treatments-xml/data/03/8B/87/038B8782B32A2B7862FAF8C1FE265153.xml
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<document id="A1BB562157CFDE23D4359B5A91C13145" ID-CLB-Dataset="26152" ID-DOI="10.11646/zootaxa.4646.1.4" ID-GBIF-Dataset="ba3f3745-b877-49ec-a6d0-04872d9e0506" ID-ISSN="1175-5326" ID-Zenodo-Dep="3346339" ID-ZooBank="86237AB4-2C4C-412B-A6D7-E44843DA9967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1563862723493" checkinUser="plazi" docAuthor="Inoue, Shota, Maruyama, Munetoshi &amp; Nomura, Shûhei" docDate="2019" docId="038B8782B32A2B7862FAF8C1FE265153" docLanguage="en" docName="zootaxa.4646.1.4.pdf" docOrigin="Zootaxa 4646 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tmesiphorus kinomurai Inoue &amp; Maruyama &amp; Nomura 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="81" masterDocId="FFB2FFFAB3212B76626DFFE4FFD25404" masterDocTitle="Revision of the genus Tmesiphorus LeConte, 1849 (Coleoptera: Staphylinidae: Pselaphinae) from Japan" masterLastPageNumber="86" masterPageNumber="67" pageNumber="78" updateTime="1698746963731" updateUser="ExternalLinkService">
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<mods:title id="E7B1FB03A9D41AFF5D4BD1A66B0C4A07">Revision of the genus Tmesiphorus LeConte, 1849 (Coleoptera: Staphylinidae: Pselaphinae) from Japan</mods:title>
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<mods:namePart id="AE71CF40D1CD75D18D6BED6D572B6AEC">Inoue, Shota</mods:namePart>
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<mods:namePart id="9B71A922831862D73113F39A88A59E3B">Maruyama, Munetoshi</mods:namePart>
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<mods:namePart id="FD68FA0559E1CE728945E6E04682743E">Nomura, Shûhei</mods:namePart>
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<mods:title id="8FE6506572EB47AFF2A8BBF50F115C05">Zootaxa</mods:title>
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<mods:date id="B3C7C5EE3F567AC2E5F1B20DEE72EFC5">2019</mods:date>
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<treatment id="038B8782B32A2B7862FAF8C1FE265153" ID-GBIF-Taxon="158849582" LSID="urn:lsid:plazi:treatment:038B8782B32A2B7862FAF8C1FE265153" httpUri="http://treatment.plazi.org/id/038B8782B32A2B7862FAF8C1FE265153" lastPageId="14" lastPageNumber="81" pageId="11" pageNumber="78">
<subSubSection id="C338651FB32A2B7D62FAF8C1FD715360" pageId="11" pageNumber="78" type="nomenclature">
<paragraph id="8B9D3694B32A2B7D62FAF8C1FDF35344" blockId="11.[151,675,1829,1928]" box="[151,545,1829,1856]" pageId="11" pageNumber="78">
<heading id="D0D581F8B32A2B7D62FAF8C1FDF35344" bold="true" box="[151,545,1829,1856]" fontSize="11" level="1" pageId="11" pageNumber="78" reason="1">
<emphasis id="B956EA86B32A2B7D62FAF8C1FDF35344" bold="true" box="[151,545,1829,1856]" pageId="11" pageNumber="78">
<taxonomicName id="4C224D17B32A2B7D62FAF8C1FE6E5344" ID-CoL="8YBWG" authority="Inoue &amp; Maruyama &amp; Nomura, 2019" authorityName="Inoue &amp; Maruyama &amp; Nomura" authorityYear="2019" box="[151,444,1829,1856]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="78" phylum="Arthropoda" rank="species" species="kinomurai" status="sp. nov.">
<emphasis id="B956EA86B32A2B7D62FAF8C1FE6E5344" bold="true" box="[151,444,1829,1856]" italics="true" pageId="11" pageNumber="78">Tmesiphorus kinomurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557FDB32A2B7D63AEF8C2FDF35344" box="[451,545,1830,1856]" pageId="11" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D3694B32A2B7D62FAF8AEFD715360" blockId="11.[151,675,1829,1928]" box="[151,675,1865,1892]" pageId="11" pageNumber="78">[Japanese name: Hoso-higekata-arizukamushi]</paragraph>
</subSubSection>
<subSubSection id="C338651FB32A2B7A62FAF889FAF85547" lastPageId="12" lastPageNumber="79" pageId="11" pageNumber="78" type="description">
<paragraph id="8B9D3694B32A2B7D62FAF889FEF5538C" blockId="11.[151,675,1829,1928]" box="[151,295,1901,1928]" pageId="11" pageNumber="78">
(
