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<document id="5A84C66F7189B9DD82CD1F46DFCD04D0" ID-DOI="10.15553/c2012v672a12" ID-ISSN="2235-3658" ID-Zenodo-Dep="5790002" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637611591098" checkinUser="carolina" docAuthor="Callmander, Martin W., Lowry Ii, Porter P., Forest, Félix, Devey, Dion S., Beentje, Henk &amp; Buerki, Sven" docDate="2012" docId="03F20C491D1A2B435D5DFEEA75ED54E1" docLanguage="en" docName="Candollea.67.2.323-345.pdf.imf" docOrigin="Candollea 67 (2)" docStyle="DocumentStyle:FB35ED2BCB231A5F446DBDA23BDBADFC.2:Candollea.2012-2014.journal_article" docStyleId="FB35ED2BCB231A5F446DBDA23BDBADFC" docStyleName="Candollea.2012-2014.journal_article" docStyleVersion="2" docTitle="Benstonea affinis Callm. &amp; Buerki, comb.nova." docType="treatment" docVersion="8" lastPageNumber="330" masterDocId="FFCB74311D1D2B445D3AFFF3771C5134" masterDocTitle="Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species" masterLastPageNumber="345" masterPageNumber="323" pageNumber="330" updateTime="1699425018902" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="7ABBAE2004869332192DF9056D2B965D">Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species</mods:title>
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<mods:namePart id="BAAE936CDF107AFBD5542CD459C11AC8">Callmander, Martin W.</mods:namePart>
<mods:affiliation id="CFF52DD560F3100C4DFA7D709CC8948D">Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Conservatoire et Jardin botaniques de la Ville de Genève, CP 60, CH- 1292 Chambésy, Switzerland.</mods:affiliation>
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<mods:namePart id="CF502894D5EEAF48FD696D4E298EEA9B">Lowry Ii, Porter P.</mods:namePart>
<mods:affiliation id="5B48FDB48FF26BE1BA114B7E9D399EC2">Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Département Systématique et Evolution (UMR 7205), Muséum National d’Histoire Naturelle, case postale 39, rue Cuvier 57, 75231 Paris (Cedex) 05, France.</mods:affiliation>
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<mods:namePart id="57A4514B36544C978759817D2717335A">Forest, Félix</mods:namePart>
<mods:affiliation id="FCE550DEB43E1DB7A4489ACD44448AD2">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:namePart id="2B30ED5711D7A9E097EC9C8082466FA2">Devey, Dion S.</mods:namePart>
<mods:affiliation id="502FDB24233BC23DC72D3A9AA3BF8CC0">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:namePart id="AAED1C63D0E19B02B5C54639811A29B0">Beentje, Henk</mods:namePart>
<mods:affiliation id="1910DA042CE399911ECE36BFD4F9FB40">Herbarium, Library, Art &amp; Archives, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:namePart id="894A02A33B801C587EE476B16133D82F">Buerki, Sven</mods:namePart>
<mods:affiliation id="85361678CD940C0445BCFB4802814245">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:title id="0EA669D33E575D893A28F771ECA742AA">Candollea</mods:title>
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<mods:date id="84C03A110F32B634A3DCA2B3F68B5A79">2012</mods:date>
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<mods:number id="8445C3A765131CDD92668A95872D11FB">2</mods:number>
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<mods:identifier id="60EC5B16AA59D0B51CD756A7A926C2DD" type="ISSN">2235-3658</mods:identifier>
<mods:identifier id="122AD621BEFF200447C453FB866E2532" type="Zenodo-Dep">5790002</mods:identifier>
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<treatment id="03F20C491D1A2B435D5DFEEA75ED54E1" ID-DOI="http://doi.org/10.5281/zenodo.5796815" ID-Zenodo-Dep="5796815" LSID="urn:lsid:plazi:treatment:03F20C491D1A2B435D5DFEEA75ED54E1" httpUri="http://treatment.plazi.org/id/03F20C491D1A2B435D5DFEEA75ED54E1" lastPageNumber="330" pageId="7" pageNumber="330">
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<heading id="D0AC0A331D1A2B435D5DFEEA75F65005" box="[103,746,281,305]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="330" reason="3">
2.
