101 lines
17 KiB
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101 lines
17 KiB
XML
<document id="0DAD4AE936ADE6A01111BC277A26EB10" ID-CLB-Dataset="89524" ID-DOI="10.11646/zootaxa.5162.1.1" ID-GBIF-Dataset="11422c4e-920a-4969-ad5d-3b820ffc2101" ID-ISSN="1175-5326" ID-Zenodo-Dep="6796605" ID-ZooBank="94DAF833-90C6-4AC8-B92A-0313F173064B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657005055755" checkinUser="plazi" docAuthor="Ulitzka, Manfred R." docDate="2022" docId="038B87FEA64A77015A86FAEEA650A8E3" docLanguage="en" docName="zootaxa.5162.1.1.pdf" docOrigin="Zootaxa 5162 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rohrthrips Nel, Penalver, Azar, Hodebert & Nel 2010" docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="FFB2FF86A640770B5A11FFF8A424AE1C" masterDocTitle="New genera and species of Rohrthripidae (Thysanoptera: Tubulifera) from Burmese Cretaceous amber" masterLastPageNumber="36" masterPageNumber="1" pageNumber="11" updateTime="1699356523688" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C31F746E5091697C86E9099FE239AEAA">New genera and species of Rohrthripidae (Thysanoptera: Tubulifera) from Burmese Cretaceous amber</mods:title>
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<mods:namePart id="B4C43F5624F49E878828951904E50DD1">Ulitzka, Manfred R.</mods:namePart>
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<mods:affiliation id="58A1B6C521EC0F5BA55179EC491D86C6">Thrips-iD, Strassburger Strasse 37 A, 77652 Offenburg, Germany.</mods:affiliation>
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<mods:title id="2FF42231495C138F70F3A6E55A66B409">Zootaxa</mods:title>
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<mods:date id="0301B2797B2A6BF2C49F8321489C8985">2022</mods:date>
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<treatment id="038B87FEA64A77015A86FAEEA650A8E3" ID-DOI="http://doi.org/10.5281/zenodo.6798806" ID-GBIF-Taxon="196390473" ID-Zenodo-Dep="6798806" LSID="urn:lsid:plazi:treatment:038B87FEA64A77015A86FAEEA650A8E3" httpUri="http://treatment.plazi.org/id/038B87FEA64A77015A86FAEEA650A8E3" lastPageNumber="11" pageId="10" pageNumber="11">
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<subSubSection id="C3386563A64A77015A86FAEEA715AB2C" box="[151,817,1302,1329]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8B9D36E8A64A77015A86FAEEA715AB2C" blockId="10.[151,817,1302,1329]" box="[151,817,1302,1329]" pageId="10" pageNumber="11">
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<taxonomicName id="4C224D6BA64A77015A86FAEEA715AB2C" ID-CoL="6444K" authority="Nel, Penalver, Azar, Hodebert & Nel, 2010" authorityName="Nel, Penalver, Azar, Hodebert & Nel" authorityYear="2010" box="[151,817,1302,1329]" class="Insecta" family="Rohrthripidae" genus="Rohrthrips" kingdom="Animalia" order="Thysanoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B956EAFAA64A77015A86FAEEA715AB2C" bold="true" box="[151,817,1302,1329]" pageId="10" pageNumber="11">
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<emphasis id="B956EAFAA64A77015A86FAEEA53FAB2D" bold="true" box="[151,283,1302,1329]" italics="true" pageId="10" pageNumber="11">Rohrthrips</emphasis>
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<bibRefCitation id="EFB34B19A64A77015B32FAEFA715AB2C" author="Nel, P. & Penalver, E. & Azar, D. & Hodebert, G. & Nel, A." box="[291,817,1302,1329]" pageId="10" pageNumber="11" pagination="154 - 163" refId="ref29076" refString="Nel, P., Penalver, E., Azar, D., Hodebert, G. & Nel, A. (2010) Modern thrips families Thripidae and Phlaeothripidae in Early Cretaceous amber (Insecta: Thysanoptera). Annales de la Societ entomologique de France, New Series, 46 (1 - 2), 154 - 163. https: // doi. org / 10.1080 / 00379271.2010.10697651" type="journal article" year="2010">Nel, Peñalver, Azar, Hodebert & Nel, 2010</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3386563A64A77015A86FAA4A77EAB6F" box="[151,858,1371,1396]" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="8B9D36E8A64A77015A86FAA4A77EAB6F" blockId="10.[151,858,1371,1396]" box="[151,858,1371,1396]" pageId="10" pageNumber="11">
