560 lines
85 KiB
XML
560 lines
85 KiB
XML
<document id="D5C90EC35D6B52D2B3D18DC56C5D172C" ID-DOI="10.11646/zootaxa.5476.1.31" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681863" ID-ZooBank="B3C36A64-00AF-4D48-B2B3-74AFE9793CB0" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720375684251" checkinUser="plazi" docAuthor="Takeda, Masatsune & Komatsu, Hironori" docDate="2024" docId="0382346DFF81FFEBA8C9FDAE1F2F8D0F" docLanguage="en" docName="zootaxa.5476.1.31.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.31" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pilumnus swajayai Ng & Rahayu 2021" docType="treatment" docVersion="4" lastPageNumber="432" masterDocId="FFBB4C15FF85FFE3A85EFFC41D438F6E" masterDocTitle="Actumnus ngankeeae sp. nov. and Pilumnus swajayai Ng & Rahayu, 2021, pilumnid crabs from Japan (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="433" masterPageNumber="424" pageNumber="428" updateTime="1721079689823" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="180AB2F86924F1522AF490FFA69A6E89">
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<mods:title id="E0C4E7C686E04AB3AD44661825953648">Actumnus ngankeeae sp. nov. and Pilumnus swajayai Ng & Rahayu, 2021, pilumnid crabs from Japan (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:name id="E814C4A76209572E44D6B03423391B20" type="personal">
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<mods:role id="C8D75604117D0D3802C034AC088F06F7">
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<mods:roleTerm id="E3A50980713835AD8F7D7CA9D1834308">Author</mods:roleTerm>
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<mods:namePart id="EC2DEBE71CADDE6A64459C3DCDCB4332">Takeda, Masatsune</mods:namePart>
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<mods:nameIdentifier id="E338F5BE517EB20C4E8ED82A27DF27A2" type="ORCID">0000-0002-0028-1397</mods:nameIdentifier>
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<mods:affiliation id="D916C402F1049D1553B787ABFFB1759E">Department of Zoology, National Museum of Nature and Science, Tokyo, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<mods:nameIdentifier id="CA6D73F0579343CB24EF3CA92B1C178D" type="email">takeda@kahaku.go.jp</mods:nameIdentifier>
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<mods:namePart id="6DD4061E2E5AFCBDC2318F496CBB9EBD">Komatsu, Hironori</mods:namePart>
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<mods:nameIdentifier id="70CA59C8BC434FDCE9D0A067B63C263A" type="ORCID">0000-0002-7573-4127</mods:nameIdentifier>
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<mods:affiliation id="EB2EE799BED51F64F77923267D0BE4EF">Department of Zoology, National Museum of Nature and Science, Tokyo, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<mods:nameIdentifier id="AD5D48ABE402D5890C916866BE6EF2D8" type="email">h-komatu@kahaku.go.jp</mods:nameIdentifier>
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<mods:title id="A138D21539CA7794AC4DF4499B561910">Zootaxa</mods:title>
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<mods:date id="7EAAF58C979BA3EA982AFB18B83EAF1C">2024</mods:date>
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<mods:number id="6FA2A6F96388A3B2F0B86FD9761BFA31">2024-07-04</mods:number>
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<mods:number id="218093945FF42EA3099E58C4C488ECCB">5476</mods:number>
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<mods:url id="A41C75B77396C0165407B639E2501F1B">http://dx.doi.org/10.11646/zootaxa.5476.1.31</mods:url>
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<treatment id="0382346DFF81FFEBA8C9FDAE1F2F8D0F" ID-DOI="http://doi.org/10.5281/zenodo.12746902" ID-Zenodo-Dep="12746902" LSID="urn:lsid:plazi:treatment:0382346DFF81FFEBA8C9FDAE1F2F8D0F" httpUri="http://treatment.plazi.org/id/0382346DFF81FFEBA8C9FDAE1F2F8D0F" lastPageId="8" lastPageNumber="432" pageId="4" pageNumber="428">
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<subSubSection id="C331D6F0FF81FFE7A8C9FDAE1F398DEA" box="[151,634,618,645]" pageId="4" pageNumber="428" type="nomenclature">
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<paragraph id="8B94857BFF81FFE7A8C9FDAE1F398DEA" blockId="4.[151,634,618,681]" box="[151,634,618,645]" pageId="4" pageNumber="428">
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<heading id="D0DC3217FF81FFE7A8C9FDAE1F398DEA" bold="true" box="[151,634,618,645]" fontSize="11" level="1" pageId="4" pageNumber="428" reason="1">
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<taxonomicName id="4C2BFEF8FF81FFE7A8C9FDAE1F398DEA" ID-CoL="8QCCC" authority="Ng & Rahayu, 2021" authorityName="Ng & Rahayu" authorityYear="2021" box="[151,634,618,645]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="428" phylum="Arthropoda" rank="species" species="swajayai">
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<emphasis id="B95F5969FF81FFE7A8C9FDAE1F398DEA" bold="true" box="[151,634,618,645]" pageId="4" pageNumber="428">
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<emphasis id="B95F5969FF81FFE7A8C9FDAE1C3F8DEB" bold="true" box="[151,380,618,645]" italics="true" pageId="4" pageNumber="428">Pilumnus swajayai</emphasis>
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<bibRefCitation id="EFBAF88AFF81FFE7A9DAFDAF1F398DEA" author="Ng, P. K. L. & Rahayu, D. L." box="[388,634,618,645]" pageId="4" pageNumber="428" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu, 2021</bibRefCitation>
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||
</emphasis>
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||
</taxonomicName>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C331D6F0FF81FFE7A8C9FD4B1C658DC7" box="[151,294,654,681]" pageId="4" pageNumber="428" type="description">
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<paragraph id="8B94857BFF81FFE7A8C9FD4B1C658DC7" blockId="4.[151,634,618,681]" box="[151,294,654,681]" pageId="4" pageNumber="428">
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(
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<figureCitation id="131099FEFF81FFE7A8C1FD4B1C408DC7" box="[159,259,655,681]" captionStart="FIGURE 3" captionStartId="5.[151,250,1035,1059]" captionTargetBox="[160,1427,190,1001]" captionTargetId="figure-20@5.[151,1436,181,1011]" captionTargetPageId="5" captionText="FIGURE 3. A–C, Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D, Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; C–D, left G1, inner view." figureDoi="http://doi.org/10.5281/zenodo.12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="4" pageNumber="428">Figs. 3D</figureCitation>
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,
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<figureCitation id="131099FEFF81FFE7A951FD4A1C5E8DC7" box="[271,285,654,681]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">5</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C331D6F0FF81FFE7A8C9FD121E8B8CEF" pageId="4" pageNumber="428" type="materials_examined">
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||
<paragraph id="8B94857BFF81FFE7A8C9FD121E8B8CEF" blockId="4.[151,1437,726,2013]" pageId="4" pageNumber="428">
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||
<emphasis id="B95F5969FF81FFE7A8C9FD121C3B8D9E" bold="true" box="[151,376,726,752]" pageId="4" pageNumber="428">Material examined</emphasis>
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||
.
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<materialsCitation id="3B438F26FF81FFE7A9DBFD131CDF8C33" collectingDate="2003-06-20" collectorName="Daiyu Maru No." country="Iceland" county="northern Ryukyu Islands" elevation="155" latitude="28.5015" location="Kado" longLatPrecision="12" longitude="129.56216" municipality="Amami-Oshima" pageId="4" pageNumber="428" specimenCount="2">
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||
<specimenCount id="9D2D4EF2FF81FFE7A9DBFD131CFC8D9F" box="[389,447,727,753]" pageId="4" pageNumber="428" type="male">Male</specimenCount>
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||
(cb 5.8 mm, cl 5.0 mm; NSMT-Cr 31015),
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<specimenCount id="9D2D4EF2FF81FFE7ABC0FD13191D8D9F" box="[926,1118,727,753]" pageId="4" pageNumber="428">ovigerous female</specimenCount>
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(cb 5.5 mm, cl 4.6 mm, with some remaining eggs after releasing larvae; NSMT-Cr 31016), off
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||
<location id="8EF4D3A0FF81FFE7ABD8FD3F1E868C7B" LSID="urn:lsid:plazi:treatment:0382346DFF81FFEBA8C9FDAE1F2F8D0F:8EF4D3A0FF81FFE7ABD8FD3F1E868C7B" box="[902,965,763,789]" country="Iceland" county="northern Ryukyu Islands" latitude="28.5015" longLatPrecision="12" longitude="129.56216" municipality="Amami-Oshima" name="Kado" pageId="4" pageNumber="428">Kado</location>
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||
,
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||
<collectingMunicipality id="6BF01F01FF81FFE7AB8EFD3F19C78C7B" box="[976,1156,763,789]" pageId="4" pageNumber="428">Amami-Oshima</collectingMunicipality>
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||
Island, northern Ryukyu Islands, tugboat
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<emphasis id="B95F5969FF81FFE7A90EFCDB1F6F8C57" box="[336,556,799,825]" italics="true" pageId="4" pageNumber="428">Daiyu Maru No. 38</emphasis>
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||
, sta. DY-03-10,
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<geoCoordinate id="EE1FE3BCFF81FFE7AAB8FCDB1E258C57" box="[742,870,799,825]" degrees="28" direction="north" minutes="29.90" orientation="latitude" pageId="4" pageNumber="428" precision="9" value="28.498333">28°29.90'N</geoCoordinate>
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||
<geoCoordinate id="EE1FE3BCFF81FFE7AB33FCDB1EBB8C57" box="[877,1016,799,825]" degrees="129" direction="east" minutes="33.44" orientation="longitude" pageId="4" pageNumber="428" precision="9" value="129.55733">129°33.44'E</geoCoordinate>
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to 28°30.09'N 129°33.73'E, 155⎼
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||
<elevation id="00066248FF81FFE7AD2CFCDA1DB58C33" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="428" unit="m" value="155.0">
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||
<quantity id="4CD3289EFF81FFE7AD2CFCDA1DEE8C33" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="428" unit="m" value="155.0">155 m</quantity>
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depth
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</elevation>
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,
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<date id="FF95A3BBFF81FFE7A95FFC871CDF8C33" box="[257,412,835,861]" pageId="4" pageNumber="428" value="2003-06-20">
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<collectingDate id="EFD15A53FF81FFE7A95FFC871CDF8C33" box="[257,412,835,861]" pageId="4" pageNumber="428" value="2003-06-20">20 June 2003</collectingDate>
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||
</date>
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</materialsCitation>
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||
.
