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<mods:title id="3CB6D33B249DD575666238ECB83267BE">Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species</mods:title>
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<mods:namePart id="1C6D8A034D1CC83F795A9729ACE30BCB">Callmander, Martin W.</mods:namePart>
<mods:affiliation id="484D751BB326F6B85530C25CC72A7720">Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Conservatoire et Jardin botaniques de la Ville de Genève, CP 60, CH- 1292 Chambésy, Switzerland.</mods:affiliation>
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<mods:namePart id="EE9C265A5E077E177C42639157E57165">Lowry Ii, Porter P.</mods:namePart>
<mods:affiliation id="E0FC6A4FE1F6FA8CCC654F54922977C5">Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Département Systématique et Evolution (UMR 7205), Muséum National dHistoire Naturelle, case postale 39, rue Cuvier 57, 75231 Paris (Cedex) 05, France.</mods:affiliation>
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<mods:namePart id="3A952F5B87E07FFB7EC5A70AD8C097EC">Forest, Félix</mods:namePart>
<mods:affiliation id="60C2DDD9DF2A741348660B8163BB508E">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:namePart id="BE374B91F83621245A8911091879D82F">Devey, Dion S.</mods:namePart>
<mods:affiliation id="AC310D49C34D48E2E41A58C4FE7DC458">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:namePart id="76B5305BE8CBBA99D43B394EDDBDFD87">Beentje, Henk</mods:namePart>
<mods:affiliation id="63642AD57F98ADE45A288E822582067E">Herbarium, Library, Art &amp; Archives, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:namePart id="1505C5D97186959709A8AE58924BA5BA">Buerki, Sven</mods:namePart>
<mods:affiliation id="C5D8B35ADEAA2640326ED848E8F5902D">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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32.
<taxonomicName id="4C5BC6DC1D132B4A5E43FC35729A52EA" ID-CoL="LHJL" authority="(Martelli) Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" baseAuthorityName="Martelli" box="[889,1414,966,990]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="14" pageNumber="337" phylum="Tracheophyta" rank="species" species="pectinata" status="comb. nova.">
<emphasis id="B92F614D1D132B4A5E43FC35734552EA" bold="true" box="[889,1113,966,990]" italics="true" pageId="14" pageNumber="337">Benstonea pectinata</emphasis>
(Martelli) Callm. &amp; Buerki
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,
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<taxonomicNameLabel id="A21CDC361D132B4A58AAFC3574AA52CA" pageId="14" pageNumber="337">comb. nova.</taxonomicNameLabel>
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<treatmentCitation id="0AFA9B4E1D132B4A5EADFBFD73575572" author="Martelli" journal="Boll. Soc. Bot. Ital." page="304" pageId="14" pageNumber="337" score="43.846824794448175" volume="1904" year="1904">
<taxonomicName id="4C5BC6DC1D132B4A5EADFBFD73C05512" ID-CoL="4CB22" authority="Martelli" authorityName="Martelli" authorityYear="1904" box="[919,1244,1038,1062]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="14" pageNumber="337" phylum="Tracheophyta" rank="species" species="pectinatus">
<emphasis id="B92F614D1D132B4A5EADFBFD73615512" box="[919,1149,1038,1062]" italics="true" pageId="14" pageNumber="337">Pandanus pectinatus</emphasis>
Martelli
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in Boll. Soc. Bot. Ital. 1904: 304. 1904
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.
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<emphasis id="B92F614D1D132B4A5E43FBA6720D5559" bold="true" box="[889,1297,1109,1133]" pageId="14" pageNumber="337">
Typus:
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.
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(
<collectingRegion id="499F73BD1D132B4A5994FBA672185559" box="[1198,1284,1109,1133]" pageId="14" pageNumber="337">Borneo</collectingRegion>
):
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<collectorName id="26AED8891D132B4A582FFBA672785559" box="[1301,1380,1109,1133]" pageId="14" pageNumber="337">Beccari</collectorName>
s.n.
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(
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:
<collectionCode id="ED4A259A1D132B4A5E43FB86748C55B9" box="[889,912,1141,1165]" country="Italy" name="Natural History Museum" pageId="14" pageNumber="337" type="Herbarium">FI</collectionCode>
[
<specimenCode id="DBFD15241D132B4A5EA6FB86731855B9" box="[924,1028,1141,1165]" pageId="14" pageNumber="337">FI003941</specimenCode>
, carpo.] image seen).
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</paragraph>
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<paragraph id="8BE4BD5F1D132B4A5E43FB6E728855C1" blockId="14.[843,1494,1181,1468]" pageId="14" pageNumber="337">
<emphasis id="B92F614D1D132B4A5E43FB6E73945581" box="[889,1160,1181,1205]" italics="true" pageId="14" pageNumber="337">Distribution and ecology</emphasis>
.
<taxonomicName id="4C5BC6DC1D132B4A5991FB6E72945581" box="[1195,1416,1181,1205]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="14" pageNumber="337" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis id="B92F614D1D132B4A5991FB6E72945581" box="[1195,1416,1181,1205]" italics="true" pageId="14" pageNumber="337">Benstonea pectinata</emphasis>
</taxonomicName>
occurs in the montane rainforests (
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) of Borneo and has been collected twice in
<collectingRegion id="499F73BD1D132B4A597DFB2E73BC55C1" box="[1095,1184,1245,1269]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="337">Sumatra</collectingRegion>
(
<bibRefCitation id="EFCAC0AE1D132B4A5995FB2E725855C1" author="STONE" box="[1199,1348,1245,1269]" firstAuthor="STONE" pageId="14" pageNumber="337" pagination="69 - 74" refId="ref16327" refString="STONE, B. C. (1975 a). Studies in Malesian Pandanaceae XV. Two new species of Pandanus from Mt. Kinabalu, Sabah (Borneo) and notes on the Pandanaceae of Mt. Kinabalu. Malay. J. Sci. 3: 69 - 74." type="journal article" year="1975" yearSuffix="a">STONE, 1975a</bibRefCitation>
,
<bibRefCitation id="EFCAC0AE1D132B4A5875FB2E729455C1" author="STONE" box="[1359,1416,1245,1269]" firstAuthor="STONE" pageId="14" pageNumber="337" pagination="1 - 74" refId="ref16436" refString="STONE, B. C. (1978). Revisio Pandanacearum, Part I. Pandanus subgenera Coronata and Acrostigma. Flora Malesiana precursores. Fed. Mus. J. 23: 1 - 74." type="journal article" year="1978">1978</bibRefCitation>
).
</paragraph>
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<subSubSection id="C341EED41D132B4A5E43FAF773025488" pageId="14" pageNumber="337" type="discussion">
<paragraph id="8BE4BD5F1D132B4A5E43FAF773025488" blockId="14.[843,1494,1181,1468]" pageId="14" pageNumber="337">
<emphasis id="B92F614D1D132B4A5E43FAF7730C5428" box="[889,1040,1284,1308]" italics="true" pageId="14" pageNumber="337">Observations</emphasis>
. This species is strictly epiphytic and easily distinguished from other epiphytic members of the genus (e.g.
<taxonomicName id="4C5BC6DC1D132B4A5EF4FAB773585468" box="[974,1092,1348,1372]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="14" pageNumber="337" phylum="Tracheophyta" rank="species" species="alticola">
<emphasis id="B92F614D1D132B4A5EF4FAB773585468" box="[974,1092,1348,1372]" italics="true" pageId="14" pageNumber="337">B. alticola</emphasis>
</taxonomicName>
) by its lobed auricles at the base of the leaves.
<taxonomicName id="4C5BC6DC1D132B4A5EFCFA9773B85448" box="[966,1188,1380,1404]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="14" pageNumber="337" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis id="B92F614D1D132B4A5EFCFA9773B85448" box="[966,1188,1380,1404]" italics="true" pageId="14" pageNumber="337">Benstonea pectinata</emphasis>
</taxonomicName>
has often been collected on fallen trees in montane rainforests, but seems to be a strict epiphyte (
<figureCitation id="1360A1DA1D132B4A5E80FA57730C5488" box="[954,1040,1444,1468]" captionStart="Fig" captionStartId="11.[103,129,1870,1890]" captionTargetBox="[102,1479,279,1835]" captionTargetPageId="11" captionText="Fig. 5. Infructescences and details of stigmas of species of Benstonea Callm. &amp; Buerki. A. Benstonea parva (Ridl.) Callm. &amp; Buerki; B. Benstonea pectinata (Martelli) Callm. &amp; Buerki; C. Benstonea rupestris (. C. Stone) Callm. &amp; Buerki; D. Benstonea thomissophylla (. C. Stone) Callm. &amp; Buerki. [Photos: M. W. Callmander]" figureDoi="http://doi.org/10.5281/zenodo.5790012" httpUri="https://zenodo.org/record/5790012/files/figure.png" pageId="14" pageNumber="337">Fig. 5B</figureCitation>
).
</paragraph>
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