<figureCitation id="13192A11B32A2B7D62F2F889FECD538C" box="[159,287,1901,1928]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="12.[151,250,1935,1960]" captionStartId-1="13.[151,250,1562,1587]" captionStartId-2="14.[151,250,1990,2015]" captionTargetBox-0="[190,1412,583,1908]" captionTargetBox-1="[189,1398,181,1542]" captionTargetBox-2="[212,1379,1402,1955]" captionTargetId-0="figure@12.[164,1424,561,1916]" captionTargetId-1="figure@13.[189,1398,181,1542]" captionTargetId-2="figure@14.[197,1391,1386,1968]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 10. A, B: Habitus of Tmesiphorus kinomurai sp. nov., C, D: left antenna. A, male; B, female; C, antenna in dorsal view; D, antennal club in ventral view by SEM." captionText-1="FIGURE 11. Tmesiphorus kinomurai sp. nov., male. A, head; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, right protarsus." captionText-2="FIGURE 12. Male genitalia of Tmesiphorus kinomurai sp. nov., A, ventral view; B, lateral view; C, dorsal view." httpUri-0="https://zenodo.org/record/3346361/files/figure.png" httpUri-1="https://zenodo.org/record/3346363/files/figure.png" httpUri-2="https://zenodo.org/record/3346365/files/figure.png" pageId="11" pageNumber="78">Figs 1012</figureCitation>
)
</paragraph>
<paragraph id="8B9D3694B32A2B7A62FAF856FAF85547" blockId="11.[151,866,1969,1994]" lastBlockId="12.[151,1437,152,503]" lastPageId="12" lastPageNumber="79" pageId="11" pageNumber="78">
<taxonomicName id="4C224D17B32A2B7D62FAF856FEC853CE" authorityName="LeConte" authorityYear="1849" box="[151,282,1970,1994]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B956EA86B32A2B7D62FAF856FEC853CE" box="[151,282,1970,1994]" italics="true" pageId="11" pageNumber="78">Tmesiphorus</emphasis>
</taxonomicName>
sp. 1,
<bibRefCitation id="EFB34B65B32A2B7D6330F856FDBB53CE" author="Maruyama, M. &amp; Komatsu, T. &amp; Kudo, S. &amp; Shimada, T. &amp; Kinomura, K." box="[349,617,1970,1994]" pageId="11" pageNumber="78" refId="ref10086" refString="Maruyama, M., Komatsu, T., Kudo, S., Shimada, T. &amp; Kinomura, K. (2013) The guests of Japanese ants. Tokai University Press, Minamiyana, 208 pp." type="book" year="2013">
Maruyama
<emphasis id="B956EA86B32A2B7D63BCF857FDD853CE" box="[465,522,1970,1994]" italics="true" pageId="11" pageNumber="78">et al.,</emphasis>
2013: 33
</bibRefCitation>
(illustration, diagnosis).
<emphasis id="B956EA86B32D2B7A62FAFF7DFE9A54B7" bold="true" box="[151,328,152,179]" pageId="12" pageNumber="79">Type material.</emphasis>
<materialsCitation id="3B4A3CC9B32D2B7A633FFF7DFD8854D3" ID-GBIF-Occurrence="2413160237" collectingDate="2003-04-20" collectingDateMax="34-03-20" collectingDateMin="2003-04-20" collectionCode="AJ" collectorName="K. Kinomura" country="Japan" county="Senbiki" location="Gifu-ken" municipality="Seki-shi" pageId="12" pageNumber="79" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54998836B32D2B7A633FFF7DFE6D54B7" box="[338,447,153,179]" pageId="12" pageNumber="79" type="holotype">Holotype</typeStatus>
</emphasis>
(placed in NSMT):
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,
<collectingCountry id="F3357604B32D2B7A60B9FF7DFCCA54B7" box="[724,792,153,179]" name="Japan" pageId="12" pageNumber="79">Japan</collectingCountry>
: [Honshû],
<location id="8EFD604FB32D2B7A61CCFF7DFBDB54B7" LSID="urn:lsid:plazi:treatment:038B8782B32A2B7862FAF8C1FE265153:8EFD604FB32D2B7A61CCFF7DFBDB54B7" box="[929,1033,153,179]" country="Japan" county="Senbiki" municipality="Seki-shi" name="Gifu-ken" pageId="12" pageNumber="79">Gifu-ken</location>
,
<collectingMunicipality id="6BF9ACEEB32D2B7A667AFF7DFBA454B7" box="[1047,1142,153,179]" pageId="12" pageNumber="79">Seki-shi</collectingMunicipality>
,
<collectingCounty id="62FC4E18B32D2B7A66E9FF7DFB0E54B7" box="[1156,1244,153,179]" pageId="12" pageNumber="79">Senbiki</collectingCounty>
,
<collectingDate id="EFD8E9BCB32D2B7A6686FF7DFAA154B7" box="[1259,1395,153,179]" pageId="12" pageNumber="79" value="2003-04-20">
<date id="FF9C1054B32D2B7A6686FF7DFAA154B7" box="[1259,1395,153,179]" pageId="12" pageNumber="79" value="2003-04-20">20 IV 2003</date>
</collectingDate>
,
<collectorName id="26D75342B32D2B7A67ECFF7DFEDB54D3" pageId="12" pageNumber="79">K. Kinomura</collectorName>
leg. (
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), NSMT-I-C-
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.
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<typeStatus id="54998836B32D2B7A6032FF59FD0554D3" box="[607,727,189,215]" pageId="12" pageNumber="79" type="paratype">
<emphasis id="B956EA86B32D2B7A6032FF59FD0554D3" bold="true" box="[607,727,189,215]" pageId="12" pageNumber="79">Paratypes</emphasis>
</typeStatus>
:
<materialsCitation id="3B4A3CC9B32D2B7A6088FF59FDD554FF" ID-GBIF-Occurrence="2293867414" collectingDate="2013-04-18" collectionCode="IV, AJ, NSMT" collectorName="K. Kinomura" country="Japan" county="Alt." elevation="50" location="Gifu-ken" municipality="Mt. Kinka-zan" pageId="12" pageNumber="79" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D24FD1DB32D2B7A6088FF59FCDB54D3" box="[741,777,189,215]" pageId="12" pageNumber="79" type="male">1♂</specimenCount>
,
<location id="8EFD604FB32D2B7A6179FF59FCAE54D3" LSID="urn:lsid:plazi:treatment:038B8782B32A2B7862FAF8C1FE265153:8EFD604FB32D2B7A6179FF59FCAE54D3" box="[788,892,189,215]" country="Japan" county="Alt." municipality="Mt. Kinka-zan" name="Gifu-ken" pageId="12" pageNumber="79">Gifu-ken</location>
,
<location id="8EFD604FB32D2B7A61EBFF59FC3454D3" LSID="urn:lsid:plazi:treatment:038B8782B32A2B7862FAF8C1FE265153:8EFD604FB32D2B7A61EBFF59FC3454D3" box="[902,998,189,215]" country="Japan" county="Alt." municipality="Mt. Kinka-zan" name="Gifu-shi" pageId="12" pageNumber="79">Gifu-shi</location>
,
<collectingMunicipality id="6BF9ACEEB32D2B7A619DFF59FB4054D3" box="[1008,1170,189,215]" pageId="12" pageNumber="79">Mt. Kinka-zan</collectingMunicipality>
(
<collectingCounty id="62FC4E18B32D2B7A66CDFF59FB1E54D3" box="[1184,1228,189,215]" pageId="12" pageNumber="79">Alt.</collectingCounty>
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),
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<date id="FF9C1054B32D2B7A6774FF59FA4A54D3" box="[1305,1432,189,215]" pageId="12" pageNumber="79" value="2013-04-18">
18
<collectionCode id="ED33AE51B32D2B7A6756FF59FA8A54D3" box="[1339,1368,189,215]" pageId="12" pageNumber="79">IV</collectionCode>
2013
</date>
</collectingDate>
,
<collectorName id="26D75342B32D2B7A62FAFF05FEFF54FF" box="[151,301,225,251]" pageId="12" pageNumber="79">K. Kinomura</collectorName>
leg. (
<collectionCode id="ED33AE51B32D2B7A631DFF05FE4054FF" box="[368,402,225,251]" country="Japan" name="Central Research Laboratories" pageId="12" pageNumber="79">AJ</collectionCode>
) (
<collectionCode id="ED33AE51B32D2B7A63C6FF05FE2D54FF" box="[427,511,225,251]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="12" pageNumber="79">NSMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4A3CC9B32D2B7A6079FF05FC7154FF" ID-GBIF-Occurrence="2293867428" box="[532,931,225,251]" collectingDate="2013-04-18" collectionCode="KUM" collectorName="K. Kinomura" country="Japan" county="Alt." elevation="50" location="Gifu-ken" municipality="Mt. Kinka-zan" pageId="12" pageNumber="79" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D24FD1DB32D2B7A6079FF05FDEA54FF" box="[532,568,225,251]" pageId="12" pageNumber="79" type="female">2♀</specimenCount>
, same data as holotype (
<collectionCode id="ED33AE51B32D2B7A6139FF05FC4954FF" box="[852,923,225,251]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="12" pageNumber="79" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4A3CC9B32D2B7A61DCFF05FD96551B" ID-GBIF-Occurrence="2293867412" collectingDate="2003-06-07" collectionCode="VI, KUM" collectorName="K. Kinomura" country="Japan" county="Nagaraobusa" location="Gifu-ken" municipality="Gifu-shi" pageId="12" pageNumber="79" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D24FD1DB32D2B7A61DCFF05FC0654FF" box="[945,980,225,251]" pageId="12" pageNumber="79" type="female">1♀</specimenCount>
,
<location id="8EFD604FB32D2B7A6189FF05FB9E54FF" LSID="urn:lsid:plazi:treatment:038B8782B32A2B7862FAF8C1FE265153:8EFD604FB32D2B7A6189FF05FB9E54FF" box="[996,1100,225,251]" country="Japan" county="Nagaraobusa" municipality="Gifu-shi" name="Gifu-ken" pageId="12" pageNumber="79">Gifu-ken</location>
,
<collectingMunicipality id="6BF9ACEEB32D2B7A6634FF05FB6B54FF" box="[1113,1209,225,251]" pageId="12" pageNumber="79">Gifu-shi</collectingMunicipality>
,
<collectingCounty id="62FC4E18B32D2B7A66ABFF05FA8954FF" box="[1222,1371,225,251]" pageId="12" pageNumber="79">Nagaraobusa</collectingCounty>
,
<collectingDate id="EFD8E9BCB32D2B7A6705FF05FF00551B" pageId="12" pageNumber="79" value="2003-06-07">
<date id="FF9C1054B32D2B7A6705FF05FF00551B" pageId="12" pageNumber="79" value="2003-06-07">
7
<collectionCode id="ED33AE51B32D2B7A6712FF05FA4F54FF" box="[1407,1437,225,251]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="79">VI</collectionCode>
2003
</date>
</collectingDate>
,
<collectorName id="26D75342B32D2B7A62B0FEE1FEA0551B" box="[221,370,261,287]" pageId="12" pageNumber="79">K. Kinomura</collectorName>
leg. (LM) (
<collectionCode id="ED33AE51B32D2B7A6398FEE1FDEE551B" box="[501,572,261,287]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="12" pageNumber="79" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4A3CC9B32D2B7A603DFEE1FF3D5547" ID-GBIF-Occurrence="2293867438" collectingDate="2006-04-22" collectionCode="IV, AJ, KUM" collectorName="K. Kinomura" country="Japan" county="Mt. Kinka-zan" location="Gifu-ken" municipality="Gifu-shi" pageId="12" pageNumber="79" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D24FD1DB32D2B7A603DFEE1FDA6551B" box="[592,628,261,287]" pageId="12" pageNumber="79" type="female">2♀</specimenCount>