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<materialsCitation id="3B438F26FF81FFE7A9F8FC871E808CEF" collectingDate="1979-08-05" collectionCode="TV" collectorName="Tokai Daigaku Maru II" country="Iceland" county="central Ryukyu Islands" elevation="80" location="Okinawa-jima Island" municipality="Okinawa-jima Island" pageId="4" pageNumber="428" specimenCount="1">
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<specimenCount id="9D2D4EF2FF81FFE7A9F8FC871CA38C33" box="[422,480,835,861]" pageId="4" pageNumber="428" type="male">Male</specimenCount>
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||
(cb
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||
<quantity id="4CD3289EFF81FFE7AA4DFC871F2F8C33" box="[531,620,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="4" pageNumber="428" unit="mm" value="8.4">8.4 mm</quantity>
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||
, cl
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||
<quantity id="4CD3289EFF81FFE7AACDFC871FAD8C33" box="[659,750,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="4" pageNumber="428" unit="mm" value="6.3">6.3 mm</quantity>
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), NSMT-Cr 31017, off
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<collectingMunicipality id="6BF01F01FF81FFE7ABA8FC8719A68C33" box="[1014,1253,835,861]" pageId="4" pageNumber="428">Okinawa-jima Island</collectingMunicipality>
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||
, central Ryukyu Islands, 60⎼
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||
<elevation id="00066248FF81FFE7A943FCA31CDD8CEF" box="[285,414,871,897]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="428" unit="m" value="80.0">
|
||
<quantity id="4CD3289EFF81FFE7A943FCA31C158CEF" box="[285,342,871,897]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="428" unit="m" value="80.0">80 m</quantity>
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||
depth
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||
</elevation>
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||
, TV
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||
<emphasis id="B95F5969FF81FFE7A98BFCA31F9B8CEE" box="[469,728,871,897]" italics="true" pageId="4" pageNumber="428">
|
||
<collectorName id="26DEE0ADFF81FFE7A98BFCA31F9B8CEE" box="[469,728,871,897]" pageId="4" pageNumber="428">Tokai Daigaku Maru II</collectorName>
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||
</emphasis>
|
||
, 02⎼
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||
<date id="FF95A3BBFF81FFE7AB51FCA21E808CEF" box="[783,963,870,897]" pageId="4" pageNumber="428" value="1979-08-05">
|
||
<collectingDate id="EFD15A53FF81FFE7AB51FCA21E808CEF" box="[783,963,870,897]" pageId="4" pageNumber="428" value="1979-08-05">05 August 1979</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C331D6F0FF81FFE6A899FC4E1F2689BE" lastPageId="5" lastPageNumber="429" pageId="4" pageNumber="428" type="description">
|
||
<paragraph id="8B94857BFF81FFE7A899FC4E1E9B8943" blockId="4.[151,1437,726,2013]" pageId="4" pageNumber="428">
|
||
<emphasis id="B95F5969FF81FFE7A899FC4E1FD48CCA" bold="true" box="[199,663,906,933]" pageId="4" pageNumber="428">Description of male (NSMT-Cr 31015)</emphasis>
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||
. Carapace dorsal surface (
|
||
<figureCitation id="131099FEFF81FFE7AB96FC4F19068CCB" box="[968,1093,906,933]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Fig. 5A, D</figureCitation>
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) wholly covered with velvety tomentum, moderately convex in both directions, not granulated, but uneven as a whole, weakly divided into regions; median longitudinal, bifurcating furrow distinct, dividing frontal, epigastric and protogastric regions on both sides; transverse furrow from post-orbital furrow to anterior end of mesogastric region; transverse furrow from last anterolateral tooth to median part of protogastric region; branchial posterior region divided into two by transverse furrow. Hepatic region produced as dentiform lobe (
|
||
<figureCitation id="131099FEFF81FFE7AB3EFBFB1EF88B37" box="[864,955,1086,1113]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Fig. 5D</figureCitation>
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). Cardiac region not distinctly demarcated from branchial regions. Anterolateral teeth (
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<figureCitation id="131099FEFF81FFE7AACDFBA71E5F8B13" box="[659,796,1122,1149]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Figs. 5C, D</figureCitation>
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) strong, with small horny tip; first tooth placed a little lower plane than line from external orbital tooth to second tooth, being distinctly tuberculated with thick basal part, directed anterolaterally; second tooth prominent, elongated, weakly curved dorsally; third tooth directed dorsally. Posterolateral margin (
|
||
<figureCitation id="131099FEFF81FFE7A9C4FB0B1F078B87" box="[410,580,1230,1257]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Figs. 5A, C, D</figureCitation>
|
||
) not strongly convergent toward carapace posterior margin, weakly bulged behind last anterolateral tooth. Subhepatic region (
|
||
<figureCitation id="131099FEFF81FFE7AABBFB371E028A63" box="[741,833,1266,1293]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Fig. 5D</figureCitation>
|
||
) armed with conical tubercle, smaller than external orbital tooth, subequal to hepatic dentiform lobe. Front (
|
||
<figureCitation id="131099FEFF81FFE7AB7DFAD31EDD8A5F" box="[803,926,1302,1329]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Fig. 5B–D</figureCitation>
|
||
) strongly deflexed, margin divided into two with median wide, large V-shaped notch; each lobe weakly convex forward for inner two-thirds, deeply concave for outer one-third. Supraorbital margin (
|
||
<figureCitation id="131099FEFF81FFE7AA20FA9B1FB88A17" box="[638,763,1374,1401]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Fig. 5B–D</figureCitation>
|
||
) moderately raised, without granules; inner part directed posteriorly, main part transverse toward external orbital tooth, with two deep interruptions; small triangular lobe at inside of inner interruption; triangular tooth tipped with spinule between two interruptions; external orbital tooth (
|
||
<figureCitation id="131099FEFF81FFE7A8C1FA0F1DB48A8B" box="[159,247,1482,1509]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Fig. 5D</figureCitation>
|
||
) well developed, nearly triangular, directed forward, with obtusely angulated tip and weakly convex outer margin. Infraorbital margin sharply incised below external orbital angle, deeply concave at median part; inner end of infraorbital margin strongly developed as triangular lobe with sharp tip.
|
||
</paragraph>
|
||
<paragraph id="8B94857BFF81FFE7A899F9F31F5289F7" blockId="4.[151,1437,726,2013]" pageId="4" pageNumber="428">Third maxilliped wide, sparsely hairy. Ischium becomes slightly narrower anteriorly, with oblique shallow furrow from anterior inner part to basal outer part. Merus widens anteriorly, leaving narrow space between both sides at articulation with ischium.</paragraph>
|
||
<paragraph id="8B94857BFF81FFE7A899F9671C8A88B3" blockId="4.[151,1437,726,2013]" pageId="4" pageNumber="428">
|
||
Chelipeds (
|
||
<figureCitation id="131099FEFF81FFE7A919F9671CD689D3" box="[327,405,1698,1725]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Figs. 5</figureCitation>
|
||
) stout, right being larger, with general shape being basically same in both chelipeds. Upper margin of ischium armed with three spines. Upper margin of merus with some spines of irregular size. Entire surface of carpus uneven, with several dispersed granules of good size; submarginal deep furrow on outer surface along distal margin; inner angle of carpus with recurved tubercle. Larger palm (
|
||
<figureCitation id="131099FEFF81FFE7AB94F8CB195D8847" box="[970,1054,1806,1833]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Fig. 5E</figureCitation>
|
||
) with outer upper surface covered with tomentum from distal end of upper margin to basal part of lower margin, bare for lower surface; sharp tips of several tuberculate granules through dense tomentum, some granules on bare surface. Smaller palm (
|
||
<figureCitation id="131099FEFF81FFE7AD5EF8931815881F" box="[1280,1366,1878,1905]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="4" pageNumber="428">Fig. 5E</figureCitation>
|
||
), with outer surface nearly covered with tomentum. Both chelae with cutting edges toothed, fingers with teeth closely occluding; larger chela with fingers thickened; smaller chela with outer surfaces of both fingers each narrowly grooved throughout length.
|
||
</paragraph>
|
||
<caption id="DF54D5F3FF80FFE6A8C9FBCF1EDB8B05" ID-DOI="http://doi.org/10.5281/zenodo.12681871" ID-Zenodo-Dep="12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="5" pageNumber="429" startId="5.[151,250,1035,1059]" targetBox="[160,1427,190,1001]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B94857BFF80FFE6A8C9FBCF1EDB8B05" blockId="5.[151,1437,1035,1131]" pageId="5" pageNumber="429">
|
||
<emphasis id="B95F5969FF80FFE6A8C9FBCF1C518B4D" bold="true" box="[151,274,1035,1059]" pageId="5" pageNumber="429">FIGURE 3.</emphasis>
|
||
A–C,
|
||
<taxonomicName id="4C2BFEF8FF80FFE6A90AFBCF1F698B4D" authorityName="Takeda & Komatsu" authorityYear="2024" box="[340,554,1035,1059]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
|
||
<emphasis id="B95F5969FF80FFE6A90AFBCF1F698B4D" box="[340,554,1035,1059]" italics="true" pageId="5" pageNumber="429">Actumnus ngankeeae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F5969FF80FFE6AA6EFBCF1FC38B4D" bold="true" box="[560,640,1035,1059]" pageId="5" pageNumber="429">
|
||
<taxonomicNameLabel id="A26CE412FF80FFE6AA6EFBCF1FC38B4D" box="[560,640,1035,1059]" pageId="5" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D,
|
||
<taxonomicName id="4C2BFEF8FF80FFE6A8EBFBEB1F078B29" authority="Ng & Rahayu, 2021" authorityName="Ng & Rahayu" authorityYear="2021" box="[181,580,1071,1095]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="swajayai">
|
||
<emphasis id="B95F5969FF80FFE6A8EBFBEB1C378B29" box="[181,372,1071,1095]" italics="true" pageId="5" pageNumber="429">Pilumnus swajayai</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF80FFE6A927FBEB1F078B29" author="Ng, P. K. L. & Rahayu, D. L." box="[377,580,1071,1095]" pageId="5" pageNumber="429" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; C–D, left G1, inner view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B94857BFF80FFE6A899FB5D1EF28B95" blockId="5.[151,1437,1177,2032]" pageId="5" pageNumber="429">
|
||
Ambulatory legs (
|
||
<figureCitation id="131099FEFF80FFE6A9C5FB5D1CB58BDA" box="[411,502,1177,1204]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="5" pageNumber="429">Fig. 5A</figureCitation>
|
||
) stout, last pair little shorter than first three pairs, covered tightly with velvety tomentum on surfaces of all articles, margins of distal three articles with long silky or club-shaped setae; meri unarmed, carpi and propodi each with some spinules on dorsal margin.
|
||
</paragraph>
|
||
<paragraph id="8B94857BFF80FFE6A899FAC119378A2D" blockId="5.[151,1437,1177,2032]" pageId="5" pageNumber="429">
|
||
Male pleon (
|
||
<figureCitation id="131099FEFF80FFE6A909FAC11CEF8A4E" box="[343,428,1285,1312]" captionStart="FIGURE 5" captionStartId="7.[151,250,1554,1578]" captionTargetBox="[215,1371,181,1530]" captionTargetId="figure-20@7.[215,1371,181,1530]" captionTargetPageId="7" captionText="FIGURE 5. Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="5" pageNumber="429">Fig. 5E</figureCitation>
|
||
) with free somites covered with short setae. G1 (
|
||
<figureCitation id="131099FEFF80FFE6AB86FAC119738A71" box="[984,1072,1285,1311]" captionStart="FIGURE 3" captionStartId="5.[151,250,1035,1059]" captionTargetBox="[160,1427,190,1001]" captionTargetId="figure-20@5.[151,1436,181,1011]" captionTargetPageId="5" captionText="FIGURE 3. A–C, Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D, Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; C–D, left G1, inner view." figureDoi="http://doi.org/10.5281/zenodo.12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="5" pageNumber="429">Fig. 3D</figureCitation>
|
||
) strongly curved dorsally for its median one-third, ventrally for its distal one-third, with long, subtruncated tip. G2 short.
|
||
</paragraph>
|
||
<paragraph id="8B94857BFF80FFE6A899FA89180E892E" blockId="5.[151,1437,1177,2032]" pageId="5" pageNumber="429">
|
||
<emphasis id="B95F5969FF80FFE6A899FA8919038A09" bold="true" box="[199,1088,1357,1383]" pageId="5" pageNumber="429">Notes on the male (NSMT-Cr 31017) and the female (NSMT-Cr 31016)</emphasis>
|
||
. In the male, the carapace is somewhat decalcified and depressed, and the right cheliped and all the right ambulatory legs are missing. The velvety tomentum of the carapace was partly removed along the anterolateral margins. The carapace anterolateral teeth are sharper than those of the other male (NSMT-Cr 31015), and the ambulatory carpi and propodi are armed each with one or two spinules on dorsal margin. The remaining left cheliped is the larger, with the smooth palm from the basal part of the lower margin to the distal part of the upper margin; both fingers are stout, with thick teeth. The G1 is well developed and strongly curved, being similar to the G1 (
|
||
<figureCitation id="131099FEFF80FFE6ABD7F9E11EA38951" box="[905,992,1573,1599]" captionStart="FIGURE 3" captionStartId="5.[151,250,1035,1059]" captionTargetBox="[160,1427,190,1001]" captionTargetId="figure-20@5.[151,1436,181,1011]" captionTargetPageId="5" captionText="FIGURE 3. A–C, Actumnus ngankeeae sp. nov., holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. D, Pilumnus swajayai Ng & Rahayu, 2021, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015.A, right chela, outer view; B, right fourth ambulatory leg, dorsal view; C–D, left G1, inner view." figureDoi="http://doi.org/10.5281/zenodo.12681871" httpUri="https://zenodo.org/record/12681871/files/figure.png" pageId="5" pageNumber="429">Fig. 3D</figureCitation>
|
||
) of the male (NSMT-Cr 31015).
|
||
</paragraph>
|
||
<paragraph id="8B94857BFF80FFE6A899F98E1F2689BE" blockId="5.[151,1437,1177,2032]" pageId="5" pageNumber="429">In the female, the ambulatory legs other than the last leg of left side are detached, and the left cheliped is missing. The carapace dorsal surface, chelipeds and ambulatory legs are similarly covered with velvety tomentum as in the male specimens. The arrangement of the spinules of the ambulatory carpi and propodi is also close to that of the male specimen (NSMT-Cr 31015).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331D6F0FF80FFEBA899F91D194D8E9B" lastPageId="8" lastPageNumber="432" pageId="5" pageNumber="429" type="discussion">
|
||
<paragraph id="8B94857BFF80FFE5A899F91D1CB48FBB" blockId="5.[151,1437,1177,2032]" lastBlockId="6.[151,1437,151,861]" lastPageId="6" lastPageNumber="430" pageId="5" pageNumber="429">
|
||
<emphasis id="B95F5969FF80FFE6A899F91D1C77899D" bold="true" box="[199,308,1753,1779]" pageId="5" pageNumber="429">Remarks</emphasis>
|
||
. Based on the velvety tomentum covering the carapace, chelipeds and ambulatory legs, the taxonomic position of this species was first considered to be in the genus
|
||
<taxonomicName id="4C2BFEF8FF80FFE6AB22F93918DB8876" authority="Turkay & Schuhmacher, 1985" authorityName="Turkay & Schuhmacher" authorityYear="1985" box="[892,1432,1789,1816]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF80FFE6AB22F939196D8879" box="[892,1070,1789,1815]" italics="true" pageId="5" pageNumber="429">Serenepilumnus</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF80FFE6AC67F93918DB8876" author="Turkay, M. & Schuhmacher, H." box="[1081,1432,1789,1816]" pageId="5" pageNumber="429" refId="ref6836" refString="Turkay, M. & Schuhmacher, H. (1985) Latopilumnus tubicolus n. gen. n. sp., eine neue korallenassoziierte Krebbe, die Bildung einer Wohnhohle induziert (Crustacea: Decapoda: Pilumnidae). Senckenbergiana maritima, 17, 55 - 63." type="journal volume" year="1985">Türkay & Schuhmacher, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
, which is a replacement name for
|
||
<taxonomicName id="4C2BFEF8FF80FFE6AA40F8E51E7F8855" authority="Serene, 1971" authorityName="Serene" authorityYear="1971" box="[542,828,1825,1851]" class="Malacostraca" family="Pilumnidae" genus="Leopoldius" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF80FFE6AA40F8E51FD98855" box="[542,666,1825,1851]" italics="true" pageId="5" pageNumber="429">Leopoldius</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF80FFE6AAFAF8E51E7F8855" author="Serene, R." box="[676,828,1825,1851]" pageId="5" pageNumber="429" refId="ref6494" refString="Serene, R. (1971) Observations preliminaires sur des brachyoures nouveaux ou mal connus du sudest asiatique (Crustacea Decapoda). Bulletin du Museum national d'Histoire naturelle, Paris, Series 2, 42, 903 ⎼ 918, pls. 1 ⎼ 6." type="journal volume" year="1971">Serène, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
. The presence of the hepatic dentiform lobe and the toothed anterolateral margin of the carapace were also indicative of
|
||
<taxonomicName id="4C2BFEF8FF80FFE6ABD3F881197C8831" authorityName="Turkay & Schuhmacher" authorityYear="1985" box="[909,1087,1861,1887]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF80FFE6ABD3F881197C8831" box="[909,1087,1861,1887]" italics="true" pageId="5" pageNumber="429">Serenepilumnus</emphasis>
|
||
</taxonomicName>
|
||
. According to
|
||
<bibRefCitation id="EFBAF88AFF80FFE6ACBAF88218DB8831" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1252,1432,1861,1888]" pageId="5" pageNumber="429" refId="ref6104" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17, 1 ⎼ 286." type="journal volume" year="2008">
|
||
Ng
|
||
<emphasis id="B95F5969FF80FFE6AD52F882187E8831" box="[1292,1341,1861,1887]" italics="true" pageId="5" pageNumber="429">et al</emphasis>
|
||
. (2008)
|
||
</bibRefCitation>
|
||
, the genus
|
||
<taxonomicName id="4C2BFEF8FF80FFE6A951F8AD1C8288ED" authorityName="Turkay & Schuhmacher" authorityYear="1985" box="[271,449,1897,1923]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF80FFE6A951F8AD1C8288ED" box="[271,449,1897,1923]" italics="true" pageId="5" pageNumber="429">Serenepilumnus</emphasis>
|
||
</taxonomicName>
|
||
is represented by four species,
|
||
<taxonomicName id="4C2BFEF8FF80FFE6AB6AF8AE19DB88ED" authority="(De Man, 1902)" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[820,1176,1897,1924]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="kuekenthali">
|
||
<emphasis id="B95F5969FF80FFE6AB6AF8AE1E9788ED" box="[820,980,1897,1924]" italics="true" pageId="5" pageNumber="429">S. kuekenthali</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFBAF88AFF80FFE6AC4CF8AE19CC88ED" author="Man, J. G. De" box="[1042,1167,1897,1924]" pageId="5" pageNumber="429" refId="ref5953" refString="Man, J. G. De (1902) Die von Herrn Professor Kukenthal im Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: Kukenthal, W. (Ed.), Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 25, pp. 467 ⎼ 929. https: // doi. org / 10.5962 / bhl. title. 10409" type="journal volume" year="1902">Man, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
[originally
|
||
<taxonomicName id="4C2BFEF8FF80FFE6AD7CF8AD18CF88ED" authorityName="Leach" authorityYear="1815" box="[1314,1420,1897,1923]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF80FFE6AD7CF8AD18CF88ED" box="[1314,1420,1897,1923]" italics="true" pageId="5" pageNumber="429">Pilumnus</emphasis>
|
||
</taxonomicName>
|
||
],
|
||
<taxonomicName id="4C2BFEF8FF80FFE6A8C9F84A1C8C88C9" authority="(Gordon, 1934)" baseAuthorityName="Gordon" baseAuthorityYear="1934" box="[151,463,1933,1960]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="leopoldi">
|
||
<emphasis id="B95F5969FF80FFE6A8C9F84A1C5088C9" box="[151,275,1933,1960]" italics="true" pageId="5" pageNumber="429">S. leopoldi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBAF88AFF80FFE6A97BF8491C8588C9" author="Gordon, I." box="[293,454,1933,1959]" pageId="5" pageNumber="429" refId="ref5535" refString="Gordon, I. (1934) Crustacea Brachyura. Resultats Scientifiques du Voyage aux Indes Orientales Neerlandaises de LL. AA. RR. le Prince et la Princesse Leopold de Belgique. Memoires du Musee Royal d'Histoire Naturelle de Belgique, Hors Serie, 3 (15), 1 - 78." type="journal volume" year="1934">Gordon, 1934</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
[originally
|
||
<taxonomicName id="4C2BFEF8FF80FFE6AA04F8491FBA88C9" authorityName="Kossmann" authorityYear="1877" box="[602,761,1933,1959]" class="Malacostraca" family="Acidopsidae" genus="Parapilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF80FFE6AA04F8491FBA88C9" box="[602,761,1933,1959]" italics="true" pageId="5" pageNumber="429">Parapilumnus</emphasis>
|
||
</taxonomicName>
|
||
],
|
||
<taxonomicName id="4C2BFEF8FF80FFE6AB4DF84A190A88C9" authority="(MacLeay, 1838)" baseAuthorityName="MacLeay" baseAuthorityYear="1838" box="[787,1097,1933,1960]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="pisifer">
|
||
<emphasis id="B95F5969FF80FFE6AB4DF84A1E3888C9" box="[787,891,1933,1960]" italics="true" pageId="5" pageNumber="429">S. pisifer</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBAF88AFF80FFE6ABD2F849190388C9" author="MacLeay, W. S." box="[908,1088,1933,1959]" pageId="5" pageNumber="429" pagination="53 - 71" refId="ref5696" refString="MacLeay, W. S. (1838) Illustrations of the Annulosa of South Africa; being a Portion of the Objects of Natural History chiefly collected during an Expedition into the Interior of South Africa, under the Direction of Dr. Andrew Smith, in the Years 1834, 1835, and 1836; fitted out by " The Cape of Good Hope Association for Exploring Central Africa. " On the brachyurous decapod Crustacea. Brought from the Cape by Dr. Smith. In: Smith, A. (Ed.), Illustrations of the Zoology of South Africa; consisting chiefly of Figures and Descriptions of the Objects of Natural History collected during an Expedition into the Interior of South Africa, in the Years 1834, 1835, and 1836; fitted out by " The Cape of Good Hope Association for Exploring Central Africa: " together with a Summary of African Zoology, and an Inquiry into the Geographical Ranges of Species in that Quarter of the Globe. Published under the Authority of the Lords Commissioners of Her Majesty's Treasury. Smith, Elder & Co., London, pp. 53 - 71, pls. 2 + 3." type="book chapter" year="1838">MacLeay, 1838</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
[originally
|
||
<taxonomicName id="4C2BFEF8FF80FFE6AC8AF849180288C9" authorityName="De Haan" authorityYear="1835" box="[1236,1345,1933,1959]" class="Malacostraca" family="Galenidae" genus="Halimede" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF80FFE6AC8AF849180288C9" box="[1236,1345,1933,1959]" italics="true" pageId="5" pageNumber="429">Halimede</emphasis>
|
||
</taxonomicName>
|
||
] and
|
||
<taxonomicName id="4C2BFEF8FF80FFE6ADD9F84A1C8988A5" authority="(Serene, 1971)" baseAuthorityName="Serene" baseAuthorityYear="1971" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="velasquiezi">
|
||
<emphasis id="B95F5969FF80FFE6ADD9F84A1C5088A5" italics="true" pageId="5" pageNumber="429">S. velasquiezi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBAF88AFF80FFE6A976F8751C8188A5" author="Serene, R." box="[296,450,1969,1995]" pageId="5" pageNumber="429" refId="ref6494" refString="Serene, R. (1971) Observations preliminaires sur des brachyoures nouveaux ou mal connus du sudest asiatique (Crustacea Decapoda). Bulletin du Museum national d'Histoire naturelle, Paris, Series 2, 42, 903 ⎼ 918, pls. 1 ⎼ 6." type="journal volume" year="1971">Serène, 1971</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
[originally
|
||
<taxonomicName id="4C2BFEF8FF80FFE6AA07F8751F9688A5" authorityName="Serene" authorityYear="1971" box="[601,725,1969,1995]" class="Malacostraca" family="Pilumnidae" genus="Leopoldius" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF80FFE6AA07F8751F9688A5" box="[601,725,1969,1995]" italics="true" pageId="5" pageNumber="429">Leopoldius</emphasis>
|
||
</taxonomicName>
|
||
]. In addition,
|
||
<bibRefCitation id="EFBAF88AFF80FFE6ABDBF875192E88A5" author="Ghory, F. & Ng, P. K. L. & Kazmi, Q. B." box="[901,1133,1969,1996]" pageId="5" pageNumber="429" refId="ref5451" refString="Ghory, F., Ng, P. K. L. & Kazmi, Q. B. (2013) On the identities of Pilumnopeus convexus MacCagno, 1936, and P. salomonensis Ward, 1942, from the Indian Ocean, with a note on P. riuii Takeda, 2001 (Decapoda, Brachyura, Pilumnidae). Crustaceana, 86, 301 ⎼ 312. https: // doi. org / 10.1163 / 15685403 - 00003180" type="book" year="2013">
|
||
Ghory
|
||
<emphasis id="B95F5969FF80FFE6AB87F876194C88A5" box="[985,1039,1969,1995]" italics="true" pageId="5" pageNumber="429">et al</emphasis>
|
||
. (2013)
|
||
</bibRefCitation>
|
||
mentioned that
|
||
<taxonomicName id="4C2BFEF8FF80FFE6AD6CF8751C3F8881" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B95F5969FF80FFE6AD6CF8751DB38881" italics="true" pageId="5" pageNumber="429">Pilumnus sinensis</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF80FFE6A8A8F8111C3F8881" author="Balss, H." box="[246,380,2005,2031]" pageId="5" pageNumber="429" refId="ref5021" refString="Balss, H. (1933) Beitrage zur Kenntnis der Gattung Pilumnus (Crustacea Dekapoda) und verwandter Gattungen. Capita Zoologica, 4 (3), 1 ⎼ 47, pls. 1 - 7." type="journal volume" year="1933">Balss, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
, is probably referable to this genus. They are all small species and covered with tomentum.
