433 lines
61 KiB
XML
433 lines
61 KiB
XML
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<mods:title id="834C0EFC54C3C16ADF255E3BC45D28F7">Systematics of the Mesalina guttulata species complex (Squamata: Lacertidae) from Arabia with the description of two new species</mods:title>
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<taxonomicName id="4C224D4C5D06FFA7FF57783FDC23F9FE" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[151,358,1597,1623]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="530" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
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<emphasis id="B956EADD5D06FFA7FF57783FDC23F9FE" bold="true" box="[151,358,1597,1623]" italics="true" pageId="17" pageNumber="530">Mesalina arnoldi</emphasis>
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<taxonomicNameLabel id="A26557A65D06FFA7FEAE783FDC8EF9FE" box="[366,459,1597,1623]" pageId="17" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B9D36CF5D06FFA7FF577862DFEEF9D1" blockId="17.[151,683,1597,1657]" box="[151,683,1632,1657]" pageId="17" pageNumber="530">
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(
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<figureCitation id="13192A4A5D06FFA7FF5F7862DC4FF9D1" box="[159,266,1632,1657]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1505,1527]" captionStartId-1="5.[151,244,1878,1899]" captionStartId-2="7.[151,250,1900,1922]" captionStartId-3="8.[151,250,1951,1973]" captionTargetBox-0="[178,1390,273,1455]" captionTargetBox-1="[255,1325,222,1812]" captionTargetBox-2="[151,1434,197,1876]" captionTargetBox-3="[153,1436,945,1927]" captionTargetId-0="figure@2.[151,1436,237,1493]" captionTargetId-1="figure@5.[228,1353,195,1843]" captionTargetId-2="figure@7.[151,1436,193,1879]" captionTargetId-3="figure@8.[151,1436,941,1929]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 1. Sampling localities of the Mesalina specimens used in this study. Circles indicate samples used only in the molecular analyses, triangles indicate specimens examined and included in the morphological analyses only, and squares indicate individuals used in both molecular and morphological analyses. Colors and locality numbers correspond to Figure 2 (see also Appendix I)." captionText-1="FIGURE 2. Bayesian phylogenetic tree of the genus Mesalina based on concatenated sequences of three mitochondrial markers (12S, 16S and cytb) and one nuclear gene (MC1R). Black dots indicate posterior probability values ≥0.95 and bootstrap values ≥70% are shown next to the nodes. Color bars correspond to the five lineages recognized within the M. guttulata complex. Sample codes are followed by locality numbers (see Figure 1 and Appendix I). Taxon names correspond to changes proposed in this study and inset pictures show specimens of the two new species described (not to scale)." captionText-2="FIGURE 3. Unrooted haplotype network of the MC1R nuclear gene. Circle sizes are proportional to the number of individuals that present that particular haplotype (see Appendix I for details). White dots represent mutational steps. Colors correspond to the five lineages recognized within the M. guttulata complex." captionText-3="FIGURE 4. Results of constrained correspondence analyses (CCA) for males (A) and females (B). This plot shows the position of each species included in the multivariate analyses on the first two axes of morphological space. See material and methods for details." httpUri-0="https://zenodo.org/record/1285066/files/figure.png" httpUri-1="https://zenodo.org/record/1285068/files/figure.png" httpUri-2="https://zenodo.org/record/1285070/files/figure.png" httpUri-3="https://zenodo.org/record/1285072/files/figure.png" pageId="17" pageNumber="530">Figs. 1–4</figureCitation>
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,
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<figureCitation id="13192A4A5D06FFA7FED67862DC63F9D0" box="[278,294,1632,1657]" captionStart="FIGURE 6" captionStartId="18.[151,250,1663,1685]" captionTargetBox="[226,1360,373,1641]" captionTargetId="figure@18.[226,1360,373,1641]" captionTargetPageId="18" captionText="FIGURE 6. Pictures of the holotype of Mesalina arnoldi sp. nov. (MCCI-R890). A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen." httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="17" pageNumber="530">6</figureCitation>
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;
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<tableCitation id="C6A003745D06FFA7FEF37862DCEAF9D0" box="[307,431,1632,1657]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStart-4="TABLE 5" captionStartId-0="9.[147,233,710,734]" captionStartId-1="10.[147,220,586,607]" captionStartId-2="11.[147,220,586,607]" captionStartId-3="12.[147,223,652,674]" captionStartId-4="13.[147,223,615,639]" captionText-0="TABLE 1. Uncorrecteđ genetic đistances (p-đistances in percentage) between all Mesalina species incluđeđ in the molecular stuđy using the 12S / 16S (lower-left) anđ cytb (upper-right) mitochonđrial gene fragments. Distances among the M. guttulata species complex are highlighteđ in bolđ." captionText-1="TABLE 2. Comparison of mensural characters (means ± SE; min. anđ max. between brackets) among male Mesalina austroarabica sp. nov. (n = 6; 6 for tail length); M. sp. (n = 1); M. arnoldi sp. nov. (n = 26; 14 for tail length); M. guttulata (n = 11; 8 for tail length) anđ syntypes of M. guttulata (n = 3; 0 for tail length; after Segoli et al. 2002); M. bahaeldini (n = 5; 4 for tail length). Measurements are in percent of SVL (except SVL, heađ inđex anđ toe inđex). Maus/Mg, Maus/Mb, Ma/Mg, Ma/Mb, Maus/Ma = significance of the đifference between species pairs. Maus = M. austroarabica sp. nov., Mg = M. guttulata, Mb = M. bahaeldini, Ma = M.arnoldi sp. nov." captionText-2="TABLE 3. Comparison of mensural characters (means ± SE; min. anđ max. between brackets) among female Mesalina austroarabica sp. nov. (n = 3, 1 for tail length), M. sp. (n = 1), M. arnoldi sp. nov. (n = 17; 6 for tail length), M. guttulata (n = 7; 3 for tail length) anđ syntypes of M. guttulata (n = 2; 1 for tail length; after Segoli et al. 2002), anđ M. bahaeldini (n = 6; 3 for tail length). Measurements are in percent of SVL (except SVL, heađ inđex anđ toe inđex). Maus/Mg, Maus/Mb, Ma/Mg, Ma/Mb, Maus/Ma = significance of the đifference between species pairs. Maus = M. austroarabica sp. nov., Mg = M. guttulata, Mb = M. bahaeldini, Ma = M.arnoldi sp. nov." captionText-3="TABLE 4. Comparison of pholiđotic characters (means ± SE; min. anđ max, between brackets) among male Mesalina austroarabica sp. nov. (n = 6), M. sp. (n = 1), M. arnoldi sp. nov. (n = 26), M. guttulata (n = 11) anđ syntypes of M. guttulata (n = 3; after Segoli et al. 2002), M. bahaeldini (n = 5). Maus/Mg, Maus/Mb, Ma/Mg, Ma/Mb, Maus/Ma = significance of the đifference between species pairs. Maus = M. austroarabica sp. nov., Ma = M. arnoldi sp. nov., Mg = M. guttulata, Mb = M. bahaeldini." captionText-4="TABLE 5. Comparison of pholidotic characters (means ± SE; min. and max. between brackets) among female Mesalina austroarabica sp. nov. (n = 3), M. sp. (n = 1), M. arnoldi sp. nov. (n = 17), M. guttulata (n = 7) and syntypes of M. guttulata (n = 2; after Segoli et al. 2002), M. bahaeldini (n = 6). Maus/Mg, Maus/Mb, Ma/Mg, Ma/Mb, Maus/Ma = significance of the difference between species pairs. Maus = M. austroarabica sp. nov., Ma = M. arnoldi sp. nov., Mg = M. guttulata, Mb = M. bahaeldini." pageId="17" pageNumber="530">Tables 1–5</tableCitation>
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, Appendices I and III)
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</paragraph>
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<taxonomicName id="4C224D4C5D06FFA7FF5778AADDB0F917" authorityName=", Boulenger" authorityYear="1921" box="[151,245,1704,1726]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="530" phylum="Chordata" rank="genus">
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<emphasis id="B956EADD5D06FFA7FF5778AADDB0F917" box="[151,245,1704,1726]" italics="true" pageId="17" pageNumber="530">Mesalina</emphasis>
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</taxonomicName>
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sp. A
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<treatmentCitation id="0A8310DE5D06FFA7FE8078AADF4DF917" author="" box="[320,520,1704,1726]" page="427" pageId="17" pageNumber="530">
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<bibRefCitation id="EFB34B3E5D06FFA7FE8078AADC96F917" box="[320,467,1704,1726]" pageId="17" pageNumber="530" refId="ref12157" refString="Arnold, E. N. (1986 a) A key and annotated checklist to the lizards and amphisbaenians of Arabia. Fauna of Saudi Arabia, 8, 385 - 435." type="journal article">Arnold 1986a</bibRefCitation>
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: 427
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</treatmentCitation>
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,
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<treatmentCitation id="0A8310DE5D06FFA7FDD578AADE1AF917" author="" box="[533,863,1704,1726]" page="371" pageId="17" pageNumber="530">
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<bibRefCitation id="EFB34B3E5D06FFA7FDD578AADE6CF917" box="[533,809,1704,1726]" pageId="17" pageNumber="530" refId="ref14596" refString="Schatti, B. & Gasperetti, J. (1994) A contribution to the herpetofauna of Southwest Arabia. Fauna of Saudi Arabia, 14, 348 - 423." type="journal article">Schätti & Gasperetti 1994</bibRefCitation>
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: 371
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</treatmentCitation>
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;
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<taxonomicName id="4C224D4C5D06FFA7FCAC78AAD8D9F917" authority="Sindaco & Jeremcenko 2008: 262" authorityName="Sindaco & Jeremcenko" authorityPageNumber="262" authorityYear="2008" box="[876,1436,1704,1726]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="530" phylum="Chordata" rank="species" species="guttulata">
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<emphasis id="B956EADD5D06FFA7FCAC78AAD975F917" box="[876,1072,1704,1726]" italics="true" pageId="17" pageNumber="530">Mesalina guttulata</emphasis>
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<treatmentCitation id="0A8310DE5D06FFA7FBF978AAD8D9F917" author="" box="[1081,1436,1704,1726]" page="262" pageId="17" pageNumber="530">
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<bibRefCitation id="EFB34B3E5D06FFA7FBF978AAD82CF917" box="[1081,1385,1704,1726]" pageId="17" pageNumber="530" refId="ref14901" refString="Sindaco, R. & Jeremcenko, V. K. (2008) The reptiles of the Western Palearctic. Annotated checklist and distributional atlas of the turtles, crocodiles, amphisbaenians and lizards of Europe, North Africa, Middle East and Central Asia. Monografie della Societas Herpetologica Italica, I, 1 - 579." type="book chapter">Sindaco & Jeremcenko 2008</bibRefCitation>
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||
: 262
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</treatmentCitation>
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</taxonomicName>
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(part.).
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</paragraph>
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<subSubSection id="C33865445D06FFA4FF57790BD9CCFF7D" lastPageId="18" lastPageNumber="531" pageId="17" pageNumber="530" type="materials_examined">
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<paragraph id="8B9D36CF5D06FFA7FF57790BDC12F8C3" blockId="17.[151,1437,1801,2006]" pageId="17" pageNumber="530">
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<materialsCitation id="3B4A3C925D06FFA7FF57790BDC12F8C3" ID-GBIF-Occurrence="1848580459" collectingDate="1998-02-07" collectionCode="MCCI-R" collectorName="Sindaco & C. Sindaco" country="Yemen" county="Adult" elevation="2700" latitude="15.51" location="Hababah" longLatPrecision="771" longitude="43.86" municipality="Zakatin village" pageId="17" pageNumber="530" specimenCode="MCCI-R890" specimenCount="1" stateProvince="Amran" typeStatus="holotype">
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<typeStatus id="5499886D5D06FFA7FF57790BDC41F88B" box="[151,260,1801,1826]" pageId="17" pageNumber="530" type="holotype">
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<emphasis id="B956EADD5D06FFA7FF57790BDC41F88B" bold="true" box="[151,260,1801,1826]" pageId="17" pageNumber="530">Holotype</emphasis>
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</typeStatus>
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.
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<collectingCounty id="62FC4E435D06FFA7FED4790BDC11F88B" box="[276,340,1801,1826]" pageId="17" pageNumber="530">Adult</collectingCounty>
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female
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<specimenCode id="DB849EB45D06FFA7FE717908DF07F88B" box="[433,578,1802,1826]" collectionCode="MCCI-R" pageId="17" pageNumber="530">MCCI-R890</specimenCode>
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,
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<collectingCountry id="F335765F5D06FFA7FD8E7908DFDAF888" box="[590,671,1802,1825]" name="Yemen" pageId="17" pageNumber="530">Yemen</collectingCountry>
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,
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<collectingRegion id="49E6F82D5D06FFA7FD6C7908DFBFF88B" box="[684,762,1802,1826]" country="Yemen" name="'Amran" pageId="17" pageNumber="530">Amran</collectingRegion>
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Governatorate, plateau between
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<collectingMunicipality id="6BF9ACB55D06FFA7FBB3790BD858F88B" box="[1139,1309,1801,1826]" pageId="17" pageNumber="530">Zakatin village</collectingMunicipality>
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(
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) to
|
||
<location id="8EFD60145D06FFA7FF74792FDC69F8EF" LSID="urn:lsid:plazi:treatment:038B87D95D06FFA5FF57783FD80AF84C:8EFD60145D06FFA7FF74792FDC69F8EF" box="[180,300,1837,1862]" country="Yemen" county="Adult" latitude="15.51" longLatPrecision="771" longitude="43.86" municipality="Zakatin village" name="Kawkaban" pageId="17" pageNumber="530" stateProvince="Amran">Kawkaban</location>
|
||
(
|
||
<location id="8EFD60145D06FFA7FEFA792CDCEAF8EF" LSID="urn:lsid:plazi:treatment:038B87D95D06FFA5FF57783FD80AF84C:8EFD60145D06FFA7FEFA792CDCEAF8EF" box="[314,431,1838,1862]" country="Yemen" county="Adult" latitude="15.51" longLatPrecision="771" longitude="43.86" municipality="Zakatin village" name="Haraz Mt." pageId="17" pageNumber="530" stateProvince="Amran">Haraz Mt.</location>
|
||
) (about
|
||
<geoCoordinate id="EE1650085D06FFA7FDCC792FDF29F8EF" box="[524,620,1837,1862]" direction="north" orientation="latitude" pageId="17" pageNumber="530" precision="555" value="15.51">15.51°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1650085D06FFA7FDB8792FDF96F8EF" box="[632,723,1837,1862]" direction="east" orientation="longitude" pageId="17" pageNumber="530" precision="555" value="43.86">43.86°E</geoCoordinate>
|
||
WGS84),
|
||
<elevation id="000FD1FC5D06FFA7FC8C792FD968F8EF" box="[844,1069,1837,1862]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="17" pageNumber="530" unit="m" value="2700.0" valueMax="2800.0" valueMin="2600.0">
|
||
<quantity id="4CDA9B2A5D06FFA7FC8C792FDEB5F8EC" box="[844,1008,1837,1862]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="17" pageNumber="530" unit="m" value="2700.0" valueMax="2800.0" valueMin="2600.0">2,600-2,800 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, R.
|
||
<collectorName id="26D753195D06FFA7FB9A792FD9F1F8EF" box="[1114,1204,1837,1862]" pageId="17" pageNumber="530">Sindaco</collectorName>
|
||
and
|
||
<collectorName id="26D753195D06FFA7FB2A792CD820F8EF" box="[1258,1381,1837,1862]" pageId="17" pageNumber="530">C. Sindaco</collectorName>
|
||
leg.,
|
||
<collectingDate id="EFD8E9E75D06FFA7FF577950DC16F8C3" box="[151,339,1873,1898]" pageId="17" pageNumber="530" value="1998-02-07">
|
||
<date id="FF9C100F5D06FFA7FF577950DC16F8C3" box="[151,339,1873,1898]" pageId="17" pageNumber="530" value="1998-02-07">7 February 1998</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9D36CF5D06FFA7FF077977D923F81B" blockId="17.[151,1437,1801,2006]" pageId="17" pageNumber="530">
|
||
<materialsCitation id="3B4A3C925D06FFA7FF077977D923F81B" ID-GBIF-Occurrence="1848580458" collectingDate="1984-01-31" collectorName="Poggesi & Borri & Manetti & M. Sammicheli" country="Yemen" latitude="15.5" location="Kawkaban" longLatPrecision="771" longitude="43.9" municipality="Adult" pageId="17" pageNumber="530" specimenCount="1" stateProvince="Al Mahwit" typeStatus="paratype">
|
||
<typeStatus id="5499886D5D06FFA7FF077977DC71F827" box="[199,308,1909,1934]" pageId="17" pageNumber="530" type="paratype">
|
||
<emphasis id="B956EADD5D06FFA7FF077977DC71F827" bold="true" box="[199,308,1909,1934]" pageId="17" pageNumber="530">Paratype</emphasis>
|
||
</typeStatus>
|
||
.
|
||
<collectingMunicipality id="6BF9ACB55D06FFA7FE837977DCC6F827" box="[323,387,1909,1934]" pageId="17" pageNumber="530">Adult</collectingMunicipality>
|
||
male MZUF-28670,
|
||
<collectingCountry id="F335765F5D06FFA7FDB57974DF83F824" box="[629,710,1910,1933]" name="Yemen" pageId="17" pageNumber="530">Yemen</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E6F82D5D06FFA7FD127977DE08F827" box="[722,845,1909,1934]" country="Yemen" name="Al Mahwit" pageId="17" pageNumber="530">Al Mahwit</collectingRegion>
|
||
Governatorate,
|
||
<location id="8EFD60145D06FFA7FBC47977D939F827" LSID="urn:lsid:plazi:treatment:038B87D95D06FFA5FF57783FD80AF84C:8EFD60145D06FFA7FBC47977D939F827" box="[1028,1148,1909,1934]" country="Yemen" latitude="15.5" longLatPrecision="771" longitude="43.9" municipality="Adult" name="Kawkaban" pageId="17" pageNumber="530" stateProvince="Al Mahwit">Kawkaban</location>
|
||
(about
|
||
<geoCoordinate id="EE1650085D06FFA7FB147977D870F827" box="[1236,1333,1909,1934]" direction="north" orientation="latitude" pageId="17" pageNumber="530" precision="555" value="15.5">15.50°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1650085D06FFA7FA817974D8D9F827" box="[1345,1436,1910,1934]" direction="east" orientation="longitude" pageId="17" pageNumber="530" precision="555" value="43.9">43.90°E</geoCoordinate>
|
||
WGS84), M.
|
||
<collectorName id="26D753195D06FFA7FEEF799BDCCEF81B" box="[303,395,1945,1970]" pageId="17" pageNumber="530">Poggesi</collectorName>
|
||
, M.
|
||
<collectorName id="26D753195D06FFA7FE7D799BDCBEF81B" box="[445,507,1945,1970]" pageId="17" pageNumber="530">Borri</collectorName>
|
||
, M.
|
||
<collectorName id="26D753195D06FFA7FDED799BDFC1F81B" box="[557,644,1945,1970]" pageId="17" pageNumber="530">Manetti</collectorName>
|
||
and
|
||
<collectorName id="26D753195D06FFA7FD7A7998DE22F81B" box="[698,871,1945,1970]" pageId="17" pageNumber="530">M. Sammicheli</collectorName>
|
||
leg.,
|
||
<collectingDate id="EFD8E9E75D06FFA7FC657998D924F81B" box="[933,1121,1946,1970]" pageId="17" pageNumber="530" value="1984-01-31">
|
||
<date id="FF9C100F5D06FFA7FC657998D924F81B" box="[933,1121,1946,1970]" pageId="17" pageNumber="530" value="1984-01-31">31 January 1984</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9D36CF5D06FFA4FF0779BFD9CCFF7D" blockId="17.[151,1437,1801,2006]" lastBlockId="18.[151,1436,151,320]" lastPageId="18" lastPageNumber="531" pageId="17" pageNumber="530">
|
||
<emphasis id="B956EADD5D06FFA7FF0779BFDF51F87F" bold="true" box="[199,532,1981,2006]" pageId="17" pageNumber="530">Other specimens examined</emphasis>
|
||
. Forty-four specimens in the collections of the Natural History Museum in London and in the Museum “La Specola” in Florence (see Appendix I) plus four specimens used only for genetic analyses (no vouchers available, juvenile or damaged specimens); all listed in Appendix I.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33865445D05FFA4FF077EDDD9BCFEE9" pageId="18" pageNumber="531" type="etymology">
|
||
<paragraph id="8B9D36CF5D05FFA4FF077EDDD9BCFEE9" blockId="18.[151,1436,151,320]" pageId="18" pageNumber="531">
|
||
<emphasis id="B956EADD5D05FFA4FF077EDDDC03FF51" bold="true" box="[199,326,223,248]" pageId="18" pageNumber="531">Etymology</emphasis>
|
||
. The species epithet “
|
||
<taxonomicName id="4C224D4C5D05FFA4FD8A7EDDDFD8FF51" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[586,669,223,248]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="531" phylum="Chordata" rank="species" species="arnoldi">
|
||
<emphasis id="B956EADD5D05FFA4FD8A7EDDDFD8FF51" box="[586,669,223,248]" italics="true" pageId="18" pageNumber="531">arnoldi</emphasis>
|
||
</taxonomicName>
|
||
” is a genitive Latin noun to honor the British herpetologist Dr E. Nicholas Arnold for his life-long dedication and contribution to Arabian herpetology, including the recognition of this taxon as a distinct species that he provisionally referred to as
|
||
<taxonomicName id="4C224D4C5D05FFA4FCB67F25DE9AFEE9" authorityName=", Boulenger" authorityYear="1921" box="[886,991,295,320]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="531" phylum="Chordata" rank="genus">
|
||
<emphasis id="B956EADD5D05FFA4FCB67F25DE9AFEE9" box="[886,991,295,320]" italics="true" pageId="18" pageNumber="531">Mesalina</emphasis>
|
||
</taxonomicName>
|
||
sp. A in
|
||
<bibRefCitation id="EFB34B3E5D05FFA4FB857F25D9B0FEE9" box="[1093,1269,295,320]" pageId="18" pageNumber="531" refId="ref12157" refString="Arnold, E. N. (1986 a) A key and annotated checklist to the lizards and amphisbaenians of Arabia. Fauna of Saudi Arabia, 8, 385 - 435." type="journal article">Arnold (1986a)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5D66475D05FFA4FF57787DDC8CF97B" httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="18" pageNumber="531" startId="18.[151,250,1663,1685]" targetBox="[226,1360,373,1641]" targetPageId="18">
|
||
<paragraph id="8B9D36CF5D05FFA4FF57787DDC8CF97B" blockId="18.[151,1436,1662,1746]" pageId="18" pageNumber="531">
|
||
<emphasis id="B956EADD5D05FFA4FF57787DDEB8F93C" bold="true" box="[151,1021,1662,1685]" pageId="18" pageNumber="531">
|
||
FIGURE 6. Pictures of the holotype of
|
||
<taxonomicName id="4C224D4C5D05FFA4FD86787CDFBEF93D" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[582,763,1662,1684]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="531" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
|
||
<emphasis id="B956EADD5D05FFA4FD86787CDFBEF93D" bold="true" box="[582,763,1662,1684]" italics="true" pageId="18" pageNumber="531">Mesalina arnoldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D05FFA4FCC4787DDE13F93C" box="[772,854,1663,1685]" pageId="18" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(MCCI-R890).
|
||
</emphasis>
|
||
A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33865445D05FFA5FF0778FDDF99FCF6" lastPageId="19" lastPageNumber="532" pageId="18" pageNumber="531" type="diagnosis">
|
||
<paragraph id="8B9D36CF5D05FFA5FF0778FDDCD3FEB4" blockId="18.[151,1437,1791,2035]" lastBlockId="19.[151,1437,151,2021]" lastPageId="19" lastPageNumber="532" pageId="18" pageNumber="531">
|
||
<emphasis id="B956EADD5D05FFA4FF0778FDDC7FF8B1" bold="true" box="[199,314,1791,1816]" pageId="18" pageNumber="531">Diagnosis</emphasis>
|
||
. A relatively large-sized
|
||
<taxonomicName id="4C224D4C5D05FFA4FDA378FDDF8EF8B1" authorityName=", Boulenger" authorityYear="1921" box="[611,715,1791,1816]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="531" phylum="Chordata" rank="genus">
|
||
<emphasis id="B956EADD5D05FFA4FDA378FDDF8EF8B1" box="[611,715,1791,1816]" italics="true" pageId="18" pageNumber="531">Mesalina</emphasis>
|
||
</taxonomicName>
|
||
characterized by the following combination of morphological characters: (1) well-developed occipital scale in contact with the interparietal (with rare exceptions) (
|
||
<figureCitation id="13192A4A5D05FFA4FACE7921D823F895" box="[1294,1382,1827,1852]" captionStart="FIGURE 6" captionStartId="18.[151,250,1663,1685]" captionTargetBox="[226,1360,373,1641]" captionTargetId="figure@18.[226,1360,373,1641]" captionTargetPageId="18" captionText="FIGURE 6. Pictures of the holotype of Mesalina arnoldi sp. nov. (MCCI-R890). A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen." httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="18" pageNumber="531">Fig. 6E</figureCitation>
|
||
); (2) lower eyelid with a window made of up two large scales (in 57% of examined specimens) or fragmented into smaller scales (43%) (
|
||
<figureCitation id="13192A4A5D05FFA4FE507969DCACF82D" box="[400,489,1899,1924]" captionStart="FIGURE 6" captionStartId="18.[151,250,1663,1685]" captionTargetBox="[226,1360,373,1641]" captionTargetId="figure@18.[226,1360,373,1641]" captionTargetPageId="18" captionText="FIGURE 6. Pictures of the holotype of Mesalina arnoldi sp. nov. (MCCI-R890). A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen." httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="18" pageNumber="531">Fig. 6D</figureCitation>
|
||
), often without black edges (67%); (3) curved collar (
|
||
<figureCitation id="13192A4A5D05FFA4FB857969D9DFF82D" box="[1093,1178,1899,1924]" captionStart="FIGURE 6" captionStartId="18.[151,250,1663,1685]" captionTargetBox="[226,1360,373,1641]" captionTargetId="figure@18.[226,1360,373,1641]" captionTargetPageId="18" captionText="FIGURE 6. Pictures of the holotype of Mesalina arnoldi sp. nov. (MCCI-R890). A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen." httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="18" pageNumber="531">Fig. 6F</figureCitation>
|
||
); (4) four upper labials in front of the subocular in 89% of the samples and five in 11% of the samples (
|
||
<figureCitation id="13192A4A5D05FFA4FBE9798DD9C1F801" box="[1065,1156,1935,1960]" captionStart="FIGURE 6" captionStartId="18.[151,250,1663,1685]" captionTargetBox="[226,1360,373,1641]" captionTargetId="figure@18.[226,1360,373,1641]" captionTargetPageId="18" captionText="FIGURE 6. Pictures of the holotype of Mesalina arnoldi sp. nov. (MCCI-R890). A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen." httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="18" pageNumber="531">Fig. 6D</figureCitation>
|
||
); (5) ventral plates in 10 (very rarely 8) straight longitudinal rows, the outermost much smaller (
|
||
<figureCitation id="13192A4A5D05FFA4FC7879B1D955F865" box="[952,1040,1971,1996]" captionStart="FIGURE 6" captionStartId="18.[151,250,1663,1685]" captionTargetBox="[226,1360,373,1641]" captionTargetId="figure@18.[226,1360,373,1641]" captionTargetPageId="18" captionText="FIGURE 6. Pictures of the holotype of Mesalina arnoldi sp. nov. (MCCI-R890). A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen." httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="18" pageNumber="531">Fig. 6B</figureCitation>
|
||
); (6) scales on the upper surface of the tibia keeled (
|
||
<figureCitation id="13192A4A5D05FFA4FE9379D8DCE8F85A" box="[339,429,2010,2035]" captionStart="FIGURE 6" captionStartId="18.[151,250,1663,1685]" captionTargetBox="[226,1360,373,1641]" captionTargetId="figure@18.[226,1360,373,1641]" captionTargetPageId="18" captionText="FIGURE 6. Pictures of the holotype of Mesalina arnoldi sp. nov. (MCCI-R890). A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen." httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="18" pageNumber="531">Fig. 6A</figureCitation>
|
||
); (7) lamellae under 4th toe, 19–26 (median = 22); (8) dorsal pattern usually very marked, background color brown-greyish, with many complete ocelli (i.e. a white spot completely surrounded by a black ring) or near so, ordered in irregular longitudinal and transverse rows. Dorsolateral and light stripes are usually evident, often interrupted; some specimens are clearly striped, while in others these lines are inconspicuous, only rarely absent (
|
||
<figureCitation id="13192A4A5D04FFA5FEF67F06DCCBFEB4" box="[310,398,260,285]" captionStart="FIGURE 6" captionStartId="18.[151,250,1663,1685]" captionTargetBox="[226,1360,373,1641]" captionTargetId="figure@18.[226,1360,373,1641]" captionTargetPageId="18" captionText="FIGURE 6. Pictures of the holotype of Mesalina arnoldi sp. nov. (MCCI-R890). A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen." httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="19" pageNumber="532">Fig. 6A</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9D36CF5D04FFA5FF077F25DE7FFD96" blockId="19.[151,1437,151,2021]" pageId="19" pageNumber="532">
|
||
There are no obvious morphological characters separating
|
||
<taxonomicName id="4C224D4C5D04FFA5FC507F2BD94AFEE9" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[912,1039,295,320]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FC507F2BD94AFEE9" box="[912,1039,295,320]" italics="true" pageId="19" pageNumber="532">M. arnoldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FBDF7F25D9C4FEE9" box="[1055,1153,295,320]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FBDF7F25D9C4FEE9" bold="true" box="[1055,1153,295,320]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from
|
||
<taxonomicName id="4C224D4C5D04FFA5FB157F2BD822FEE9" authorityName="Sindaco & Jeremcenko" authorityYear="2008" box="[1237,1383,295,320]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="guttulata">
|
||
<emphasis id="B956EADD5D04FFA5FB157F2BD822FEE9" box="[1237,1383,295,320]" italics="true" pageId="19" pageNumber="532">M. guttulata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C224D4C5D04FFA5FABD7F2BDC4AFECC" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="bahaeldini">
|
||
<emphasis id="B956EADD5D04FFA5FABD7F2BDC4AFECC" italics="true" pageId="19" pageNumber="532">M. bahaeldini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C224D4C5D04FFA5FE857F4FDF4DFECC" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[325,520,332,357]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="austroarabica" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FE857F4FDF4DFECC" box="[325,520,332,357]" italics="true" pageId="19" pageNumber="532">M. austroarabica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FDCF7F4EDF2CFECC" box="[527,617,332,357]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FDCF7F4EDF2CFECC" bold="true" box="[527,617,332,357]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
The statistical analyses (see Results above) show significant differences, with
|
||
<taxonomicName id="4C224D4C5D04FFA5FF107F73DC0DFE21" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[208,328,367,392]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FF107F73DC0DFE21" box="[208,328,367,392]" italics="true" pageId="19" pageNumber="532">M. arnoldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FE8E7F6DDCEDFE21" box="[334,424,367,392]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FE8E7F6DDCEDFE21" bold="true" box="[334,424,367,392]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
having more gulars (males), more dorsals at midbody (males), more femoral pores (males and females) than
|
||
<taxonomicName id="4C224D4C5D04FFA5FEA87F9ADCB1FE06" authorityName="Sindaco & Jeremcenko" authorityYear="2008" box="[360,500,406,431]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="guttulata">
|
||
<emphasis id="B956EADD5D04FFA5FEA87F9ADCB1FE06" box="[360,500,406,431]" italics="true" pageId="19" pageNumber="532">M. guttulata</emphasis>
|
||
</taxonomicName>
|
||
. Moreover,
|
||
<taxonomicName id="4C224D4C5D04FFA5FD407F9ADFBCFE06" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[640,761,406,431]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FD407F9ADFBCFE06" box="[640,761,406,431]" italics="true" pageId="19" pageNumber="532">M. arnoldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FCC17F94DE19FE06" box="[769,860,406,431]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FCC17F94DE19FE06" bold="true" box="[769,860,406,431]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has smaller %HD (males), smaller %4th toe length (males), smaller toe-index (males), lesser value of Lamellae percSVL (males).
|
||
<taxonomicName id="4C224D4C5D04FFA5FBE37FB8D9A2FE7A" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[1059,1255,442,467]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FBE37FB8D9A2FE7A" box="[1059,1255,442,467]" italics="true" pageId="19" pageNumber="532">Mesalina arnoldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FB317FB8D808FE7A" box="[1265,1357,442,467]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FB317FB8D808FE7A" bold="true" box="[1265,1357,442,467]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
shows significant differences from
|
||
<taxonomicName id="4C224D4C5D04FFA5FE147FE2DF37FE5E" box="[468,626,478,503]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="bahaeldini">
|
||
<emphasis id="B956EADD5D04FFA5FE147FE2DF37FE5E" box="[468,626,478,503]" italics="true" pageId="19" pageNumber="532">M. bahaeldini</emphasis>
|
||
</taxonomicName>
|
||
in having more gulars (males and females), more femoral pores (males) and more supralabials (females). Differences between
|
||
<taxonomicName id="4C224D4C5D04FFA5FCD57C06DED7FDB2" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[789,914,514,539]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FCD57C06DED7FDB2" box="[789,914,514,539]" italics="true" pageId="19" pageNumber="532">M. arnoldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FC5E7C00DEB9FDB2" box="[926,1020,514,539]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FC5E7C00DEB9FDB2" bold="true" box="[926,1020,514,539]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C224D4C5D04FFA5FBFC7C06D841FDB2" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[1084,1284,514,539]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="austroarabica" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FBFC7C06D841FDB2" box="[1084,1284,514,539]" italics="true" pageId="19" pageNumber="532">M. austroarabica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FAD07C00D82BFDB2" box="[1296,1390,514,539]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FAD07C00D82BFDB2" bold="true" box="[1296,1390,514,539]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are discussed in the description of the latter species (see above).
|
||
</paragraph>
|
||
<paragraph id="8B9D36CF5D04FFA5FF077C48DF99FCF6" blockId="19.[151,1437,151,2021]" pageId="19" pageNumber="532">
|
||
<emphasis id="B956EADD5D04FFA5FF077C48DF33FDCA" bold="true" box="[199,630,586,611]" pageId="19" pageNumber="532">Genetic and phylogenetic remarks</emphasis>
|
||
. This species had not been included in any previous phylogenetic analyses, not even the comprehensive study by
|
||
<bibRefCitation id="EFB34B3E5D04FFA5FD7A7C6CDED6FD2E" box="[698,915,622,647]" pageId="19" pageNumber="532" refId="ref13499" refString="Kapli, P., Lymberakis, P., Crochet, P., Geniez, P., Brito, J. C., Almutairi, M., Ahmadzadeh, F., Schmitz, A., Wilms, T. & Pouyani, N. R. (2015) Historical biogeography of the lacertid lizard Mesalina in North Africa and the Middle East. Journal of Biogeography, 42 (2), 267 - 279. https: // doi. org / 10.1111 / jbi. 12420" type="journal article">
|
||
Kapli
|
||
<emphasis id="B956EADD5D04FFA5FCC27C72DE70FD2E" box="[770,821,622,647]" italics="true" pageId="19" pageNumber="532">et al</emphasis>
|
||
. (2015)
|
||
</bibRefCitation>
|
||
. The phylogenetic analyses performed in this study (
|
||
<figureCitation id="13192A4A5D04FFA5FF267C90DC6AFD02" box="[230,303,658,683]" captionStart="FIGURE 2" captionStartId="5.[151,244,1878,1899]" captionTargetBox="[255,1325,222,1812]" captionTargetId="figure@5.[228,1353,195,1843]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian phylogenetic tree of the genus Mesalina based on concatenated sequences of three mitochondrial markers (12S, 16S and cytb) and one nuclear gene (MC1R). Black dots indicate posterior probability values ≥0.95 and bootstrap values ≥70% are shown next to the nodes. Color bars correspond to the five lineages recognized within the M. guttulata complex. Sample codes are followed by locality numbers (see Figure 1 and Appendix I). Taxon names correspond to changes proposed in this study and inset pictures show specimens of the two new species described (not to scale)." httpUri="https://zenodo.org/record/1285068/files/figure.png" pageId="19" pageNumber="532">Fig. 2</figureCitation>
|
||
;
|
||
<tableCitation id="C6A003745D04FFA5FEFE7C90DCD7FD02" box="[318,402,658,683]" captionStart="TABLE 1" captionStartId="9.[147,233,710,734]" captionText="TABLE 1. Uncorrecteđ genetic đistances (p-đistances in percentage) between all Mesalina species incluđeđ in the molecular stuđy using the 12S / 16S (lower-left) anđ cytb (upper-right) mitochonđrial gene fragments. Distances among the M. guttulata species complex are highlighteđ in bolđ." pageId="19" pageNumber="532">Table 1</tableCitation>
|
||
) support the hypothesis that
|
||
<taxonomicName id="4C224D4C5D04FFA5FD2A7C96DE21FD02" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[746,868,658,683]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FD2A7C96DE21FD02" box="[746,868,658,683]" italics="true" pageId="19" pageNumber="532">M. arnoldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FCAF7C90DE89FD02" box="[879,972,658,683]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FCAF7C90DE89FD02" bold="true" box="[879,972,658,683]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is an independent species. The level of genetic differentiation (
|
||
<emphasis id="B956EADD5D04FFA5FE677CBADCF0FD66" box="[423,437,696,719]" italics="true" pageId="19" pageNumber="532">p</emphasis>
|
||
-distance) between the new species and the other members of the
|
||
<taxonomicName id="4C224D4C5D04FFA5FB037CB4D8D9FD66" authorityName="Sindaco & Jeremcenko" authorityYear="2008" box="[1219,1436,694,719]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="guttulata">
|
||
<emphasis id="B956EADD5D04FFA5FB037CB4D8D9FD66" box="[1219,1436,694,719]" italics="true" pageId="19" pageNumber="532">Mesalina guttulata</emphasis>
|
||
</taxonomicName>
|
||
species complex ranges between 5.2–6.6% in the
|
||
<emphasis id="B956EADD5D04FFA5FD1D7CD9DE48FD5A" box="[733,781,731,755]" italics="true" pageId="19" pageNumber="532">12S,</emphasis>
|
||
6.1–7.1% in the
|
||
<emphasis id="B956EADD5D04FFA5FC1C7CD8D943FD5A" box="[988,1030,730,755]" italics="true" pageId="19" pageNumber="532">16S</emphasis>
|
||
and 11.9–15.7% in the
|
||
<taxonomicName id="4C224D4C5D04FFA5FAE37CD8D814FD5A" box="[1315,1361,730,755]" class="Reptilia" family="Lacertidae" genus="Sauria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="cytb">
|
||
<emphasis id="B956EADD5D04FFA5FAE37CD8D814FD5A" box="[1315,1361,730,755]" italics="true" pageId="19" pageNumber="532">cytb</emphasis>
|
||
</taxonomicName>
|
||
genes (
|
||
<tableCitation id="C6A003745D04FFA5FF5E7CFCDDB7FCBE" box="[158,242,766,791]" captionStart="TABLE 1" captionStartId="9.[147,233,710,734]" captionText="TABLE 1. Uncorrecteđ genetic đistances (p-đistances in percentage) between all Mesalina species incluđeđ in the molecular stuđy using the 12S / 16S (lower-left) anđ cytb (upper-right) mitochonđrial gene fragments. Distances among the M. guttulata species complex are highlighteđ in bolđ." pageId="19" pageNumber="532">Table 1</tableCitation>
|
||
). A network analysis of the nuclear gene
|
||
<emphasis id="B956EADD5D04FFA5FD0A7CFDDE56FCBE" box="[714,787,767,791]" italics="true" pageId="19" pageNumber="532">MC1R</emphasis>
|
||
indicates that, despite the large number of samples of the
|
||
<taxonomicName id="4C224D4C5D04FFA5FF577D26DC64FC92" authorityName="Sindaco & Jeremcenko" authorityYear="2008" box="[151,289,802,827]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="guttulata">
|
||
<emphasis id="B956EADD5D04FFA5FF577D26DC64FC92" box="[151,289,802,827]" italics="true" pageId="19" pageNumber="532">M. guttulata</emphasis>
|
||
</taxonomicName>
|
||
species complex included in the analysis (
|
||
<specimenCount id="9D24FD465D04FFA5FD3E7D20DEDFFC92" box="[766,922,802,827]" pageId="19" pageNumber="532" type="generic">36 specimens</specimenCount>
|
||
; 72 alleles), all four haplotypes (10 alleles) of
|
||
<taxonomicName id="4C224D4C5D04FFA5FF577D4ADC4BFCF6" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[151,270,838,863]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FF577D4ADC4BFCF6" box="[151,270,838,863]" italics="true" pageId="19" pageNumber="532">M. arnoldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FED57D44DC2AFCF6" box="[277,367,838,863]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FED57D44DC2AFCF6" bold="true" box="[277,367,838,863]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are private (
|
||
<figureCitation id="13192A4A5D04FFA5FE3C7D44DF04FCF6" box="[508,577,838,863]" captionStart="FIGURE 3" captionStartId="7.[151,250,1900,1922]" captionTargetBox="[151,1434,197,1876]" captionTargetId="figure@7.[151,1436,193,1879]" captionTargetPageId="7" captionText="FIGURE 3. Unrooted haplotype network of the MC1R nuclear gene. Circle sizes are proportional to the number of individuals that present that particular haplotype (see Appendix I for details). White dots represent mutational steps. Colors correspond to the five lineages recognized within the M. guttulata complex." httpUri="https://zenodo.org/record/1285070/files/figure.png" pageId="19" pageNumber="532">Fig. 3</figureCitation>
|
||
; Appendix I).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33865445D04FFA5FF077D68DF36F8FC" pageId="19" pageNumber="532" type="description">
|
||
<paragraph id="8B9D36CF5D04FFA5FF077D68DC87FB2C" blockId="19.[151,1437,151,2021]" pageId="19" pageNumber="532">
|
||
<emphasis id="B956EADD5D04FFA5FF077D68DF49FC2A" bold="true" box="[199,524,874,899]" pageId="19" pageNumber="532">
|
||
Description of the
|
||
<typeStatus id="5499886D5D04FFA5FE657D68DF49FC2A" box="[421,524,874,899]" pageId="19" pageNumber="532" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. An adult female with partly regenerated tail. Measurements, meristic characters and indexes: SVL = 53.0 mm, HL-1 =
|
||
<quantity id="4CDA9B2A5D04FFA5FD897D8DDFEEFC0E" box="[585,683,911,935]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="19" pageNumber="532" unit="mm" value="11.7">11.7 mm</quantity>
|
||
(22% of SVL), HL-2 = 5.0 mm (9% of SVL), HL-3 =
|
||
<quantity id="4CDA9B2A5D04FFA5FAD17D8DD822FC0E" box="[1297,1383,911,935]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="19" pageNumber="532" unit="mm" value="4.3">4.3 mm</quantity>
|
||
(8% of SVL), Head width =
|
||
<quantity id="4CDA9B2A5D04FFA5FE6B7DB1DF46FC63" box="[427,515,947,971]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="19" pageNumber="532" unit="mm" value="7.3">7.3 mm</quantity>
|
||
(14% of SVL), Head depth =
|
||
<quantity id="4CDA9B2A5D04FFA5FCA57DB1DEF8FC63" box="[869,957,947,971]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="19" pageNumber="532" unit="mm" value="4.9">4.9 mm</quantity>
|
||
(9% of SVL), pileus =
|
||
<quantity id="4CDA9B2A5D04FFA5FB107DB1D873FC63" box="[1232,1334,947,971]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="19" pageNumber="532" unit="mm" value="10.4">10.4 mm</quantity>
|
||
(20% of SVL), Forelimb length =
|
||
<quantity id="4CDA9B2A5D04FFA5FE777DDBDF59FC58" box="[439,540,985,1010]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="19" pageNumber="532" unit="mm" value="16.3">16.3 mm</quantity>
|
||
(31% of SVL), Hindlimb length =
|
||
<quantity id="4CDA9B2A5D04FFA5FC6C7DDBD954FC58" box="[940,1041,985,1010]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="19" pageNumber="532" unit="mm" value="26.6">26.6 mm</quantity>
|
||
(50% of SVL), 4th toe length =
|
||
<quantity id="4CDA9B2A5D04FFA5FAB97DDBDD86FBBC" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="19" pageNumber="532" unit="mm" value="8.6">8.6 mm</quantity>
|
||
(16% of SVL), Tail length = 62.0 mm (partly regenerated), supralabials 9/9, subocular = 5/5, gulars = 23, enlarged plates in collar = 9, midbody scales = 48, longitudinal rows of ventrals = 8+2 (smaller), transversal rows of ventrals = 36, femoral pores = 14+14, lamellae under the 4th toe = 21. Head index = 160, Toe index = 32, Lamellae percSVL = 0.77.
|
||
</paragraph>
|
||
<paragraph id="8B9D36CF5D04FFA5FF077A92DDA6FA44" blockId="19.[151,1437,151,2021]" pageId="19" pageNumber="532">Coloration in alcohol: numerous ocelli, each one formed by several whitish scales forming a dot and surrounded by an almost complete ring of black colored scales (ocelli are reduced to black dots on the neck). These ocelli form 4-6 rather regular longitudinal series (the paravertebral and lateral ones more marked) and about 13 very irregular transverse series, between fore- and hindlimbs; black and white dots are present on the tail and hindlimbs. The pileus is grey without specks (only the outer margin of the parietals is bordered with black). On the sides of the head a continuous dark stripe is present from near the upper border of the ear opening, across the eye, to the loreal scale. Another well-defined dark stripe lies between the mid- ear opening and the subocular scale. Flanks with two series of ocelli, without evident stripes. The ventral side is creamy-white, immaculate, with the exception of infralabial scales, irregularly sprinkled with gray, as well as outer ventrals and the anterior margin of thighs.</paragraph>
|
||
<paragraph id="8B9D36CF5D04FFA5FF077BFADFFAF8A4" blockId="19.[151,1437,151,2021]" pageId="19" pageNumber="532">
|
||
<emphasis id="B956EADD5D04FFA5FF077BFADC70F9B8" bold="true" box="[199,309,1528,1553]" pageId="19" pageNumber="532">Variation</emphasis>
|
||
. In specimens MZUF-28132 the occipital scale is almost absent. The two lateral rows of ventrals are usually much smaller than the inner ventrals, sometimes subequal in size, and absent in specimen MZUF-28132. The dorsal pattern is very variable; specimens with the pattern similar to the
|
||
<typeStatus id="5499886D5D04FFA5FBC27842D921F9F0" box="[1026,1124,1600,1625]" pageId="19" pageNumber="532" type="holotype">holotype</typeStatus>
|
||
are frequent, but in several specimens the white dots of outer dorsal ocelli tend to form a whitish, more or less interrupted, supraciliar stripe along the sides of the back. In several specimens, instead of small ocelli, there are dark blotches on the back, parallel to the light supraciliar stripes, forming a distinct striped pattern (BM1938.8.1.27, BM1977.423). In specimens BM1977.425 and MZUF-28673 there are four uninterrupted white stripes: two supraciliar stripes and two subocular stripes along the sides of the body.
|
||
</paragraph>
|
||
<paragraph id="8B9D36CF5D04FFA5FF07791ADF36F8FC" blockId="19.[151,1437,151,2021]" pageId="19" pageNumber="532">
|
||
<emphasis id="B956EADD5D04FFA5FF07791ADCD7F898" bold="true" box="[199,402,1816,1841]" pageId="19" pageNumber="532">Coloration in life</emphasis>
|
||
. Ground color brownish with more or less intense shades of gray. Ocelli whitish surrounded by dark brown incomplete rings (
|
||
<figureCitation id="13192A4A5D04FFA5FDCC793EDF23F8FC" box="[524,614,1852,1877]" captionStart="FIGURE 6" captionStartId="18.[151,250,1663,1685]" captionTargetBox="[226,1360,373,1641]" captionTargetId="figure@18.[226,1360,373,1641]" captionTargetPageId="18" captionText="FIGURE 6. Pictures of the holotype of Mesalina arnoldi sp. nov. (MCCI-R890). A) dorsal view of the body and tail; B) ventral view, C) detail of the femoral pores; D) right side of the head; E) upper (dorsal) part of the head; F) ventral (gular) side of the head; G) live specimen." httpUri="https://zenodo.org/record/1285078/files/figure.png" pageId="19" pageNumber="532">Fig. 6G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33865445D04FFA5FF077962D80AF84C" pageId="19" pageNumber="532" type="distribution">
|
||
<paragraph id="8B9D36CF5D04FFA5FF077962D80AF84C" blockId="19.[151,1437,151,2021]" pageId="19" pageNumber="532">
|
||
<emphasis id="B956EADD5D04FFA5FF077962DCA9F8D0" bold="true" box="[199,492,1888,1913]" pageId="19" pageNumber="532">Distribution and habitat</emphasis>
|
||
. Specimens referable to
|
||
<taxonomicName id="4C224D4C5D04FFA5FCC67962DE8CF8D0" authority="Sindaco & Simó-Riudalbas & Sacchi & Carranza, 2018" authorityName="Sindaco & Simó-Riudalbas & Sacchi & Carranza" authorityYear="2018" box="[774,969,1888,1913]" class="Reptilia" family="Lacertidae" genus="Mesalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="532" phylum="Chordata" rank="species" species="arnoldi" status="sp. nov.">
|
||
<emphasis id="B956EADD5D04FFA5FCC67962DE8CF8D0" box="[774,969,1888,1913]" italics="true" pageId="19" pageNumber="532">Mesalina arnoldi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26557A65D04FFA5FC127962D969F8D0" box="[978,1068,1888,1913]" pageId="19" pageNumber="532" rank="species">
|
||
<emphasis id="B956EADD5D04FFA5FC127962D969F8D0" bold="true" box="[978,1068,1888,1913]" pageId="19" pageNumber="532">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are widespread in the highlands of southwestern
|
||
<collectingCountry id="F335765F5D04FFA5FE8D7986DC9BF834" box="[333,478,1924,1949]" name="Saudi Arabia" pageId="19" pageNumber="532">Saudi Arabia</collectingCountry>
|
||
and
|
||
<collectingRegion id="49E6F82D5D04FFA5FDD47987DF28F835" box="[532,621,1925,1948]" country="American Samoa" name="Western" pageId="19" pageNumber="532">Western</collectingRegion>
|
||
<collectingCountry id="F335765F5D04FFA5FDB47987DF81F835" box="[628,708,1925,1948]" name="Yemen" pageId="19" pageNumber="532">Yemen</collectingCountry>
|
||
. The
|
||
<typeStatus id="5499886D5D04FFA5FCC17986DE24F834" box="[769,865,1924,1949]" pageId="19" pageNumber="532" type="holotype">holotype</typeStatus>
|
||
was collected in a stony plateau with basaltic rocks and scarce vegetation, at an altitude of
|
||
<quantity id="4CDA9B2A5D04FFA5FD9379AADFB3F869" box="[595,758,1960,1985]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="19" pageNumber="532" unit="m" value="2700.0" valueMax="2800.0" valueMin="2600.0">2,600-2,800 m</quantity>
|
||
a.s.l. The
|
||
<typeStatus id="5499886D5D04FFA5FCB079ABDE8BF868" box="[880,974,1961,1985]" pageId="19" pageNumber="532" type="paratype">paratype</typeStatus>
|
||
was collected in the same area, between 1,950 and
|
||
<quantity id="4CDA9B2A5D04FFA5FEC979CFDC26F84D" box="[265,355,1997,2021]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="19" pageNumber="532" unit="m" value="2300.0">2,300 m</quantity>
|
||
a.s.l. According to
|
||
<bibRefCitation id="EFB34B3E5D04FFA5FDF979CEDE2DF84C" box="[569,872,1996,2021]" pageId="19" pageNumber="532" refId="ref14596" refString="Schatti, B. & Gasperetti, J. (1994) A contribution to the herpetofauna of Southwest Arabia. Fauna of Saudi Arabia, 14, 348 - 423." type="journal article">Schätti & Gasperetti (1994)</bibRefCitation>
|
||
this species is found as low as
|
||
<quantity id="4CDA9B2A5D04FFA5FB7D79CFD852F84D" box="[1213,1303,1997,2021]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="19" pageNumber="532" unit="m" value="1300.0">1,300 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |