285 lines
43 KiB
XML
285 lines
43 KiB
XML
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<mods:title id="FC30CC1ABF922EBB9BF4455505F59D18">Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)</mods:title>
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<mods:namePart id="DA644ECB5575D772CB5344851B882FFE">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="668240ECC2242FECC38A030B6B58450F">Zootaxa</mods:title>
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<mods:date id="1522E9B102684B47006162E0EF90C5CC">2023</mods:date>
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<mods:number id="012E80344904EA5B43F2425F6627024B">5238</mods:number>
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<treatment id="03C79010FFC7D72DFF707EC22736F852" ID-DOI="http://doi.org/10.5281/zenodo.7621919" ID-GBIF-Taxon="209295623" ID-Zenodo-Dep="7621919" LSID="urn:lsid:plazi:treatment:03C79010FFC7D72DFF707EC22736F852" httpUri="http://treatment.plazi.org/id/03C79010FFC7D72DFF707EC22736F852" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
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<subSubSection id="C374728DFFC7D72EFF707EC221DAFA08" box="[151,732,1481,1508]" pageId="49" pageNumber="50" type="nomenclature">
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<paragraph id="8BD12106FFC7D72EFF707EC221DAFA08" blockId="49.[151,732,1481,1543]" box="[151,732,1481,1508]" pageId="49" pageNumber="50">
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<heading id="D099966AFFC7D72EFF707EC221DAFA08" bold="true" box="[151,732,1481,1508]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
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<emphasis id="B91AFD14FFC7D72EFF707EC221DAFA08" bold="true" box="[151,732,1481,1508]" pageId="49" pageNumber="50">
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<taxonomicName id="4C6E5A85FFC7D72EFF707EC2215EFA08" ID-CoL="52W8Y" authority="de Quatrefages, 1866" authorityName="de Quatrefages" authorityYear="1866" box="[151,600,1481,1508]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="49" pageNumber="50" phylum="Annelida" rank="species" species="fucata">
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<emphasis id="B91AFD14FFC7D72EFF707EC2224CFA08" bold="true" box="[151,330,1481,1508]" italics="true" pageId="49" pageNumber="50">Chloeia fucata</emphasis>
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<bibRefCitation id="EFFF5CF7FFC7D72EFEB67EC2215EFA08" author="De Quatrefages, A." box="[337,600,1481,1507]" pageId="49" pageNumber="50" pagination="578 - 600" refId="ref105083" refString="De Quatrefages, A. (1866 (1865 )) Histoire naturelle des Anneles marins et d'eau douce. Annelides et Gephyriens. Vol. 1. Librarie Encyclopedique de Roret, Paris, 588 pp. [publication date after Wright, E. P. (1866). Section 6 (Annelida, etc.). Zoological Record, 1866, 578 - 600] https: // doi. org / 10.5962 / bhl. title. 122818" type="book chapter" year="1866">de Quatrefages, 1866</bibRefCitation>
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</taxonomicName>
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||
reinstated
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C374728DFFC7D72EFF707EE623E1F9EC" box="[151,231,1517,1543]" pageId="49" pageNumber="50" type="description">
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<paragraph id="8BD12106FFC7D72EFF707EE623E1F9EC" blockId="49.[151,732,1481,1543]" box="[151,231,1517,1543]" pageId="49" pageNumber="50">
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<figureCitation id="13553D83FFC7D72EFF707EE623E1F9EC" box="[151,231,1517,1543]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="49" pageNumber="50">Fig. 21</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C374728DFFC7D72EFF707D3927EEF9A1" box="[151,1256,1585,1610]" pageId="49" pageNumber="50" type="reference_group">
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<paragraph id="8BD12106FFC7D72EFF707D3927EEF9A1" blockId="49.[151,1256,1585,1610]" box="[151,1256,1585,1610]" pageId="49" pageNumber="50">
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<treatmentCitationGroup id="AB7E0628FFC7D72EFF707D3927EEF9A1" box="[151,1256,1585,1610]" pageId="49" pageNumber="50">
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<treatmentCitation id="0ACF0717FFC7D72EFF707D392144F9A1" author="De Quatrefages, A." box="[151,578,1585,1610]" page="390" pageId="49" pageNumber="50" year="1866">
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<taxonomicName id="4C6E5A85FFC7D72EFF707D392144F9A1" ID-CoL="52W8Y" authority="de Quatrefages, 1866: 390" authorityName="de Quatrefages" authorityPageNumber="390" authorityYear="1866" box="[151,578,1585,1610]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="49" pageNumber="50" phylum="Annelida" rank="species" species="fucata">
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<emphasis id="B91AFD14FFC7D72EFF707D39222BF9A1" box="[151,301,1586,1610]" italics="true" pageId="49" pageNumber="50">Chloeia fucata</emphasis>
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<bibRefCitation id="EFFF5CF7FFC7D72EFED47D392144F9A1" author="De Quatrefages, A." box="[307,578,1585,1610]" pageId="49" pageNumber="50" pagination="578 - 600" refId="ref105083" refString="De Quatrefages, A. (1866 (1865 )) Histoire naturelle des Anneles marins et d'eau douce. Annelides et Gephyriens. Vol. 1. Librarie Encyclopedique de Roret, Paris, 588 pp. [publication date after Wright, E. P. (1866). Section 6 (Annelida, etc.). Zoological Record, 1866, 578 - 600] https: // doi. org / 10.5962 / bhl. title. 122818" type="book chapter" year="1866">de Quatrefages, 1866: 390</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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;
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||
<treatmentCitation id="0ACF0717FFC7D72EFDAB7D3921F2F9A1" author="Baird, W." box="[588,756,1585,1610]" page="232" pageId="49" pageNumber="50" year="1868">
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<bibRefCitation id="EFFF5CF7FFC7D72EFDAB7D3921F2F9A1" author="Baird, W." box="[588,756,1585,1610]" pageId="49" pageNumber="50" pagination="215 - 250" refId="ref103011" refString="Baird, W. (1868) Contributions towards a monograph of the species of annelides belonging to the Amphinomacea, with a list of the known species, and a description of several new species (belonging to the group) contained in the National Collection of the British Museum. To which is appended a short account of two hitherto nondescript annulose animals of a larval character. The Journal of the Linnean Society of London, Zoology, 10 (44), 215 - 250, pls. 4 - 6. https: // doi. org / 10.1111 / j. 1096 - 3642.1868. tb 02233. x" type="journal article" year="1868">Baird 1868: 232</bibRefCitation>
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</treatmentCitation>
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;
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||
<treatmentCitation id="0ACF0717FFC7D72EFD197D3920C0F9A1" author="Hartman, O." box="[766,966,1585,1610]" page="131" pageId="49" pageNumber="50" year="1959">
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<bibRefCitation id="EFFF5CF7FFC7D72EFD197D3920C0F9A1" author="Hartman, O." box="[766,966,1585,1610]" pageId="49" pageNumber="50" pagination="1 - 353" refId="ref107655" refString="Hartman, O. (1959) Catalogue of the polychaetous annelids of the World. Part 1. Allan Hancock Foundation Publications, Occasional Paper, 23, 1 - 353." type="journal article" year="1959">Hartman 1959: 131</bibRefCitation>
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</treatmentCitation>
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;
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||
<bibRefCitation id="EFFF5CF7FFC7D72EFC367D3927B8F9A1" author="Solis-Weiss, V. & Bertrand, Y. & Helleouet, M. - N. & Pleijel, F." box="[977,1214,1586,1610]" pageId="49" pageNumber="50" pagination="377 - 384" refId="ref113040" refString="Solis-Weiss, V., Bertrand, Y., Helleouet, M. - N. & Pleijel, F. (2004) Types of polychaetous annelids at the Museum national d'Histoire naturelle, Paris. Zoosystema, 26, 377 - 384, Supplement 21 pp." type="book chapter" year="2004">
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Solís-Weiss
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||
<emphasis id="B91AFD14FFC7D72EFBA97D392787F9A1" box="[1102,1153,1586,1610]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
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2004
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</bibRefCitation>
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||
: S2.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C374728DFFC7D72EFF707D73210AF95D" pageId="49" pageNumber="50" type="materials_examined">
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||
<paragraph id="8BD12106FFC7D72EFF707D73210AF95D" blockId="49.[151,1437,1655,2042]" pageId="49" pageNumber="50">
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||
<emphasis id="B91AFD14FFC7D72EFF707D732245F97A" bold="true" box="[151,323,1655,1682]" pageId="49" pageNumber="50">Type material</emphasis>
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||
.