|
||
<bibRefCitation id="EFBAF88AFF80FFE5AD24F8121C1A8FDF" author="Ng, P. K. L. & Rahayu, D. L." lastPageId="6" lastPageNumber="430" pageId="5" pageNumber="429" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu (2021)</bibRefCitation>
|
||
examined the
|
||
<typeStatus id="54903BD9FF83FFE5A9A4FF531F698FDF" box="[506,554,151,177]" pageId="6" pageNumber="430">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C2BFEF8FF83FFE5AAE3FF5319088FDF" authority="Serene, 1971" authorityName="Serene" authorityYear="1971" box="[701,1099,151,177]" class="Malacostraca" family="Pilumnidae" genus="Leopoldius" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="velasquezi">
|
||
<emphasis id="B95F5969FF83FFE5AAE3FF531EF78FDF" box="[701,948,151,177]" italics="true" pageId="6" pageNumber="430">Leopoldius velasquezi</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF83FFE5ABE4FF5319088FDF" author="Serene, R." box="[954,1099,151,177]" pageId="6" pageNumber="430" refId="ref6494" refString="Serene, R. (1971) Observations preliminaires sur des brachyoures nouveaux ou mal connus du sudest asiatique (Crustacea Decapoda). Bulletin du Museum national d'Histoire naturelle, Paris, Series 2, 42, 903 ⎼ 918, pls. 1 ⎼ 6." type="journal volume" year="1971">Serène, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
and indicated that it may even be conspecific with
|
||
<taxonomicName id="4C2BFEF8FF83FFE5A926FF7F1CB38FBB" baseAuthorityName="Gordon" baseAuthorityYear="1934" box="[376,496,187,213]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="leopoldi">
|
||
<emphasis id="B95F5969FF83FFE5A926FF7F1CB38FBB" box="[376,496,187,213]" italics="true" pageId="6" pageNumber="430">S. leopoldi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B94857BFF83FFE5A899FF1B1C138EBF" blockId="6.[151,1437,151,861]" pageId="6" pageNumber="430">
|
||
The genus
|
||
<taxonomicName id="4C2BFEF8FF83FFE5A91BFF1B1CB48F97" authorityName="Turkay & Schuhmacher" authorityYear="1985" box="[325,503,223,249]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF83FFE5A91BFF1B1CB48F97" box="[325,503,223,249]" italics="true" pageId="6" pageNumber="430">Serenepilumnus</emphasis>
|
||
</taxonomicName>
|
||
was noted by
|
||
<bibRefCitation id="EFBAF88AFF83FFE5AAC0FF1B1E888F97" author="Ng, P. K. L. & Rahayu, D. L." box="[670,971,223,249]" pageId="6" pageNumber="430" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu (2021: 270)</bibRefCitation>
|
||
in the paper describing a new species of
|
||
<taxonomicName id="4C2BFEF8FF83FFE5A8C9FEC71C428E73" authorityName="Leach" authorityYear="1815" box="[151,257,259,285]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF83FFE5A8C9FEC71C428E73" box="[151,257,259,285]" italics="true" pageId="6" pageNumber="430">Pilumnus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2BFEF8FF83FFE5A951FEC71CCD8E72" authorityName="Ng & Rahayu" authorityYear="2021" box="[271,398,259,284]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="swajayai">
|
||
<emphasis id="B95F5969FF83FFE5A951FEC71CCD8E72" box="[271,398,259,284]" italics="true" pageId="6" pageNumber="430">P. swajayai</emphasis>
|
||
</taxonomicName>
|
||
from Indonesian waters: a dense tomentum completely obscures the surfaces and margins of the carapace, chelipeds and ambulatory legs, the carapace is prominently wide, the subhepatic region is unarmed; the carpus of the cheliped is without grooves, the cutting edges of the fingers of the chela are blade-like, and the ambulatory legs are very short. Although, at present, it may be difficult to determine whether all of these features are definitely generic, or may be specific or individual, the Indonesian species was referred not to
|
||
<taxonomicName id="4C2BFEF8FF83FFE5ACBDFE5718D68EC3" authorityName="Turkay & Schuhmacher" authorityYear="1985" box="[1251,1429,403,429]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF83FFE5ACBDFE5718D68EC3" box="[1251,1429,403,429]" italics="true" pageId="6" pageNumber="430">Serenepilumnus</emphasis>
|
||
</taxonomicName>
|
||
, but to
|
||
<taxonomicName id="4C2BFEF8FF83FFE5A881FE731C0A8EBF" authorityName="Leach" authorityYear="1815" box="[223,329,439,465]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF83FFE5A881FE731C0A8EBF" box="[223,329,439,465]" italics="true" pageId="6" pageNumber="430">Pilumnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B94857BFF83FFE5A899FE1F1EA88C33" blockId="6.[151,1437,151,861]" pageId="6" pageNumber="430">
|
||
The specimens from Japanese waters seem to be more densely covered with tomentum than the Indonesian specimens of
|
||
<taxonomicName id="4C2BFEF8FF83FFE5A968FE3B1CF48D76" authorityName="Ng & Rahayu" authorityYear="2021" box="[310,439,511,536]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="swajayai">
|
||
<emphasis id="B95F5969FF83FFE5A968FE3B1CF48D76" box="[310,439,511,536]" italics="true" pageId="6" pageNumber="430">P. swajayai</emphasis>
|
||
</taxonomicName>
|
||
as reported by
|
||
<bibRefCitation id="EFBAF88AFF83FFE5AA32FE3B1E218D77" author="Ng, P. K. L. & Rahayu, D. L." box="[620,866,511,537]" pageId="6" pageNumber="430" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu (2021)</bibRefCitation>
|
||
, but on denudation, the fronto-orbital margin, the carapace anterolateral teeth, the chelipeds, and the ambulatory legs are close in the forms from both areas. They are considered to be the same species, with some minor variations. However, most species of
|
||
<taxonomicName id="4C2BFEF8FF83FFE5ACF4FD8318578D0F" authorityName="Leach" authorityYear="1815" box="[1194,1300,583,609]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF83FFE5ACF4FD8318578D0F" box="[1194,1300,583,609]" italics="true" pageId="6" pageNumber="430">Pilumnus</emphasis>
|
||
</taxonomicName>
|
||
are covered with setae of various lengths, without velvety tomentum, the carapace dorsal surface is shallowly divided into regions and sparsely granulated or minutely tuberculated, and the carapace anterolateral margin is usually armed with spines or spiniform teeth with broad base.
|
||
<bibRefCitation id="EFBAF88AFF83FFE5AAEEFD771EDC8DA3" author="Ng, P. K. L. & Rahayu, D. L." box="[688,927,691,717]" pageId="6" pageNumber="430" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu (2021)</bibRefCitation>
|
||
mentioned of
|
||
<taxonomicName id="4C2BFEF8FF83FFE5AC1AFD7719878DA2" authorityName="Ng & Rahayu" authorityYear="2021" box="[1092,1220,691,716]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="swajayai">
|
||
<emphasis id="B95F5969FF83FFE5AC1AFD7719878DA2" box="[1092,1220,691,716]" italics="true" pageId="6" pageNumber="430">P. swajayai</emphasis>
|
||
</taxonomicName>
|
||
that its appearance is close to many Indo-West Pacific species of
|
||
<taxonomicName id="4C2BFEF8FF83FFE5AACBFD131FBC8D9F" authorityName="Leach" authorityYear="1815" box="[661,767,727,753]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF83FFE5AACBFD131FBC8D9F" box="[661,767,727,753]" italics="true" pageId="6" pageNumber="430">Pilumnus</emphasis>
|
||
</taxonomicName>
|
||
, and thus referred it to that genus. The species was also said to be distinct in possessing a hepatic dentiform lobe, the presence of a distinct supraorbital tooth, and the carinate and lobiform infraorbital margin. The validity of velvety tomentum tightly covering the carapace, chelipeds and ambulatory legs was not mentioned, but might be also peculiar in
|
||
<taxonomicName id="4C2BFEF8FF83FFE5AB24FC871EA78C33" authorityName="Leach" authorityYear="1815" box="[890,996,835,861]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF83FFE5AB24FC871EA78C33" box="[890,996,835,861]" italics="true" pageId="6" pageNumber="430">Pilumnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF54D5F3FF83FFE5A8C9F8491D9988A7" ID-DOI="http://doi.org/10.5281/zenodo.12681873" ID-Zenodo-Dep="12681873" httpUri="https://zenodo.org/record/12681873/files/figure.png" pageId="6" pageNumber="430" startId="6.[151,250,1933,1957]" targetBox="[272,1315,885,1909]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B94857BFF83FFE5A8C9F8491D9988A7" blockId="6.[151,1436,1933,1993]" pageId="6" pageNumber="430">
|
||
<emphasis id="B95F5969FF83FFE5A8C9F8491C5188CB" bold="true" box="[151,274,1933,1957]" pageId="6" pageNumber="430">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C2BFEF8FF83FFE5A946F8491CAE88CB" authorityName="Takeda & Komatsu" authorityYear="2024" box="[280,493,1933,1957]" class="Malacostraca" family="Pilumnidae" genus="Actumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
|
||
<emphasis id="B95F5969FF83FFE5A946F8491CAE88CB" box="[280,493,1933,1957]" italics="true" pageId="6" pageNumber="430">Actumnus ngankeeae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F5969FF83FFE5A9ADF8491F0088CB" bold="true" box="[499,579,1933,1957]" pageId="6" pageNumber="430">
|
||
<taxonomicNameLabel id="A26CE412FF83FFE5A9ADF8491F0088CB" box="[499,579,1933,1957]" pageId="6" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male (cb 4.6 mm, cl 3.5 mm), Ogasawara Islands, NSMT-Cr 31014. Color in life.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF54D5F3FF82FFE4A8C9F9D61E6D891C" ID-DOI="http://doi.org/10.5281/zenodo.12681875" ID-Zenodo-Dep="12681875" httpUri="https://zenodo.org/record/12681875/files/figure.png" pageId="7" pageNumber="431" startId="7.[151,250,1554,1578]" targetBox="[215,1371,181,1530]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B94857BFF82FFE4A8C9F9D61E6D891C" blockId="7.[151,1436,1554,1650]" pageId="7" pageNumber="431">
|
||
<emphasis id="B95F5969FF82FFE4A8C9F9D61C558944" bold="true" box="[151,278,1554,1578]" pageId="7" pageNumber="431">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C2BFEF8FF82FFE4A941F9D61F818944" authority="Ng & Rahayu, 2021" authorityName="Ng & Rahayu" authorityYear="2021" box="[287,706,1554,1578]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="swajayai">
|
||
<emphasis id="B95F5969FF82FFE4A941F9D61CA18944" box="[287,482,1554,1578]" italics="true" pageId="7" pageNumber="431">Pilumnus swajayai</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF82FFE4A9B5F9D61F818944" author="Ng, P. K. L. & Rahayu, D. L." box="[491,706,1554,1578]" pageId="7" pageNumber="431" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, male (cb 5.8 mm, cl 5.0 mm), Ryukyu Islands, NSMT-Cr 31015: A, habitus, dorsal view; habitus, anterior view; C, habitus, oblique dorsal view (carapace, right-half denuded); D, habitus, dorsal view (carapace, right-half denuded); E, both chelae, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B94857BFF82FFEBA899F965194D8E9B" blockId="7.[151,1437,1696,2011]" lastBlockId="8.[151,1437,151,609]" lastPageId="8" lastPageNumber="432" pageId="7" pageNumber="431">
|
||
<bibRefCitation id="EFBAF88AFF82FFE4A899F9651CF889D4" author="Ng, P. K. L. & Rahayu, D. L." box="[199,443,1696,1723]" pageId="7" pageNumber="431" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu (2021)</bibRefCitation>
|
||
also discussed the affinity of
|
||
<taxonomicName id="4C2BFEF8FF82FFE4AB4BF9651ED489D4" authorityName="Ng & Rahayu" authorityYear="2021" box="[789,919,1697,1722]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="swajayai">
|
||
<emphasis id="B95F5969FF82FFE4AB4BF9651ED489D4" box="[789,919,1697,1722]" italics="true" pageId="7" pageNumber="431">P. swajayai</emphasis>
|
||
</taxonomicName>
|
||
with the genus
|
||
<taxonomicName id="4C2BFEF8FF82FFE4AC0EF96418DB89D4" authority="Takeda, 1974" authorityName="Takeda" authorityYear="1974" box="[1104,1432,1696,1722]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF82FFE4AC0EF96419B089D4" box="[1104,1267,1696,1722]" italics="true" pageId="7" pageNumber="431">Nanopilumnus</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF82FFE4ACA3F96418DB89D4" author="Takeda, M." box="[1277,1432,1696,1722]" pageId="7" pageNumber="431" refId="ref6637" refString="Takeda, M. (1974) Pilumnid crabs of the family Xanthidae from the West Pacific. V. Definition of a new genus, with description of its type-species. Bulletin of the National Science Museum, Tokyo, 17, 215 ⎼ 219." type="journal volume" year="1974">Takeda, 1974</bibRefCitation>
|
||
</taxonomicName>
|
||
, which was erected based on
|
||
<taxonomicName id="4C2BFEF8FF82FFE4A98BF9001E4489B1" authority="Balss, 1935" authorityName="Balss" authorityYear="1935" box="[469,775,1732,1759]" class="Malacostraca" family="Xanthidae" genus="Medaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="B95F5969FF82FFE4A98BF9001F3889B0" box="[469,635,1732,1758]" italics="true" pageId="7" pageNumber="431">Medaeus rouxi</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF82FFE4AADFF9001E4489B1" author="Balss, H." box="[641,775,1732,1759]" pageId="7" pageNumber="431" refId="ref5060" refString="Balss, H. (1935) On three Indian crabs (Decapoda, Brachyura) of the Madras Museum. Records of the Indian Museum, 37, 45 ⎼ 48. https: // doi. org / 10.26515 / rzsi / v 37 / i 1 / 1935 / 163005" type="journal volume" year="1935">Balss, 1935</bibRefCitation>
|
||
</taxonomicName>
|
||
, and at present includes six species (
|
||
<bibRefCitation id="EFBAF88AFF82FFE4ACC3F900180289B0" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1181,1345,1732,1759]" pageId="7" pageNumber="431" refId="ref6104" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17, 1 ⎼ 286." type="journal volume" year="2008">
|
||
Ng
|
||
<emphasis id="B95F5969FF82FFE4AC96F90119BB89B0" box="[1224,1272,1732,1758]" italics="true" pageId="7" pageNumber="431">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBAF88AFF82FFE4AD12F9001C20886C" author="Takeda, M. & Komatsu, H." pageId="7" pageNumber="431" refId="ref6684" refString="Takeda, M. & Komatsu, H. (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 52, 153 ⎼ 189." type="journal volume" year="2018">Takeda & Komatsu 2018</bibRefCitation>
|
||
):
|
||
<taxonomicName id="4C2BFEF8FF82FFE4A924F92D1E7F886C" authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[378,828,1768,1794]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="barbatus">
|
||
<emphasis id="B95F5969FF82FFE4A924F92D1F42886C" box="[378,513,1768,1794]" italics="true" pageId="7" pageNumber="431">N. barbatus</emphasis>
|
||
(A.
|
||
<bibRefCitation id="EFBAF88AFF82FFE4AA64F92C1E77886C" author="Milne-Edwards, A." box="[570,820,1768,1794]" pageId="7" pageNumber="431" refId="ref6025" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 ⎼ 332, pls. 4 ⎼ 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal volume" year="1873">Milne-Edwards, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2BFEF8FF82FFE4AB15F92D190D886D" authority="(Balss, 1935)" baseAuthorityName="Balss" baseAuthorityYear="1935" box="[843,1102,1768,1795]" class="Malacostraca" family="Xanthidae" genus="Medaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="B95F5969FF82FFE4AB15F92D1EEA886C" box="[843,937,1769,1794]" italics="true" pageId="7" pageNumber="431">N. rouxi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBAF88AFF82FFE4ABE2F92C1906886D" author="Balss, H." box="[956,1093,1768,1795]" pageId="7" pageNumber="431" refId="ref5060" refString="Balss, H. (1935) On three Indian crabs (Decapoda, Brachyura) of the Madras Museum. Records of the Indian Museum, 37, 45 ⎼ 48. https: // doi. org / 10.26515 / rzsi / v 37 / i 1 / 1935 / 163005" type="journal volume" year="1935">Balss, 1935</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2BFEF8FF82FFE4AC03F92D18DB886C" authority="(Sakai, 1934)" baseAuthorityName="Sakai" baseAuthorityYear="1934" box="[1117,1432,1768,1794]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="heterodon">
|
||
<emphasis id="B95F5969FF82FFE4AC03F92D19B1886C" box="[1117,1266,1768,1794]" italics="true" pageId="7" pageNumber="431">N. heterodon</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBAF88AFF82FFE4AD5DF92C18D3886C" author="Sakai, T." box="[1283,1424,1768,1794]" pageId="7" pageNumber="431" refId="ref6456" refString="Sakai, T. (1934) Brachyura from the coast of Kyushu, Japan. Science Reports of Tokyo Bunrika Diagaku, Series B, 1, 281 ⎼ 330, pls. 17, 18." type="journal volume" year="1934">Sakai, 1934</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2BFEF8FF82FFE4A8C9F8C91FD78848" authority="(Takeda & Miyake, 1969)" baseAuthorityName="Takeda & Miyake" baseAuthorityYear="1969" box="[151,660,1804,1830]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="coralliophilus">
|
||
<emphasis id="B95F5969FF82FFE4A8C9F8C91C1A8848" box="[151,345,1804,1830]" italics="true" pageId="7" pageNumber="431">N. coralliophilus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBAF88AFF82FFE4A935F8C81FCF8848" author="Takeda, M. & Miyake, S." box="[363,652,1804,1830]" pageId="7" pageNumber="431" refId="ref6738" refString="Takeda, M. & Miyake, S. (1969) Pilumnid crabs of the family Xanthidae from the West Pacific. II. Twenty-one species of four genera, with descriptions of four new species. Ohmu, 2, 93 - 156." type="journal volume" year="1969">Takeda & Miyake, 1969</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2BFEF8FF82FFE4AAFDF8C919128848" authority="(Takeda & Miyake, 1969)" baseAuthorityName="Takeda & Miyake" baseAuthorityYear="1969" box="[675,1105,1804,1830]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="hondai">
|
||
<emphasis id="B95F5969FF82FFE4AAFDF8C91E558848" box="[675,790,1804,1830]" italics="true" pageId="7" pageNumber="431">N. hondai</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBAF88AFF82FFE4AB76F8C8190A8848" author="Takeda, M. & Miyake, S." box="[808,1097,1804,1830]" pageId="7" pageNumber="431" refId="ref6738" refString="Takeda, M. & Miyake, S. (1969) Pilumnid crabs of the family Xanthidae from the West Pacific. II. Twenty-one species of four genera, with descriptions of four new species. Ohmu, 2, 93 - 156." type="journal volume" year="1969">Takeda & Miyake, 1969</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2BFEF8FF82FFE4ACCAF8C91C0B8824" authority="Takeda & Komatsu, 2018" authorityName="Takeda & Komatsu" authorityYear="2018" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="modestus">
|
||
<emphasis id="B95F5969FF82FFE4ACCAF8C918638848" box="[1172,1312,1804,1830]" italics="true" pageId="7" pageNumber="431">N. modestus</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF82FFE4AD75F8C81C0B8824" author="Takeda, M. & Komatsu, H." pageId="7" pageNumber="431" refId="ref6684" refString="Takeda, M. & Komatsu, H. (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 52, 153 ⎼ 189." type="journal volume" year="2018">Takeda & Komatsu, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C2BFEF8FF82FFE4A90BF8F41E438825" authority="Monod, 1956" authorityName="Monod" authorityYear="1956" box="[341,768,1840,1867]" class="Malacostraca" family="Acidopsidae" genus="Parapilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="boletifer">
|
||
<emphasis id="B95F5969FF82FFE4A90BF8F41F188824" box="[341,603,1840,1866]" italics="true" pageId="7" pageNumber="431">Parapilumnus boletifer</emphasis>
|
||
<bibRefCitation id="EFBAF88AFF82FFE4AA3AF8F41E438825" author="Monod, Th." box="[612,768,1840,1867]" pageId="7" pageNumber="431" refId="ref6079" refString="Monod, Th. (1956) Hippidea et Brachyura oust-africains. Memoires de l'Institut Francais d'Afrique Noire, 45, 1 ⎼ 674." type="journal volume" year="1956">Monod, 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
, formerly included in
|
||
<taxonomicName id="4C2BFEF8FF82FFE4AC5FF8F4182A8824" authority="(Ng et al. 2008)" baseAuthorityName="Ng" baseAuthorityYear="2008" box="[1025,1385,1840,1867]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF82FFE4AC5FF8F419E78824" box="[1025,1188,1840,1866]" italics="true" pageId="7" pageNumber="431">Nanopilumnus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBAF88AFF82FFE4ACEBF8F418228824" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1205,1377,1840,1867]" pageId="7" pageNumber="431" refId="ref6104" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17, 1 ⎼ 286." type="journal volume" year="2008">
|
||
Ng
|
||
<emphasis id="B95F5969FF82FFE4ACBCF8F518558824" box="[1250,1302,1840,1866]" italics="true" pageId="7" pageNumber="431">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, has been designated as the
|
||
<typeStatus id="54903BD9FF82FFE4A9C9F8911C848801" box="[407,455,1877,1903]" pageId="7" pageNumber="431">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="4C2BFEF8FF82FFE4AAEAF8901E268800" authorityName="Stevcic" authorityYear="2011" box="[692,869,1876,1902]" class="Malacostraca" family="Pilumnidae" genus="Balssopilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF82FFE4AAEAF8901E268800" box="[692,869,1876,1902]" italics="true" pageId="7" pageNumber="431">Balssopilumnus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFBAF88AFF82FFE4ABD0F89219778800" author="Stevcic, Z." box="[910,1076,1876,1905]" pageId="7" pageNumber="431" refId="ref6599" refString="Stevcic, Z. (2011) Addition to the reclassification of the brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica, 20, 125 - 139." type="journal volume" year="2011">Števčić (2011)</bibRefCitation>
|
||
. As mentioned by
|
||
<bibRefCitation id="EFBAF88AFF82FFE4AD5CF8911DA788FC" author="Ng, P. K. L. & Rahayu, D. L." pageId="7" pageNumber="431" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu (2021)</bibRefCitation>
|
||
, species of
|
||
<taxonomicName id="4C2BFEF8FF82FFE4A92FF8BC1F5788FC" box="[369,532,1912,1938]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="431" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF82FFE4A92FF8BC1F5788FC" box="[369,532,1912,1938]" italics="true" pageId="7" pageNumber="431">Nanopilumnus</emphasis>
|
||
</taxonomicName>
|
||
share the common characters of a small adult size with the carapace smaller than cb
|
||
<quantity id="4CD3289EFF82FFE4A8ADF8581C7488D9" box="[243,311,1948,1975]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="431" unit="mm" value="6.0">6 mm</quantity>
|
||
, the relatively dense, soft setae covering the carapace, chelipeds and ambulatory legs, the presence of three distinct, stout anterolateral teeth of the carapace, and no subhepatic tooth. However, the presence of a distinct subhepatic tooth and the deep transverse groove along the distal part of the cheliped carpus argue against its placement in
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBA911FF7F1CB18FBB" box="[335,498,187,213]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF8DFFEBA911FF7F1CB18FBB" box="[335,498,187,213]" italics="true" pageId="8" pageNumber="432">Nanopilumnus</emphasis>
|
||
</taxonomicName>
|
||
. In the genus
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBAAD1FF7F1FBA8FBB" authorityName="Leach" authorityYear="1815" box="[655,761,187,213]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF8DFFEBAAD1FF7F1FBA8FBB" box="[655,761,187,213]" italics="true" pageId="8" pageNumber="432">Pilumnus</emphasis>
|
||
</taxonomicName>
|
||
, the subhepatic surface of the carapace is unarmed or armed with a prominent tooth or cluster of some granules, and the cheliped merus is usually provided with a shallow marginal furrow. The
|
||
<typeStatus id="54903BD9FF8DFFEBA9D2FEC71CFF8E73" box="[396,444,259,285]" pageId="8" pageNumber="432">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBAAF3FEC71E138E73" box="[685,848,259,285]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF8DFFEBAAF3FEC71E138E73" box="[685,848,259,285]" italics="true" pageId="8" pageNumber="432">Nanopilumnus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBAB01FEC71EF98E72" baseAuthorityName="Balss" baseAuthorityYear="1935" box="[863,954,259,284]" class="Malacostraca" family="Xanthidae" genus="Medaeus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="B95F5969FF8DFFEBAB01FEC71EF98E72" box="[863,954,259,284]" italics="true" pageId="8" pageNumber="432">N. rouxi</emphasis>
|
||
</taxonomicName>
|
||
, was well documented by
|
||
<bibRefCitation id="EFBAF88AFF8DFFEBACB8FEC71DA78E2F" author="Serene, R. & Umali, A. F." pageId="8" pageNumber="432" refId="ref6542" refString="Serene, R. & Umali, A. F. (1972) The family Raninidae and other new and rare species of brachyuran decapods from the Philippines and adjacent regions. Philippine Journal of Science, 99, 21 ⎼ 105, pls. 1 ⎼ 9. [dated 1970, published 1972]" type="journal volume" year="1972">Serène & Umali (1972)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFBAF88AFF8DFFEBA8AFFEE31F408E2F" author="Tirmizi, N. M. & Ghani, N." box="[241,515,295,321]" pageId="8" pageNumber="432" refId="ref6784" refString="Tirmizi, N. M. & Ghani, N. (1986) Range extension of a rare pilumnid crab into the northern Arabian Sea (Decapoda, Brachyura). Crustaceana, 51, 293 ⎼ 295. https: // doi. org / 10.1163 / 156854086 X 00458" type="journal volume" year="1986">Tirmizi & Ghani (1986)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="EFBAF88AFF8DFFEBAA1EFEE31E708E2F" author="Asada, Y. & Sato, T." box="[576,819,295,321]" pageId="8" pageNumber="432" refId="ref4970" refString="Asada, Y. & Sato, T. (2020) New record of pilumnid crab Nanopilumnus rouxi (Balss, 1935) (Decapoda) from Okayama Prefecture, western Japan. Cancer, 29, e 113 ⎼ e 117. [in Japanese with English title]" type="journal volume" year="2020">Asada & Sato (2020)</bibRefCitation>
|
||
, and the Japanese species of
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBACDEFEE318608E2F" box="[1152,1315,295,321]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF8DFFEBACDEFEE318608E2F" box="[1152,1315,295,321]" italics="true" pageId="8" pageNumber="432">Nanopilumnus</emphasis>
|
||
</taxonomicName>
|
||
, including
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBA8C9FE8F1DB78E0A" baseAuthorityName="Balss" baseAuthorityYear="1935" box="[151,244,331,356]" class="Malacostraca" family="Xanthidae" genus="Medaeus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="B95F5969FF8DFFEBA8C9FE8F1DB78E0A" box="[151,244,331,356]" italics="true" pageId="8" pageNumber="432">N. rouxi</emphasis>
|
||
</taxonomicName>
|
||
, were taxonomically clarified by
|
||
<bibRefCitation id="EFBAF88AFF8DFFEBAA29FE8F1E138E0B" author="Maenosono, T." box="[631,848,331,357]" pageId="8" pageNumber="432" pagination="26 - 28" refId="ref5910" refString="Maenosono, T. (2022) On species of Nanopilumnus Takeda, 1974 (Pilumnidae) collected from southern Japan, with new distributional records. Niche Life, 9, 26 - 28. [in Japanese with English abstract]" type="journal article" year="2022">Maenosono (2022)</bibRefCitation>
|
||
, but it is still insufficient to delimit the genus from some related genera. As pointed by
|
||
<bibRefCitation id="EFBAF88AFF8DFFEBAA6EFEAB1ED08EE7" author="Turkay, M. & Schuhmacher, H." box="[560,915,366,393]" pageId="8" pageNumber="432" refId="ref6836" refString="Turkay, M. & Schuhmacher, H. (1985) Latopilumnus tubicolus n. gen. n. sp., eine neue korallenassoziierte Krebbe, die Bildung einer Wohnhohle induziert (Crustacea: Decapoda: Pilumnidae). Senckenbergiana maritima, 17, 55 - 63." type="journal volume" year="1985">Türkay & Schuhmacher (1985)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFBAF88AFF8DFFEBAB93FEAB19268EE7" author="Davie, P. J. F." box="[973,1125,367,393]" pageId="8" pageNumber="432" refId="ref5342" refString="Davie, P. J. F. (1989) Two new genera of the family Pilumnidae (Crustacea: Decapoda: Brachyura) from Queensland, Australia. Journal of Natural History, 23, 1353 ⎼ 1365. https: // doi. org / 10.1080 / 00222938900770721" type="journal volume" year="1989">Davie (1989)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="EFBAF88AFF8DFFEBACFDFEAB18D48EE7" author="Ng, P. K. L. & Rahayu, D. L." box="[1187,1431,367,393]" pageId="8" pageNumber="432" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu (2021)</bibRefCitation>
|
||
, the definition of
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBA90BFE571CBB8EC3" box="[341,504,403,429]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF8DFFEBA90BFE571CBB8EC3" box="[341,504,403,429]" italics="true" pageId="8" pageNumber="432">Nanopilumnus</emphasis>
|
||
</taxonomicName>
|
||
should be clarified. As mentioned above, the present species from Indonesian and Japanese waters has some mosaic characters among
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBAA87FE731EC88EBF" authorityName="Turkay & Schuhmacher" authorityYear="1985" box="[729,907,439,465]" class="Malacostraca" family="Pilumnidae" genus="Serenepilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF8DFFEBAA87FE731EC88EBF" box="[729,907,439,465]" italics="true" pageId="8" pageNumber="432">Serenepilumnus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBABC9FE7319798EBF" box="[919,1082,439,465]" class="Malacostraca" family="Pilumnidae" genus="Nanopilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF8DFFEBABC9FE7319798EBF" box="[919,1082,439,465]" italics="true" pageId="8" pageNumber="432">Nanopilumnus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBAC31FE73199A8EBF" authorityName="Leach" authorityYear="1815" box="[1135,1241,439,465]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF8DFFEBAC31FE73199A8EBF" box="[1135,1241,439,465]" italics="true" pageId="8" pageNumber="432">Pilumnus</emphasis>
|
||
</taxonomicName>
|
||
. In this paper, the species is referred to
|
||
<taxonomicName id="4C2BFEF8FF8DFFEBA9DBFE1F1CAC8E9B" authorityName="Leach" authorityYear="1815" box="[389,495,475,501]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F5969FF8DFFEBA9DBFE1F1CAC8E9B" box="[389,495,475,501]" italics="true" pageId="8" pageNumber="432">Pilumnus</emphasis>
|
||
</taxonomicName>
|
||
for the present, following
|
||
<bibRefCitation id="EFBAF88AFF8DFFEBAB47FE1F19498E9B" author="Ng, P. K. L. & Rahayu, D. L." box="[793,1034,475,501]" pageId="8" pageNumber="432" pagination="265 - 276" refId="ref6157" refString="Ng, P. K. L. & Rahayu, D. L. (2021) On the Pilumnidae (Crustacea: Brachyura: Pilumnidae) collected during the SJADES 2018 biodiversity cruise in Indonesia, with description of a new species of Pilumnus. Raffles Bulletin of Zoology, Supplement 36, 265 - 276." type="journal article" year="2021">Ng & Rahayu (2021)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331D6F0FF8DFFEBA899FE3A1F2F8D0F" pageId="8" pageNumber="432" type="distribution">
|
||
<paragraph id="8B94857BFF8DFFEBA899FE3A1F2F8D0F" blockId="8.[151,1437,151,609]" pageId="8" pageNumber="432">
|
||
<emphasis id="B95F5969FF8DFFEBA899FE3A1C228D76" bold="true" box="[199,353,510,536]" pageId="8" pageNumber="432">Distribution.</emphasis>
|
||
This species has originally been reported from the Sunda Strait,
|
||
<collectingRegion id="49EF4B99FF8DFFEBAC03FE3B19D28D77" box="[1117,1169,511,537]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="432">Java</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33CC5EBFF8DFFEBACFFFE3B184D8D77" box="[1185,1294,511,537]" name="Indonesia" pageId="8" pageNumber="432">Indonesia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE1FE3BCFF8DFFEBAD7EFE3B18DF8D77" box="[1312,1436,511,537]" degrees="6" direction="south" minutes="09.803" orientation="latitude" pageId="8" pageNumber="432" precision="1" value="-6.1633835">6°09.803'S</geoCoordinate>
|
||
<geoCoordinate id="EE1FE3BCFF8DFFEBA8C9FDE61C738D53" box="[151,304,546,573]" degrees="104" direction="east" minutes="57.976" orientation="longitude" pageId="8" pageNumber="432" precision="1" value="104.96627">104°57.976'E</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="EE1FE3BCFF8DFFEBA910FDE71C8A8D53" box="[334,457,547,573]" degrees="6" direction="south" minutes="09.606" orientation="latitude" pageId="8" pageNumber="432" precision="1" value="-6.1601">6°09.606'S</geoCoordinate>
|
||
<geoCoordinate id="EE1FE3BCFF8DFFEBA990FDE61F2A8D53" box="[462,617,546,573]" degrees="104" direction="east" minutes="58.208" orientation="longitude" pageId="8" pageNumber="432" precision="1" value="104.97013">104°58.208'E</geoCoordinate>
|
||
; 92⎼
|
||
<quantity id="4CD3289EFF8DFFEBAAC2FDE71FA78D53" box="[668,740,547,573]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="8" pageNumber="432" unit="m" value="103.0">103 m</quantity>
|
||
), and south of Cilacap, southern
|
||
<collectingRegion id="49EF4B99FF8DFFEBAC1BFDE719348D53" box="[1093,1143,547,573]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="432">Java</collectingRegion>
|
||
(
|
||
<geoCoordinate id="EE1FE3BCFF8DFFEBACDDFDE719BC8D53" box="[1155,1279,547,573]" degrees="8" direction="south" minutes="07.462" orientation="latitude" pageId="8" pageNumber="432" precision="1" value="-8.124367">8°07.462'S</geoCoordinate>
|
||
<geoCoordinate id="EE1FE3BCFF8DFFEBAD5AFDE618DF8D53" box="[1284,1436,546,573]" degrees="109" direction="east" minutes="05.639" orientation="longitude" pageId="8" pageNumber="432" precision="1" value="109.09399">109°05.639'E</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="EE1FE3BCFF8DFFEBA8EAFD831C6C8D0F" box="[180,303,583,609]" degrees="8" direction="south" minutes="07.864" orientation="latitude" pageId="8" pageNumber="432" precision="1" value="-8.131066">8°07.864'S</geoCoordinate>
|
||
<geoCoordinate id="EE1FE3BCFF8DFFEBA968FD831C928D0F" box="[310,465,583,609]" degrees="109" direction="east" minutes="06.470" orientation="longitude" pageId="8" pageNumber="432" precision="1" value="109.107834">109°06.470'E</geoCoordinate>
|
||
; 163⎼
|
||
<quantity id="4CD3289EFF8DFFEBAA4BFD831F238D0F" box="[533,608,583,609]" metricMagnitude="2" metricUnit="m" metricValue="1.66" pageId="8" pageNumber="432" unit="m" value="166.0">166 m</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |