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<document id="7981FCD632439C9175EE3AF468F58863" ID-CLB-Dataset="32292" ID-DOI="10.5852/ejt.2017.257" ID-GBIF-Dataset="5183353a-133b-4453-a274-1f75169c9002" ID-Zenodo-Dep="851561" ID-ZooBank="urn:lsid:zoobank.org:pub:D7142011-F22B-48C0-987D-FD8015A0E985" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503855107135" checkinUser="plazi" docAuthor="Devanshu Gupta, Kailash Chandra &amp; Aleš Bezděk" docDate="2017" docId="0382CE3EFFCCFFCF3918FA26523131E1" docLanguage="en" docName="EJT-257_Gupta.pdf" docOrigin="European Journal of Taxonomy 257" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pukupuku arunachalensis Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk 2017, sp. nov." docType="treatment" docUuid="4C196777-31DC-4BEC-A847-7E36ED9E01C5" docUuidSource="ZooBank" docVersion="10" lastPageNumber="10" masterDocId="FFBBB646FFCDFFC63B51FFC550423022" masterDocTitle="Pukupuku arunachalensis sp. nov. (Coleoptera, Scarabaeidae, Rutelinae) from Arunachal Pradesh, India" masterLastPageNumber="11" masterPageNumber="1" pageNumber="2" updateTime="1698467356779" updateUser="ExternalLinkService">
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<mods:title id="35F13EFAD939723D4F8F90507EE7DFFD">Pukupuku arunachalensis sp. nov. (Coleoptera, Scarabaeidae, Rutelinae) from Arunachal Pradesh, India</mods:title>
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<mods:namePart id="D7EE3D250970FD507166744E1680280E">Devanshu Gupta</mods:namePart>
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<mods:namePart id="6C1135A7122039BC6935744C58AF3529">Kailash Chandra</mods:namePart>
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<mods:namePart id="9F273FFF0A09E39A3E25723CBE343576">Aleš Bezděk</mods:namePart>
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<mods:title id="22AC0C9D51AA415A30C8E4F1DBA1B8BC">European Journal of Taxonomy</mods:title>
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<mods:date id="3247E583FAB40FF7463FA3A18FF8A704">2017</mods:date>
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<mods:number id="33FE6126FAC4021B43698F0B7EFDDA54">2017-01-03</mods:number>
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<mods:number id="C1507AF0229C7C53459BA07FC5C2E3BC">257</mods:number>
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<subSubSection id="C3312CA3FFCCFFC73918FA2654853600" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8B947F28FFCCFFC73918FA2653A935DC" blockId="1.[364,1223,1507,1604]" box="[585,1003,1507,1534]" pageId="1" pageNumber="2">
<heading id="D0DCC844FFCCFFC73918FA2653A935DC" box="[585,1003,1507,1534]" centered="true" fontSize="11" level="3" pageId="1" pageNumber="2" reason="4">
<taxonomicName id="4C2B04ABFFCCFFC73918FA2653CB35DC" authority="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk, 2017" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[585,905,1507,1534]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="B95FA33AFFCCFFC73918FA2653CB35DC" bold="true" box="[585,905,1507,1534]" italics="true" pageId="1" pageNumber="2">Pukupuku arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C1E41FFCCFFC738C1FA2153A935DC" box="[912,1003,1508,1534]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B947F28FFCCFFC73A3DF9C254853600" blockId="1.[364,1223,1507,1604]" box="[364,1223,1543,1570]" pageId="1" pageNumber="2">urn:lsid:zoobank.org:act:4C196777-31DC-4BEC-A847-7E36ED9E01C5</paragraph>
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<subSubSection id="C3312CA3FFCCFFC7398EF9EF53163666" box="[735,852,1578,1604]" pageId="1" pageNumber="2" type="description">
<paragraph id="8B947F28FFCCFFC7398EF9EF53163666" blockId="1.[364,1223,1507,1604]" box="[735,852,1578,1604]" pageId="1" pageNumber="2">
<figureCitation id="131063ADFFCCFFC7398EF9EF53163666" box="[735,852,1578,1604]" captionStart-0="Figs 1 2" captionStart-1="Figs 3 10" captionStart-2="Figs 11 19" captionStart-3="Figs 20 24" captionStart-4="Figs 25 33" captionStartId-0="2.[189,241,1954,1980]" captionStartId-1="4.[189,241,1925,1951]" captionStartId-2="5.[189,241,1876,1902]" captionStartId-3="6.[189,241,1925,1951]" captionStartId-4="7.[189,241,1881,1907]" captionTargetBox-0="[514,1076,1390,1915]" captionTargetBox-1="[225,1361,275,1885]" captionTargetBox-2="[230,1358,265,1836]" captionTargetBox-3="[195,1398,364,1800]" captionTargetBox-4="[215,1372,265,1841]" captionTargetId-0="figure@2.[512,1076,1377,1915]" captionTargetId-1="figure@4.[215,1373,265,1885]" captionTargetId-2="figure@5.[226,1362,265,1836]" captionTargetId-3="figure@6.[189,1398,349,1800]" captionTargetId-4="figure@7.[215,1372,265,1841]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="Figs 1 2. Pukupuku arunachalensis sp. nov. 1. Holotype, ♂, habitus, dorsal view. 2. Paratype, ♀, habitus, dorsal view." captionText-1="Figs 3 10. Pukupuku arunachalensis sp. nov., holotype, ♂. 3. Clypeus. 4. Mandibular horns (dorsal view). 5. Mandibular horns (lateral view). 6. Mentum (arrow indicates the character separating Pukupuku from the closely related genera). 7. Pronotum. 8. Scutellar plate. 9. Right elytron (dorsal view). 10. Elytra." captionText-2="Figs 11 19. Pukupuku arunachalensis sp. nov., holotype, ♂. 11. Protibia. 12. Apical part of mesotibia. 13. Apical part of metatibia. 14. 4 th mesotarsomere. 15. 4 th metatarsomere. 16. Pygidium. 17. Prosternum. 18. Abdomen. 19. Meso- and metasternum. (Arrows indicate the character separating Pukupuku from the closely related genera)." captionText-3="Figs 20 24. Pukupuku arunachalensis sp. nov, holotype, ♂. 20. Aedeagus (dorsal view). 21. Parameres (dorsal view). 22. Aedeagus (lateral view). 23. Map of India. 24. Map of the state of Arunachal Pradesh showing the type locality." captionText-4="Figs 25 33. Pukupuku arunachalensis sp. nov., paratype, ♀. 25. Clypeus. 26. Abdomen. 27. Pronotum. 28. Pygidium. 29. Mentum. 30. Apical part of mesotibia. 31. Apical part of metatibia. 32. 4 th mesotarsomere. 33. 4 th metatarsomere. (Arrows indicate the character separating Pukupuku from the closely related genera)." httpUri-0="https://zenodo.org/record/851563/files/figure.png" httpUri-1="https://zenodo.org/record/851565/files/figure.png" httpUri-2="https://zenodo.org/record/851567/files/figure.png" httpUri-3="https://zenodo.org/record/851569/files/figure.png" httpUri-4="https://zenodo.org/record/851571/files/figure.png" pageId="1" pageNumber="2">Figs 133</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3312CA3FFCCFFC73BECF9AE537F3763" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8B947F28FFCCFFC73BECF9AE517436A7" blockId="1.[189,310,1643,1669]" box="[189,310,1643,1669]" pageId="1" pageNumber="2">
<heading id="D0DCC844FFCCFFC73BECF9AE517436A7" bold="true" box="[189,310,1643,1669]" fontSize="11" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis id="B95FA33AFFCCFFC73BECF9AE517436A7" bold="true" box="[189,310,1643,1669]" pageId="1" pageNumber="2">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B947F28FFCCFFC73BECF95F537F3763" blockId="1.[189,1399,1690,1857]" pageId="1" pageNumber="2">
The newly described species can be easily distinguished by its unique structure of aedeagus, shape and size of mandibles which are sharply pointed at end with a small tooth in middle at outer edge (in lateral view), and four times as long as clypeus, pronotum smooth without setae, pygidium smooth (setae absent in male, present in female). See
<tableCitation id="C6A94A93FFCCFFC739B3F8C15379373C" box="[738,827,1796,1822]" captionStart="Table 1" captionStartId="2.[189,258,268,294]" captionText="Table 1. Differential characters (males) of Pukupuku arunachalensis sp. nov., Pukupuku curtus (Arrow, 1919) and Pukupuku katsurai (Muramoto, 2002)." pageId="1" pageNumber="2">Table 1</tableCitation>
for detailed differential characters separating
<taxonomicName id="4C2B04ABFFCCFFC73E30F8C051323763" authority="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk, 2017" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="B95FA33AFFCCFFC73E30F8C051323763" italics="true" pageId="1" pageNumber="2">P. arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C1E41FFCCFFC73A29F8E251903763" box="[376,466,1831,1857]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
from
<taxonomicName id="4C2B04ABFFCCFFC73948F8ED523D3763" authorityName="Curved" baseAuthorityName="Arrow" baseAuthorityYear="1919" box="[537,639,1832,1857]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B95FA33AFFCCFFC73948F8ED523D3763" box="[537,639,1832,1857]" italics="true" pageId="1" pageNumber="2">P. curtus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2B04ABFFCCFFC739E8F8ED53773763" authorityName="Curved" baseAuthorityName="Muramoto" baseAuthorityYear="2002" box="[697,821,1831,1857]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="katsurai">
<emphasis id="B95FA33AFFCCFFC739E8F8ED53773763" box="[697,821,1831,1857]" italics="true" pageId="1" pageNumber="2">P. katsurai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3312CA3FFCCFFC73BECF8AD55783793" box="[189,1338,1896,1969]" pageId="1" pageNumber="2" type="etymology">
<paragraph id="8B947F28FFCCFFC73BECF8AD510637A0" blockId="1.[189,324,1896,1922]" box="[189,324,1896,1922]" pageId="1" pageNumber="2">
<heading id="D0DCC844FFCCFFC73BECF8AD510637A0" bold="true" box="[189,324,1896,1922]" fontSize="11" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis id="B95FA33AFFCCFFC73BECF8AD510637A0" bold="true" box="[189,324,1896,1922]" pageId="1" pageNumber="2">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B947F28FFCCFFC73BECF85255783793" blockId="1.[189,1338,1943,1969]" box="[189,1338,1943,1969]" pageId="1" pageNumber="2">
The name
<taxonomicName id="4C2B04ABFFCCFFC73A68F85251AE3793" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[313,492,1943,1969]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="arunachalensis">
<emphasis id="B95FA33AFFCCFFC73A68F85251AE3793" box="[313,492,1943,1969]" italics="true" pageId="1" pageNumber="2">arunachalensis</emphasis>
</taxonomicName>
refers to the Northeastern Himalayan state of India, Arunachal Pradesh.
</paragraph>
</subSubSection>
<caption id="DF542FA0FFCFFFC43BECFEC952BD316B" pageId="2" pageNumber="3">
<paragraph id="8B947F28FFCFFFC43BECFEC952BD316B" blockId="2.[189,1398,267,329]" pageId="2" pageNumber="3">
<emphasis id="B95FA33AFFCFFFC43BECFEC9515D3107" bold="true" box="[189,287,267,294]" pageId="2" pageNumber="3">Table 1.</emphasis>
Differential characters (males) of
<taxonomicName id="4C2B04ABFFCFFFC439E1FECE539D3107" authority="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk, 2017" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[688,991,267,293]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="B95FA33AFFCFFFC439E1FECE539D3107" box="[688,991,267,293]" italics="true" pageId="2" pageNumber="3">Pukupuku arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C1E41FFCFFFC438B6FEC954013104" box="[999,1091,268,294]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C2B04ABFFCFFFC43F1FFECE5143316B" authority="(Arrow, 1919)" authorityName="Curved" authorityYear="1919" baseAuthorityName="Arrow" baseAuthorityYear="1919" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B95FA33AFFCFFFC43F1FFECE55513107" box="[1102,1299,267,293]" italics="true" pageId="2" pageNumber="3">Pukupuku curtus</emphasis>
(Arrow, 1919)
</taxonomicName>
and
<taxonomicName id="4C2B04ABFFCFFFC43A6BFEEA52B9316B" authority="(Muramoto, 2002)" authorityName="Curved" authorityYear="2002" baseAuthorityName="Muramoto" baseAuthorityYear="2002" box="[314,763,303,329]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="katsurai">
<emphasis id="B95FA33AFFCFFFC43A6BFEEA5255316B" box="[314,535,303,329]" italics="true" pageId="2" pageNumber="3">Pukupuku katsurai</emphasis>
(Muramoto, 2002)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B947F28FFCFFFC43BECFEB355723523" pageId="2" pageNumber="3">
<table id="F92B8D88FFCF00393BECFEB3552C3523" box="[189,1390,374,1281]" gridcols="4" gridrows="9" pageId="2" pageNumber="3">
<tr id="351B7D6AFFCF00393BECFEB3552C3219" box="[189,1390,374,571]" gridrow="0" pageId="2" pageNumber="3">
<th id="76CA1416FFCF00393BECFEB351023219" box="[189,320,374,571]" gridcol="0" gridrow="0" pageId="2" pageNumber="3">
<emphasis id="B95FA33AFFCFFFC43BECFEB35169318D" bold="true" pageId="2" pageNumber="3">Species / character</emphasis>
Shape of mandibles
</th>
<th id="76CA1416FFCF00393A1FFEB352EA3219" box="[334,680,374,571]" gridcol="1" gridrow="0" pageId="2" pageNumber="3">
<emphasis id="B95FA33AFFCFFFC43A3BFEB35264318D" bold="true" pageId="2" pageNumber="3">
<taxonomicName id="4C2B04ABFFCFFFC43A3BFEB352CF31AC" authority="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk, 2017" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[362,653,374,398]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="B95FA33AFFCFFFC43A3BFEB352CF31AC" bold="true" box="[362,653,374,398]" italics="true" pageId="2" pageNumber="3">Pukupuku arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C1E41FFCFFFC43A80FE525264318D" box="[465,550,407,431]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Sharply pointed at end, with a small tooth in middle at outer edge (in lateral view) (Fig. 5)
</th>
<th id="76CA1416FFCF003939E2FEB354483219" box="[691,1034,374,571]" gridcol="2" gridrow="0" pageId="2" pageNumber="3">
<taxonomicName id="4C2B04ABFFCFFFC439AEFEB3534631F9" authority="(Arrow, 1919) Curved" authorityName="Curved" baseAuthorityName="Arrow" baseAuthorityYear="1919" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B95FA33AFFCFFFC439AEFEB353EF318C" bold="true" pageId="2" pageNumber="3">
<emphasis id="B95FA33AFFCFFFC439AEFEB353FD31AC" bold="true" box="[767,959,374,398]" italics="true" pageId="2" pageNumber="3">Pukupuku curtus</emphasis>
(Arrow, 1919)
</emphasis>
Curved
</taxonomicName>
upward toothed at outer edge (Fig. 34)
</th>
<th id="76CA1416FFCF00393F46FEB3552C3219" box="[1047,1390,374,571]" gridcol="3" gridrow="0" pageId="2" pageNumber="3">
<taxonomicName id="4C2B04ABFFCFFFC43F09FEB3543F31F9" authority="(Muramoto, 2002) Curved" authorityName="Curved" baseAuthorityName="Muramoto" baseAuthorityYear="2002" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="katsurai">
<emphasis id="B95FA33AFFCFFFC43F09FEB35568318C" bold="true" pageId="2" pageNumber="3">
<emphasis id="B95FA33AFFCFFFC43F09FEB3556E31AC" bold="true" box="[1112,1324,374,398]" italics="true" pageId="2" pageNumber="3">Pukupuku katsurai</emphasis>
(Muramoto, 2002)
</emphasis>
Curved
</taxonomicName>
upward and slightly inward near apex, with a long tooth before middle of upper surface (Muramoto 2002: fig. 2)
</th>
</tr>
<tr id="351B7D6AFFCF00393BECFD96552C328E" box="[189,1390,595,684]" gridrow="1" pageId="2" pageNumber="3">
<th id="76CA1416FFCF00393BECFD965102328E" box="[189,320,595,684]" gridcol="0" gridrow="1" pageId="2" pageNumber="3">Size of mandibles</th>
<td id="76CA1416FFCF00393A1FFD9652EA328E" box="[334,680,595,684]" gridcol="1" gridrow="1" pageId="2" pageNumber="3">Four times as long as clypeus, 16.0 mm (Figs 45)</td>
<td id="76CA1416FFCF003939E2FD965448328E" box="[691,1034,595,684]" gridcol="2" gridrow="1" pageId="2" pageNumber="3">Not long, 3.0 mm in length (Fig. 34)</td>
<td id="76CA1416FFCF00393F46FD96552C328E" box="[1047,1390,595,684]" gridcol="3" gridrow="1" pageId="2" pageNumber="3">Twice as long as length of clypeus, 4.04.5 mm (Muramoto 2002: figs 12)</td>
</tr>
<tr id="351B7D6AFFCF00393BECFD01552C32DE" box="[189,1390,708,764]" gridrow="2" pageId="2" pageNumber="3">
<th id="76CA1416FFCF00393BECFD01510232DE" box="[189,320,708,764]" gridcol="0" gridrow="2" pageId="2" pageNumber="3">Sculpture of pronotum</th>
<td id="76CA1416FFCF00393A1FFD0152EA32DE" box="[334,680,708,764]" gridcol="1" gridrow="2" pageId="2" pageNumber="3">Entirely smooth, without setae (Fig. 7)</td>
<td id="76CA1416FFCF003939E2FD01544832DE" box="[691,1034,708,764]" gridcol="2" gridrow="2" pageId="2" pageNumber="3">Densely covered with minute setae</td>
<td id="76CA1416FFCF00393F46FD01552C32DE" box="[1047,1390,708,764]" gridcol="3" gridrow="2" pageId="2" pageNumber="3">Densely covered with minute setae</td>
</tr>
<tr id="351B7D6AFFCF00393BECFCD0552C336F" box="[189,1390,789,845]" gridrow="3" pageId="2" pageNumber="3">
<th id="76CA1416FFCF00393BECFCD05102336F" box="[189,320,789,845]" gridcol="0" gridrow="3" pageId="2" pageNumber="3">Pygidium</th>
<td id="76CA1416FFCF00393A1FFCD052EA336F" box="[334,680,789,845]" gridcol="1" gridrow="3" pageId="2" pageNumber="3">Smooth, setae absent in male; in female with long setae</td>
<td id="76CA1416FFCF003939E2FCD05448336F" box="[691,1034,789,845]" gridcol="2" gridrow="3" pageId="2" pageNumber="3">Impunctate covered with long setae</td>
<td id="76CA1416FFCF00393F46FCD0552C336F" box="[1047,1390,789,845]" gridcol="3" gridrow="3" pageId="2" pageNumber="3">Coriaceous, posterior margin with a row of setae</td>
</tr>
<tr id="351B7D6AFFCF00393BECFCA3552C339C" box="[189,1390,870,958]" gridrow="4" pageId="2" pageNumber="3">
<th id="76CA1416FFCF00393BECFCA35102339C" box="[189,320,870,958]" gridcol="0" gridrow="4" pageId="2" pageNumber="3">Aedeagus</th>
<td id="76CA1416FFCF00393A1FFCA352EA339C" box="[334,680,870,958]" gridcol="1" gridrow="4" pageId="2" pageNumber="3">Symmetrical, differently shaped (Figs 2022)</td>
<td id="76CA1416FFCF003939E2FCA35448339C" box="[691,1034,870,958]" gridcol="2" gridrow="4" pageId="2" pageNumber="3">Symmetrical, differently shaped (Muramoto 2002: fig. 6)</td>
<td id="76CA1416FFCF00393F46FCA3552C339C" box="[1047,1390,870,958]" gridcol="3" gridrow="4" pageId="2" pageNumber="3">Symmetrical, differently shaped (Muramoto 2002: fig. 3)</td>
</tr>
<tr id="351B7D6AFFCF00393BECFC13552C340C" box="[189,1390,982,1070]" gridrow="5" pageId="2" pageNumber="3">
<th id="76CA1416FFCF00393BECFC135102340C" box="[189,320,982,1070]" gridcol="0" gridrow="5" pageId="2" pageNumber="3">Length (including mandibles)</th>
<td id="76CA1416FFCF00393A1FFC1352EA340C" box="[334,680,982,1070]" gridcol="1" gridrow="5" pageId="2" pageNumber="3">32.0 mm</td>
<td id="76CA1416FFCF003939E2FC135448340C" box="[691,1034,982,1070]" gridcol="2" gridrow="5" pageId="2" pageNumber="3">1819.5 mm</td>
<td id="76CA1416FFCF00393F46FC13552C340C" box="[1047,1390,982,1070]" gridcol="3" gridrow="5" pageId="2" pageNumber="3">20.521 mm</td>
</tr>
<tr id="351B7D6AFFCF00393BECFB82552C34BD" box="[189,1390,1095,1183]" gridrow="6" pageId="2" pageNumber="3">
<th id="76CA1416FFCF00393BECFB82510234BD" box="[189,320,1095,1183]" gridcol="0" gridrow="6" pageId="2" pageNumber="3">Length (excluding mandibles)</th>
<td id="76CA1416FFCF00393A1FFB8252EA34BD" box="[334,680,1095,1183]" gridcol="1" gridrow="6" pageId="2" pageNumber="3">20.0 mm</td>
<td id="76CA1416FFCF003939E2FB82544834BD" box="[691,1034,1095,1183]" gridcol="2" gridrow="6" pageId="2" pageNumber="3">1516.5 mm</td>
<td id="76CA1416FFCF00393F46FB82552C34BD" box="[1047,1390,1095,1183]" gridcol="3" gridrow="6" pageId="2" pageNumber="3">16.5 mm</td>
</tr>
<tr id="351B7D6AFFCF00393BECFB7D552C34F2" box="[189,1390,1208,1232]" gridrow="7" pageId="2" pageNumber="3">
<th id="76CA1416FFCF00393BECFB7D510234F2" box="[189,320,1208,1232]" gridcol="0" gridrow="7" pageId="2" pageNumber="3">Width</th>
<td id="76CA1416FFCF00393A1FFB7D52EA34F2" box="[334,680,1208,1232]" gridcol="1" gridrow="7" pageId="2" pageNumber="3">19.0 mm</td>
<td id="76CA1416FFCF003939E2FB7D544834F2" box="[691,1034,1208,1232]" gridcol="2" gridrow="7" pageId="2" pageNumber="3">99.5 mm</td>
<td id="76CA1416FFCF00393F46FB7D552C34F2" box="[1047,1390,1208,1232]" gridcol="3" gridrow="7" pageId="2" pageNumber="3">11 mm</td>
</tr>
<tr id="351B7D6AFFCF00393BECFB2D552C3523" box="[189,1390,1256,1281]" gridrow="8" pageId="2" pageNumber="3">
<th id="76CA1416FFCF00393BECFB2D51023523" box="[189,320,1256,1281]" gridcol="0" gridrow="8" pageId="2" pageNumber="3">Distribution</th>
<td id="76CA1416FFCF00393A1FFB2D52EA3523" box="[334,680,1256,1281]" gridcol="1" gridrow="8" pageId="2" pageNumber="3">Arunachal Pradesh (Figs 2324)</td>
<td id="76CA1416FFCF003939E2FB2D54483523" box="[691,1034,1256,1281]" gridcol="2" gridrow="8" pageId="2" pageNumber="3">Laos, North Thailand</td>
<td id="76CA1416FFCF00393F46FB2D552C3523" box="[1047,1390,1256,1281]" gridcol="3" gridrow="8" pageId="2" pageNumber="3">Vietnam (Cao Bang)</td>
</tr>
</table>
</paragraph>
<caption id="DF542FA0FFCFFFC43BECF86751EC37C2" httpUri="https://zenodo.org/record/851563/files/figure.png" pageId="2" pageNumber="3" targetBox="[514,1076,1390,1915]" targetPageId="2">
<paragraph id="8B947F28FFCFFFC43BECF86751EC37C2" blockId="2.[189,1398,1954,2016]" pageId="2" pageNumber="3">
<emphasis id="B95FA33AFFCFFFC43BECF8675173379E" bold="true" box="[189,305,1954,1980]" pageId="2" pageNumber="3">Figs 12.</emphasis>
<taxonomicName id="4C2B04ABFFCFFFC43A6CF8665233379F" authority="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk, 2017" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[317,625,1955,1981]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="B95FA33AFFCFFFC43A6CF8665233379F" box="[317,625,1955,1981]" italics="true" pageId="2" pageNumber="3">Pukupuku arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C1E41FFCFFFC4392FF866529F379F" box="[638,733,1955,1981]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B95FA33AFFCFFFC439B8F86752BA379E" bold="true" box="[745,760,1954,1980]" pageId="2" pageNumber="3">1</emphasis>
. Holotype, ♂, habitus, dorsal view.
<emphasis id="B95FA33AFFCFFFC43FECF867548E379E" bold="true" box="[1213,1228,1954,1980]" pageId="2" pageNumber="3">2</emphasis>
. Paratype, ♀, habitus, dorsal view.
</paragraph>
</caption>
<subSubSection id="C3312CA3FFCEFFC53BECFECE54EE3307" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8B947F28FFCEFFC53BECFECE51E83107" blockId="3.[189,426,267,293]" box="[189,426,267,293]" pageId="3" pageNumber="4">
<heading id="D0DCC844FFCEFFC53BECFECE51E83107" bold="true" box="[189,426,267,293]" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<emphasis id="B95FA33AFFCEFFC53BECFECE51E83107" bold="true" box="[189,426,267,293]" pageId="3" pageNumber="4">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B437575FFCEFFC53BBBFEFF53D431B9" ID-GBIF-Occurrence="1587001884" collectingDate="1985-10-04" collectionCode="ZSI" collectorName="S. Biswas" country="India" location="Namdapha / Tirap / Mizo / Vijay Nagar" municipality="Namdapha / Tirap / Mizo / Vijay Nagar" pageId="3" pageNumber="4" specimenCode="21630/H4A" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<paragraph id="8B947F28FFCEFFC53BBBFEFF511E3176" blockId="3.[189,1398,314,411]" box="[234,348,314,340]" pageId="3" pageNumber="4">
<heading id="D0DCC844FFCEFFC53BBBFEFF511E3176" bold="true" box="[234,348,314,340]" fontSize="11" level="2" pageId="3" pageNumber="4" reason="6">
<emphasis id="B95FA33AFFCEFFC53BBBFEFF511E3176" bold="true" box="[234,348,314,340]" pageId="3" pageNumber="4">
<typeStatus id="5490C18AFFCEFFC53BBBFEFF511E3176" box="[234,348,314,340]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B947F28FFCEFFC53BECFE9853D431B9" blockId="3.[189,1398,314,411]" pageId="3" pageNumber="4">
<collectingCountry id="F33C3FB8FFCEFFC53BECFE9851573155" box="[189,277,349,375]" name="India" pageId="3" pageNumber="4">INDIA</collectingCountry>
:
<specimenCount id="9D2DB4A1FFCEFFC53A4EFE9851773155" box="[287,309,349,375]" pageId="3" pageNumber="4" type="male"></specimenCount>
labeled: “
<collectingCountry id="F33C3FB8FFCEFFC53AFBFE9852413155" box="[426,515,349,375]" name="India" pageId="3" pageNumber="4">INDIA</collectingCountry>
,
<collectingRegion id="49EFB1CAFFCEFFC5395DFE9852A83155" box="[524,746,349,375]" country="India" name="Arunachal Pradesh" pageId="3" pageNumber="4">Arunachal Pradesh</collectingRegion>
/
<collectingMunicipality id="6BF0E552FFCEFFC53851FE98548B3155" box="[768,1225,349,375]" pageId="3" pageNumber="4">Namdapha / Tirap / Mizo / Vijay Nagar</collectingMunicipality>
/
<date id="FF9559E8FFCEFFC53F8EFE98552A3155" box="[1247,1384,348,375]" pageId="3" pageNumber="4" value="1985-10-04">
<collectingDate id="EFD1A000FFCEFFC53F8EFE98552A3155" box="[1247,1384,348,375]" pageId="3" pageNumber="4" value="1985-10-04">4 Oct. 1985</collectingDate>
</date>
/ collected
<collectorName id="26DE1AFEFFCEFFC53A7FFE4551E631B8" box="[302,420,384,410]" pageId="3" pageNumber="4">S. Biswas</collectorName>
” (
<collectionCode id="ED3AE7EDFFCEFFC53AEFFE4551AE31B8" box="[446,492,384,410]" collectionName="ZSI" pageId="3" pageNumber="4">ZSI</collectionCode>
Registration Number:
<specimenCode id="DB8DD753FFCEFFC539ABFE4553C831B9" box="[762,906,384,411]" pageId="3" pageNumber="4">21630/H4A</specimenCode>
).
</paragraph>
</materialsCitation>
<materialsCitation id="3B437575FFCEFFC53BBBFE04511B3200" ID-GBIF-Occurrence="1587001883" collectionCode="ZSI" country="India" location="INDIA" pageId="3" pageNumber="4" specimenCode="21630/H4A, 21635/H4A" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<paragraph id="8B947F28FFCEFFC53BBBFE04512531F9" blockId="3.[189,1399,449,546]" box="[234,359,449,475]" pageId="3" pageNumber="4">
<heading id="D0DCC844FFCEFFC53BBBFE04512531F9" bold="true" box="[234,359,449,475]" fontSize="11" level="2" pageId="3" pageNumber="4" reason="6">
<emphasis id="B95FA33AFFCEFFC53BBBFE04512531F9" bold="true" box="[234,359,449,475]" pageId="3" pageNumber="4">
<typeStatus id="5490C18AFFCEFFC53BBBFE04512531F9" box="[234,359,449,475]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B947F28FFCEFFC53BECFE20511B3200" blockId="3.[189,1399,449,546]" pageId="3" pageNumber="4">
<collectingCountry id="F33C3FB8FFCEFFC53BECFE20515731DD" box="[189,277,485,511]" name="India" pageId="3" pageNumber="4">INDIA</collectingCountry>
:
<specimenCount id="9D2DB4A1FFCEFFC53A72FE20512B31DD" box="[291,361,485,511]" pageId="3" pageNumber="4" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="9D2DB4A1FFCEFFC53AF8FE2051B231DD" box="[425,496,485,511]" pageId="3" pageNumber="4" type="female">3 ♀♀</specimenCount>
, same locality data as holotype (
<collectionCode id="ED3AE7EDFFCEFFC538D4FE2053F131DD" box="[901,947,485,511]" collectionName="ZSI" pageId="3" pageNumber="4">ZSI</collectionCode>
Registration Number:
<specimenCode id="DB8DD753FFCEFFC53F9DFE21551731DC" box="[1228,1365,484,510]" pageId="3" pageNumber="4">21630/H4A</specimenCode>
to
<specimenCode id="DB8DD753FFCEFFC53BECFDC2510F3200" box="[189,333,519,546]" pageId="3" pageNumber="4">21635/H4A</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8B947F28FFCEFFC53BECFD8C511F3241" blockId="3.[189,349,585,611]" box="[189,349,585,611]" pageId="3" pageNumber="4">
<heading id="D0DCC844FFCEFFC53BECFD8C511F3241" bold="true" box="[189,349,585,611]" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<emphasis id="B95FA33AFFCEFFC53BECFD8C511F3241" bold="true" box="[189,349,585,611]" pageId="3" pageNumber="4">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B947F28FFCEFFC53BECFDBD51253297" blockId="3.[189,1398,632,693]" pageId="3" pageNumber="4">INDIA: Arunachal Pradesh state, Tirap district (now in Changlang district), Vijay Nagar, Namdapha National Park.</paragraph>
<paragraph id="8B947F28FFCEFFC53BECFD1951C432D4" blockId="3.[189,390,732,758]" box="[189,390,732,758]" pageId="3" pageNumber="4">
<heading id="D0DCC844FFCEFFC53BECFD1951C432D4" bold="true" box="[189,390,732,758]" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<emphasis id="B95FA33AFFCEFFC53BECFD1951C432D4" bold="true" box="[189,390,732,758]" pageId="3" pageNumber="4">Type depository</emphasis>
</heading>
</paragraph>
<paragraph id="8B947F28FFCEFFC53BECFCCE54EE3307" blockId="3.[189,1196,779,805]" box="[189,1196,779,805]" pageId="3" pageNumber="4">Deposited in the collections of the Zoological Survey of India, New Alipore, Kolkata.</paragraph>
</subSubSection>
<subSubSection id="C3312CA3FFCEFFCE3BECFC8953DA33A4" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4" type="description">
<paragraph id="8B947F28FFCEFFC53BECFC8952323344" blockId="3.[189,1399,844,952]" box="[189,624,844,870]" pageId="3" pageNumber="4">
<emphasis id="B95FA33AFFCEFFC53BECFC89510C3344" bold="true" box="[189,334,844,870]" pageId="3" pageNumber="4">Description</emphasis>
(holotype,
<figureCitation id="131063ADFFCEFFC53A85FC89525F3344" box="[468,541,844,870]" captionStart="Figs 1 2" captionStartId="2.[189,241,1954,1980]" captionTargetBox="[514,1076,1390,1915]" captionTargetId="figure@2.[512,1076,1377,1915]" captionTargetPageId="2" captionText="Figs 1 2. Pukupuku arunachalensis sp. nov. 1. Holotype, ♂, habitus, dorsal view. 2. Paratype, ♀, habitus, dorsal view." httpUri="https://zenodo.org/record/851563/files/figure.png" pageId="3" pageNumber="4">Figs 1</figureCitation>
,
<figureCitation id="131063ADFFCEFFC5397DFC89522A3344" box="[556,616,844,870]" captionStart-0="Figs 3 10" captionStart-1="Figs 11 19" captionStart-2="Figs 20 24" captionStartId-0="4.[189,241,1925,1951]" captionStartId-1="5.[189,241,1876,1902]" captionStartId-2="6.[189,241,1925,1951]" captionTargetBox-0="[225,1361,275,1885]" captionTargetBox-1="[230,1358,265,1836]" captionTargetBox-2="[195,1398,364,1800]" captionTargetId-0="figure@4.[215,1373,265,1885]" captionTargetId-1="figure@5.[226,1362,265,1836]" captionTargetId-2="figure@6.[189,1398,349,1800]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Figs 3 10. Pukupuku arunachalensis sp. nov., holotype, ♂. 3. Clypeus. 4. Mandibular horns (dorsal view). 5. Mandibular horns (lateral view). 6. Mentum (arrow indicates the character separating Pukupuku from the closely related genera). 7. Pronotum. 8. Scutellar plate. 9. Right elytron (dorsal view). 10. Elytra." captionText-1="Figs 11 19. Pukupuku arunachalensis sp. nov., holotype, ♂. 11. Protibia. 12. Apical part of mesotibia. 13. Apical part of metatibia. 14. 4 th mesotarsomere. 15. 4 th metatarsomere. 16. Pygidium. 17. Prosternum. 18. Abdomen. 19. Meso- and metasternum. (Arrows indicate the character separating Pukupuku from the closely related genera)." captionText-2="Figs 20 24. Pukupuku arunachalensis sp. nov, holotype, ♂. 20. Aedeagus (dorsal view). 21. Parameres (dorsal view). 22. Aedeagus (lateral view). 23. Map of India. 24. Map of the state of Arunachal Pradesh showing the type locality." httpUri-0="https://zenodo.org/record/851565/files/figure.png" httpUri-1="https://zenodo.org/record/851567/files/figure.png" httpUri-2="https://zenodo.org/record/851569/files/figure.png" pageId="3" pageNumber="4">324</figureCitation>
)
</paragraph>
<paragraph id="8B947F28FFCEFFC53BECFCBE528A339A" blockId="3.[189,1399,844,952]" pageId="3" pageNumber="4">MEASUREMENTS. Total body length of males (excluding mandibles): 20.021.0 mm (holotype 20.0 mm), total body length of females: 19.020.0 mm.</paragraph>
<paragraph id="8B947F28FFCEFFC53BECFC25512B34A5" blockId="3.[189,1399,991,1159]" pageId="3" pageNumber="4">DETAILED MEASUREMENTS. Mandibular horn: length 16.0 mm. Clypeus: length 4.0 mm, inter ocular distance (distance between inner sides of each eye) 5.8 mm. Pronotum: length 9.9 mm, maximum width 17.5 mm. Pygidium: length 4.5 mm, width 12.5 mm. Elytra: length 18.4 mm, maximum width 19.3 mm. Aedeagus: length 7.6 mm, maximum width 2.5 mm; Parameres: length 3.0 mm, maximum width 2.2 mm.</paragraph>
<paragraph id="8B947F28FFCEFFC53BECFB6B527834EA" blockId="3.[189,570,1198,1224]" box="[189,570,1198,1224]" pageId="3" pageNumber="4">BODY. Broadly oval and convex.</paragraph>
<paragraph id="8B947F28FFCEFFC53BECFB3553EB35DD" blockId="3.[189,1399,1263,1536]" pageId="3" pageNumber="4">
HEAD (
<figureCitation id="131063ADFFCEFFC53A5AFB2A51363528" box="[267,372,1263,1290]" captionStart="Figs 3 10" captionStartId="4.[189,241,1925,1951]" captionTargetBox="[225,1361,275,1885]" captionTargetId="figure@4.[215,1373,265,1885]" captionTargetPageId="4" captionText="Figs 3 10. Pukupuku arunachalensis sp. nov., holotype, ♂. 3. Clypeus. 4. Mandibular horns (dorsal view). 5. Mandibular horns (lateral view). 6. Mentum (arrow indicates the character separating Pukupuku from the closely related genera). 7. Pronotum. 8. Scutellar plate. 9. Right elytron (dorsal view). 10. Elytra." httpUri="https://zenodo.org/record/851565/files/figure.png" pageId="3" pageNumber="4">Figs 36</figureCitation>
). Glabrous, entirely smooth without any hairs dorsally. Clypeus trapezoidal, convergent in basal half, anterior clypeal margin slightly arcuate with few short and long hairs, anterior angles rounded, surface with few scattered punctures in middle and at sides, slightly convex in middle, clypeofrontal suture nearly absent with only small carina at each side, vertex with moderately large rugopunctation. Area near each eye smooth and shining. Eye canthus spatulate, wide near end and small at base, surface smooth. Labrum straight with long hairs at margin. Mentum with a rather large rounded hollow at middle. Mandibles flat and broad at base, sharply pointed at end with a small tooth in middle (in lateral view), almost four times as long as length of clypeus.
</paragraph>
<paragraph id="8B947F28FFCEFFC53BECF9E251673688" blockId="3.[189,1399,1575,1706]" pageId="3" pageNumber="4">
PRONOTUM (
<figureCitation id="131063ADFFCEFFC53A1BF9E251D73663" box="[330,405,1575,1601]" captionStart="Figs 3 10" captionStartId="4.[189,241,1925,1951]" captionTargetBox="[225,1361,275,1885]" captionTargetId="figure@4.[215,1373,265,1885]" captionTargetPageId="4" captionText="Figs 3 10. Pukupuku arunachalensis sp. nov., holotype, ♂. 3. Clypeus. 4. Mandibular horns (dorsal view). 5. Mandibular horns (lateral view). 6. Mentum (arrow indicates the character separating Pukupuku from the closely related genera). 7. Pronotum. 8. Scutellar plate. 9. Right elytron (dorsal view). 10. Elytra." httpUri="https://zenodo.org/record/851565/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
). Convex, 1.9 times as wide as long, widest before middle, without any hairs or setae on dorsal surface, with rather small but indistinct punctures in middle, laterally smooth, narrowly marginate except near base, base distinctly lobed before scutellum, anterior, lateral, and hind angles rounded.
</paragraph>
<paragraph id="8B947F28FFCEFFC53BECF91752033732" blockId="3.[189,1398,1746,1808]" pageId="3" pageNumber="4">
SCUTELLAR PLATE (
<figureCitation id="131063ADFFCEFFC53ADFF917519136CE" box="[398,467,1746,1772]" captionStart="Figs 3 10" captionStartId="4.[189,241,1925,1951]" captionTargetBox="[225,1361,275,1885]" captionTargetId="figure@4.[215,1373,265,1885]" captionTargetPageId="4" captionText="Figs 3 10. Pukupuku arunachalensis sp. nov., holotype, ♂. 3. Clypeus. 4. Mandibular horns (dorsal view). 5. Mandibular horns (lateral view). 6. Mentum (arrow indicates the character separating Pukupuku from the closely related genera). 7. Pronotum. 8. Scutellar plate. 9. Right elytron (dorsal view). 10. Elytra." httpUri="https://zenodo.org/record/851565/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
). Triangular, base carinate, lateral sides curved, apex rounded, densely punctate on sides with a line of punctures.
</paragraph>
<paragraph id="8B947F28FFCEFFC53BECF8F2533C37FF" blockId="3.[189,1398,1846,2014]" pageId="3" pageNumber="4">
ELYTRA (
<figureCitation id="131063ADFFCEFFC53A76F8F351DD3773" box="[295,415,1846,1873]" captionStart="Figs 3 10" captionStartId="4.[189,241,1925,1951]" captionTargetBox="[225,1361,275,1885]" captionTargetId="figure@4.[215,1373,265,1885]" captionTargetPageId="4" captionText="Figs 3 10. Pukupuku arunachalensis sp. nov., holotype, ♂. 3. Clypeus. 4. Mandibular horns (dorsal view). 5. Mandibular horns (lateral view). 6. Mentum (arrow indicates the character separating Pukupuku from the closely related genera). 7. Pronotum. 8. Scutellar plate. 9. Right elytron (dorsal view). 10. Elytra." httpUri="https://zenodo.org/record/851565/files/figure.png" pageId="3" pageNumber="4">Figs 910</figureCitation>
). Convex, almost as wide as long, widest at middle; rather densely and closely punctate near sutural margins from base to apex, rather sparsely and finely punctate in middle and at lateral sides, entirely yellow with a brown spot behind scutellar apex near sutural margin, brown pit at base in middle, two transverse bands, one large and one small, at lateral side on humeral umbone (in lateral view), apex with a large brown band on each elytron.
</paragraph>
<caption id="DF542FA0FFC9FFC23BECF840551937C7" httpUri="https://zenodo.org/record/851565/files/figure.png" pageId="4" pageNumber="5" targetBox="[225,1361,275,1885]" targetPageId="4">
<paragraph id="8B947F28FFC9FFC23BECF840551937C7" blockId="4.[189,1399,1925,2022]" pageId="4" pageNumber="5">
<emphasis id="B95FA33AFFC9FFC23BECF840517537BD" bold="true" box="[189,311,1925,1951]" pageId="4" pageNumber="5">Figs 310.</emphasis>
<taxonomicName id="4C2B04ABFFC9FFC23A6DF840522A37BD" authority="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk, 2017" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[316,616,1925,1951]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="B95FA33AFFC9FFC23A6DF840522A37BD" box="[316,616,1925,1951]" italics="true" pageId="4" pageNumber="5">Pukupuku arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C1E41FFC9FFC2393CF84352853782" box="[621,711,1926,1952]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂.
<emphasis id="B95FA33AFFC9FFC23835F840533137BD" bold="true" box="[868,883,1925,1951]" pageId="4" pageNumber="5">3</emphasis>
. Clypeus.
<emphasis id="B95FA33AFFC9FFC238BBF84053BB37BD" bold="true" box="[1002,1017,1925,1951]" pageId="4" pageNumber="5">4</emphasis>
. Mandibular horns (dorsal view).
<emphasis id="B95FA33AFFC9FFC23BECF86D508E37E0" bold="true" box="[189,204,1960,1986]" pageId="4" pageNumber="5">5</emphasis>
. Mandibular horns (lateral view).
<emphasis id="B95FA33AFFC9FFC2390AF86D522837E0" bold="true" box="[603,618,1960,1986]" pageId="4" pageNumber="5">6</emphasis>
. Mentum (arrow indicates the character separating
<taxonomicName id="4C2B04ABFFC9FFC23F90F86D557437E0" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[1217,1334,1960,1986]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B95FA33AFFC9FFC23F90F86D557437E0" box="[1217,1334,1960,1986]" italics="true" pageId="4" pageNumber="5">Pukupuku</emphasis>
</taxonomicName>
from the closely related genera).
<emphasis id="B95FA33AFFC9FFC23951F809524D37C4" bold="true" box="[512,527,1996,2022]" pageId="4" pageNumber="5">7</emphasis>
. Pronotum.
<emphasis id="B95FA33AFFC9FFC239F1F80E52ED37C7" bold="true" box="[672,687,1995,2021]" pageId="4" pageNumber="5">8</emphasis>
. Scutellar plate.
<emphasis id="B95FA33AFFC9FFC23824F80E53C637C7" bold="true" box="[885,900,1995,2021]" pageId="4" pageNumber="5">9</emphasis>
. Right elytron (dorsal view).
<emphasis id="B95FA33AFFC9FFC23F8EF80E54BE37C7" bold="true" box="[1247,1276,1995,2021]" pageId="4" pageNumber="5">10</emphasis>
. Elytra.
</paragraph>
</caption>
<caption id="DF542FA0FFC8FFC33BECF89151B837FA" httpUri="https://zenodo.org/record/851567/files/figure.png" pageId="5" pageNumber="6" targetBox="[230,1358,265,1836]" targetPageId="5">
<paragraph id="8B947F28FFC8FFC33BECF89151B837FA" blockId="5.[189,1399,1876,2008]" pageId="5" pageNumber="6">
<emphasis id="B95FA33AFFC8FFC33BECF891510A374C" bold="true" box="[189,328,1876,1902]" pageId="5" pageNumber="6">Figs 1119.</emphasis>
<taxonomicName id="4C2B04ABFFC8FFC33A01F89152C3374C" authority="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk, 2017" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[336,641,1876,1902]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="B95FA33AFFC8FFC33A01F89152C3374C" box="[336,641,1876,1902]" italics="true" pageId="5" pageNumber="6">Pukupuku arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C1E41FFC8FFC339D8F89152A5374C" box="[649,743,1876,1902]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂.
<emphasis id="B95FA33AFFC8FFC338DFF89153EA374C" bold="true" box="[910,936,1876,1902]" pageId="5" pageNumber="6">11</emphasis>
. Protibia.
<emphasis id="B95FA33AFFC8FFC33F77F8915401374C" bold="true" box="[1062,1091,1876,1902]" pageId="5" pageNumber="6">12</emphasis>
. Apical part of mesotibia.
<emphasis id="B95FA33AFFC8FFC33BECF8B2509837B3" bold="true" box="[189,218,1911,1937]" pageId="5" pageNumber="6">13</emphasis>
. Apical part of metatibia.
<emphasis id="B95FA33AFFC8FFC33954F8B2526037B3" bold="true" box="[517,546,1911,1937]" pageId="5" pageNumber="6">14</emphasis>
. 4th mesotarsomere.
<emphasis id="B95FA33AFFC8FFC3385EF8B2536E37B3" bold="true" box="[783,812,1911,1937]" pageId="5" pageNumber="6">15</emphasis>
. 4th metatarsomere.
<emphasis id="B95FA33AFFC8FFC33F45F8B2547337B3" bold="true" box="[1044,1073,1911,1937]" pageId="5" pageNumber="6">16</emphasis>
. Pygidium.
<emphasis id="B95FA33AFFC8FFC33FEFF8B2549937B3" bold="true" box="[1214,1243,1911,1937]" pageId="5" pageNumber="6">17</emphasis>
. Prosternum.
<emphasis id="B95FA33AFFC8FFC33BECF85F50983796" bold="true" box="[189,218,1946,1972]" pageId="5" pageNumber="6">18</emphasis>
. Abdomen.
<emphasis id="B95FA33AFFC8FFC33A3CF85F51C83796" bold="true" box="[365,394,1946,1972]" pageId="5" pageNumber="6">19</emphasis>
. Meso- and metasternum. (Arrows indicate the character separating
<taxonomicName id="4C2B04ABFFC8FFC33FEEF85F55763796" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[1215,1332,1946,1972]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B95FA33AFFC8FFC33FEEF85F55763796" box="[1215,1332,1946,1972]" italics="true" pageId="5" pageNumber="6">Pukupuku</emphasis>
</taxonomicName>
from the closely related genera).
</paragraph>
</caption>
<caption id="DF542FA0FFCBFFC03BECF84051A937C7" httpUri="https://zenodo.org/record/851569/files/figure.png" pageId="6" pageNumber="7" targetBox="[195,1398,364,1800]" targetPageId="6">
<paragraph id="8B947F28FFCBFFC03BECF84051A937C7" blockId="6.[189,1399,1925,2022]" pageId="6" pageNumber="7">
<emphasis id="B95FA33AFFCBFFC03BECF840510B37BD" bold="true" box="[189,329,1925,1951]" pageId="6" pageNumber="7">Figs 2024.</emphasis>
<taxonomicName id="4C2B04ABFFCBFFC03A01F840523D37BD" authority="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk, 2017" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[336,639,1925,1951]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="B95FA33AFFCBFFC03A01F840523D37BD" box="[336,639,1925,1951]" italics="true" pageId="6" pageNumber="7">Pukupuku arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C1E41FFCBFFC039D6F843529E3782" box="[647,732,1926,1952]" pageId="6" pageNumber="7" rank="species">sp. nov</taxonomicNameLabel>
, holotype, ♂.
<emphasis id="B95FA33AFFCBFFC038D3F84053DD37BD" bold="true" box="[898,927,1925,1951]" pageId="6" pageNumber="7">20</emphasis>
. Aedeagus (dorsal view).
<emphasis id="B95FA33AFFCBFFC03F81F84054AF37BD" bold="true" box="[1232,1261,1925,1951]" pageId="6" pageNumber="7">21</emphasis>
. Parameres (dorsal view).
<emphasis id="B95FA33AFFCBFFC03A36F86D51C637E0" bold="true" box="[359,388,1960,1986]" pageId="6" pageNumber="7">22</emphasis>
. Aedeagus (lateral view).
<emphasis id="B95FA33AFFCBFFC039E6F86D529637E0" bold="true" box="[695,724,1960,1986]" pageId="6" pageNumber="7">23</emphasis>
. Map of India.
<emphasis id="B95FA33AFFCBFFC038DBF86D53E537E0" bold="true" box="[906,935,1960,1986]" pageId="6" pageNumber="7">24</emphasis>
. Map of the state of Arunachal Pradesh showing the type locality.
</paragraph>
</caption>
<caption id="DF542FA0FFCAFFC13BECF89C51BB37FF" httpUri="https://zenodo.org/record/851571/files/figure.png" pageId="7" pageNumber="8" targetBox="[215,1372,265,1841]" targetPageId="7">
<paragraph id="8B947F28FFCAFFC13BECF89C51BB37FF" blockId="7.[189,1399,1881,2013]" pageId="7" pageNumber="8">
<emphasis id="B95FA33AFFCAFFC13BECF89C511B3751" bold="true" box="[189,345,1881,1907]" pageId="7" pageNumber="8">Figs 2533.</emphasis>
<taxonomicName id="4C2B04ABFFCAFFC13A21F89C52ED3751" authority="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk, 2017" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[368,687,1881,1907]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="B95FA33AFFCAFFC13A21F89C52ED3751" box="[368,687,1881,1907]" italics="true" pageId="7" pageNumber="8">Pukupuku arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C1E41FFCAFFC13996F89F53763756" box="[711,820,1882,1908]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀.
<emphasis id="B95FA33AFFCAFFC13F56F89C54663751" bold="true" box="[1031,1060,1881,1907]" pageId="7" pageNumber="8">25</emphasis>
. Clypeus.
<emphasis id="B95FA33AFFCAFFC13F90F89C549C3751" bold="true" box="[1217,1246,1881,1907]" pageId="7" pageNumber="8">26</emphasis>
. Abdomen.
<emphasis id="B95FA33AFFCAFFC13BECF8B9509837B4" bold="true" box="[189,218,1916,1942]" pageId="7" pageNumber="8">27</emphasis>
. Pronotum.
<emphasis id="B95FA33AFFCAFFC13A21F8B951CF37B4" bold="true" box="[368,397,1916,1942]" pageId="7" pageNumber="8">28</emphasis>
. Pygidium.
<emphasis id="B95FA33AFFCAFFC13970F8B9527C37B4" bold="true" box="[545,574,1916,1942]" pageId="7" pageNumber="8">29</emphasis>
. Mentum.
<emphasis id="B95FA33AFFCAFFC13995F8B952A337B4" bold="true" box="[708,737,1916,1942]" pageId="7" pageNumber="8">30</emphasis>
. Apical part of mesotibia.
<emphasis id="B95FA33AFFCAFFC13F75F8B9540337B4" bold="true" box="[1060,1089,1916,1942]" pageId="7" pageNumber="8">31</emphasis>
. Apical part of metatibia.
<emphasis id="B95FA33AFFCAFFC13BECF85A5098379B" bold="true" box="[189,218,1951,1977]" pageId="7" pageNumber="8">32</emphasis>
. 4th mesotarsomere.
<emphasis id="B95FA33AFFCAFFC13A9DF85A51AB379B" bold="true" box="[460,489,1951,1977]" pageId="7" pageNumber="8">33</emphasis>
. 4th metatarsomere. (Arrows indicate the character separating
<taxonomicName id="4C2B04ABFFCAFFC13F90F86555743798" authorityName="Devanshu Gupta &amp; Kailash Chandra &amp; Aleš Bezděk" authorityYear="2017" box="[1217,1334,1952,1978]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B95FA33AFFCAFFC13F90F86555743798" box="[1217,1334,1952,1978]" italics="true" pageId="7" pageNumber="8">Pukupuku</emphasis>
</taxonomicName>
from the closely related genera).
</paragraph>
</caption>
<paragraph id="8B947F28FFC5FFCE3BECFEC95348314E" blockId="8.[189,1399,267,364]" pageId="8" pageNumber="9">
LEGS (
<figureCitation id="131063ADFFC5FFCE3A5BFECE51CD3104" box="[266,399,267,294]" captionStart="Figs 11 19" captionStartId="5.[189,241,1876,1902]" captionTargetBox="[230,1358,265,1836]" captionTargetId="figure@5.[226,1362,265,1836]" captionTargetPageId="5" captionText="Figs 11 19. Pukupuku arunachalensis sp. nov., holotype, ♂. 11. Protibia. 12. Apical part of mesotibia. 13. Apical part of metatibia. 14. 4 th mesotarsomere. 15. 4 th metatarsomere. 16. Pygidium. 17. Prosternum. 18. Abdomen. 19. Meso- and metasternum. (Arrows indicate the character separating Pukupuku from the closely related genera)." httpUri="https://zenodo.org/record/851567/files/figure.png" pageId="8" pageNumber="9">Figs 1115</figureCitation>
). Protibae tridentate, claws not cleft. Meso- and metatibiae unidentate, outer claw with a ventral tooth respectively, 4th tarsomere of both meso- and metatibiae with a single acuminate ventroapical process with a pair of fine long setae.
</paragraph>
<paragraph id="8B947F28FFC5FFCE3BECFE5153A7318F" blockId="8.[189,997,403,430]" box="[189,997,403,430]" pageId="8" pageNumber="9">
PYGIDIUM (
<figureCitation id="131063ADFFC5FFCE3A6BFE5651D2318C" box="[314,400,403,430]" captionStart="Figs 11 19" captionStartId="5.[189,241,1876,1902]" captionTargetBox="[230,1358,265,1836]" captionTargetId="figure@5.[226,1362,265,1836]" captionTargetPageId="5" captionText="Figs 11 19. Pukupuku arunachalensis sp. nov., holotype, ♂. 11. Protibia. 12. Apical part of mesotibia. 13. Apical part of metatibia. 14. 4 th mesotarsomere. 15. 4 th metatarsomere. 16. Pygidium. 17. Prosternum. 18. Abdomen. 19. Meso- and metasternum. (Arrows indicate the character separating Pukupuku from the closely related genera)." httpUri="https://zenodo.org/record/851567/files/figure.png" pageId="8" pageNumber="9">Fig. 16</figureCitation>
). Triangular, thrice as broad as long, without hairs.
</paragraph>
<paragraph id="8B947F28FFC5FFCE3BECFE1055133217" blockId="8.[189,1399,469,566]" pageId="8" pageNumber="9">
VENTER (
<figureCitation id="131063ADFFC5FFCE3A74FE1051E431CD" box="[293,422,469,495]" captionStart="Figs 11 19" captionStartId="5.[189,241,1876,1902]" captionTargetBox="[230,1358,265,1836]" captionTargetId="figure@5.[226,1362,265,1836]" captionTargetPageId="5" captionText="Figs 11 19. Pukupuku arunachalensis sp. nov., holotype, ♂. 11. Protibia. 12. Apical part of mesotibia. 13. Apical part of metatibia. 14. 4 th mesotarsomere. 15. 4 th metatarsomere. 16. Pygidium. 17. Prosternum. 18. Abdomen. 19. Meso- and metasternum. (Arrows indicate the character separating Pukupuku from the closely related genera)." httpUri="https://zenodo.org/record/851567/files/figure.png" pageId="8" pageNumber="9">Figs 1719</figureCitation>
). Ventral surface of thorax and metasternum densely hairy. Prosternum with a small process in middle of posterior margin, posterior face of it flat. Meso- and metasternal processes absent. Abdominal sternites brown black, not abbreviated, smooth without hairs, segments curved in shape.
</paragraph>
<paragraph id="8B947F28FFC5FFCE3BECFD9854213254" blockId="8.[189,1123,604,631]" box="[189,1123,604,631]" pageId="8" pageNumber="9">
AEDEAGUS (
<figureCitation id="131063ADFFC5FFCE3A12FD9951843255" box="[323,454,604,631]" captionStart="Figs 20 24" captionStartId="6.[189,241,1925,1951]" captionTargetBox="[195,1398,364,1800]" captionTargetId="figure@6.[189,1398,349,1800]" captionTargetPageId="6" captionText="Figs 20 24. Pukupuku arunachalensis sp. nov, holotype, ♂. 20. Aedeagus (dorsal view). 21. Parameres (dorsal view). 22. Aedeagus (lateral view). 23. Map of India. 24. Map of the state of Arunachal Pradesh showing the type locality." httpUri="https://zenodo.org/record/851569/files/figure.png" pageId="8" pageNumber="9">Figs 2022</figureCitation>
). Symmetrical, parameres fused at base, rather flat, bare.
</paragraph>
<paragraph id="8B947F28FFC5FFCE3BECFD5B52053367" blockId="8.[189,1399,670,837]" pageId="8" pageNumber="9">
SEXUAL DIMORPHISM (
<figureCitation id="131063ADFFC5FFCE3AFEFD5B5270329B" box="[431,562,670,697]" captionStart="Figs 25 33" captionStartId="7.[189,241,1881,1907]" captionTargetBox="[215,1372,265,1841]" captionTargetId="figure@7.[215,1372,265,1841]" captionTargetPageId="7" captionText="Figs 25 33. Pukupuku arunachalensis sp. nov., paratype, ♀. 25. Clypeus. 26. Abdomen. 27. Pronotum. 28. Pygidium. 29. Mentum. 30. Apical part of mesotibia. 31. Apical part of metatibia. 32. 4 th mesotarsomere. 33. 4 th metatarsomere. (Arrows indicate the character separating Pukupuku from the closely related genera)." httpUri="https://zenodo.org/record/851571/files/figure.png" pageId="8" pageNumber="9">Figs 2533</figureCitation>
). Females differ from males in the following characters: oval in shape, feeble shining; anterior margin of clypeus bilobed, surface rather strongly rugopunctate, mandibles normally developed (
<figureCitation id="131063ADFFC5FFCE3A95FD21526132DD" box="[452,547,740,767]" captionStart="Figs 25 33" captionStartId="7.[189,241,1881,1907]" captionTargetBox="[215,1372,265,1841]" captionTargetId="figure@7.[215,1372,265,1841]" captionTargetPageId="7" captionText="Figs 25 33. Pukupuku arunachalensis sp. nov., paratype, ♀. 25. Clypeus. 26. Abdomen. 27. Pronotum. 28. Pygidium. 29. Mentum. 30. Apical part of mesotibia. 31. Apical part of metatibia. 32. 4 th mesotarsomere. 33. 4 th metatarsomere. (Arrows indicate the character separating Pukupuku from the closely related genera)." httpUri="https://zenodo.org/record/851571/files/figure.png" pageId="8" pageNumber="9">Fig. 25</figureCitation>
); pronotum rather distinctly and finely punctate (
<figureCitation id="131063ADFFC5FFCE3FC1FD2154AD32DC" box="[1168,1263,740,766]" captionStart="Figs 25 33" captionStartId="7.[189,241,1881,1907]" captionTargetBox="[215,1372,265,1841]" captionTargetId="figure@7.[215,1372,265,1841]" captionTargetPageId="7" captionText="Figs 25 33. Pukupuku arunachalensis sp. nov., paratype, ♀. 25. Clypeus. 26. Abdomen. 27. Pronotum. 28. Pygidium. 29. Mentum. 30. Apical part of mesotibia. 31. Apical part of metatibia. 32. 4 th mesotarsomere. 33. 4 th metatarsomere. (Arrows indicate the character separating Pukupuku from the closely related genera)." httpUri="https://zenodo.org/record/851571/files/figure.png" pageId="8" pageNumber="9">Fig. 27</figureCitation>
); abdomen yellowish and shining, densely hairy with segments straight, 5th segment broadest in middle (
<figureCitation id="131063ADFFC5FFCE3E5EFCC2552A3300" box="[1295,1384,775,802]" captionStart="Figs 25 33" captionStartId="7.[189,241,1881,1907]" captionTargetBox="[215,1372,265,1841]" captionTargetId="figure@7.[215,1372,265,1841]" captionTargetPageId="7" captionText="Figs 25 33. Pukupuku arunachalensis sp. nov., paratype, ♀. 25. Clypeus. 26. Abdomen. 27. Pronotum. 28. Pygidium. 29. Mentum. 30. Apical part of mesotibia. 31. Apical part of metatibia. 32. 4 th mesotarsomere. 33. 4 th metatarsomere. (Arrows indicate the character separating Pukupuku from the closely related genera)." httpUri="https://zenodo.org/record/851571/files/figure.png" pageId="8" pageNumber="9">Fig. 26</figureCitation>
); pygidium densely hairy (
<figureCitation id="131063ADFFC5FFCE3AB3FCEE52783367" box="[482,570,811,837]" captionStart="Figs 25 33" captionStartId="7.[189,241,1881,1907]" captionTargetBox="[215,1372,265,1841]" captionTargetId="figure@7.[215,1372,265,1841]" captionTargetPageId="7" captionText="Figs 25 33. Pukupuku arunachalensis sp. nov., paratype, ♀. 25. Clypeus. 26. Abdomen. 27. Pronotum. 28. Pygidium. 29. Mentum. 30. Apical part of mesotibia. 31. Apical part of metatibia. 32. 4 th mesotarsomere. 33. 4 th metatarsomere. (Arrows indicate the character separating Pukupuku from the closely related genera)." httpUri="https://zenodo.org/record/851571/files/figure.png" pageId="8" pageNumber="9">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="8B947F28FFC5FFCE3BECFCA953DA33A4" blockId="8.[189,920,876,903]" box="[189,920,876,903]" pageId="8" pageNumber="9">COLOUR. Yellow with brown marking on lateral sides of elytra.</paragraph>
</subSubSection>
<caption id="DF542FA0FFC5FFCE3BECF843527E37C5" httpUri="https://zenodo.org/record/851573/files/figure.png" pageId="8" pageNumber="9" targetBox="[343,1245,926,1883]" targetPageId="8">
<paragraph id="8B947F28FFC5FFCE3BECF843527E37C5" blockId="8.[189,1398,1926,2023]" pageId="8" pageNumber="9">
<emphasis id="B95FA33AFFC5FFCE3BECF843510A3782" bold="true" box="[189,328,1926,1952]" pageId="8" pageNumber="9">Figs 3435.</emphasis>
<taxonomicName id="4C2B04ABFFC5FFCE3A01F84352853782" authority="(Arrow, 1919)" authorityName="Curved" authorityYear="1919" baseAuthorityName="Arrow" baseAuthorityYear="1919" box="[336,711,1926,1952]" class="Insecta" family="Rutelidae" genus="Pukupuku" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B95FA33AFFC5FFCE3A01F84352573782" box="[336,533,1926,1952]" italics="true" pageId="8" pageNumber="9">Pukupuku curtus</emphasis>
(Arrow, 1919)
</taxonomicName>
, syntype,
<emphasis id="B95FA33AFFC5FFCE386AF84353133782" box="[827,849,1926,1952]" italics="true" pageId="8" pageNumber="9">3</emphasis>
.
<emphasis id="B95FA33AFFC5FFCE3831F843533F3782" bold="true" box="[864,893,1926,1952]" pageId="8" pageNumber="9">34</emphasis>
. Habitus (dorsal view).
<emphasis id="B95FA33AFFC5FFCE3FC6F84354F63782" bold="true" box="[1175,1204,1926,1952]" pageId="8" pageNumber="9">35</emphasis>
. Attached labels [Source: Muséum national dHistoire naturelle, Paris (France). Collection: Insects-Coleoptera (EC) Specimen MNHN-EC-EC1464].
</paragraph>
</caption>
<subSubSection id="C3312CA3FFC4FFCF3BECFECE54A53176" box="[189,1255,267,340]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8B947F28FFC4FFCF3BECFECE51143107" blockId="9.[189,342,267,293]" box="[189,342,267,293]" pageId="9" pageNumber="10">
<heading id="D0DCC844FFC4FFCF3BECFECE51143107" bold="true" box="[189,342,267,293]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B95FA33AFFC4FFCF3BECFECE51143107" bold="true" box="[189,342,267,293]" pageId="9" pageNumber="10">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B947F28FFC4FFCF3BECFEFF54A53176" blockId="9.[189,1255,313,340]" box="[189,1255,313,340]" pageId="9" pageNumber="10">
So far known only from Vijay Nagar in the state of Arunachal Pradesh, India (
<figureCitation id="131063ADFFC4FFCF3F00FEFC54983176" box="[1105,1242,313,340]" captionStart="Figs 20 24" captionStartId="6.[189,241,1925,1951]" captionTargetBox="[195,1398,364,1800]" captionTargetId="figure@6.[189,1398,349,1800]" captionTargetPageId="6" captionText="Figs 20 24. Pukupuku arunachalensis sp. nov, holotype, ♂. 20. Aedeagus (dorsal view). 21. Parameres (dorsal view). 22. Aedeagus (lateral view). 23. Map of India. 24. Map of the state of Arunachal Pradesh showing the type locality." httpUri="https://zenodo.org/record/851569/files/figure.png" pageId="9" pageNumber="10">Figs 2324</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3312CA3FFC4FFCF3BECFEBE523131E1" pageId="9" pageNumber="10" type="description">
<paragraph id="8B947F28FFC4FFCF3BECFEBE51B631B7" blockId="9.[189,627,379,452]" box="[189,500,379,405]" pageId="9" pageNumber="10">
<heading id="D0DCC844FFC4FFCF3BECFEBE51B631B7" bold="true" box="[189,500,379,405]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="6">
<emphasis id="B95FA33AFFC4FFCF3BECFEBE51B631B7" bold="true" box="[189,500,379,405]" pageId="9" pageNumber="10">Collecting circumstances</emphasis>
</heading>
</paragraph>
<paragraph id="8B947F28FFC4FFCF3BECFE6C523131E1" blockId="9.[189,627,379,452]" box="[189,627,425,452]" pageId="9" pageNumber="10">Collected at night by light after dusk.</paragraph>
</subSubSection>
</treatment>
</document>