149 lines
19 KiB
XML
149 lines
19 KiB
XML
<document id="824441BD4FB01F9E6B134CE48D946E16" ID-DOI="10.15298/rusentj.29.3.02" ID-ISSN="0132-8069" ID-Zenodo-Dep="13375227" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722580952034" checkinUser="felipe" docAuthor="Emeljanov, A. F. & Shcherbakov, D. E." docDate="2020" docId="038B87A2FFFFFF9CCB3BFC24AE33FCD7" docLanguage="en" docName="RussEntomolJ.29.3.237-246.pdf" docOrigin="Russian Entomological Journal 29 (3)" docSource="https://kmkjournals.com/journals/REJ/REJ_Index_Volumes/REJ_29/REJ_29_3_237_246" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Derbachile hochae Emeljanov et Shcherbakov 2020, sp.n." docType="treatment" docVersion="4" lastPageNumber="240" masterDocId="FFB2FFDAFFFEFF9FC803FFB1AF40FFE6" masterDocTitle="The first Mesozoic Derbidae (Homoptera: Fulgoroidea) from Cretaceous Burmese amber" masterLastPageNumber="246" masterPageNumber="237" pageNumber="238" updateTime="1724677411652" updateUser="ExternalLinkService">
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<mods:titleInfo id="99D608E704AA89C866AB74DD244EEB9F">
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<mods:title id="964946B333D9352CB006B3432EBB9310">The first Mesozoic Derbidae (Homoptera: Fulgoroidea) from Cretaceous Burmese amber</mods:title>
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<mods:namePart id="FEF775E2FB7556164B59BFCDB5A55E56">Emeljanov, A. F.</mods:namePart>
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<mods:roleTerm id="62EF807668629A92AF683BB7B02985A9">Author</mods:roleTerm>
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<mods:namePart id="695087770863FBD46148CABF2E8EE8D9">Shcherbakov, D. E.</mods:namePart>
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<mods:typeOfResource id="5364C62FDF02DE805CECA53B5ECD0E4C">text</mods:typeOfResource>
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<mods:title id="76EE32C2449E62544D284CDBA53E2DA5">Russian Entomological Journal</mods:title>
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<mods:part id="4974664BC03A2DAA74E0CEEE7885A052">
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<mods:date id="64AB9F204222FF62813CF8FD439F4495">2020</mods:date>
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<mods:number id="97A0A068B83B3C893102385432AE0277">2020-12-31</mods:number>
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<mods:number id="AA6F931CADE2403EA610BDFA4AA60BDE">29</mods:number>
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<mods:url id="DEAF75A88D7374C95201E6A53F91C800">https://kmkjournals.com/journals/REJ/REJ_Index_Volumes/REJ_29/REJ_29_3_237_246</mods:url>
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<mods:classification id="95FA9802B77BFB977FEFD9E31570BCBF">journal article</mods:classification>
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<mods:identifier id="7F782E185AADC908CBDC7E3939979D6D" type="DOI">10.15298/rusentj.29.3.02</mods:identifier>
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<mods:identifier id="AEA6D7950366FAB6D052EF4087B8D58A" type="ISSN">0132-8069</mods:identifier>
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<treatment id="038B87A2FFFFFF9CCB3BFC24AE33FCD7" LSID="urn:lsid:plazi:treatment:038B87A2FFFFFF9CCB3BFC24AE33FCD7" httpUri="http://treatment.plazi.org/id/038B87A2FFFFFF9CCB3BFC24AE33FCD7" lastPageId="3" lastPageNumber="240" pageId="1" pageNumber="238">
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<subSubSection id="C338653FFFFFFF9ECB3BFC24AA2BFC4B" box="[824,1387,917,941]" pageId="1" pageNumber="238" type="nomenclature">
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<paragraph id="8B9D36B4FFFFFF9ECB3BFC24AA2BFC4B" blockId="1.[812,1399,917,2064]" box="[824,1387,917,941]" pageId="1" pageNumber="238">
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<heading id="D0D581D8FFFFFF9ECB3BFC24AA2BFC4B" box="[824,1387,917,941]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="238" reason="8">
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<taxonomicName id="4C224D37FFFFFF9ECB3BFC24AA6CFC4B" ID-CoL="9F85P" authority="Emeljanov et Shcherbakov" authorityName="Emeljanov et Shcherbakov" authorityYear="2020" box="[824,1324,917,941]" class="Insecta" family="Derbidae" genus="Derbachile" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="238" phylum="Arthropoda" rank="species" species="hochae" status="sp. nov.">
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<emphasis id="B956EAA6FFFFFF9ECB3BFC24AB40FC4B" box="[824,1024,917,941]" italics="true" pageId="1" pageNumber="238">Derbachile hochae</emphasis>
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Emeljanov et Shcherbakov
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</taxonomicName>
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,
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<emphasis id="B956EAA6FFFFFF9ECD35FC24AA2BFC4B" bold="true" box="[1334,1387,917,941]" pageId="1" pageNumber="238">
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<taxonomicNameLabel id="A26557DDFFFFFF9ECD35FC24AA2BFC4B" box="[1334,1387,917,941]" pageId="1" pageNumber="238" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C338653FFFFFFF9ECC07FC03ABDFFC2C" box="[1028,1183,946,970]" pageId="1" pageNumber="238" type="description">
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<paragraph id="8B9D36B4FFFFFF9ECC07FC03ABDFFC2C" blockId="1.[812,1399,917,2064]" box="[1028,1183,946,970]" pageId="1" pageNumber="238">
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<figureCitation id="13192A31FFFFFF9ECC07FC03AB31FC2C" box="[1028,1137,946,970]" captionStart="Figs 1–10" captionStartId="2.[226,263,1897,1916]" captionTargetBox="[189,1399,318,1862]" captionTargetId="figure-179@2.[189,1399,318,1862]" captionTargetPageId="2" captionText="Figs 1–10. Derbachile hochae gen. et sp.n., holotype male, Burmese amber: 1 — habitus, dorsal; 2 — habitus, ventral; 3 — head and thorax, dorsal; 4 — head and thorax, ventral (arrows, subantennal ridges); 5 — tegmen, anterodorsal; 6–7 — C and stigmal cell: 6 — right, 7 — left tegmen; 8 — hind legs; 9 — genitalia, ventral; 10 — genitalia, dorsal. Scale bars: 0.5 mm. Рис. 1–10. Derbachile hochae gen. et sp.n., голотип, самец, бирманский Янтарь: 1 — обЩий вид, сверху; 2 — обЩий вид, сниЗу; 3 — голова и грудь, сверху; 4 — голова и грудь, сниЗу (стрелки — подусиковые кили); 5 — переднее крыло, спереди-сверху; 6–7 — C и стигмальнаЯ Ячейка: 6 — правое переднее крыло, 7 — левое переднее крыло; 8 — Задние ноги; 9 — гениталии, сниЗу; 10 — гениталии, сверху. Длина масШтабной линейки: 0,5 мм." figureDoi="http://doi.org/10.5281/zenodo.13375229" httpUri="https://zenodo.org/record/13375229/files/figure.png" pageId="1" pageNumber="238">Figs 1–10</figureCitation>
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,
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<figureCitation id="13192A31FFFFFF9ECC7EFC03ABDBFC2C" box="[1149,1179,946,970]" captionStart="Figs 25–28" captionStartId="7.[226,263,1969,1988]" captionTargetBox="[189,1399,1354,1939]" captionTargetId="figure-684@7.[189,1399,1353,1939]" captionTargetPageId="7" captionText="Figs 25–28. Venation of Derbachilini: 25–26 — Derbachile aschei sp.n.: 25 — tegmen; 26 — hind wing; 27 — Derbachile hochae gen. et sp.n., tegmen; 28 — Achiderbe obrienae gen. et sp.n., tegmen. Scale bars: 1.0 mm. Рис. 25–28. Жилкование Derbachilini: 25–26 — Derbachile aschei sp.n.: 25 — переднее крыло; 26 — Заднее крыло; 27 — Derbachile hochae gen. et sp.n., переднее крыло; 28 — Achiderbe obrienae gen. et sp.n., переднее крыло. МасШтабные линейки: 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13375233" httpUri="https://zenodo.org/record/13375233/files/figure.png" pageId="1" pageNumber="238">27</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C338653FFFFFFF9ECB51FC66AB52FBFC" pageId="1" pageNumber="238" type="materials_examined">
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<paragraph id="8B9D36B4FFFFFF9ECB51FC66AB52FBFC" blockId="1.[812,1399,917,2064]" pageId="1" pageNumber="238">
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<materialsCitation id="3B4A3CE9FFFFFF9ECB51FC66AB52FBFC" collectionCode="PIN" country="Myanmar" location="Burmese" municipality="Hukawng Valley" pageId="1" pageNumber="238" specimenCode="PIN 5608" specimenCount="1" specimenCount-male="1" stateProvince="Kachin" typeStatus="holotype">
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MATERIAL.
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<typeStatus id="54998816FFFFFF9ECBCDFC66AB5EFC0C" box="[974,1054,983,1002]" pageId="1" pageNumber="238" type="holotype">Holotype</typeStatus>
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<specimenCode id="DB849ECFFFFFFF9ECC25FC69AB3DFC0C" box="[1062,1149,983,1003]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="238">PIN 5608</specimenCode>
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/134,
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<specimenCount id="9D24FD3DFFFFFF9ECCACFC6BAB83FC0C" box="[1199,1219,986,1002]" count="1" pageId="1" pageNumber="238" type="male">♂</specimenCount>
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—
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<location id="8EFD606FFFFFFF9ECCE4FC69AA73FC0D" LSID="urn:lsid:plazi:treatment:038B87A2FFFFFF9CCB3BFC24AE33FCD7:8EFD606FFFFFFF9ECCE4FC69AA73FC0D" box="[1255,1331,984,1003]" country="Myanmar" municipality="Hukawng Valley" name="Burmese" pageId="1" pageNumber="238" stateProvince="Kachin">Burmese</location>
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amber,
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<collectingMunicipality id="6BF9ACCEFFFFFF9ECB2FFC5EAC82FBE4" box="[812,962,1007,1026]" pageId="1" pageNumber="238">Hukawng Valley</collectingMunicipality>
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,
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<collectingRegion id="49E6F856FFFFFF9ECBCFFC5EAB7EFBE4" box="[972,1086,1007,1026]" country="Myanmar" name="Kachin" pageId="1" pageNumber="238">Kachin State</collectingRegion>
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,
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<collectingCountry id="F3357624FFFFFF9ECC4BFC41ABDEFBE5" box="[1096,1182,1008,1027]" name="Myanmar" pageId="1" pageNumber="238">Myanmar</collectingCountry>
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; mid-Cretaceous (probably Albian–Cenomanian).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C338653FFFFFFF9ECB51FBAEAA52FB8D" pageId="1" pageNumber="238" type="diagnosis">
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<paragraph id="8B9D36B4FFFFFF9ECB51FBAEAA52FB8D" blockId="1.[812,1399,917,2064]" pageId="1" pageNumber="238">DIAGNOSIS. Wings hyaline; body pale, disc of mesonotum dark; legs pale with dark streaks; median carina of mesonotum absent; hind tibia with 2 lateral teeth.</paragraph>
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</subSubSection>
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<subSubSection id="C338653FFFFFFF9CCB51FBDEAD82FD07" lastPageId="3" lastPageNumber="240" pageId="1" pageNumber="238" type="description">
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<paragraph id="8B9D36B4FFFFFF9CCB51FBDEAD82FD07" blockId="1.[812,1399,917,2064]" lastBlockId="3.[189,775,315,817]" lastPageId="3" lastPageNumber="240" pageId="1" pageNumber="238">
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DESCRIPTION. Tegmen
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<quantity id="4CDA9B51FFFFFF9ECC4FFBDEABD5FB63" box="[1100,1173,1135,1157]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="238" unit="mm" value="2.6">2.6 mm</quantity>
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long, ca. 1.0 mm wide, elongate, much widened distally, 2.5:1, hyaline, slightly infuscate along anterior margin, veins and stigmal cell fuscous. Anterior margin shallowly convex, with slight concavity at nodus; apical margin rounded. Costal vein turning flat, transversely corrugate distad of ScR fork; C and ambient vein with sensory pits up to apex of tegmen. ScR fork slightly proximad of, and CuA fork distad of, union of claval veins. Nodal
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<emphasis id="B956EAA6FFFFFF9ECD57FA9AAA36FAA6" box="[1364,1398,1323,1344]" italics="true" pageId="1" pageNumber="238">r-m</emphasis>
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distal to nodal
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<emphasis id="B956EAA6FFFFFF9ECBB6FAF4ACA4FABC" box="[949,996,1349,1370]" italics="true" pageId="1" pageNumber="238">m-cu</emphasis>
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. MA fork about halfway from nodal
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<emphasis id="B956EAA6FFFFFF9ECD3EFAF4AA1FFABC" box="[1341,1375,1349,1370]" italics="true" pageId="1" pageNumber="238">r-m</emphasis>
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to
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<emphasis id="B956EAA6FFFFFF9ECB2FFAEEAC1AFA92" box="[812,858,1375,1396]" italics="true" pageId="1" pageNumber="238">2r-m</emphasis>
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. Medial cell long and narrow, about as large as cell distal to it and twice larger than cell anterodistal to it. Crossvein on clavus between CuP and Pcu before middle. Clavus truncate at apex. Hind wing
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<quantity id="4CDA9B51FFFFFF9ECBD2FA1EAB5AFA23" box="[977,1050,1455,1477]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="238" unit="mm" value="2.2">2.2 mm</quantity>
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long,
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<quantity id="4CDA9B51FFFFFF9ECC54FA1EABE0FA23" box="[1111,1184,1455,1477]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="238" unit="mm" value="1.1">1.1 mm</quantity>
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wide, relatively large, 0.8 as long as tegmen, and moderately broad, 2.0:1, apex distinctly not reaching apex of tegmen in repose; membrane transparent, veins infuscate. Costal margin biconvex, coupling lobe in concavity before wing midlength. ScRA apex level with M fork. M fork 1.2 times as long as CuA1 fork. Body
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<quantity id="4CDA9B51FFFFFF9ECD5BF984AC11F983" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="238" unit="mm" value="2.2">2.2 mm</quantity>
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long; body and appendages pale fuscous, disc of mesonotum brown, femora and tibiae with brown longitudinal streaks. Head 1/2 as wide as pronotum; eyes large, hemispherical, emarginate below; head and coryphe deeply arcuately excavate posteriorly; posterior angles of coryphe at eye midlength. Coryphe trapezoidal, anteriorly less than 1/3 head width and as wide as long in middle, depressed between high lateral carinae converging anteriorly, with strong median carina; coryphe curved into, and median carina continued onto, metope. Metope tricarinate, narrow and more depressed dorsally, more than twice widened ventrally; epistomal suture depressed. Scape elongate, obliquely produced anterodorsally, pedicel twice longer than scape, oval, ear-shaped, flattened dorsoventrally, scape + pedicel 2/3 as long as head width. Pedicel covered with cuticular denticles, with more than ten plaque organs on each side. Low subantennal ridge running from epistomal suture to below antennal fovea. Postclypeus tricarinate, strongly narrowed towards unicarinate, laterally compressed anteclypeus. Rostrum reaching hind coxae, apical segment at least 4 times as long as wide. Pronotum 0.85 as wide as mesonotum, 1/3 as long in middle as wide, inverted Vshaped, lateral margins acute, posterior margin sinuate medially; strongly elevated triangular tricarinate disc 1.4 as wide as long in middle, almost 1/2 pronotum width. Mesonotum with nearly diamond-shaped disc, lateral carinae of disc strongly converging anteriorly, median carina absent. Legs very slen- der, femora and tibiae carinate. Hind tarsus nearly 2/3 as long as tibia, tarsomere I longer than II+III; hind tibia and tarsomeres I and II weakly widened at apices; tibia with minute lateral tooth near base and larger one before midlength, and 8 apical teeth in uneven pecten; tarsomere I with 6, II with 4 apical teeth. Claws slender. Male genital block small, anal tube rather short, styles directed caudally, with slender processes.
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</paragraph>
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</subSubSection>
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<caption id="DF5D663CFFFCFF9DC8E1F8D8AD85F7F7" ID-DOI="http://doi.org/10.5281/zenodo.13375229" ID-Zenodo-Dep="13375229" httpUri="https://zenodo.org/record/13375229/files/figure.png" pageId="2" pageNumber="239" startId="2.[226,263,1897,1916]" targetBox="[189,1399,318,1862]" targetPageId="2" targetType="figure">
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<paragraph id="8B9D36B4FFFCFF9DC8E1F8D8AB6CF84B" blockId="2.[189,1399,1897,2065]" pageId="2" pageNumber="239">
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Figs 1–10.
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<taxonomicName id="4C224D37FFFCFF9DC946F8D8AEAAF89A" authorityName="Polevoi & Maximova & Subbotina" authorityYear="2020" box="[325,490,1897,1916]" class="Insecta" family="Derbidae" genus="Derbachile" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="239" phylum="Arthropoda" rank="species" species="hochae" status="gen. et sp. n.">
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<emphasis id="B956EAA6FFFCFF9DC946F8D8AEAAF89A" box="[325,490,1897,1916]" italics="true" pageId="2" pageNumber="239">Derbachile hochae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26557DDFFFCFF9DC9F3F8D8AD1DF89A" box="[496,605,1897,1917]" pageId="2" pageNumber="239" rank="species">
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<emphasis id="B956EAA6FFFCFF9DC9F3F8D8AD55F89A" bold="true" box="[496,533,1897,1916]" pageId="2" pageNumber="239">gen.</emphasis>
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||
et
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<emphasis id="B956EAA6FFFCFF9DCA31F8D8AD1DF89A" bold="true" box="[562,605,1897,1916]" pageId="2" pageNumber="239">sp.n.</emphasis>
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</taxonomicNameLabel>
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, holotype male, Burmese amber: 1 — habitus, dorsal; 2 — habitus, ventral; 3 — head and thorax, dorsal; 4 — head and thorax, ventral (arrows, subantennal ridges); 5 — tegmen, anterodorsal; 6–7 — C and stigmal cell: 6 — right, 7 — left tegmen; 8 — hind legs; 9 — genitalia, ventral; 10 — genitalia, dorsal. Scale bars: 0.5 mm.
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</paragraph>
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<paragraph id="8B9D36B4FFFCFF9DC8E1F807AD85F7F7" blockId="2.[189,1399,1897,2065]" pageId="2" pageNumber="239">
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Рис. 1–10.
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<taxonomicName id="4C224D37FFFCFF9DC946F804AEAAF82E" authorityName="Polevoi & Maximova & Subbotina" authorityYear="2020" box="[325,490,1973,1992]" class="Insecta" family="Derbidae" genus="Derbachile" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="239" phylum="Arthropoda" rank="species" species="hochae" status="gen. et sp. n.">
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<emphasis id="B956EAA6FFFCFF9DC946F804AEAAF82E" box="[325,490,1973,1992]" italics="true" pageId="2" pageNumber="239">Derbachile hochae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26557DDFFFCFF9DC9F3F804AD1CF82E" box="[496,604,1973,1993]" pageId="2" pageNumber="239" rank="species">
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||
<emphasis id="B956EAA6FFFCFF9DC9F3F804AD55F82E" bold="true" box="[496,533,1973,1992]" pageId="2" pageNumber="239">gen.</emphasis>
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et
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<emphasis id="B956EAA6FFFCFF9DCA32F804AD1CF82E" bold="true" box="[561,604,1973,1992]" pageId="2" pageNumber="239">sp.n.</emphasis>
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</taxonomicNameLabel>
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||
, голотип, самец, бирманский Янтарь: 1 — обЩий вид, сверху; 2 — обЩий вид, сниЗу; 3 — голова и грудь, сверху; 4 — голова и грудь, сниЗу (стрелки — подусиковые кили); 5 — переднее крыло, спереди- сверху; 6–7 — C и стигмальнаЯ Ячейка: 6 — правое переднее крыло, 7 — левое переднее крыло; 8 — Задние ноги; 9 — гениталии, сниЗу; 10 — гениталии, сверху. Длина масШтабной линейки: 0,5 мм.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C338653FFFFDFF9CC8E1FD57AE33FCD7" pageId="3" pageNumber="240" type="etymology">
|
||
<paragraph id="8B9D36B4FFFDFF9CC8E1FD57AE33FCD7" blockId="3.[189,775,315,817]" pageId="3" pageNumber="240">
|
||
ETYMOLOGY. Named in honour of Prof. Hannelore Hoch, a brilliant entomologist, specialist on
|
||
<taxonomicName id="4C224D37FFFDFF9CCA84FCB0AD99FCF1" authorityName="Spinola" authorityYear="1839" box="[647,729,769,791]" class="Insecta" family="Cixiidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="240" phylum="Arthropoda" rank="family">Cixiidae</taxonomicName>
|
||
and other Fulgoroidea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |