228 lines
27 KiB
XML
228 lines
27 KiB
XML
<document id="90598F9D41281D136992BDB88D24E505" ID-DOI="10.5281/zenodo.193356" ID-GBIF-Dataset="25d23af4-b9c8-47be-b68e-33ae68076464" ID-ISSN="1175-5326" ID-Zenodo-Dep="193356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460135747041" checkinUser="plazi" docAuthor="Tarasov, Sergey I. & Kabakov, Oleg N." docDate="2010" docId="038B87DEFFDD1E39FF02F89F3170F843" docLanguage="en" docName="zt02344p028.pdf" docOrigin="Zootaxa 2344" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus (Sinonthophagus) nampatensis Tarasov & Kabakov, sp.n." docType="treatment" docVersion="10" lastPageNumber="22" masterDocId="FFB2FFA6FFD91E3CFF95FFD23424FFCF" masterDocTitle="Two new species of Onthophagus (Coleoptera: Scarabaeidae) from Indochina, with a discussion of some problems with the classification of Serrophorus and similar subgenera" masterLastPageNumber="28" masterPageNumber="17" pageNumber="21" updateTime="1698235207373" updateUser="plazi">
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<mods:titleInfo id="B4456FA478F27ED3E23779E472F1D79A">
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<mods:title id="FA142B4D7DD4B80895D560326B834A8B">Two new species of Onthophagus (Coleoptera: Scarabaeidae) from Indochina, with a discussion of some problems with the classification of Serrophorus and similar subgenera</mods:title>
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</mods:titleInfo>
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<mods:name id="FA7FA01D77F14DD09EDE2125C6B4BEA1" type="personal">
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<mods:roleTerm id="6FCCF82D55B5F43823B160B2114BF435">Author</mods:roleTerm>
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<mods:namePart id="13C1AE8C3662D872CFE6D8C393FDAE41">Tarasov, Sergey I.</mods:namePart>
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<mods:roleTerm id="ACFE4A0C9676CCF461B1F3F73681F979">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1D5CE723455B52527C60B0D4DB430C64">Kabakov, Oleg N.</mods:namePart>
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<mods:typeOfResource id="03ED40BCC67E47903E329F072E263D2A">text</mods:typeOfResource>
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<mods:title id="AEF30C77739BF57FFC0B408735E4A630">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DAD9D3AA43E8D456C6081CAEAEBD6E1F">
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<mods:date id="C1E1B207E2E68B71220EF0D6F88F6559">2010</mods:date>
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<mods:detail id="A37537FA1B7FEC6C21A8842EAD903C96" type="volume">
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<mods:number id="DB54A897126BA81E39AA83D1722ED8D3">2344</mods:number>
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<mods:identifier id="AA86EDA22CDD76882809C0F6CCB0CE20" type="DOI">10.5281/zenodo.193356</mods:identifier>
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<mods:identifier id="D6423CED708A93C0491E3DE8202A1696" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="35ABBEBD345B47963190CEEDE8F2FDA6" type="Zenodo-Dep">193356</mods:identifier>
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<treatment id="038B87DEFFDD1E39FF02F89F3170F843" ID-DOI="http://doi.org/10.5281/zenodo.5687354" ID-GBIF-Taxon="119383042" ID-Zenodo-Dep="5687354" LSID="urn:lsid:plazi:treatment:038B87DEFFDD1E39FF02F89F3170F843" httpUri="http://treatment.plazi.org/id/038B87DEFFDD1E39FF02F89F3170F843" lastPageId="5" lastPageNumber="22" pageId="4" pageNumber="21">
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<subSubSection id="C3386543FFDD1E38FF02F89F3548F845" pageId="4" pageNumber="21" type="nomenclature">
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<paragraph id="8B9D36C8FFDD1E38FF02F89F3026F8A8" blockId="4.[151,1026,1869,1930]" box="[151,1026,1869,1895]" pageId="4" pageNumber="21">
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<heading id="D0D581A4FFDD1E38FF02F89F3026F8A8" bold="true" box="[151,1026,1869,1895]" fontSize="11" level="1" pageId="4" pageNumber="21" reason="1">
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<emphasis id="B956EADAFFDD1E38FF02F89F3026F8A8" bold="true" box="[151,1026,1869,1895]" pageId="4" pageNumber="21">
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||
<taxonomicName id="4C224D4BFFDD1E38FF02F89F37E4F8A8" ID-CoL="49MVR" authority="Tarasov & Kabakov" authorityName="Tarasov & Kabakov" box="[151,960,1869,1895]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="nampatensis" status="sp. nov." subGenus="Sinonthophagus">
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<emphasis id="B956EADAFFDD1E38FF02F89F369FF8A8" bold="true" box="[151,699,1869,1895]" italics="true" pageId="4" pageNumber="21">Onthophagus (Sinonthophagus) nampatensis</emphasis>
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Tarasov & Kabakov
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</taxonomicName>
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<taxonomicNameLabel id="A26557A1FFDD1E38FC52F89F3026F8A8" box="[967,1026,1869,1895]" pageId="4" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8B9D36C8FFDD1E38FF02F8A23548F845" blockId="4.[151,1026,1869,1930]" box="[151,364,1904,1930]" pageId="4" pageNumber="21">
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(
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<figureCitation id="13192A4DFFDD1E38FF0AF8A234CAF845" box="[159,238,1904,1930]" captionStart="FIGURE 1" captionStartId="2.[151,257,1711,1735]" captionTargetBox="[152,1414,315,1710]" captionTargetId="figure@2.[151,1436,269,1712]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Onthophagus. a, b— O. streltsovi; c, d— O. nampatensis; a, c—males, holotypes; b—female, paratype; d—small male paratype." httpUri="https://zenodo.org/record/193357/files/figure.png" pageId="4" pageNumber="21">Figs. 1</figureCitation>
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c, d; 3a–d)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3386543FFDD1E39FF02F8683770FE5D" lastPageId="5" lastPageNumber="22" pageId="4" pageNumber="21" type="materials_examined">
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<paragraph id="8B9D36C8FFDD1E39FF02F868312BFF18" blockId="4.[151,1036,1978,2005]" lastBlockId="5.[151,1438,152,1932]" lastPageId="5" lastPageNumber="22" pageId="4" pageNumber="21">
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<emphasis id="B956EADAFFDD1E38FF02F86835AFF81B" bold="true" box="[151,395,1978,2004]" pageId="4" pageNumber="21">Material examined.</emphasis>
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<emphasis id="B956EADAFFDD1E38FE07F86935DAF81B" box="[402,510,1979,2004]" italics="true" pageId="4" pageNumber="21">
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<typeStatus id="5499886AFFDD1E38FE07F86935DAF81B" box="[402,510,1979,2004]" pageId="4" pageNumber="21" type="holotype">Holotype</typeStatus>
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</emphasis>
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(
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<collectionCode id="ED33AE0DFFDD1E38FD98F869364CF81A" box="[525,616,1979,2005]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="21">ZMUC</collectionCode>
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), male bearing the following labels: 1. White, printed:
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<collectingCountry id="F3357658FFDC1E39FE20FF4A362FFF7D" box="[437,523,152,178]" name="Laos" pageId="5" pageNumber="22">LAOS</collectingCountry>
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, Khammouane prov.
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<quantity id="4CDA9B2DFFDC1E39FC8BFF4A3743FF7D" box="[798,871,152,178]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="22" unit="km" value="10.0">10km</quantity>
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W of Ban Khun Kham (Nahin)
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<geoCoordinate id="EE16500FFFDC1E39FA9FFF4A31BAFF7D" box="[1290,1438,152,178]" direction="north" orientation="latitude" pageId="5" pageNumber="22" precision="1" value="18.1487">N18°8.922'</geoCoordinate>
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<geoCoordinate id="EE16500FFFDC1E39FF02FF6F351EFF18" box="[151,314,189,215]" direction="east" orientation="longitude" pageId="5" pageNumber="22" precision="1" value="104.47715">E104°28.629'</geoCoordinate>
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, h
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<quantity id="4CDA9B2DFFDC1E39FEC1FF6F35BDFF18" box="[340,409,189,215]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="22" unit="m" value="280.0">280m</quantity>
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, limestone tropical forest, dung trap (human). 4–
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<date id="FF9C1008FFDC1E39FC43FF6F307EFF18" box="[982,1114,189,215]" pageId="5" pageNumber="22" value="2008-06-18">18.06.2008</date>
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leg. S. Tarasov
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</paragraph>
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<paragraph id="8B9D36C8FFDC1E39FF51FF303520FEED" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
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2. Orange, printed:
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<typeStatus id="5499886AFFDC1E39FE33FF303671FF33" box="[422,597,226,252]" pageId="5" pageNumber="22" type="holotype">HOLOTYPUS</typeStatus>
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<taxonomicName id="4C224D4BFFDC1E39FDCEFF3031B8FF33" authority="S. Tarasov & O. Kabakov" authorityName="S. Tarasov & O. Kabakov" box="[603,1436,226,252]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="nampatensis" subGenus="Sinonthophagus">
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<emphasis id="B956EADAFFDC1E39FDCEFF30304EFF34" box="[603,1130,226,251]" italics="true" pageId="5" pageNumber="22">Onthophagus (Sinonthophagus) nampatensis</emphasis>
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S. Tarasov & O. Kabakov
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</taxonomicName>
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det. 2009
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</paragraph>
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<paragraph id="8B9D36C8FFDC1E39FF51FEFF3770FE5D" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
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<emphasis id="B956EADAFFDC1E39FF51FEFF3562FE89" box="[196,326,301,326]" italics="true" pageId="5" pageNumber="22">
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<typeStatus id="5499886AFFDC1E39FF51FEFF3562FE89" box="[196,326,301,326]" pageId="5" pageNumber="22" type="paratype">Paratypes</typeStatus>
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</emphasis>
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: 1 3,
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<collectingCountry id="F3357658FFDC1E39FE0EFEFF35F9FE88" box="[411,477,301,327]" name="Laos" pageId="5" pageNumber="22">Laos</collectingCountry>
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, Khammouane province
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<quantity id="4CDA9B2DFFDC1E39FCB5FEFF374CFE88" box="[800,872,301,327]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="22" unit="km" value="10.0">10km</quantity>
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W of Ban Khun Kham (Nahin)
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<geoCoordinate id="EE16500FFFDC1E39FA9EFEFF31BAFE88" box="[1291,1438,301,327]" direction="north" orientation="latitude" pageId="5" pageNumber="22" precision="1" value="18.1487">N18°8.922'</geoCoordinate>
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<geoCoordinate id="EE16500FFFDC1E39FF02FE80351EFEA3" box="[151,314,338,364]" direction="east" orientation="longitude" pageId="5" pageNumber="22" precision="1" value="104.47715">E104°28.629'</geoCoordinate>
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, h
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<quantity id="4CDA9B2DFFDC1E39FEC1FE8035BDFEA3" box="[340,409,338,364]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="22" unit="m" value="280.0">280m</quantity>
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, limestone tropical forest, dung trap 4–
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<date id="FF9C1008FFDC1E39FCF4FE8037C1FEA3" box="[865,997,338,364]" pageId="5" pageNumber="22" value="2008-06-18">18.06.2008</date>
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leg. S. Tarasov; 1 3, North
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<collectingCountry id="F3357658FFDC1E39FAB9FE8031B3FEA3" box="[1324,1431,338,364]" name="Thailand" pageId="5" pageNumber="22">Thailand</collectingCountry>
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, Doi Suthep,
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<date id="FF9C1008FFDC1E39FEBFFEAA3586FE5D" box="[298,418,376,402]" pageId="5" pageNumber="22" value="1958-06-19">19.6.1958</date>
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, h
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<quantity id="4CDA9B2DFFDC1E39FE28FEAA3634FE5D" box="[445,528,376,402]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="22" unit="m" value="1100.0">1100m</quantity>
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, leg. B. Degerbøl (
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<collectionCode id="ED33AE0DFFDC1E39FD78FEAA376DFE5D" box="[749,841,376,402]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="5" pageNumber="22">ZMUC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3386543FFDC1E39FF51FE4F37EAFB6D" pageId="5" pageNumber="22" type="description">
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<paragraph id="8B9D36C8FFDC1E39FF51FE4F3713FDCD" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
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<emphasis id="B956EADAFFDC1E39FF51FE4F3579FE78" bold="true" box="[196,349,413,439]" pageId="5" pageNumber="22">Description.</emphasis>
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Body black, slightly lustrous, elytra with stronger luster; antennae and mouthparts orange; elytra and abdomen with short black setae turning white depending on light angle; rest of abdominal side and pygidium with long, orange hairs. Length 10.0–
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<quantity id="4CDA9B2DFFDC1E39FD5DFE3A3717FDCD" box="[712,819,488,514]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="5" pageNumber="22" unit="mm" value="12.8">12.8 mm</quantity>
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.
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</paragraph>
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<paragraph id="8B9D36C8FFDC1E39FF51FDDF3793FDBD" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
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<emphasis id="B956EADAFFDC1E39FF51FDDF3572FDE9" box="[196,342,525,550]" italics="true" pageId="5" pageNumber="22">Male major</emphasis>
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(
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<figureCitation id="13192A4DFFDC1E39FEFDFDDF3594FDE8" box="[360,432,525,551]" captionStart="FIGURE 1" captionStartId="2.[151,257,1711,1735]" captionTargetBox="[152,1414,315,1710]" captionTargetId="figure@2.[151,1436,269,1712]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Onthophagus. a, b— O. streltsovi; c, d— O. nampatensis; a, c—males, holotypes; b—female, paratype; d—small male paratype." httpUri="https://zenodo.org/record/193357/files/figure.png" pageId="5" pageNumber="22">Fig. 1</figureCitation>
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c). Head rounded; frontoclypeal ridge slightly expressed, in smaller males more distinct; vertex with a pair of long, tapering and inwardly-rounded horns connected by ridge; eyes completely divided by genal appendage; clypeus densely, transversely rugulose.
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</paragraph>
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<paragraph id="8B9D36C8FFDC1E39FF51FDAF316CFCE3" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">Pronotum with distinct excavations near anterior angles, anterior part protruding forward between cephalic horns; postlateral section strongly sinuate; anterior and lateral edges marginate, basal edge shortly marginate only near median basal depression; anterior angles slightly protruded forward, widely rounded; pronotal disc covered with simple, sparse punctures (separated by 1–2 puncture diameters) becoming denser and rasplike anteriorly; anterior part of propleura with ridge almost reaching propleural lateral margin.</paragraph>
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<paragraph id="8B9D36C8FFDC1E39FF51FCEA3738FC9D" blockId="5.[151,1438,152,1932]" box="[196,796,824,850]" pageId="5" pageNumber="22">Elytra shagreened, sparsely and densely punctured.</paragraph>
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<paragraph id="8B9D36C8FFDC1E39FF51FC8F375CFC0D" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">Protibia with four outer teeth, median part of anterior edge slightly produced into triangle tooth, internal apical angle with small tooth; apical spur short, acuminate; metatibiae strongly triangularly broadened toward apex; mesotibial and metatibial apex with short and long setae.</paragraph>
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<paragraph id="8B9D36C8FFDC1E39FF51FC1F374DFBC3" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
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Aedeagus as in
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<figureCitation id="13192A4DFFDC1E39FEE8FC1F35EAFC28" box="[381,462,973,999]" captionStart="FIGURE 3" captionStartId="6.[151,255,1352,1376]" captionTargetBox="[193,1393,209,1293]" captionTargetId="figure@6.[151,1436,165,1337]" captionTargetPageId="6" captionText="FIGURE 3. Elements of genitalia of Onthophagus nampatensis. a—lamella copulatrix; b—lamella copulatrix, AIS and ALS removed; c—aedeagus, lateral view; d—aedeagus, apical view." httpUri="https://zenodo.org/record/193359/files/figure.png" pageId="5" pageNumber="22">Figs. 3</figureCitation>
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c, d. Lamella copulatrix with additional internal sclerite (AIS) and small additional lateral sclerite (ALS) (
|
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<figureCitation id="13192A4DFFDC1E39FE0BFC2035C0FBC3" box="[414,484,1010,1036]" captionStart="FIGURE 3" captionStartId="6.[151,255,1352,1376]" captionTargetBox="[193,1393,209,1293]" captionTargetId="figure@6.[151,1436,165,1337]" captionTargetPageId="6" captionText="FIGURE 3. Elements of genitalia of Onthophagus nampatensis. a—lamella copulatrix; b—lamella copulatrix, AIS and ALS removed; c—aedeagus, lateral view; d—aedeagus, apical view." httpUri="https://zenodo.org/record/193359/files/figure.png" pageId="5" pageNumber="22">Fig. 3</figureCitation>
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a); superior part simple (
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<figureCitation id="13192A4DFFDC1E39FC91FC20376EFBC3" box="[772,842,1010,1036]" captionStart="FIGURE 3" captionStartId="6.[151,255,1352,1376]" captionTargetBox="[193,1393,209,1293]" captionTargetId="figure@6.[151,1436,165,1337]" captionTargetPageId="6" captionText="FIGURE 3. Elements of genitalia of Onthophagus nampatensis. a—lamella copulatrix; b—lamella copulatrix, AIS and ALS removed; c—aedeagus, lateral view; d—aedeagus, apical view." httpUri="https://zenodo.org/record/193359/files/figure.png" pageId="5" pageNumber="22">Fig. 3</figureCitation>
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b).
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</paragraph>
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<paragraph id="8B9D36C8FFDC1E39FF51FBCA3582FBFD" blockId="5.[151,1438,152,1932]" box="[196,422,1048,1074]" pageId="5" pageNumber="22">
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<emphasis id="B956EADAFFDC1E39FF51FBCA3538FBFE" box="[196,284,1048,1073]" italics="true" pageId="5" pageNumber="22">Female</emphasis>
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. Unknown.
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</paragraph>
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<paragraph id="8B9D36C8FFDC1E39FF51FBEF37EAFB6D" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
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<emphasis id="B956EADAFFDC1E39FF51FBEF350BFB99" box="[196,303,1085,1110]" italics="true" pageId="5" pageNumber="22">Va r i a t i o n</emphasis>
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. Males vary allometrically in the degree of expression of such body parts as horns, pronotum and punctation. Small males (
|
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<figureCitation id="13192A4DFFDC1E39FE60FBB0361CFBB3" box="[501,568,1122,1148]" captionStart="FIGURE 1" captionStartId="2.[151,257,1711,1735]" captionTargetBox="[152,1414,315,1710]" captionTargetId="figure@2.[151,1436,269,1712]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Onthophagus. a, b— O. streltsovi; c, d— O. nampatensis; a, c—males, holotypes; b—female, paratype; d—small male paratype." httpUri="https://zenodo.org/record/193357/files/figure.png" pageId="5" pageNumber="22">Fig. 1</figureCitation>
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d) usually have small horns; anterior part of pronotum with two tubercles medially; pronotum flatter, not concave; disc more densely punctured.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C3386543FFDC1E39FF51FB7F37CFF9D8" pageId="5" pageNumber="22" type="materials_examined">
|
||
<paragraph id="8B9D36C8FFDC1E39FF51FB7F3557FA4D" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
|
||
<emphasis id="B956EADAFFDC1E39FF51FB7F3510FB09" box="[196,308,1197,1222]" italics="true" pageId="5" pageNumber="22">
|
||
<typeStatus id="5499886AFFDC1E39FF51FB7F3510FB09" box="[196,308,1197,1222]" pageId="5" pageNumber="22" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13192A4DFFDC1E39FED3FB7F35A9FB08" box="[326,397,1197,1223]" captionStart="FIGURE 1" captionStartId="2.[151,257,1711,1735]" captionTargetBox="[152,1414,315,1710]" captionTargetId="figure@2.[151,1436,269,1712]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Onthophagus. a, b— O. streltsovi; c, d— O. nampatensis; a, c—males, holotypes; b—female, paratype; d—small male paratype." httpUri="https://zenodo.org/record/193357/files/figure.png" pageId="5" pageNumber="22">Fig. 1</figureCitation>
|
||
c). Major male, Length
|
||
<quantity id="4CDA9B2DFFDC1E39FD23FB7F370CFB08" box="[694,808,1197,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="5" pageNumber="22" unit="mm" value="12.8">12.8 mm</quantity>
|
||
. Body black, slightly lustrous, elytra with stronger luster; antennae and mouthparts orange. Head rounded; frontoclypeal ridge weakly expressed (more distinct in smaller males); vertex with a pair of long, tapering and inwardly-rounded horns connected by ridge. Pronotum with distinct excavations near anterior angles, anterior part protruding forward between cephalic horns; disc covered with simple, sparse punctures (separated by 1–2 puncture diameters) becoming denser and rasplike anteriorly.
|
||
</paragraph>
|
||
<paragraph id="8B9D36C8FFDC1E39FF51FA5F37CFF9D8" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
|
||
<emphasis id="B956EADAFFDC1E39FF51FA5F35C5FA68" bold="true" box="[196,481,1421,1447]" pageId="5" pageNumber="22">Differential diagnosis.</emphasis>
|
||
The new species is most similar to
|
||
<taxonomicName id="4C224D4BFFDC1E39FC0CFA5F3517FA03" authority="Arrow, 1907" authorityName="Arrow" authorityYear="1907" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="productus" subGenus="Sinonthophagus">
|
||
<emphasis id="B956EADAFFDC1E39FC0CFA5F31B8FA69" box="[921,1436,1421,1446]" italics="true" pageId="5" pageNumber="22">Onthophagus (Sinonthophagus) productus</emphasis>
|
||
Arrow, 1907
|
||
</taxonomicName>
|
||
but differs by slightly sparser punctation of the pronotum and distinctly different shape of parameres. It can be also separated from all other species of the subgenus
|
||
<emphasis id="B956EADAFFDC1E39FB95FA0A3099FA3E" box="[1024,1213,1496,1521]" italics="true" pageId="5" pageNumber="22">Sinonthophagus</emphasis>
|
||
<bibRefCitation id="EFB34B39FFDC1E39FB50FA0A3153FA3D" author="Kabakov" box="[1221,1399,1496,1522]" pageId="5" pageNumber="28" refString="Kabakov, O. N. (2006) Beetles of the subfamily Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) of the fauna of Russia and adjacent countries. Scientific Publications KMK, Moscow, 374 pp. (in Russian)" type="book" year="2006">Kabakov, 2006</bibRefCitation>
|
||
by the unique shape of parameres with protruding lateral lobes (
|
||
<figureCitation id="13192A4DFFDC1E39FCCAFA2F3794F9D8" box="[863,944,1533,1559]" captionStart="FIGURE 3" captionStartId="6.[151,255,1352,1376]" captionTargetBox="[193,1393,209,1293]" captionTargetId="figure@6.[151,1436,165,1337]" captionTargetPageId="6" captionText="FIGURE 3. Elements of genitalia of Onthophagus nampatensis. a—lamella copulatrix; b—lamella copulatrix, AIS and ALS removed; c—aedeagus, lateral view; d—aedeagus, apical view." httpUri="https://zenodo.org/record/193359/files/figure.png" pageId="5" pageNumber="22">Figs. 3</figureCitation>
|
||
c, d).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3386543FFDC1E39FF51F9F036DAF88D" pageId="5" pageNumber="22" type="description">
|
||
<paragraph id="8B9D36C8FFDC1E39FF51F9F036DAF88D" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
|
||
<emphasis id="B956EADAFFDC1E39FF51F9F035B9F9F3" bold="true" box="[196,413,1570,1596]" pageId="5" pageNumber="22">Taxonomic notes.</emphasis>
|
||
This species has been already cited in two papers as
|
||
<emphasis id="B956EADAFFDC1E39FB99F9F13105F9F3" box="[1036,1313,1571,1596]" italics="true" pageId="5" pageNumber="22">nomen novum in litteris</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB34B39FFDC1E39FAA5F9F0357AF9AD" author="Kabakov" pageId="5" pageNumber="28" refString="Kabakov, O. N. & Napolov, A. (1999) Fauna and ecology of Lamellicornia of subfamily Scarabaeinae of Vietnam and some parts of adjacent countries: South China, Laos, and Thailand. Latvijas Entomologs, 37, 58 - 96." type="journal article" year="1999">Kabakov & Napolov 1999</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB34B39FFDC1E39FEFFF99A3633F9AD" author="Kabakov" box="[362,535,1608,1634]" pageId="5" pageNumber="28" refString="Kabakov, O. N. (2006) Beetles of the subfamily Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) of the fauna of Russia and adjacent countries. Scientific Publications KMK, Moscow, 374 pp. (in Russian)" type="book" year="2006">Kabakov 2006</bibRefCitation>
|
||
) but has not been described. Moreover, the specimen previously indicated as the
|
||
<typeStatus id="5499886AFFDC1E39FF51F9BF3509F948" box="[196,301,1645,1671]" pageId="5" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
(
|
||
<bibRefCitation id="EFB34B39FFDC1E39FEA8F9BF35D4F948" author="Kabakov" box="[317,496,1645,1671]" pageId="5" pageNumber="28" refString="Kabakov, O. N. & Napolov, A. (1999) Fauna and ecology of Lamellicornia of subfamily Scarabaeinae of Vietnam and some parts of adjacent countries: South China, Laos, and Thailand. Latvijas Entomologs, 37, 58 - 96." type="journal article" year="1999">Kabakov 1999</bibRefCitation>
|
||
) is not included in the
|
||
<typeStatus id="5499886AFFDC1E39FC9CF9BF3719F948" box="[777,829,1645,1671]" pageId="5" pageNumber="22">type</typeStatus>
|
||
series in this paper because its aedeagus was lost. Though we are certain that the specimen previously chosen as the
|
||
<typeStatus id="5499886AFFDC1E39FC28F9403002F963" box="[957,1062,1682,1708]" pageId="5" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
is conspecific with the current
|
||
<typeStatus id="5499886AFFDC1E39FF02F96A34EEF91D" box="[151,202,1720,1746]" pageId="5" pageNumber="22">type</typeStatus>
|
||
specimens (also it was collected in the same area with the
|
||
<typeStatus id="5499886AFFDC1E39FCE2F96A37C6F91D" box="[887,994,1720,1746]" pageId="5" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
), we deliberately excluded it from the
|
||
<typeStatus id="5499886AFFDC1E39FF02F90F34E9F938" box="[151,205,1757,1783]" pageId="5" pageNumber="22">type</typeStatus>
|
||
series because the new species can be reliably separated from the close
|
||
<taxonomicName id="4C224D4BFFDC1E39FBF7F90F3121F939" box="[1122,1285,1757,1782]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="productus">
|
||
<emphasis id="B956EADAFFDC1E39FBF7F90F3121F939" box="[1122,1285,1757,1782]" italics="true" pageId="5" pageNumber="22">O. productus</emphasis>
|
||
</taxonomicName>
|
||
only by the characters of structure of parameres. For practical reasons we use the same species name as the
|
||
<emphasis id="B956EADAFFDC1E39FB62F8D131B8F8D3" box="[1271,1436,1795,1820]" italics="true" pageId="5" pageNumber="22">nomen nudum</emphasis>
|
||
in
|
||
<bibRefCitation id="EFB34B39FFDC1E39FF20F8FA35DAF88D" author="Kabakov" box="[181,510,1832,1858]" pageId="5" pageNumber="28" refString="Kabakov, O. N. & Napolov, A. (1999) Fauna and ecology of Lamellicornia of subfamily Scarabaeinae of Vietnam and some parts of adjacent countries: South China, Laos, and Thailand. Latvijas Entomologs, 37, 58 - 96." type="journal article" year="1999">Kabakov & Napolov (1999)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFB34B39FFDC1E39FDA2F8FA36DFF88D" author="Kabakov" box="[567,763,1832,1858]" pageId="5" pageNumber="28" refString="Kabakov, O. N. (2006) Beetles of the subfamily Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) of the fauna of Russia and adjacent countries. Scientific Publications KMK, Moscow, 374 pp. (in Russian)" type="book" year="2006">Kabakov (2006)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3386543FFDC1E39FF51F89F3170F843" pageId="5" pageNumber="22" type="distribution">
|
||
<paragraph id="8B9D36C8FFDC1E39FF51F89F3170F843" blockId="5.[151,1438,152,1932]" pageId="5" pageNumber="22">
|
||
<emphasis id="B956EADAFFDC1E39FF51F89F3627F8A8" bold="true" box="[196,515,1869,1895]" pageId="5" pageNumber="22">Distribution and Ecology.</emphasis>
|
||
The new species is known only from north
|
||
<collectingCountry id="F3357658FFDC1E39FB90F89F3049F8A8" box="[1029,1133,1869,1895]" name="Thailand" pageId="5" pageNumber="22">Thailand</collectingCountry>
|
||
and central
|
||
<collectingCountry id="F3357658FFDC1E39FB68F89F3112F8A8" box="[1277,1334,1869,1895]" name="Laos" pageId="5" pageNumber="22">Laos</collectingCountry>
|
||
(
|
||
<figureCitation id="13192A4DFFDC1E39FAD0F89F31AAF8A8" box="[1349,1422,1869,1895]" captionStart="FIGURE 4" captionStartId="7.[151,255,1747,1771]" captionText="FIGURE 4. Distribution map of O. streltsovi and O. nanpatensis." pageId="5" pageNumber="22">Fig. 4</figureCitation>
|
||
). In
|
||
<collectingCountry id="F3357658FFDC1E39FF23F8A034D6F843" box="[182,242,1906,1932]" name="Laos" pageId="5" pageNumber="22">Laos</collectingCountry>
|
||
, this species was collected by pitfall traps baited with human dung installed in limestone forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |