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<document id="4FFE646D88DEF8B3DE0E9805FADEE0EE" ID-DOI="10.11646/zootaxa.5361.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10150590" ID-ZooBank="33F819C3-F34D-431B-B262-C8FDBFD5FF4F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700247603070" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi, Geshev, Georgi &amp; Pathomwattananurak, Wuttipon" docDate="2023" docId="03B8222866725506FF75FDF9FBD2FC63" docLanguage="en" docName="zootaxa.5361.1.6.pdf" docOrigin="Zootaxa 5361 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudohomonyx crassus Arrow 1911" docType="treatment" docVersion="2" lastPageNumber="121" masterDocId="FF815A5066755501FFE2FFDBFFD6FFBF" masterDocTitle="A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake &amp; Yamaya, 1997 male and a catalogue of the genus" masterLastPageNumber="124" masterPageNumber="114" pageNumber="121" updateTime="1700247997433" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B611BB49D04748E31E951F5864844CDE">A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake &amp; Yamaya, 1997 male and a catalogue of the genus</mods:title>
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<mods:namePart id="32F0588B6C5C3F3D43D5BD43C5B63952">Yang, Qiao-Zhi</mods:namePart>
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<mods:namePart id="0E3769E96708B3F7FB4B7316B8DA0710">Geshev, Georgi</mods:namePart>
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<treatment id="03B8222866725506FF75FDF9FBD2FC63" LSID="urn:lsid:plazi:treatment:03B8222866725506FF75FDF9FBD2FC63" httpUri="http://treatment.plazi.org/id/03B8222866725506FF75FDF9FBD2FC63" lastPageNumber="121" pageId="7" pageNumber="121">
<subSubSection id="C30BC0B566725506FF75FDF9FD89FD83" box="[151,607,546,573]" pageId="7" pageNumber="121" type="nomenclature">
<paragraph id="8BAE933E66725506FF75FDF9FD89FD83" blockId="7.[151,607,546,573]" box="[151,607,546,573]" pageId="7" pageNumber="121">
<heading id="D0E6245266725506FF75FDF9FD89FD83" bold="true" box="[151,607,546,573]" fontSize="11" level="1" pageId="7" pageNumber="121" reason="1">
<taxonomicName id="4C11E8BD66725506FF75FDF9FD89FD83" ID-CoL="78G33" authority="Arrow, 1911" authorityName="Arrow" authorityYear="1911" box="[151,607,546,573]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="121" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B9654F2C66725506FF75FDF9FD89FD83" bold="true" box="[151,607,546,573]" pageId="7" pageNumber="121">
<emphasis id="B9654F2C66725506FF75FDF9FE17FD82" bold="true" box="[151,449,546,573]" italics="true" pageId="7" pageNumber="121">Pseudohomonyx crassus</emphasis>
<bibRefCitation id="EF80EECF66725506FE24FDF9FD89FD83" author="Arrow, G. J." box="[454,607,546,572]" pageId="7" pageNumber="121" pagination="151 - 176" refId="ref5218" refString="Arrow, G. J. (1911) Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (8), 151 - 176. https: // doi. org / 10.1080 / 00222931108693008" type="journal article" year="1911">Arrow, 1911</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566725506FF75FDBCFBD3FD03" pageId="7" pageNumber="121" type="reference_group">
<paragraph id="8BAE933E66725506FF75FDBCFBD3FD03" blockId="7.[151,1436,614,700]" pageId="7" pageNumber="121">
<treatmentCitationGroup id="AB01B41066725506FF75FDBCFBD3FD03" pageId="7" pageNumber="121">
<treatmentCitation id="0AB0B52F66725506FF75FDBCFD98FDC1" author="Arrow, G. J." box="[151,590,614,639]" page="159" pageId="7" pageNumber="121" year="1911">
<taxonomicName id="4C11E8BD66725506FF75FDBCFD98FDC1" ID-CoL="78G33" authority="Arrow, 1911: 159" authorityName="Arrow" authorityPageNumber="159" authorityYear="1911" box="[151,590,614,639]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="121" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B9654F2C66725506FF75FDBCFE46FDC1" box="[151,400,615,639]" italics="true" pageId="7" pageNumber="121">Pseudohomonyx crassus</emphasis>
<bibRefCitation id="EF80EECF66725506FE75FDBCFD98FDC1" author="Arrow, G. J." box="[407,590,614,639]" pageId="7" pageNumber="121" pagination="151 - 176" refId="ref5218" refString="Arrow, G. J. (1911) Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (8), 151 - 176. https: // doi. org / 10.1080 / 00222931108693008" type="journal article" year="1911">Arrow, 1911: 159</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB0B52F66725506FDBAFDBCFD28FDC0" author="Arrow, G. J." box="[600,766,615,639]" page="32" pageId="7" pageNumber="121" year="1937">
<bibRefCitation id="EF80EECF66725506FDBAFDBCFD28FDC0" author="Arrow, G. J." box="[600,766,615,639]" pageId="7" pageNumber="121" pagination="1 - 124" refId="ref5276" refString="Arrow, G. J. (1937) Scarabaeidae: Dynastinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 156. W. Junk, Gravenhage, pp. 1 - 124." type="book chapter" year="1937">Arrow 1937: 32</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB0B52F66725506FCEBFDBCFC1CFDC0" author="Endrodi, S." box="[777,970,615,639]" page="441" pageId="7" pageNumber="121" year="1969">
<bibRefCitation id="EF80EECF66725506FCEBFDBCFC1CFDC0" author="Endrodi, S." box="[777,970,615,639]" pageId="7" pageNumber="121" pagination="403 - 479" refId="ref5370" refString="Endrodi, S. (1969) Monographie der Dynastinae 4 Tribus Pentodontini der orientalische Region. Entomologische Arbeiten aus dem Museum G. Frey, 20, 403 - 479." type="journal article" year="1969">Endrődi 1969: 441</bibRefCitation>
</treatmentCitation>
, figs. 3637 (parameres);
<treatmentCitation id="0AB0B52F66725506FB35FDBCFA4EFDC0" author="Endrodi, S." box="[1239,1432,614,639]" page="316" pageId="7" pageNumber="121" year="1985">
<bibRefCitation id="EF80EECF66725506FB35FDBCFA4EFDC0" author="Endrodi, S." box="[1239,1432,614,639]" pageId="7" pageNumber="121" refId="ref5402" refString="Endrodi, S. (1985) The Dynastinae of the World. W. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrődi 1985: 316</bibRefCitation>
</treatmentCitation>
, figs. 12201221 (parameres);
<treatmentCitation id="0AB0B52F66725506FE19FD5EFCC9FD22" author="Miyake, Y. &amp; Yamaya, S." box="[507,799,645,670]" page="10" pageId="7" pageNumber="121" year="1997">
<bibRefCitation id="EF80EECF66725506FE19FD5EFCC9FD22" author="Miyake, Y. &amp; Yamaya, S." box="[507,799,645,670]" pageId="7" pageNumber="121" pagination="5 - 20" refId="ref5570" refString="Miyake, Y. &amp; Yamaya, S. (1997) New or little known Scarabaeid beetles from Southeast Asia, preserved in Nagaoka Municipal Science Museum. Bulletin of the Nagaoka municipal Science Museum, 32, 5 - 20" type="journal article" year="1997">Miyake &amp; Yamaya 1997: 10</bibRefCitation>
</treatmentCitation>
, figs. 8ab (parameres);
<treatmentCitation id="0AB0B52F66725506FBFDFD5EFB19FD22" author="Krajcik, M." box="[1055,1231,645,669]" page="55" pageId="7" pageNumber="121" year="2005">
<bibRefCitation id="EF80EECF66725506FBFDFD5EFB19FD22" author="Krajcik, M." box="[1055,1231,645,669]" pageId="7" pageNumber="121" pagination="1 - 122" refId="ref5515" refString="Krajcik, M. (2005) Dynastinae of the World Checklist (Coleoptera, Scarabaeidae, Dynastinae). Animma. x, Supplement 2, 1 - 122." type="journal article" year="2005">Krajčík 2005: 55</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB0B52F66725506FB39FD5EFA4EFD22" author="Krajcik, M." box="[1243,1432,645,669]" page="228" pageId="7" pageNumber="121" year="2012">
<bibRefCitation id="EF80EECF66725506FB39FD5EFA4EFD22" author="Krajcik, M." box="[1243,1432,645,669]" pageId="7" pageNumber="121" pagination="1 - 278" refId="ref5546" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement 5, 1 - 278." type="journal article" year="2012">Krajčík 2012: 228</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB0B52F66725506FF25FD7FFE78FD03" author="Jakl, S. &amp; Zidek, J." box="[199,430,676,700]" page="22" pageId="7" pageNumber="121" year="2017">
<bibRefCitation id="EF80EECF66725506FF25FD7FFE78FD03" author="Jakl, S. &amp; Zidek, J." box="[199,430,676,700]" pageId="7" pageNumber="121" pagination="19 - 30" refId="ref5477" refString="Jakl, S. &amp; Zidek, J. (2017) A contribution to the knowledge of Pseudohomonyx Arrow, 1908. Folia Heyrovskyana, Series A, 25 (2), 19 - 30." type="journal article" year="2017">Jákl &amp; Zídek 2017: 22</bibRefCitation>
</treatmentCitation>
, figs. 6 (habitus), 7 (head and pronotum), 89 (parameres).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566725506FF75FD31FD76FC97" pageId="7" pageNumber="121" type="materials_examined">
<paragraph id="8BAE933E66725506FF75FD31FD86FCBB" blockId="7.[151,1436,746,989]" box="[151,592,746,772]" pageId="7" pageNumber="121">
<emphasis id="B9654F2C66725506FF75FD31FEEFFCBB" bold="true" box="[151,313,746,772]" pageId="7" pageNumber="121">
<typeStatus id="54AA2D9C66725506FF75FD31FF04FCBB" box="[151,210,746,772]" pageId="7" pageNumber="121">Type</typeStatus>
locality:
</emphasis>
<taxonomicName id="4C11E8BD66725506FEA8FD31FE30FCBB" authority="Balu, N." authorityName="Balu, N." box="[330,486,746,772]" class="Echinoidea" family="Schizasteridae" genus="Kina" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spatangoida" pageId="7" pageNumber="121" phylum="Echinodermata" rank="genus">Kina Balu, N.</taxonomicName>
-Borneo”
</paragraph>
<paragraph id="8BAE933E66725506FF25FCD5FD76FC97" blockId="7.[151,1436,746,989]" box="[199,672,782,811]" pageId="7" pageNumber="121">
<emphasis id="B9654F2C66725506FF25FCD5FE46FC97" bold="true" box="[199,400,782,808]" pageId="7" pageNumber="121">Type depository:</emphasis>
<materialsCitation id="3B79996366725506FE75FCD4FD76FC97" box="[407,672,782,811]" collectionCode="BMNH" pageId="7" pageNumber="121" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54AA2D9C66725506FE75FCD4FDDDFC96" box="[407,523,783,809]" pageId="7" pageNumber="121" type="lectotype">Lectotype</typeStatus>
: ♁ (
<collectionCode id="ED000BFB66725506FDD8FCD5FD42FC97" box="[570,660,782,808]" pageId="7" pageNumber="121">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566725506FF25FCE9FC90FCF3" box="[199,838,818,845]" pageId="7" pageNumber="121" type="distribution">
<paragraph id="8BAE933E66725506FF25FCE9FC90FCF3" blockId="7.[151,1436,746,989]" box="[199,838,818,845]" pageId="7" pageNumber="121">
<emphasis id="B9654F2C66725506FF25FCE9FEB5FCF3" bold="true" box="[199,355,818,844]" pageId="7" pageNumber="121">Distribution:</emphasis>
<collectingCountry id="F306D3AE66725506FE88FCE9FE03FCF3" box="[362,469,818,844]" name="Malaysia" pageId="7" pageNumber="121">Malaysia</collectingCountry>
: Borneo;
<collectingCountry id="F306D3AE66725506FDA2FCE9FD67FCF3" box="[576,689,818,844]" name="Indonesia" pageId="7" pageNumber="121">Indonesia</collectingCountry>
:
<collectingRegion id="49D55DDC66725506FD5EFCE9FC94FCF3" box="[700,834,818,844]" country="Indonesia" name="Kalimantan Utara" pageId="7" pageNumber="121">Kalimantan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566725506FF25FC8DFBD2FC63" pageId="7" pageNumber="121" type="discussion">
<paragraph id="8BAE933E66725506FF25FC8DFBD2FC63" blockId="7.[151,1436,746,989]" pageId="7" pageNumber="121">
<emphasis id="B9654F2C66725506FF25FC8DFEEDFCCF" bold="true" box="[199,315,854,880]" pageId="7" pageNumber="121">Remarks.</emphasis>
As mentioned by
<bibRefCitation id="EF80EECF66725506FDF3FC8DFD17FCCF" author="Endrodi, S." box="[529,705,854,880]" pageId="7" pageNumber="121" pagination="403 - 479" refId="ref5370" refString="Endrodi, S. (1969) Monographie der Dynastinae 4 Tribus Pentodontini der orientalische Region. Entomologische Arbeiten aus dem Museum G. Frey, 20, 403 - 479." type="journal article" year="1969">Endrődi (1969)</bibRefCitation>
, a “
<taxonomicName id="4C11E8BD66725506FD10FC8CFCB0FCCF" authorityName="Arrow" authorityYear="1911" box="[754,870,855,880]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="121" phylum="Arthropoda" rank="species" species="craceus">
<emphasis id="B9654F2C66725506FD10FC8CFCB0FCCF" box="[754,870,855,880]" italics="true" pageId="7" pageNumber="121">P. craceus</emphasis>
</taxonomicName>
♁-
<typeStatus id="54AA2D9C66725506FC64FC83FC11FCCC" box="[902,967,856,883]" pageId="7" pageNumber="121">Type</typeStatus>
” was found in Prells collection (now in MFNB) which is presented in the section of
<taxonomicName id="4C11E8BD66725506FD68FCA1FBE5FC2B" authority="Arrow, 1911" authorityName="Arrow" authorityYear="1911" box="[650,1075,890,916]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="121" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B9654F2C66725506FD68FCA1FC4BFC2B" box="[650,925,890,916]" italics="true" pageId="7" pageNumber="121">Pseudohomonyx crassus</emphasis>
<bibRefCitation id="EF80EECF66725506FC46FCA1FBE5FC2B" author="Arrow, G. J." box="[932,1075,890,916]" pageId="7" pageNumber="121" pagination="151 - 176" refId="ref5218" refString="Arrow, G. J. (1911) Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (8), 151 - 176. https: // doi. org / 10.1080 / 00222931108693008" type="journal article" year="1911">Arrow, 1911</bibRefCitation>
</taxonomicName>
. However, it has been identified by Endrődi as
<taxonomicName id="4C11E8BD66725506FEDEFC44FDBCFC07" authority="Arrow, 1908" authorityName="Arrow" authorityYear="1908" box="[316,618,926,952]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="121" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="B9654F2C66725506FEDEFC44FE04FC07" box="[316,466,926,952]" italics="true" pageId="7" pageNumber="121">P. borneensis</emphasis>
<bibRefCitation id="EF80EECF66725506FE3AFC45FDBCFC07" author="Arrow, G. J." box="[472,618,926,952]" pageId="7" pageNumber="121" pagination="321 - 358" refId="ref5162" refString="Arrow, G. J. (1908) A contribution to the classification of the Coleopterous family Dynastidae. Transactions of the Entomological Society of London, 56 (2), 321 - 358. https: // doi. org / 10.1111 / j. 1365 - 2311.1908. tb 02151. x" type="journal article" year="1908">Arrow, 1908</bibRefCitation>
</taxonomicName>
, with dubious locality of “
<collectingRegion id="49D55DDC66725506FC74FC45FC18FC07" box="[918,974,926,952]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="121">Java</collectingRegion>
”; and without any publication from Prell regarding this specimen, it was excluded from any
<typeStatus id="54AA2D9C66725506FD31FC18FCD5FC62" box="[723,771,963,989]" pageId="7" pageNumber="121">type</typeStatus>
series (
<bibRefCitation id="EF80EECF66725506FCBAFC19FC21FC63" author="Endrodi, S." box="[856,1015,962,988]" pageId="7" pageNumber="121" pagination="403 - 479" refId="ref5370" refString="Endrodi, S. (1969) Monographie der Dynastinae 4 Tribus Pentodontini der orientalische Region. Entomologische Arbeiten aus dem Museum G. Frey, 20, 403 - 479." type="journal article" year="1969">Endrődi 1969</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>