample from Primorsky Kray." figureDoi="http://doi.org/10.5281/zenodo.7965102" httpUri="https://zenodo.org/record/7965102/files/figure.png" pageId="38" pageNumber="39">12</figureCitation>
by the absence of semicircular furrow on the lamella postvaginalis (
<figureCitation id="13602A0A6937FF83FD664429FCD40D09" box="[709,810,259,285]" captionStart="FIGURE 37" captionStartId="82.[151,250,1862,1886]" captionTargetBox="[153,1423,192,1832]" captionTargetId="figure-25@82.[152,1435,181,1838]" captionTargetPageId="82" captionText="FIGURE 37. Female genitalia of Siberian Geometrinae. A—Geometra papilionaria; B—Thetidia chlorophyllaria; C—T. smaragdaria; D—Comibaena amoenaria; E—Hemistola chrysoprasaria; F, G—H. zimmermanni; H—Hemithea aestivaria. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965188" httpUri="https://zenodo.org/record/7965188/files/figure.png" pageId="38" pageNumber="39">Fig. 37E</figureCitation>
).
</paragraph>
<paragraph id="8BE4368F6937FF83FF64440DFCDD0ED9" blockId="38.[151,1437,151,825]" pageId="38" pageNumber="39">
It was previously shown that the moths corresponding to the features of 
<taxonomicName id="4C5B4D0C6937FF83FBBF440DFB240D54" authorityName="Hedemann" authorityYear="1879" box="[1052,1242,295,320]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6937FF83FBBF440DFB240D54" box="[1052,1242,295,320]" italics="true" pageId="38" pageNumber="39">H. zimmermanni</emphasis>
</taxonomicName>
from Irkutskaya Oblast, Buryatia and Zabaikalsky Kray were genetically homogeneous (
<bibRefCitation id="EFCA4B7E6937FF83FC1E4461FB020D71" author="Makhov, I. A. &amp; Lukhtanov, V. A." box="[957,1276,331,357]" pageId="38" pageNumber="39" pagination="1154 - 1172" refId="ref72922" refString="Makhov, I. A. &amp; Lukhtanov, V. A. (2021) Geometrid moths (Lepidoptera, Geometridae) of the Baikal region: additions to the species list and results of DNA barcoding. Entomological Review, 101 (8), 1154 - 1172. https: // doi. org / 10.1134 / S 001387382108011 X" type="journal article" year="2021">Makhov &amp; Lukhtanov 2021</bibRefCitation>
). Later, I have obtained three 
<emphasis id="B92FEA9D6937FF83FEE04445FE8D0D9D" box="[323,371,367,393]" italics="true" pageId="38" pageNumber="39">COI</emphasis>
sequences of the moths collected in the 
<typeStatus id="54E0882D6937FF83FCEC4445FC810D9D" box="[847,895,367,393]" pageId="38" pageNumber="39">type</typeStatus>
locality of 
<taxonomicName id="4C5B4D0C6937FF83FBAB4445FB580D9D" authority="Djakonov, 1926" box="[1032,1190,367,393]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6937FF83FBAB4445FB580D9D" box="[1032,1190,367,393]" italics="true" pageId="38" pageNumber="39">H. intermedia</emphasis>
</taxonomicName>
(Krasnoyarsky Kray, see “Material examined”). Analysis of these sequences shows that they are almost identical to the sequences we analysed earlier (moths from Irkutskaya Oblast, Buryatia and Zabaikalsky Kray). New genetic data from moths in Minusinsky District of Krasnoyarsky Kray and from Ust’-Abakansky District of Khakassia was included in the 
<emphasis id="B92FEA9D6937FF83FACF44F1FA620DE1" box="[1388,1436,475,501]" italics="true" pageId="38" pageNumber="39">COI</emphasis>
dataset (the data obtained by us and the data available in the GenBank and BoLD) and used to build a Bayesian tree (
<figureCitation id="13602A0A6937FF83FF724709FED80E29" box="[209,294,547,573]" captionStart="FIGURE 13" captionStartId="36.[151,250,1892,1916]" captionTargetBox="[155,1432,185,1863]" captionTargetId="figure-25@36.[151,1436,181,1868]" captionTargetPageId="36" captionText="FIGURE 13. Bayesian tree of Hemistola based on analysis of COI gene sequences. The numbers above the branches are Bayesian posterior probabilities. Support values less than 0.7 are not shown. The sequences of specimens from type locality of Hemistola intermedia and adjacent area (Khakassia) are highlighted in red." figureDoi="http://doi.org/10.5281/zenodo.7965104" httpUri="https://zenodo.org/record/7965104/files/figure.png" pageId="38" pageNumber="39">Fig. 13</figureCitation>
). The mentioned specimens Hml1, Hml2 and Hml3 (highlighted in red) fall into the same clade with the samples from the Siberian regions and from 
<collectingCountry id="F34C761F6937FF83FD12476DFD0A0E75" box="[689,756,583,609]" name="China" pageId="38" pageNumber="39">China</collectingCountry>
identified as 
<taxonomicName id="4C5B4D0C6937FF83FC28476DFBBB0E74" authorityName="Hedemann" authorityYear="1879" box="[907,1093,583,608]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6937FF83FC28476DFBBB0E74" box="[907,1093,583,608]" italics="true" pageId="38" pageNumber="39">H. zimmermanni</emphasis>
</taxonomicName>
. The samples Hml1 and Hml2 (Republic of Khakassia) are placed in the same subclade together with samples from Buryatia, Zabaikalsky Kray and Irkutskaya Oblast. Thus, DNA barcoding of “ 
<taxonomicName id="4C5B4D0C6937FF83FD6047A5FCA00EBD" authority="Djakonov, 1926" box="[707,862,655,681]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6937FF83FD6047A5FCA00EBD" box="[707,862,655,681]" italics="true" pageId="38" pageNumber="39">H. intermedia</emphasis>
</taxonomicName>
” from the 
<typeStatus id="54E0882D6937FF83FC7B47A5FBF60EBD" box="[984,1032,655,681]" pageId="38" pageNumber="39">type</typeStatus>
locality and adjacent areas does not confirm the assumption of the independence of this taxon.
</paragraph>
<paragraph id="8BE4368F6937FF83FF6447FDFC600F2D" blockId="38.[151,1437,151,825]" pageId="38" pageNumber="39">
Taking into account the morphological aspects considered above and the lack of 
<emphasis id="B92FEA9D6937FF83FBFD47FDFB700EE5" box="[1118,1166,727,753]" italics="true" pageId="38" pageNumber="39">COI</emphasis>
differentiation between “ 
<taxonomicName id="4C5B4D0C6937FF83FF0747D1FEC00F01" authority="Djakonov, 1926" box="[164,318,763,789]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6937FF83FF0747D1FEC00F01" box="[164,318,763,789]" italics="true" pageId="38" pageNumber="39">H. intermedia</emphasis>
</taxonomicName>
” and 
<taxonomicName id="4C5B4D0C6937FF83FEDC47D1FDC60F00" authorityName="Hedemann" authorityYear="1879" box="[383,568,763,788]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6937FF83FEDC47D1FDC60F00" box="[383,568,763,788]" italics="true" pageId="38" pageNumber="39">H. zimmermanni</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5B4D0C6937FF83FDE547D1FD1E0F01" authority="Djakonov, 1926" box="[582,736,763,789]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B92FEA9D6937FF83FDE547D1FD1E0F01" box="[582,736,763,789]" italics="true" pageId="38" pageNumber="39">H. intermedia</emphasis>
</taxonomicName>
is considered here a synonym of 
<taxonomicName id="4C5B4D0C6937FF83FBF147D1FAF50F00" authorityName="Hedemann" authorityYear="1879" box="[1106,1291,763,788]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6937FF83FBF147D1FAF50F00" box="[1106,1291,763,788]" italics="true" pageId="38" pageNumber="39">H. zimmermanni</emphasis>
</taxonomicName>
<emphasis id="B92FEA9D6937FF83FAB247D1FAA00F01" bold="true" box="[1297,1374,763,789]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="A21C57E66937FF83FAB247D1FAA00F01" box="[1297,1374,763,789]" pageId="38" pageNumber="39">syn. n.</taxonomicNameLabel>
</emphasis>
, only being a small form of 
<taxonomicName id="4C5B4D0C6937FF83FE2C4635FDB70F2C" authorityName="Hedemann" authorityYear="1879" box="[399,585,799,824]" class="Insecta" family="Geometridae" genus="Hemistola" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B92FEA9D6937FF83FE2C4635FDB70F2C" box="[399,585,799,824]" italics="true" pageId="38" pageNumber="39">H. zimmermanni</emphasis>
</taxonomicName>
in the western part of its area.
</paragraph>
</subSubSection>
</treatment>
</document>