,
<location id="8EFD604FB32D2B7A60ECFEE1FD3B551B" LSID="urn:lsid:plazi:treatment:038B8782B32A2B7862FAF8C1FE265153:8EFD604FB32D2B7A60ECFEE1FD3B551B" box="[641,745,261,287]" country="Japan" county="Mt. Kinka-zan" municipality="Gifu-shi" name="Gifu-ken" pageId="12" pageNumber="79">Gifu-ken</location>
,
<collectingMunicipality id="6BF9ACEEB32D2B7A6099FEE1FC86551B" box="[756,852,261,287]" pageId="12" pageNumber="79">Gifu-shi</collectingMunicipality>
,
<collectingCounty id="62FC4E18B32D2B7A610DFEE1FBD5551B" box="[864,1031,261,287]" pageId="12" pageNumber="79">Mt. Kinka-zan</collectingCounty>
,
<collectingDate id="EFD8E9BCB32D2B7A667CFEE1FB46551B" box="[1041,1172,261,287]" pageId="12" pageNumber="79" value="2006-04-22">
<date id="FF9C1054B32D2B7A667CFEE1FB46551B" box="[1041,1172,261,287]" pageId="12" pageNumber="79" value="2006-04-22">
22
<collectionCode id="ED33AE51B32D2B7A6658FEE1FB80551B" box="[1077,1106,261,287]" pageId="12" pageNumber="79">IV</collectionCode>
2006
</date>
</collectingDate>
,
<collectorName id="26D75342B32D2B7A66F2FEE1FAE1551B" box="[1183,1331,261,287]" pageId="12" pageNumber="79">K. Kinomura</collectorName>
leg. (
<collectionCode id="ED33AE51B32D2B7A671FFEE1FA46551B" box="[1394,1428,261,287]" country="Japan" name="Central Research Laboratories" pageId="12" pageNumber="79">AJ</collectionCode>
) (
<collectionCode id="ED33AE51B32D2B7A62F2FECDFF345547" box="[159,230,297,323]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="12" pageNumber="79" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4A3CC9B32D2B7A6297FECDFAF45547" ID-GBIF-Occurrence="2293867430" box="[250,1318,296,323]" collectingDate="2013-05-05" collectionCode="AJ, KUM" collectorName="K. Kinomura" country="Japan" county="Isshiki" location="Gifu-ken" municipality="Shoukawa-cho" pageId="12" pageNumber="79" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D24FD1DB32D2B7A6297FECDFECF5547" box="[250,285,297,323]" pageId="12" pageNumber="79" type="female">1♀</specimenCount>
,
<location id="8EFD604FB32D2B7A6346FECDFE415547" LSID="urn:lsid:plazi:treatment:038B8782B32A2B7862FAF8C1FE265153:8EFD604FB32D2B7A6346FECDFE415547" box="[299,403,297,323]" country="Japan" county="Isshiki" municipality="Shoukawa-cho" name="Gifu-ken" pageId="12" pageNumber="79">Gifu-ken</location>
,
<location id="8EFD604FB32D2B7A63F3FECDFDEC5547" LSID="urn:lsid:plazi:treatment:038B8782B32A2B7862FAF8C1FE265153:8EFD604FB32D2B7A63F3FECDFDEC5547" box="[414,574,297,323]" country="Japan" county="Isshiki" municipality="Shoukawa-cho" name="Takayama-shi" pageId="12" pageNumber="79">Takayama-shi</location>
,
<collectingMunicipality id="6BF9ACEEB32D2B7A6025FECDFD235547" box="[584,753,297,323]" pageId="12" pageNumber="79">Shoukawa-chô</collectingMunicipality>
,
<collectingCounty id="62FC4E18B32D2B7A6090FECDFC995547" box="[765,843,297,323]" pageId="12" pageNumber="79">Isshiki</collectingCounty>
,
<collectingDate id="EFD8E9BCB32D2B7A6138FECCFC6D5547" box="[853,959,296,323]" pageId="12" pageNumber="79" value="2013-05-05">
<date id="FF9C1054B32D2B7A6138FECCFC6D5547" box="[853,959,296,323]" pageId="12" pageNumber="79" value="2013-05-05">5 V 2013</date>
</collectingDate>
,
<collectorName id="26D75342B32D2B7A61A7FECDFB8D5547" box="[970,1119,297,323]" pageId="12" pageNumber="79">K. Kinomura</collectorName>
leg. (
<collectionCode id="ED33AE51B32D2B7A66F0FECDFB6D5547" box="[1181,1215,297,323]" country="Japan" name="Central Research Laboratories" pageId="12" pageNumber="79">AJ</collectionCode>
) (
<collectionCode id="ED33AE51B32D2B7A66BBFECDFACF5547" box="[1238,1309,297,323]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="12" pageNumber="79" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338651FB32D2B7862AAFEA8FE265153" lastPageId="14" lastPageNumber="81" pageId="12" pageNumber="79" type="materials_examined">
<paragraph id="8B9D3694B32D2B7A62AAFEA8FB7055D7" blockId="12.[151,1437,152,503]" pageId="12" pageNumber="79">
<emphasis id="B956EA86B32D2B7A62AAFEA8FE905563" bold="true" box="[199,322,332,359]" pageId="12" pageNumber="79">Diagnosis.</emphasis>
<taxonomicName id="4C224D17B32D2B7A6327FEA9FD845563" authority="Inoue &amp; Maruyama &amp; Nomura, 2019" authorityName="Inoue &amp; Maruyama &amp; Nomura" authorityYear="2019" box="[330,598,333,359]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="79" phylum="Arthropoda" rank="species" species="kinomurai" status="sp. nov.">
<emphasis id="B956EA86B32D2B7A6327FEA9FD845563" box="[330,598,333,359]" italics="true" pageId="12" pageNumber="79">Tmesiphorus kinomurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557FDB32D2B7A6033FEA9FD645563" box="[606,694,333,359]" pageId="12" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
is very similar in general appearance to
<emphasis id="B956EA86B32D2B7A66EBFEAAFAD95563" box="[1158,1291,334,359]" italics="true" pageId="12" pageNumber="79">
<taxonomicName id="4C224D17B32D2B7A66EBFEAAFAD55563" authorityName="Sharp" authorityYear="1883" box="[1158,1287,334,359]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="79" phylum="Arthropoda" rank="species" species="princeps">T. princeps</taxonomicName>
.
</emphasis>
However,
<taxonomicName id="4C224D17B32D2B7A67E5FEAAFED8558F" authorityName="Inoue &amp; Maruyama &amp; Nomura" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="79" phylum="Arthropoda" rank="species" species="kinomurai">
<emphasis id="B956EA86B32D2B7A67E5FEAAFED8558F" italics="true" pageId="12" pageNumber="79">T. kinomurai</emphasis>
</taxonomicName>
is readily distinguished from
<taxonomicName id="4C224D17B32D2B7A6035FE96FD01558F" authorityName="Sharp" authorityYear="1883" box="[600,723,370,395]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="79" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="B956EA86B32D2B7A6035FE96FD01558F" box="[600,723,370,395]" italics="true" pageId="12" pageNumber="79">T. princeps</emphasis>
</taxonomicName>
by the following states: head longer than wide, weakly shining; eyes small; postocular margin longer than eyes; antennae slender; antennomeres IXXI each longer than wide; XI lanceolate; pronotum slender; tergite V with longitudinal carinae extending to posterior half.
</paragraph>
<paragraph id="8B9D3694B32D2B7A62AAFE38FAD055F3" blockId="12.[151,1437,152,503]" box="[199,1282,476,503]" pageId="12" pageNumber="79">
<emphasis id="B956EA86B32D2B7A62AAFE38FE8B55F3" bold="true" box="[199,345,476,503]" pageId="12" pageNumber="79">Description.</emphasis>
Male (
<figureCitation id="13192A11B32D2B7A63C5FE39FDC955F3" box="[424,539,477,503]" captionStart="FIGURE 10" captionStartId="12.[151,250,1935,1960]" captionTargetBox="[190,1412,583,1908]" captionTargetId="figure@12.[164,1424,561,1916]" captionTargetPageId="12" captionText="FIGURE 10. A, B: Habitus of Tmesiphorus kinomurai sp. nov., C, D: left antenna. A, male; B, female; C, antenna in dorsal view; D, antennal club in ventral view by SEM." httpUri="https://zenodo.org/record/3346361/files/figure.png" pageId="12" pageNumber="79">Fig. 10-A</figureCitation>
). Body length
<quantity id="4CDA9B71B32D2B7A60D3FE39FCB655F3" box="[702,868,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="2.665" metricValueMax="2.71" metricValueMin="2.62" pageId="12" pageNumber="79" unit="mm" value="2.665" valueMax="2.71" valueMin="2.62">2.622.71 mm</quantity>
. Body dark brown to reddish-brown.
</paragraph>
<caption id="DF5D661CB32D2B7A62FAF86BFDAE53C8" httpUri="https://zenodo.org/record/3346361/files/figure.png" pageId="12" pageNumber="79" startId="12.[151,250,1935,1960]" targetBox="[190,1412,583,1908]" targetPageId="12">
<paragraph id="8B9D3694B32D2B7A62FAF86BFDAE53C8" blockId="12.[151,1436,1935,1996]" pageId="12" pageNumber="79">
<emphasis id="B956EA86B32D2B7A62FAF86BFEB053AC" bold="true" box="[151,354,1935,1960]" pageId="12" pageNumber="79">FIGURE 10. A, B:</emphasis>
Habitus of
<emphasis id="B956EA86B32D2B7A63B1F874FD0353A3" box="[476,721,1935,1960]" italics="true" pageId="12" pageNumber="79">
<taxonomicName id="4C224D17B32D2B7A63B1F874FD8D53AC" authorityName="LeConte" authorityYear="1849" box="[476,607,1936,1960]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="79" phylum="Arthropoda" rank="genus">Tmesiphorus</taxonomicName>
<emphasis id="B956EA86B32D2B7A600AF86BFDA653AC" bold="true" box="[615,628,1935,1960]" italics="true" pageId="12" pageNumber="79">k</emphasis>
inomurai
</emphasis>
sp. nov., C, D: left antenna. A, male; B, female; C, antenna in dorsal view; D, antennal club in ventral view by SEM.
</paragraph>
</caption>
<caption id="DF5D661CB32C2B7B62FAF9FEFEE05252" httpUri="https://zenodo.org/record/3346363/files/figure.png" pageId="13" pageNumber="80" startId="13.[151,250,1562,1587]" targetBox="[189,1398,181,1542]" targetPageId="13">
<paragraph id="8B9D3694B32C2B7B62FAF9FEFEE05252" blockId="13.[151,1436,1562,1623]" pageId="13" pageNumber="80">
<emphasis id="B956EA86B32C2B7B62FAF9FEFECD5237" bold="true" box="[151,287,1562,1587]" pageId="13" pageNumber="80">FIGURE 11.</emphasis>
<emphasis id="B956EA86B32C2B7B6348F9FFFDCA5236" box="[293,536,1562,1587]" italics="true" pageId="13" pageNumber="80">
<taxonomicName id="4C224D17B32C2B7B6348F9FFFE7A5237" authorityName="LeConte" authorityYear="1849" box="[293,424,1563,1587]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="80" phylum="Arthropoda" rank="genus">Tmesiphorus</taxonomicName>
<emphasis id="B956EA86B32C2B7B63C2F9FEFE6E5237" bold="true" box="[431,444,1562,1587]" italics="true" pageId="13" pageNumber="80">k</emphasis>
inomurai
</emphasis>
sp. nov., male.
<emphasis id="B956EA86B32C2B7B60D7F9FEFD1E5237" bold="true" box="[698,716,1562,1587]" pageId="13" pageNumber="80">A</emphasis>
, head; B, head in dorsal view;
<emphasis id="B956EA86B32C2B7B6665F9FEFBC85237" bold="true" box="[1032,1050,1562,1587]" pageId="13" pageNumber="80">C</emphasis>
, pronotum;
<emphasis id="B956EA86B32C2B7B66F5F9FFFB785237" bold="true" box="[1176,1194,1563,1587]" pageId="13" pageNumber="80">D</emphasis>
, elytra;
<emphasis id="B956EA86B32C2B7B6693F9FFFADD5237" bold="true" box="[1278,1295,1563,1587]" pageId="13" pageNumber="80">E</emphasis>
, abdomen;
<emphasis id="B956EA86B32C2B7B67EBF9FFFA475237" bold="true" box="[1414,1429,1563,1587]" pageId="13" pageNumber="80">F</emphasis>
, right protarsus.
</paragraph>
</caption>
<paragraph id="8B9D3694B32C2B7862AAF961FD2A5547" blockId="13.[151,1437,1668,2020]" lastBlockId="14.[151,1437,153,1367]" lastPageId="14" lastPageNumber="81" pageId="13" pageNumber="80">
Head (
<figureCitation id="13192A11B32C2B7B637FF961FE7F52A4" box="[274,429,1669,1696]" captionStart="FIGURE 11" captionStartId="13.[151,250,1562,1587]" captionTargetBox="[189,1398,181,1542]" captionTargetId="figure@13.[189,1398,181,1542]" captionTargetPageId="13" captionText="FIGURE 11. Tmesiphorus kinomurai sp. nov., male. A, head; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, right protarsus." httpUri="https://zenodo.org/record/3346363/files/figure.png" pageId="13" pageNumber="80">Figs. 11-A, B</figureCitation>
) longer than wide, HL
<quantity id="4CDA9B71B32C2B7B60DBF960FC9C52A4" box="[694,846,1668,1696]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="13" pageNumber="80" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50.53 mm</quantity>
, HW
<quantity id="4CDA9B71B32C2B7B61FDF961FBE452A4" box="[912,1078,1669,1696]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="13" pageNumber="80" unit="mm" value="0.45499999999999996" valueMax="0.48" valueMin="0.43">0.430.48 mm</quantity>
, slender, almost ovoid in dorsal view, with long setae; frons narrowed, with longitudinal groove including small fovea, densely punctate, narrowed posteriorly to vertex; vertex with two small foveae, with short longitudinal carina between vertexal foveae, weakly shining, slightly depressed around the two vertexal foveae; eyes small, prominent; occiput rounded, with tufts of yellow setae; postocular margin two times as long as eyes; antennae (
<figureCitation id="13192A11B32C2B7B61AEF8F1FBB15334" box="[963,1123,1813,1840]" captionStart="FIGURE 10" captionStartId="12.[151,250,1935,1960]" captionTargetBox="[190,1412,583,1908]" captionTargetId="figure@12.[164,1424,561,1916]" captionTargetPageId="12" captionText="FIGURE 10. A, B: Habitus of Tmesiphorus kinomurai sp. nov., C, D: left antenna. A, male; B, female; C, antenna in dorsal view; D, antennal club in ventral view by SEM." httpUri="https://zenodo.org/record/3346361/files/figure.png" pageId="13" pageNumber="80">Figs. 10-C, D</figureCitation>
) more slender, with antennomeres I thick, elongate, each 2.5 times longer than II, 1.5 times wider than II; II each as long as wide; III each slightly as long as wide, narrowed at the base, broadened apically, IVVI each as long as wide; VIIVIII each transverse; IIIVIII successively broadened towards apices; IXXI enlarged to form club, slightly curved inward in outer margin; IX each longer than wide, slightly wider than I, shallowly depressed longitudinally with dense setae on ventral surface; with a depression from base to apex, 2/3 as wide as segmental width on apical margin; X each longer than wide, slightly slender than IX, with glabrous area with sparse setae on ventral surface; XI each longer than wide, about as thick as IX, slightly produced outward at middle, narrowed apically from middle to apices, lanceolate; shallowly depressed longitudinally from base to middle with sparse setae on ventral surface; maxillary palpi asymmetrical; segments II and III each with slightly long penicillate spine on outside; segment
<collectionCode id="ED33AE51B32F2B78676CFEE1FACC551B" box="[1281,1310,261,287]" pageId="14" pageNumber="81">IV</collectionCode>
thick, with rounded lobe on lateral margin, apical portion conical.
</paragraph>
<paragraph id="8B9D3694B32F2B7862AAFEA9FDCF56CB" blockId="14.[151,1437,153,1367]" pageId="14" pageNumber="81">
Pronotum (
<figureCitation id="13192A11B32F2B786325FEA9FE655563" box="[328,439,333,359]" captionStart="FIGURE 11" captionStartId="13.[151,250,1562,1587]" captionTargetBox="[189,1398,181,1542]" captionTargetId="figure@13.[189,1398,181,1542]" captionTargetPageId="13" captionText="FIGURE 11. Tmesiphorus kinomurai sp. nov., male. A, head; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, right protarsus." httpUri="https://zenodo.org/record/3346363/files/figure.png" pageId="14" pageNumber="81">Fig. 11-C</figureCitation>
) slightly longer than wide,
<collectionCode id="ED33AE51B32F2B786094FEA9FCCB5563" box="[761,793,333,359]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="14" pageNumber="81" type="Museum">PL</collectionCode>
<quantity id="4CDA9B71B32F2B78614FFEA8FC5B5563" box="[802,905,332,359]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="14" pageNumber="81" unit="mm" value="0.56">0.56 mm</quantity>
<collectionCode id="ED33AE51B32F2B7861FFFEA9FC695563" box="[914,955,333,359]" country="Thailand" name="Paleontological Collections" pageId="14" pageNumber="81">PW</collectionCode>
<quantity id="4CDA9B71B32F2B7861ABFEA8FBBC5563" box="[966,1134,332,359]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="14" pageNumber="81" unit="mm" value="0.54" valueMax="0.55" valueMin="0.53">0.530.55 mm</quantity>
; dorsum broadly rounded, widest to middle, narrowed posteriorly, weakly shining, sparsely covered with long setae, with distinct median and lateral foveae at posterior base; elytra (
<figureCitation id="13192A11B32F2B78603BFE71FD1755AB" box="[598,709,405,431]" captionStart="FIGURE 11" captionStartId="13.[151,250,1562,1587]" captionTargetBox="[189,1398,181,1542]" captionTargetId="figure@13.[189,1398,181,1542]" captionTargetPageId="13" captionText="FIGURE 11. Tmesiphorus kinomurai sp. nov., male. A, head; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, right protarsus." httpUri="https://zenodo.org/record/3346363/files/figure.png" pageId="14" pageNumber="81">Fig. 11-D</figureCitation>
) much wider than long, EL
<quantity id="4CDA9B71B32F2B786667FE71FB6355AB" box="[1034,1201,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" metricValueMax="7.3" metricValueMin="7.1" pageId="14" pageNumber="81" unit="mm" value="0.72" valueMax="0.73" valueMin="0.71">0.710.73 mm</quantity>
, EW 1.00
<quantity id="4CDA9B71B32F2B78675FFE71FA4A55AB" box="[1330,1432,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="14" pageNumber="81" unit="mm" value="1.11">1.11 mm</quantity>
, sparsely covered with long setae; each elytron with basal median and inner foveae pubescent, weakly sulcate from median fovea to posterior 2/3 of elytra, weakly carinate from between two foveae to posterior 1/2 of elytra, slightly carinate from outside of median fovea to posterior margin, with thick brush-like of yellow setae along posterior margin; profemora thickest, with white setae at anterior face; mesofemora, metafemora slender; tibiae slender; protibiae thickest, arcuate and thickened at middle, as long as mesotibiae; metatibiae longest; tibiae all with dense yellow setae at apices; tarsi (
<figureCitation id="13192A11B32F2B7863BBFD89FDED5683" box="[470,575,621,647]" captionStart="FIGURE 11" captionStartId="13.[151,250,1562,1587]" captionTargetBox="[189,1398,181,1542]" captionTargetId="figure@13.[189,1398,181,1542]" captionTargetPageId="13" captionText="FIGURE 11. Tmesiphorus kinomurai sp. nov., male. A, head; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, right protarsus." httpUri="https://zenodo.org/record/3346363/files/figure.png" pageId="14" pageNumber="81">Fig. 11-F</figureCitation>
) all slightly slender, elongate; tarsomeres
<date id="FF9C1054B32F2B786678FD89FBEE5683" box="[1045,1084,621,647]" pageId="14" pageNumber="81">II 2</date>
/3 as long as III; tarsal claws all asymmetrical; protarsal anterior claws each thicker than posterior one; mesotarsal and metatarsal posterior claws slightly longer than anterior claws.
</paragraph>
<paragraph id="8B9D3694B32F2B7862AAFD3DFE0E5787" blockId="14.[151,1437,153,1367]" pageId="14" pageNumber="81">
Abdomen (
<figureCitation id="13192A11B32F2B786329FD3DFE7C56F7" box="[324,430,729,755]" captionStart="FIGURE 11" captionStartId="13.[151,250,1562,1587]" captionTargetBox="[189,1398,181,1542]" captionTargetId="figure@13.[189,1398,181,1542]" captionTargetPageId="13" captionText="FIGURE 11. Tmesiphorus kinomurai sp. nov., male. A, head; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, right protarsus." httpUri="https://zenodo.org/record/3346363/files/figure.png" pageId="14" pageNumber="81">Fig. 11-E</figureCitation>
) large, wider than long, AL
<quantity id="4CDA9B71B32F2B786089FD3DFC5A56F7" box="[740,904,729,755]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.3" metricValueMin="8.2" pageId="14" pageNumber="81" unit="mm" value="0.875" valueMax="0.93" valueMin="0.82">0.820.93 mm</quantity>
, AW
<quantity id="4CDA9B71B32F2B7861AEFD3DFBB556F7" box="[963,1127,729,755]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" metricValueMax="1.18" metricValueMin="1.08" pageId="14" pageNumber="81" unit="mm" value="1.13" valueMax="1.18" valueMin="1.08">1.081.18 mm</quantity>
, sparsely covered with long setae, tergite
<collectionCode id="ED33AE51B32F2B786344FD19FE945713" box="[297,326,765,791]" pageId="14" pageNumber="81">IV</collectionCode>
as long as tergite
<collectionCode id="ED33AE51B32F2B786066FD19FDF95713" box="[523,555,765,791]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="14" pageNumber="81">VI</collectionCode>
, tergite V longest, declined toward apex, tergite VII small, tergite VIII smallest, semicircular; tergites
<collectionCode id="ED33AE51B32F2B7863E7FCC5FE75573F" box="[394,423,801,827]" pageId="14" pageNumber="81">IV</collectionCode>
and V with lateral carinae, those of tergite
<collectionCode id="ED33AE51B32F2B7861E2FCC5FC7E573F" box="[911,940,801,827]" pageId="14" pageNumber="81">IV</collectionCode>
extending from anterior margin to posterior margin, those of tergite V extending from anterior margin to posterior 1/2; tergite
<collectionCode id="ED33AE51B32F2B78664CFCA1FBEC575B" box="[1057,1086,837,863]" pageId="14" pageNumber="81">IV</collectionCode>
with thick brush-like of yellow setae along posterior margin.
</paragraph>
<paragraph id="8B9D3694B32F2B7862AAFC69FDC65017" blockId="14.[151,1437,153,1367]" pageId="14" pageNumber="81">
Male genitalia (
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)
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in length, well-sclerotized, more like oval; parameres symmetric, strongly projecting ventrally; median lobe asymmetric, projecting ventrally, thick, curved downwardly from middle to apex; basal bulb broad; apex slightly asymmetrical; secondary gonopore small; endophallus elongate, circular at base, weakly curved dorsally at middle.
</paragraph>
<paragraph id="8B9D3694B32F2B7862AAFBF9FCDB50A7" blockId="14.[151,1437,153,1367]" pageId="14" pageNumber="81">
Female (
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). BL
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: HL
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; HW
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;
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;
<collectionCode id="ED33AE51B32F2B786741FBF9FA875033" box="[1324,1365,1053,1079]" country="Thailand" name="Paleontological Collections" pageId="14" pageNumber="81">PW</collectionCode>
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; EL
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; EW
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; AL
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; AW
<quantity id="4CDA9B71B32F2B78619FFBA4FB48505F" box="[1010,1178,1088,1115]" metricMagnitude="-3" metricUnit="m" metricValue="1.125" metricValueMax="1.2" metricValueMin="1.05" pageId="14" pageNumber="81" unit="mm" value="1.125" valueMax="1.2" valueMin="1.05">1.051.20 mm</quantity>
. Antennae with antennomeres IIVIII slightly longer than in male, successively broadened towards apices; antennomeres IXXI not modified, smaller than in male, antennal club indistinct.
</paragraph>
<paragraph id="8B9D3694B32F2B7862AAFB49FE7250EF" blockId="14.[151,1437,153,1367]" pageId="14" pageNumber="81">
<emphasis id="B956EA86B32F2B7862AAFB49FE9A50C3" bold="true" box="[199,328,1197,1223]" pageId="14" pageNumber="81">Etymology</emphasis>
. This species is named after the collector of the
<typeStatus id="54998836B32F2B78610CFB49FC1450C3" box="[865,966,1197,1223]" pageId="14" pageNumber="81" type="holotype">holotype</typeStatus>
, Mr. Kyôichi Kinomura, who is the super- visor of the first author.
</paragraph>
<paragraph id="8B9D3694B32F2B7862AAFB10FD415114" blockId="14.[151,1437,153,1367]" box="[199,659,1268,1296]" pageId="14" pageNumber="81">
<emphasis id="B956EA86B32F2B7862AAFB10FEB3510B" bold="true" box="[199,353,1268,1295]" pageId="14" pageNumber="81">Distribution.</emphasis>
<collectingCountry id="F3357604B32F2B78630AFB11FE79510B" box="[359,427,1269,1295]" name="Japan" pageId="14" pageNumber="81">Japan</collectingCountry>
: Honshû (Gifu-ken).
</paragraph>
<paragraph id="8B9D3694B32F2B7862AAFAFCFE265153" blockId="14.[151,1437,153,1367]" pageId="14" pageNumber="81">
<emphasis id="B956EA86B32F2B7862AAFAFCFEF55137" bold="true" box="[199,295,1304,1331]" pageId="14" pageNumber="81">Biology.</emphasis>
This species has been collected from nests of
<taxonomicName id="4C224D17B32F2B78614EFAFDFBE65137" authorityName="Forel" authorityYear="1911" box="[803,1076,1305,1331]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="81" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B956EA86B32F2B78614EFAFDFBE65137" box="[803,1076,1305,1331]" italics="true" pageId="14" pageNumber="81">Aphaenogaster japonica</emphasis>
</taxonomicName>
,
<emphasis id="B956EA86B32F2B78662CFAFDFB155137" box="[1089,1223,1305,1331]" italics="true" pageId="14" pageNumber="81">
<taxonomicName id="4C224D17B32F2B78662CFAFDFB105137" authorityName="Smith" authorityYear="1874" box="[1089,1218,1305,1331]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="81" phylum="Arthropoda" rank="species" species="famelica">A. famelica</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C224D17B32F2B786696FAFEFE3F5153" authority="(Maruyama et al. 2013)" authorityName="Maruyama" authorityYear="2013" baseAuthorityName="Maruyama" baseAuthorityYear="2013" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="81" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B956EA86B32F2B786696FAFEFF095153" italics="true" pageId="14" pageNumber="81">Lasius meridionalis</emphasis>
(
<bibRefCitation id="EFB34B65B32F2B786287FAD9FE375153" author="Maruyama, M. &amp; Komatsu, T. &amp; Kudo, S. &amp; Shimada, T. &amp; Kinomura, K." box="[234,485,1341,1367]" pageId="14" pageNumber="81" refId="ref10086" refString="Maruyama, M., Komatsu, T., Kudo, S., Shimada, T. &amp; Kinomura, K. (2013) The guests of Japanese ants. Tokai University Press, Minamiyana, 208 pp." type="book" year="2013">
Maruyama
<emphasis id="B956EA86B32F2B786301FADAFE775153" box="[364,421,1341,1367]" italics="true" pageId="14" pageNumber="81">et al.</emphasis>
2013
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>