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<document id="8DB9CF1FC9D7DB2D0FEAB3D9F0B56604" ID-CLB-Dataset="32686" ID-DOI="10.11646/zootaxa.4291.2.2" ID-GBIF-Dataset="62931799-dbc5-475d-9028-b99ac00d1edb" ID-ISSN="1175-5326" ID-Zenodo-Dep="829531" ID-ZooBank="28772C8C-1E20-4A92-A9BD-1F5D016BD981" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500111698979" checkinUser="plazi" docAuthor="Schwarz, Christian J. &amp; Shcherbakov, Evgeny" docDate="2017" docId="03DC81241819A163FF6BF9A5FDD66776" docLanguage="en" docName="zootaxa.4291.2.2.pdf" docOrigin="Zootaxa 4291 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudohestiasula Schwarz &amp; Shcherbakov, 2017, gen. nov." docType="treatment" docUuid="65F02439-54F9-4810-9E86-EA892FF67DED" docUuidSource="ZooBank" docVersion="8" lastPageNumber="262" masterDocId="FFE5F95C180BA170FFFCFFB1FFEF6366" masterDocTitle="Revision of Hestiasulini Giglio-Tos, 1915 stat. rev. (Insecta: Mantodea: Hymenopodidae) of Borneo, with description of new taxa and comments on the taxonomy of the tribe" masterLastPageNumber="274" masterPageNumber="243" pageNumber="261" updateTime="1698465566927" updateUser="ExternalLinkService">
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<mods:title id="7AC9ADF4D2B04469DE57450A52B0AA4F">Revision of Hestiasulini Giglio-Tos, 1915 stat. rev. (Insecta: Mantodea: Hymenopodidae) of Borneo, with description of new taxa and comments on the taxonomy of the tribe</mods:title>
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<mods:namePart id="41ADB0E5671439DD910FC3F5DC38C494">Schwarz, Christian J.</mods:namePart>
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<mods:namePart id="EB04A6CCF567238C4BC32AEEC14D1570">Shcherbakov, Evgeny</mods:namePart>
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<mods:date id="2BE1232EC90132F653D79050DF1E61DB">2017</mods:date>
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<treatment id="03DC81241819A163FF6BF9A5FDD66776" ID-DOI="http://doi.org/10.5281/zenodo.6050962" ID-GBIF-Taxon="132549685" ID-Zenodo-Dep="6050962" LSID="urn:lsid:zoobank.org:act:65F02439-54F9-4810-9E86-EA892FF67DED" httpUri="http://treatment.plazi.org/id/03DC81241819A163FF6BF9A5FDD66776" lastPageId="19" lastPageNumber="262" pageId="18" pageNumber="261">
<subSubSection id="C36F63B91819A162FF6BF9A5FDDA6548" box="[151,565,1556,1582]" pageId="18" pageNumber="261" type="nomenclature">
<paragraph id="8BCA30321819A162FF6BF9A5FDDA6548" blockId="18.[151,565,1556,1582]" box="[151,565,1556,1582]" pageId="18" pageNumber="261">
<heading id="D082875E1819A162FF6BF9A5FDDA6548" bold="true" box="[151,565,1556,1582]" fontSize="11" level="1" pageId="18" pageNumber="261" reason="1">
<emphasis id="B901EC201819A162FF6BF9A5FDDA6548" bold="true" box="[151,565,1556,1582]" pageId="18" pageNumber="261">
Genus
<taxonomicName id="4C754BB11819A162FF12F9A5FE2F6548" box="[238,448,1556,1582]" class="Insecta" family="Empusidae" genus="Pseudohestiasula" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="261" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B901EC201819A162FF12F9A5FE2F6548" bold="true" box="[238,448,1556,1582]" italics="true" pageId="18" pageNumber="261">Pseudohestiasula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232515B1819A162FE3BF9A5FDDA6548" box="[455,565,1556,1582]" pageId="18" pageNumber="261" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36F63B91819A162FF6BF9EAFD7A6516" box="[151,661,1626,1648]" pageId="18" pageNumber="261" type="materials_examined">
<paragraph id="8BCA30321819A162FF6BF9EAFD7A6516" blockId="18.[151,661,1626,1648]" box="[151,661,1626,1648]" pageId="18" pageNumber="261">
Type species:
<taxonomicName id="4C754BB11819A162FEDAF9EBFDD16516" box="[294,574,1626,1648]" class="Insecta" family="Empusidae" genus="Pseudohestiasula" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="261" phylum="Arthropoda" rank="species" species="borneana" status="sp. nov.">
<emphasis id="B901EC201819A162FEDAF9EBFDD16516" box="[294,574,1626,1648]" italics="true" pageId="18" pageNumber="261">Pseudohestiasula borneana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232515B1819A162FDB8F9EAFD7A6516" box="[580,661,1627,1648]" pageId="18" pageNumber="261" rank="species">
<emphasis id="B901EC201819A162FDB8F9EAFD7A6516" bold="true" box="[580,661,1627,1648]" pageId="18" pageNumber="261">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C36F63B91819A163FF6BF92CFCE36391" lastPageId="19" lastPageNumber="262" pageId="18" pageNumber="261" type="diagnosis">
<paragraph id="8BCA30321819A162FF6BF92CFB9C65D0" blockId="18.[151,1436,1693,2006]" box="[151,1139,1693,1718]" pageId="18" pageNumber="261">
<emphasis id="B901EC201819A162FF6BF92CFEFE65D0" bold="true" box="[151,273,1693,1718]" pageId="18" pageNumber="261">Diagnosis.</emphasis>
Medium-sized
<taxonomicName id="4C754BB11819A162FE3DF92CFDD165D0" box="[449,574,1693,1718]" class="Insecta" family="Oxypilinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="261" phylum="Arthropoda" rank="family">Oxypilinae</taxonomicName>
with moderately dilated femora and reduced styli.
</paragraph>
<paragraph id="8BCA30321819A162FF3BF970FDC3640C" blockId="18.[151,1436,1693,2006]" pageId="18" pageNumber="261">Head with short median process. Pronotum without paramedian spines. Width of dilated part of fore femur smaller than width of non-dilated part. Fore tibia dorsally straight. Meso- and meta-femora with a ventral carina, on meso-femur starting from a very small proximal lobe. Male subgenital plate dorsally with two sclerotized carinae on right side. Styli absent. Dorsal lamina of left phallomere strongly folded along median line, ventral process of right phallomere large, pear-shaped.</paragraph>
<paragraph id="8BCA30321819A163FF3BF8C4FCE36391" blockId="18.[151,1436,1693,2006]" lastBlockId="19.[151,1437,151,1040]" lastPageId="19" lastPageNumber="262" pageId="18" pageNumber="261">
<taxonomicName id="4C754BB11819A162FF3BF8C4FE6464E8" box="[199,395,1909,1934]" class="Insecta" family="Empusidae" genus="Pseudohestiasula" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="261" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B901EC201819A162FF3BF8C4FE6464E8" box="[199,395,1909,1934]" italics="true" pageId="18" pageNumber="261">Pseudohestiasula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232515B1819A162FE6DF8C4FE1564E8" box="[401,506,1909,1934]" pageId="18" pageNumber="261" rank="genus">
<emphasis id="B901EC201819A162FE6DF8C4FE1564E8" bold="true" box="[401,506,1909,1934]" pageId="18" pageNumber="261">gen. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C754BB11819A162FD70F8C4FCE764E8" box="[652,776,1909,1934]" class="Insecta" family="Hymenopodidae" genus="Astyliasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="261" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B901EC201819A162FD70F8C4FCE764E8" box="[652,776,1909,1934]" italics="true" pageId="18" pageNumber="261">Astyliasula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232515B1819A162FCF3F8C4FC9764E8" box="[783,888,1909,1934]" pageId="18" pageNumber="261" rank="genus">
<emphasis id="B901EC201819A162FCF3F8C4FC9764E8" bold="true" box="[783,888,1909,1934]" pageId="18" pageNumber="261">gen. nov.</emphasis>
</taxonomicNameLabel>
by the presence of a process on the vertex and of a ventral lobe at the base of the meso femora, longer fore leg spines, the shorter and wider apical process, and also by the much less dilated dorsal lobe of the fore femora. It can be distinguished from
<taxonomicName id="4C754BB11819A162FBB2F80CFB2964B0" box="[1102,1222,1981,2006]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201819A162FBB2F80CFB2964B0" box="[1102,1222,1981,2006]" italics="true" pageId="18" pageNumber="261">Hestiasula</emphasis>
</taxonomicName>
,
<taxonomicName id="4C754BB11819A162FB28F80CFA8A64B0" box="[1236,1381,1981,2006]" class="Insecta" family="Hymenopodidae" genus="Ephestiasula" kingdom="Animalia" order="Mantodea" pageId="18" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201819A162FB28F80CFA8A64B0" box="[1236,1381,1981,2006]" italics="true" pageId="18" pageNumber="261">Ephestiasula</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C754BB11818A163FF6BFF26FECC63D6" box="[151,291,151,176]" class="Insecta" family="Hymenopodidae" genus="Catestiasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201818A163FF6BFF26FECC63D6" box="[151,291,151,176]" italics="true" pageId="19" pageNumber="262">Catestiasula</emphasis>
</taxonomicName>
by the absence of styli and of the membraneous gap along the left edge of the ventral phallomere, and by the presence of sclerotized carinae on the dorsal surface of subgenital plate, a sharply folded dorsal lamina of the left phallomere, and a narrow, digitiform apical process.
</paragraph>
</subSubSection>
<subSubSection id="C36F63B91818A163FF3BFEB5FDF060AE" pageId="19" pageNumber="262" type="description">
<paragraph id="8BCA30321818A163FF3BFEB5FEF16203" blockId="19.[151,1437,151,1040]" pageId="19" pageNumber="262">
<emphasis id="B901EC201818A163FF3BFEB5FEB6627B" bold="true" box="[199,345,260,285]" pageId="19" pageNumber="262">Description.</emphasis>
Male.
<emphasis id="B901EC201818A163FE4FFEB5FE00627B" box="[435,495,260,285]" italics="true" pageId="19" pageNumber="262">Head</emphasis>
pentagonal, slightly wider than long. Vertex with a small process. Eyes large, exophthalmic, ocelli large.
<taxonomicName id="4C754BB11818A163FE2FFE96FDAA6226" box="[467,581,295,320]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="19" pageNumber="251" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
pentagonal, about two times wider than high, with acute apex. Antennae moniliform.
</paragraph>
<paragraph id="8BCA30321818A163FF3BFEC0FAE4615B" blockId="19.[151,1437,151,1040]" pageId="19" pageNumber="262">
<emphasis id="B901EC201818A163FF3BFEC0FED962EE" box="[199,310,369,392]" italics="true" pageId="19" pageNumber="262">Pronotum</emphasis>
short, ovoid, lateral margin with
<taxonomicName id="4C754BB11818A163FD3EFEC1FCA962EE" box="[706,838,367,392]" class="Insecta" family="Hymenopodidae" genus="Astyliasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="251" phylum="Arthropoda" rank="species" species="triangular">a triangular</taxonomicName>
tooth, metazona slightly longer than prozona. Fore coxae longer than pronotum, with divergent apical lobes, proximal part anteriorly black, distal part pink. Furcasternum longer than metazona of pronotum, anteriorly black, posteriorly pink. Fore femora with dilated, foliaceous dorsal margin, dilated part slightly less wide than ventral part, with 4 discoidal, 4 postero-ventral, and 1314 antero-ventral spines, claw groove near base. Fore tibiae with 1416 postero-ventral and 1213 anteroventral spines, dorsal margin straight. First tarsomere about as long as remaining segments combined.
</paragraph>
<paragraph id="8BCA30321818A163FF3BFDF6FE9361CE" blockId="19.[151,1437,151,1040]" pageId="19" pageNumber="262">
<emphasis id="B901EC201818A163FF3BFDF6FE7E6106" box="[199,401,583,608]" italics="true" pageId="19" pageNumber="262">Mid and hind legs</emphasis>
rather short. Femora with a ventral carina, meso-femora also with a small elongate proximal lobe. Meso- and meta-tibiae simple, circular in cross-section. Hind metatarsus slightly shorter than remaining segments combined.
</paragraph>
<paragraph id="8BCA30321818A163FF3BFD04FEB76196" blockId="19.[151,1437,151,1040]" pageId="19" pageNumber="262">
<emphasis id="B901EC201818A163FF3BFD04FEE561AA" box="[199,266,693,716]" italics="true" pageId="19" pageNumber="262">Wings</emphasis>
macropterous, longer than abdomen, hyaline, spotted with dark and with dark apex. Costal area narrow, stigma indistinct.
</paragraph>
<paragraph id="8BCA30321818A163FF3BFD4DFBF4605E" blockId="19.[151,1437,151,1040]" pageId="19" pageNumber="262">
<emphasis id="B901EC201818A163FF3BFD4DFEDE6073" box="[199,305,764,789]" italics="true" pageId="19" pageNumber="262">Abdomen</emphasis>
simple, without lobes. Supra-anal plate trapezoidal. Cerci short, pilose, last segment conical. Subgenital plate dorsally with two sclerotized carinae on right side. Styli absent.
</paragraph>
<paragraph id="8BCA30321818A163FF3BFCF5FD8B60C3" blockId="19.[151,1437,151,1040]" pageId="19" pageNumber="262">
<emphasis id="B901EC201818A163FF3BFCF5FEDE603B" box="[199,305,836,861]" italics="true" pageId="19" pageNumber="262">Genitalia</emphasis>
with strongly pigmented processes. Dorsal lamina of left phallomere strongly folded along median line. Distal process spoon-shaped. Phalloid apophysis
<taxonomicName id="4C754BB11818A163FD03FCD6FC8160E6" box="[767,878,871,896]" class="Insecta" family="Hymenopodidae" genus="Astyliasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="262" phylum="Arthropoda" rank="species" species="triangular">triangular</taxonomicName>
, slightly bifurcate subapically. Ventral process of right phallomere very large, pear-shaped.
</paragraph>
<paragraph id="8BCA30321818A163FF3BFC1EFDF060AE" blockId="19.[151,1437,151,1040]" box="[199,543,943,968]" pageId="19" pageNumber="262">
<emphasis id="B901EC201818A163FF3BFC1EFEF360AE" box="[199,284,943,968]" italics="true" pageId="19" pageNumber="262">Female</emphasis>
and ootheca unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36F63B91818A163FF3BFC65FDD66776" pageId="19" pageNumber="262" type="etymology">
<paragraph id="8BCA30321818A163FF3BFC65FDD66776" blockId="19.[151,1437,151,1040]" pageId="19" pageNumber="262">
<emphasis id="B901EC201818A163FF3BFC65FEA1608B" bold="true" box="[199,334,980,1005]" pageId="19" pageNumber="262">Etymology.</emphasis>
The name is the combination of the ancient Greek ψευδής for “false”, and “
<taxonomicName id="4C754BB11818A163FB44FC65FADB608B" box="[1208,1332,980,1005]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="262" phylum="Arthropoda" rank="genus">Hestiasula</taxonomicName>
”. Named for its close resemblance to the latter.
</paragraph>
</subSubSection>
</treatment>
</document>