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<typeStatus id="5499884AA64A77015A86FAA4A4EDAB68" box="[151,201,1372,1396]" pageId="10" pageNumber="11">Type</typeStatus>
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species:
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<taxonomicName id="4C224D6BA64A77015B37FAA4A77EAB6F" authority="Nel, Penalver, Azar, Hodebert & Nel, 2010" authorityName="Nel, Penalver, Azar, Hodebert & Nel" authorityYear="2010" box="[294,858,1371,1395]" class="Insecta" family="Rohrthripidae" genus="Rohrthrips" kingdom="Animalia" order="Thysanoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="libanicus">
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<emphasis id="B956EAFAA64A77015B37FAA4A5BBAB6F" box="[294,415,1371,1395]" italics="true" pageId="10" pageNumber="11">R. libanicus</emphasis>
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<bibRefCitation id="EFB34B19A64A77015BB7FAA3A77EAB6F" author="Nel, P. & Penalver, E. & Azar, D. & Hodebert, G. & Nel, A." box="[422,858,1371,1395]" pageId="10" pageNumber="11" pagination="154 - 163" refId="ref29076" refString="Nel, P., Penalver, E., Azar, D., Hodebert, G. & Nel, A. (2010) Modern thrips families Thripidae and Phlaeothripidae in Early Cretaceous amber (Insecta: Thysanoptera). Annales de la Societ entomologique de France, New Series, 46 (1 - 2), 154 - 163. https: // doi. org / 10.1080 / 00379271.2010.10697651" type="journal article" year="2010">Nel, Peñalver, Azar, Hodebert & Nel, 2010</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<subSubSection id="C3386563A64A77015A86FA59A650A8E3" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="8B9D36E8A64A77015A86FA59A650A8E3" blockId="10.[151,1437,1441,1791]" pageId="10" pageNumber="11">
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<emphasis id="B956EAFAA64A77015A86FA59A536ABA7" bold="true" box="[151,274,1441,1467]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
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Abdominal segment X tubiform, without a median suture, but with a crown of long apical anal setae. Antennae nine-segmented. Maxillary stylets deeply retracted into the head capsule and close together (
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<figureCitation id="13192A6DA64A77015EE1FA3DA165ABC3" box="[1264,1345,1477,1503]" captionStart="FIGURES 35–38" captionStartId="31.[151,264,1871,1895]" captionTargetBox="[160,1429,855,1838]" captionTargetId="figure-337@31.[151,1436,849,1847]" captionTargetPageId="31" captionText="FIGURES 35–38. Line drawings of Rohrthripidae. Head, mouthparts and prothorax (md: mandibula, mx: maxillary stylets) 35–37: (35) Adstrictubothrips mirapterus gen. et sp. n. holotype female; (36) Gemineurothrips peculiaris gen. et sp. n. holotype (spa: posteroangular setae, left broken off and lying close to coxa; scx: coxal setae); (37) Rohrthrips rhamphorhynchus sp. n. holotype. (38) Rohrthrips multihamuli sp. n. holotype; left antenna, segment numbers in Roman numerals, indicated: ventral projections on segments V–VII (cs: campaniform sensillum, uncolored trichomes indicate presumptive sense cones)." figureDoi="http://doi.org/10.5281/zenodo.6796637" httpUri="https://zenodo.org/record/6796637/files/figure.png" pageId="10" pageNumber="11">Fig. 37</figureCitation>
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;
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<bibRefCitation id="EFB34B19A64A77015F5DFA3DA4F6A81F" author="Ulitzka, M. R." pageId="10" pageNumber="11" pagination="548 - 558" refId="ref30112" refString="Ulitzka, M. R. (2018) A first survey of Cretaceous thrips from Burmese amber including the establishment of a new family of Tubulifera (Insecta: Thysanoptera). Zootaxa, 4486 (4), 548 - 558. https: // doi. org / 10.11646 / zootaxa. 4486.4.8" type="journal article" year="2018">Ulitzka 2018</bibRefCitation>
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,
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<bibRefCitation id="EFB34B19A64A77015AF1FA11A539A81F" author="Ulitzka, M. R." box="[224,285,1513,1539]" pageId="10" pageNumber="11" pagination="27 - 40" refId="ref30168" refString="Ulitzka, M. R. (2019) Five new species of Rohrthrips (Thysanoptera: Rohrthripidae) from Burmese amber, and the evolution of Tubulifera wings. Zootaxa, 4585 (1), 27 - 40. https: // doi. org / 10.11646 / zootaxa. 4585.1.2" type="journal article" year="2019">2019</bibRefCitation>
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). Fore wings becoming progressively larger in distal half (
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<figureCitation id="13192A6DA64A770159AEFA11A066A81F" box="[959,1090,1513,1539]" captionStart="FIGURES 12–15" captionStartId="14.[151,275,1747,1773]" captionTargetBox="[151,1436,181,1725]" captionTargetId="figure-22@14.[151,1436,181,1725]" captionTargetPageId="14" captionText="FIGURES 12–15. Rohrthrips spp. (12) Rohrthrips brachyvenis sp. n. holotype female, dorsal view; (13) Rohrthrips breviceps male, dorsal view (Note: the median shading makes the tube appear narrow, but the lighter edges are part of the tube, which actually is broad at base); (14) Rohrthrips jiewenae male, dorsal view; (15) Rohrthrips multihamuli sp. n. holotype female, head and pronotum, antennal segments III–IV indicated with roman numerals." figureDoi="http://doi.org/10.5281/zenodo.6796617" httpUri="https://zenodo.org/record/6796617/files/figure.png" pageId="10" pageNumber="11">Figs 12–13</figureCitation>
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,
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<figureCitation id="13192A6DA64A77015E41FA11A04BA81F" box="[1104,1135,1513,1539]" captionStart="FIGURES 16–18" captionStartId="19.[151,264,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-15@19.[151,1436,181,1839]" captionTargetPageId="19" captionText="FIGURES 16–18. Rohrthrips spp. (16) Rohrthrips pandemicus sp. n. holotype female aptera, dorsal view; (17) Rohrthrips rhamphorhynchus sp. n. holotype, dorsal view, indicated: second vein at base of fore wing; mouth cone outlined white; (18) Rohrthrips setiger sp. n. holotype female, right fore wing, indicated: second wing vein at base." figureDoi="http://doi.org/10.5281/zenodo.6796619" httpUri="https://zenodo.org/record/6796619/files/figure.png" pageId="10" pageNumber="11">18</figureCitation>
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); with a first vein bearing setae and, in some species, additionally with a second much shorter vein without setae (
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<figureCitation id="13192A6DA64A77015E97F9F5A0FFA83B" box="[1158,1243,1549,1575]" captionStart="FIGURES 16–18" captionStartId="19.[151,264,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-15@19.[151,1436,181,1839]" captionTargetPageId="19" captionText="FIGURES 16–18. Rohrthrips spp. (16) Rohrthrips pandemicus sp. n. holotype female aptera, dorsal view; (17) Rohrthrips rhamphorhynchus sp. n. holotype, dorsal view, indicated: second vein at base of fore wing; mouth cone outlined white; (18) Rohrthrips setiger sp. n. holotype female, right fore wing, indicated: second wing vein at base." figureDoi="http://doi.org/10.5281/zenodo.6796619" httpUri="https://zenodo.org/record/6796619/files/figure.png" pageId="10" pageNumber="11">Fig. 18</figureCitation>
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); cross-veins not developed; wing tips broadly rounded. Fringe cilia in most species arising from sockets. Duplicated cilia present around wing apex; running parallel to normal wing fringes. Fore wing clavus with paired setiform processes at tip and hind wing with basal recurved setae, wing coupling hamulo-frenate (
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<figureCitation id="13192A6DA64A770159ACF981A034A88F" box="[957,1040,1657,1683]" captionStart="FIGURES 24–29" captionStartId="28.[151,264,1636,1660]" captionTargetBox="[151,1436,273,1613]" captionTargetId="figure-72@28.[151,1436,273,1613]" captionTargetPageId="28" captionText="FIGURES 24–29. Thysanoptera wings. Wing coupling (f: frenulum, h: hamuli, r: retinaculum, smr: submarginal ridge, indicat- ed: microtrichia) 24–27: (24) Taeniothrips inconsequens Uzel (Terebrantia: Thripidae); (25) Elaphrothrips cf. costalimai Hood (Tubulifera: Phlaeothripidae: Idolothripinae); (26) Gemineurothrips microcephalus gen. et sp. n. (Tubulifera: Rohrthripidae); (27) Rohrthrips multihamuli sp. n. (Tubulifera: Rohrthripidae). Tubulifera fore wing duplicated cilia (green to red arrow: area with duplicated cilia) 28–29: (28) Haplothrips setiger Priesner (Phlaeothripidae); (29) Gemineurothrips microcephalus gen. et sp. n. (Rohrthripidae)." figureDoi="http://doi.org/10.5281/zenodo.6796631" httpUri="https://zenodo.org/record/6796631/files/figure.png" pageId="10" pageNumber="11">Fig. 27</figureCitation>
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). Abdomen without sigmoidal wing retaining setae; abdominal tergite II with a pair of distinct dorsolateral sutures (
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<figureCitation id="13192A6DA64A77015E00F965A041A8AB" box="[1041,1125,1693,1719]" captionStart="FIGURES 30–34" captionStartId="30.[151,264,1808,1832]" captionTargetBox="[151,1436,237,1785]" captionTargetId="figure-59@30.[151,1436,237,1785]" captionTargetPageId="30" captionText="FIGURES 30–34. Abdomen of Tubulifera. Tergites (P: pelta, T: tube, II–IX: abdominal segments number) 30–31: (30) Rohrthrips c.f. burmiticus male (MU-Fos-114/1), wings broken off (remains indicated white), indicated black: dorsolateral grooves on tergum II; (31) Gynaikothrips uzeli (Zimmermann) female, indicated: wing retaining sigmoid setae. Distal sternites of males, indicated white: edge of pointed emargination at base of tube 32–34: (32) Alavathrips moralesi reinterpreted from Peñalver et al. (2022, p. 5 Figs 3c), indicated black and dashed redrawn: phallus (designated “ovipositor” in Peñalver et al. 2022); (33) Holothrips schaubergeri (Priesner); (34) Rohrthrips breviceps holotype male (T: tube; IX: hypandrium, sternite IX)." figureDoi="http://doi.org/10.5281/zenodo.6796635" httpUri="https://zenodo.org/record/6796635/files/figure.png" pageId="10" pageNumber="11">Fig. 30</figureCitation>
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). The latter feature is rarely found in
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<taxonomicName id="4C224D6BA64A77015AEFF939A557A8C7" baseAuthorityName="Ulitzka" baseAuthorityYear="2018" box="[254,371,1729,1755]" class="Insecta" kingdom="Animalia" order="Thysanoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="subOrder" subOrder="Tubulifera">Tubulifera</taxonomicName>
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and has not been documented so far in
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<taxonomicName id="4C224D6BA64A77015926F939A794A8C7" authorityName="Nel, Penalver, Azar, Hodebert & Nel" authorityYear="2010" box="[823,944,1729,1755]" class="Insecta" family="Rohrthripidae" genus="Rohrthrips" kingdom="Animalia" order="Thysanoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B956EAFAA64A77015926F939A794A8C7" box="[823,944,1729,1755]" italics="true" pageId="10" pageNumber="11">Rohrthrips</emphasis>
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</taxonomicName>
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(see discussion below); it is also present in
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<taxonomicName id="4C224D6BA64A77015A86F91DA5FCA8E3" authority="Ulitzka, 2022" authorityName="Ulitzka" authorityYear="2022" box="[151,472,1765,1791]" class="Insecta" family="Rohrthripidae" genus="Paralleloalathrips" kingdom="Animalia" order="Thysanoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bivenatus" status="gen. et sp. n.">
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<emphasis id="B956EAFAA64A77015A86F91DA5FCA8E3" box="[151,472,1765,1791]" italics="true" pageId="10" pageNumber="11">Paralleloalathrips bivenatus</emphasis>
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</taxonomicName>
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<emphasis id="B956EAFAA64A77015BCEF91DA650A8E3" bold="true" box="[479,628,1765,1791]" pageId="10" pageNumber="11">
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<taxonomicNameLabel id="A2655781A64A77015BCEF91DA650A8E3" box="[479,628,1765,1791]" pageId="10" pageNumber="11" rank="species">gen. et sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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