<taxonomicName id="4C5BC6DC1D1A2B435DB6FEEA75405005" ID-CoL="LHHD" authority="(Kurz) Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" baseAuthorityName="Kurz" box="[140,604,281,305]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="species" species="affinis" status="comb.nova.">
<emphasis id="B92F614D1D1A2B435DB6FEEA76535005" bold="true" box="[140,335,281,305]" italics="true" pageId="7" pageNumber="330">Benstonea affinis</emphasis>
(Kurz) Callm. &amp; Buerki
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,
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<paragraph id="8BE4BD5F1D1A2B435DB6FEB37583506D" blockId="7.[103,753,281,759]" box="[140,671,320,346]" pageId="7" pageNumber="330">
<treatmentCitationGroup id="AB4B9A711D1A2B435DB6FEB37583506D" box="[140,671,320,346]" pageId="7" pageNumber="330">
≡
<treatmentCitation id="0AFA9B4E1D1A2B435D88FEB27587506D" author="Kurz" box="[178,667,321,345]" journal="J. Bot." page="101" pageId="7" pageNumber="330" score="11.733904942993295" volume="5" year="1867">
<taxonomicName id="4C5BC6DC1D1A2B435D88FEB276BE506D" ID-CoL="75M3S" authority="Kurz" authorityName="Kurz" box="[178,418,321,345]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="B92F614D1D1A2B435D88FEB2767B506D" box="[178,359,321,345]" italics="true" pageId="7" pageNumber="330">Pandanus affinis</emphasis>
Kurz
</taxonomicName>
in J. Bot. 5: 101. 1867
</treatmentCitation>
.
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</paragraph>
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<materialsCitation id="3B33B7021D1A2B435DB6FE9A76B55096" collectionCode="BO" collectorName="Kurz" country="Indonesia" location="BangkaIsl" pageId="7" pageNumber="330" specimenCount="1" stateProvince="BangkaIsl" typeStatus="lectotype">
<emphasis id="B92F614D1D1A2B435DB6FE9A76C550B5" bold="true" box="[140,473,361,385]" pageId="7" pageNumber="330">
Typus:
<collectingCountry id="F34CFDCF1D1A2B435DE1FE9A764E50B4" box="[219,338,361,385]" pageId="7" pageNumber="330">INDONESIA</collectingCountry>
.
<collectingRegion id="499F73BD1D1A2B435C6DFE9A76D350B5" box="[343,463,361,385]" pageId="7" pageNumber="330">BangkaIsl</collectingRegion>
.:
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<emphasis id="B92F614D1D1A2B435CD1FE9A750250B5" box="[491,542,361,385]" italics="true" pageId="7" pageNumber="330">
<collectorName id="26AED8891D1A2B435CD1FE9A750250B5" box="[491,542,361,385]" pageId="7" pageNumber="330">Kurz</collectorName>
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(
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:
<collectionCode id="ED4A259A1D1A2B435F4FFE9A758750B5" box="[629,667,361,385]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="7" pageNumber="330" type="Herbarium">BO</collectionCode>
) (designated by
<bibRefCitation id="EFCAC0AE1D1A2B435DD6FE7976805096" author="STONE" box="[236,412,394,418]" firstAuthor="STONE" page="13" pageId="7" pageNumber="330" pagination="1 - 74" refId="ref16436" refString="STONE, B. C. (1978). Revisio Pandanacearum, Part I. Pandanus subgenera Coronata and Acrostigma. Flora Malesiana precursores. Fed. Mus. J. 23: 1 - 74." type="journal article" year="1978">STONE, 1978: 13</bibRefCitation>
).
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<subSubSection id="C341EED41D1A2B435DB6FE42768950DF" pageId="7" pageNumber="330" type="reference_group">
<paragraph id="8BE4BD5F1D1A2B435DB6FE42768950DF" blockId="7.[103,753,281,759]" pageId="7" pageNumber="330">
<treatmentCitationGroup id="AB4B9A711D1A2B435DB6FE42768950DF" pageId="7" pageNumber="330">
=
<treatmentCitation id="0AFA9B4E1D1A2B435D88FE41768950DF" author="Warb" page="50" pageId="7" pageNumber="330" year="1904">
<taxonomicName id="4C5BC6DC1D1A2B435D88FE4176C150FE" ID-CoL="6TKRJ" authority="Warb." authorityName="Warb." box="[178,477,434,458]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="species" species="merrillii">
<emphasis id="B92F614D1D1A2B435D88FE41769750FE" box="[178,395,434,458]" italics="true" pageId="7" pageNumber="330">Pandanus merrillii</emphasis>
Warb.
</taxonomicName>
in
<collectorName id="26AED8891D1A2B435F37FE41757F50FE" box="[525,611,434,458]" pageId="7" pageNumber="330">Perkins</collectorName>
,
<collectorName id="26AED8891D1A2B435F4EFE41761550DF" pageId="7" pageNumber="330">Fragm. Fl. Philipp.</collectorName>
1: 50. 1904.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="C341EED41D1A2B435CA7FE20759E5379" pageId="7" pageNumber="330" type="materials_examined">
<paragraph id="8BE4BD5F1D1A2B435CA7FE20759E5379" blockId="7.[103,753,281,759]" pageId="7" pageNumber="330">
<materialsCitation id="3B33B7021D1A2B435CA7FE2076CC5318" collectingDate="1903-02" collectionCode="B" collectorName="Merrill" country="Philippines" location="San Antognio Bay" pageId="7" pageNumber="330" specimenCode="B100279959" specimenCount="1" stateProvince="Palawan Isl." typeStatus="holotype">
<emphasis id="B92F614D1D1A2B435CA7FE2077C2533F" bold="true" pageId="7" pageNumber="330">
<typeStatus id="54E003FD1D1A2B435CA7FE2076F550DF" box="[413,489,467,491]" pageId="7" pageNumber="330">Typus</typeStatus>
:
<collectingCountry id="F34CFDCF1D1A2B435CD4FE20756250DF" box="[494,638,467,491]" pageId="7" pageNumber="330">PHILIPPINES</collectingCountry>
.
<collectingRegion id="499F73BD1D1A2B435FB9FE2077C4533F" pageId="7" pageNumber="330">Palawan Isl.</collectingRegion>
:
</emphasis>
<location id="8E84EB841D1A2B435DDEFE0076B6533F" LSID="urn:lsid:plazi:treatment:03F20C491D1A2B435D5DFEEA75ED54E1:8E84EB841D1A2B435DDEFE0076B6533F" box="[228,426,499,523]" country="Philippines" name="San Antognio Bay" pageId="7" pageNumber="330" stateProvince="Palawan Isl.">San Antognio Bay</location>
,
<date id="FFE59B9F1D1A2B435C8EFE007518533F" box="[436,516,499,523]" pageId="7" pageNumber="330" value="1903-02">
<collectingDate id="EFA162771D1A2B435C8EFE007518533F" box="[436,516,499,523]" pageId="7" pageNumber="330" value="1903-02">II.1903</collectingDate>
</date>
,
<emphasis id="B92F614D1D1A2B435F35FE007596533F" box="[527,650,499,523]" italics="true" pageId="7" pageNumber="330">
<collectorName id="26AED8891D1A2B435F35FE007540533F" box="[527,604,499,523]" pageId="7" pageNumber="330">Merrill</collectorName>
840
</emphasis>
(
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:
<collectionCode id="ED4A259A1D1A2B435FE5FE0075ED533F" box="[735,753,499,523]" pageId="7" pageNumber="330">B</collectionCode>
[
<specimenCode id="DBFD15241D1A2B435D82FDE7765B5318" box="[184,327,532,556]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="7" pageNumber="330" type="Herbarium">B100279959</specimenCode>
] image seen
</materialsCitation>
;
<materialsCitation id="3B33B7021D1A2B435CE6FDE7759E5379" collectingDate="1903-02" collectionCode="FI, NY, US" collectorName="Merrill" country="Philippines" location="San Antognio Bay" pageId="7" pageNumber="330" specimenCode="FI003946, US00086602" specimenCount="3" stateProvince="Palawan Isl." typeStatus="isotype">
<typeStatus id="54E003FD1D1A2B435CE6FDE775145318" box="[476,520,532,556]" pageId="7" pageNumber="330" type="isotype">iso-</typeStatus>
:
<collectionCode id="ED4A259A1D1A2B435F2BFDE775355318" box="[529,553,532,556]" pageId="7" pageNumber="330">FI</collectionCode>
[
<specimenCode id="DBFD15241D1A2B435F0CFDE775BF5318" box="[566,675,532,556]" pageId="7" pageNumber="330">FI003946</specimenCode>
] image seen,
<collectionCode id="ED4A259A1D1A2B435DCBFDC676055379" box="[241,281,565,589]" pageId="7" pageNumber="330">NY</collectionCode>
,
<collectionCode id="ED4A259A1D1A2B435C1EFDC6765A5379" box="[292,326,565,589]" pageId="7" pageNumber="330">US</collectionCode>
[
<specimenCode id="DBFD15241D1A2B435C69FDC676F65379" box="[339,490,565,589]" pageId="7" pageNumber="330">US00086602</specimenCode>
] image seen),
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</paragraph>
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<paragraph id="8BE4BD5F1D1A2B435FB6FDC675F65379" blockId="7.[103,753,281,759]" box="[652,746,565,589]" pageId="7" pageNumber="330">
<emphasis id="B92F614D1D1A2B435FB6FDC675F65379" bold="true" box="[652,746,565,589]" pageId="7" pageNumber="330">
<taxonomicNameLabel id="A21CDC361D1A2B435FB6FDC675F65379" box="[652,746,565,589]" pageId="7" pageNumber="330">syn.nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D1A2B435DB6FDAF76BD53A1" pageId="7" pageNumber="330" type="reference_group">
<paragraph id="8BE4BD5F1D1A2B435DB6FDAF76BD53A1" blockId="7.[103,753,281,759]" pageId="7" pageNumber="330">
<treatmentCitationGroup id="AB4B9A711D1A2B435DB6FDAF768253A1" pageId="7" pageNumber="330">
=
<treatmentCitation id="0AFA9B4E1D1A2B435D88FDAE768253A1" author="Martelli" journal="Bull. Soc. Bot. Ital." page="303" pageId="7" pageNumber="330" score="23.254043343992382" volume="1904" year="1904">
<taxonomicName id="4C5BC6DC1D1A2B435D88FDAE753E5341" ID-CoL="4CB6Y" authority="Martelli" authorityName="Martelli" box="[178,546,605,629]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="species" species="sarawakensis">
<emphasis id="B92F614D1D1A2B435D88FDAE76DC5341" box="[178,448,605,629]" italics="true" pageId="7" pageNumber="330">Pandanus sarawakensis</emphasis>
Martelli
</taxonomicName>
in Bull. Soc. Bot. Ital. 1904: 303. 1904
</treatmentCitation>
</treatmentCitationGroup>
.
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D1A2B435C93FD8E760C53C3" pageId="7" pageNumber="330" type="materials_examined">
<paragraph id="8BE4BD5F1D1A2B435C93FD8E760C53C3" blockId="7.[103,753,281,759]" pageId="7" pageNumber="330">
<materialsCitation id="3B33B7021D1A2B435C93FD8E760C53C3" collectingDate="1865-08" collectionCode="FI-B" collectorName="Beccari" country="Malaysia" location="Kuching" pageId="7" pageNumber="330" specimenCode="FI003947" specimenCount="1" stateProvince="Sarawak" typeStatus="lectotype">
<emphasis id="B92F614D1D1A2B435C93FD8E753B53A1" bold="true" box="[425,551,637,661]" pageId="7" pageNumber="330">
<typeStatus id="54E003FD1D1A2B435C93FD8E753B53A1" box="[425,551,637,661]" pageId="7" pageNumber="330" type="lectotype">Lectotypus</typeStatus>
</emphasis>
(designated here):
<emphasis id="B92F614D1D1A2B435D88FD6D751B5382" bold="true" box="[178,519,670,694]" pageId="7" pageNumber="330">
<collectingCountry id="F34CFDCF1D1A2B435D88FD6D763E5381" box="[178,290,670,694]" name="Malaysia" pageId="7" pageNumber="330">MALAYSIA</collectingCountry>
.
<collectingRegion id="499F73BD1D1A2B435C1DFD6D76935382" box="[295,399,670,694]" pageId="7" pageNumber="330">Sarawak</collectingRegion>
(
<collectingRegion id="499F73BD1D1A2B435CA5FD6D76E65382" box="[415,506,670,694]" pageId="7" pageNumber="330">Borneo</collectingRegion>
):
</emphasis>
<location id="8E84EB841D1A2B435F2CFD6D75695382" LSID="urn:lsid:plazi:treatment:03F20C491D1A2B435D5DFEEA75ED54E1:8E84EB841D1A2B435F2CFD6D75695382" box="[534,629,670,694]" country="Malaysia" municipality="Beccari" name="Kuching" pageId="7" pageNumber="330" stateProvince="Sarawak">Kuching</location>
,
<date id="FFE59B9F1D1A2B435FBAFD6D75F15382" box="[640,749,670,694]" pageId="7" pageNumber="330" value="1865-08">
<collectingDate id="EFA162771D1A2B435FBAFD6D75F15382" box="[640,749,670,694]" pageId="7" pageNumber="330" value="1865-08">VIII.1865</collectingDate>
</date>
,
<emphasis id="B92F614D1D1A2B435D88FD4C762F53E3" box="[178,307,703,727]" italics="true" pageId="7" pageNumber="330">
<collectorName id="26AED8891D1A2B435D88FD4C761953E3" box="[178,261,703,727]" pageId="7" pageNumber="330">Beccari</collectorName>
350
</emphasis>
(
<collectionCode id="ED4A259A1D1A2B435C7BFD4C766953E3" box="[321,373,703,727]" pageId="7" pageNumber="330">FI-B</collectionCode>
[
<specimenCode id="DBFD15241D1A2B435CB8FD4C76EE53E3" box="[386,498,703,727]" pageId="7" pageNumber="330">FI003947</specimenCode>
: ErbarioBeccari 11902, carpo.]!),
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D1A2B435C2DFD2C766853C3" box="[279,372,735,759]" pageId="7" pageNumber="330" type="multiple">
<paragraph id="8BE4BD5F1D1A2B435C2DFD2C766853C3" blockId="7.[103,753,281,759]" box="[279,372,735,759]" pageId="7" pageNumber="330">
<emphasis id="B92F614D1D1A2B435C2DFD2C766853C3" bold="true" box="[279,372,735,759]" pageId="7" pageNumber="330">
<taxonomicNameLabel id="A21CDC361D1A2B435C2DFD2C766853C3" box="[279,372,735,759]" pageId="7" pageNumber="330">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D1A2B435DB6FCFB76905296" pageId="7" pageNumber="330" type="distribution">
<paragraph id="8BE4BD5F1D1A2B435DB6FCFB76905296" blockId="7.[103,753,776,930]" pageId="7" pageNumber="330">
<emphasis id="B92F614D1D1A2B435DB6FCFB76865214" box="[140,410,776,800]" italics="true" pageId="7" pageNumber="330">Distribution and ecology</emphasis>
. –
<taxonomicName id="4C5BC6DC1D1A2B435C86FCFB756A5214" box="[444,630,776,800]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="B92F614D1D1A2B435C86FCFB756A5214" box="[444,630,776,800]" italics="true" pageId="7" pageNumber="330">Benstonea affinis</emphasis>
</taxonomicName>
is found in peat swamps and gallery forest/river edges from sea level to c.
<specimenCount id="9D5D76D61D1A2B435DBAFCBA77D05255" box="[128,204,841,865]" pageId="7" pageNumber="330" type="generic">
<quantity id="4CA310BA1D1A2B435DBAFCBA77D45255" box="[128,200,841,865]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="330" unit="m" value="500.0">500 m</quantity>
.
</specimenCount>
It is a widespread species, distributed from Peninsular
<collectingCountry id="F34CFDCF1D1A2B435D98FC99761852B6" box="[162,260,874,898]" name="Malaysia" pageId="7" pageNumber="330">Malaysia</collectingCountry>
to the
<collectingCountry id="F34CFDCF1D1A2B435C75FC9976D752B6" box="[335,459,874,898]" name="Philippines" pageId="7" pageNumber="330">Philippines</collectingCountry>
, including
<collectingCountry id="F34CFDCF1D1A2B435F7BFC99758352B6" box="[577,671,874,898]" name="Vietnam" pageId="7" pageNumber="330">Vietnam</collectingCountry>
, Sumatra,
<collectingCountry id="F34CFDCF1D1A2B435DAAFC7977E25296" box="[144,254,906,930]" name="Singapore" pageId="7" pageNumber="330">Singapore</collectingCountry>
, and Borneo.
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D1A2B435DB6FC4175ED54E1" pageId="7" pageNumber="330" type="discussion">
<paragraph id="8BE4BD5F1D1A2B435DB6FC4175ED54E1" blockId="7.[103,755,946,1493]" pageId="7" pageNumber="330">
<emphasis id="B92F614D1D1A2B435DB6FC41763452FE" box="[140,296,946,970]" italics="true" pageId="7" pageNumber="330">Observations.</emphasis>
–This species is characterized by its 3 to 9 ovoid fruit heads crowded at the apex of an erect peduncle (
<figureCitation id="1360A1DA1D1A2B435D55FC0777DB5538" box="[111,199,1012,1036]" captionStart="Fig" captionStartId="8.[118,143,1873,1893]" captionTargetBox="[118,1494,265,1835]" captionTargetPageId="8" captionText="Fig. 4. – Infructescences and details of stigmas of species of Benstonea Callm. &amp; Buerki. A. Benstonea affinis (Kurz) Callm. &amp; Buerki; B. Benstonea gibbsiana (Martelli) Callm. &amp; Buerki; C. Benstonea ornata (Solms) Callm. &amp; Buerki; D. Benstonea pachyphylla (Merr.) Callm. &amp; Buerki. [Photos: M. W. Callmander]" figureDoi="http://doi.org/10.5281/zenodo.5799762" httpUri="https://zenodo.org/record/5799762/files/figure.png" pageId="7" pageNumber="330">Fig. 4A</figureCitation>
).
<bibRefCitation id="EFCAC0AE1D1A2B435DE0FC0776765538" author="STONE" box="[218,362,1012,1036]" firstAuthor="STONE" pageId="7" pageNumber="330" pagination="127 - 136" refId="ref16252" refString="STONE, B. C. (1972). Studies in the Malesian Pandanaceae X. Four new Malesian species of Pandanus and observations on P. sarawakensis. Fed. Mus. J. 14: 127 - 136." type="journal article" year="1972">STONE (1972</bibRefCitation>
,
<bibRefCitation id="EFCAC0AE1D1A2B435C4CFC0776B35538" author="STONE" box="[374,431,1012,1036]" firstAuthor="STONE" pageId="7" pageNumber="330" pagination="1 - 74" refId="ref16436" refString="STONE, B. C. (1978). Revisio Pandanacearum, Part I. Pandanus subgenera Coronata and Acrostigma. Flora Malesiana precursores. Fed. Mus. J. 23: 1 - 74." type="journal article" year="1978">1978</bibRefCitation>
,
<bibRefCitation id="EFCAC0AE1D1A2B435C80FC07751C5538" author="STONE" box="[442,512,1012,1036]" firstAuthor="STONE" pageId="7" pageNumber="330" pagination="205 - 212" refId="ref16580" refString="STONE, B. C. (1983 a). Some new and critical Pandanus species of subgenus Acrostigma, I. Supplement to Revisio Pandanacearum. Gard. Bull. Singapore 36: 205 - 212." type="journal article" year="1983" yearSuffix="a">1983a</bibRefCitation>
,
<bibRefCitation id="EFCAC0AE1D1A2B435F31FC07754D5538" author="STONE, B. C." box="[523,593,1012,1036]" pageId="7" pageNumber="330" pagination="35 - 84" refId="ref16757" refString="STONE, B. C. (1993). Studies in the Malesian Pandanaceae 21. The genus Pandanus in Borneo. Sandakania 2: 35 - 84." type="journal article" year="1993">1993a</bibRefCitation>
), mentionning some differences in dimensions of the fruit, recognized the two species placed in synonymy above, albeit with doubt. He wrote: “
<taxonomicName id="4C5BC6DC1D1A2B435D81FBA576A5555A" box="[187,441,1110,1134]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="genus">
<emphasis id="B92F614D1D1A2B435D81FBA5766B555A" box="[187,375,1110,1134]" italics="true" pageId="7" pageNumber="330">Pandanus merrilli</emphasis>
Warb
</taxonomicName>
. is so similar to
<taxonomicName id="4C5BC6DC1D1A2B435F64FBA575ED555A" authority="Kurz" authorityName="Kurz" box="[606,753,1110,1134]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="B92F614D1D1A2B435F64FBA575AB555A" box="[606,695,1110,1134]" italics="true" pageId="7" pageNumber="330">P. affinis</emphasis>
Kurz
</taxonomicName>
that, on the basis of herbarium material alone, discrimination is virtually impossible” (
<bibRefCitation id="EFCAC0AE1D1A2B435CB2FB647575559B" author="STONE" box="[392,617,1175,1199]" firstAuthor="STONE" journalOrPublisher="Gard. Bull. Singapore" page="208" pageId="7" pageNumber="330" pagination="205 - 212" part="36" refId="ref16580" refString="STONE, B. C. (1983 a). Some new and critical Pandanus species of subgenus Acrostigma, I. Supplement to Revisio Pandanacearum. Gard. Bull. Singapore 36: 205 - 212." title="Some new and critical Pandanus species of subgenus Acrostigma, I. Supplement to Revisio Pandanacearum" type="journal article" year="1983" yearSuffix="a">STONE, 1983a: 208</bibRefCitation>
).
<taxonomicName id="4C5BC6DC1D1A2B435FB8FB6477EA55E4" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="species" species="sarawakensis">
<emphasis id="B92F614D1D1A2B435FB8FB6477EA55E4" italics="true" pageId="7" pageNumber="330">Pandanus sarawakensis</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFCAC0AE1D1A2B435C82FB4B757355E4" author="MARTELLI" box="[440,623,1208,1232]" firstAuthor="MARTELLI" pageId="7" pageNumber="330" pagination="298 - 304" refId="ref15762" refString="MARTELLI, U. (1904). Pandani asiatici nuovi. Bull. Soc. Bot. Ital. 1904: 298 - 304." type="journal article" year="1904">MARTELLI (1904)</bibRefCitation>
on the basis of
<specimenCount id="9D5D76D61D1A2B435DB9FB2B760C55C4" box="[131,272,1240,1264]" pageId="7" pageNumber="330" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
:
<emphasis id="B92F614D1D1A2B435C23FB2B768A55C4" box="[281,406,1240,1264]" italics="true" pageId="7" pageNumber="330">Beccari 350</emphasis>
and
<emphasis id="B92F614D1D1A2B435CF2FB2B76E055C4" box="[456,508,1240,1264]" italics="true" pageId="7" pageNumber="330">1895</emphasis>
. The first of these, misprinted as
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<emphasis id="B92F614D1D1A2B435DEEFB0A77E05425" box="[212,252,1273,1297]" italics="true" pageId="7" pageNumber="330">550</emphasis>
in
</quantity>
<bibRefCitation id="EFCAC0AE1D1A2B435C24FB0A75125425" author="MARTELLI" box="[286,526,1273,1297]" firstAuthor="MARTELLI" page="303" pageId="7" pageNumber="330" pagination="298 - 304" refId="ref15762" refString="MARTELLI, U. (1904). Pandani asiatici nuovi. Bull. Soc. Bot. Ital. 1904: 298 - 304." type="journal article" year="1904">MARTELLI (1904: 303)</bibRefCitation>
, is designated here as the
<typeStatus id="54E003FD1D1A2B435DB5FAEA77EE5405" box="[143,242,1305,1329]" pageId="7" pageNumber="330" type="lectotype">lectotype</typeStatus>
because it comprises a sheet with a leaf and an infructescence deposited in the Beccari herbarium and a spirit collection of one syncarp in the associated carpological collection. Further taxonomic work is clearly needed for a better understanding of
<taxonomicName id="4C5BC6DC1D1A2B435C26FA6F76CC5480" box="[284,464,1436,1460]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="B92F614D1D1A2B435C26FA6F76CC5480" box="[284,464,1436,1460]" italics="true" pageId="7" pageNumber="330">Benstonea affinis</emphasis>
</taxonomicName>
with respect to its relatives, especially
<taxonomicName id="4C5BC6DC1D1A2B435DEEFA4E757054E1" authority="(Warb.) Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" baseAuthorityName="Warb." box="[212,620,1469,1493]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="330" phylum="Tracheophyta" rank="species" species="ellipsoidea">
<emphasis id="B92F614D1D1A2B435DEEFA4E767E54E1" box="[212,354,1469,1493]" italics="true" pageId="7" pageNumber="330">B. ellipsoidea</emphasis>
(Warb.) Callm. &amp; Buerki
</taxonomicName>
(see below).
</paragraph>
</subSubSection>
</treatment>
</document>
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