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<emphasis id="B91AFD14FFC7D72EFEB17D7C21AFF97A" bold="true" box="[342,681,1655,1681]" pageId="49" pageNumber="50">Indian Ocean, Arabian Sea</emphasis>
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||
.
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<materialsCitation id="3B062B5BFFC7D72EFD5B7D73210AF95D" ID-GBIF-Occurrence="4029882374" collectionCode="MNHN" collectorName="M. Leclancher" country="Oman" location="Mascate" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Masqat" typeStatus="holotype">
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||
<typeStatus id="54D59FA4FFC7D72EFD5B7D732022F979" box="[700,804,1656,1682]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
|
||
(
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||
<collectionCode id="ED7FB9C3FFC7D72EFCDF7D732089F979" box="[824,911,1656,1682]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="49" pageNumber="50">MNHN</collectionCode>
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||
IA-TYPE 80,
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||
<collectingRegion id="49AAEFE4FFC7D72EFBA47D7327A6F979" box="[1091,1184,1656,1682]" country="Oman" name="Masqat" pageId="49" pageNumber="50">Mascate</collectingRegion>
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||
(Muscat),
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||
<collectingCountry id="F3796196FFC7D72EFAC27D73266AF979" box="[1317,1388,1656,1682]" name="Oman" pageId="49" pageNumber="50">Oman</collectingCountry>
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||
,
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<collectorName id="269B44D0FFC7D72EFA9B7D732213F95D" pageId="49" pageNumber="50">M. Leclancher</collectorName>
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, coll., no further data.
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C374728DFFC7D72EFF207DCB2061F8C9" pageId="49" pageNumber="50" type="diagnosis">
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||
<paragraph id="8BD12106FFC7D72EFF207DCB2061F8C9" blockId="49.[151,1437,1655,2042]" pageId="49" pageNumber="50">
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||
<emphasis id="B91AFD14FFC7D72EFF207DCB223CF931" bold="true" box="[199,314,1728,1754]" pageId="49" pageNumber="50">Diagnosis</emphasis>
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||
.
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<taxonomicName id="4C6E5A85FFC7D72EFEAC7DCB22A2F931" authorityName="Savigny" authorityYear="1818" box="[331,420,1728,1754]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="49" pageNumber="50" phylum="Annelida" rank="genus">
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||
<emphasis id="B91AFD14FFC7D72EFEAC7DCB22A2F931" box="[331,420,1728,1754]" italics="true" pageId="49" pageNumber="50">Chloeia</emphasis>
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||
</taxonomicName>
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||
with tiny bipinnate branchiae from chaetiger 3, gradually larger medially, progressively smaller posteriorly; middorsal spots subrectangular, longer than wide, sometimes wider medially; notochaetae furcates and harpoon-chaetae without spurs; neurochaetae furcates.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C374728DFFC7D72DFF207C2727B3FBFA" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="description">
|
||
<paragraph id="8BD12106FFC7D72EFF207C27225FF8AD" blockId="49.[151,1437,1655,2042]" box="[199,345,1836,1862]" pageId="49" pageNumber="50">
|
||
<emphasis id="B91AFD14FFC7D72EFF207C272257F8AD" bold="true" box="[199,337,1836,1862]" pageId="49" pageNumber="50">Description</emphasis>
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||
.
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||
</paragraph>
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||
<paragraph id="8BD12106FFC7D72EFF207C5B227EF865" blockId="49.[151,1437,1655,2042]" pageId="49" pageNumber="50">
|
||
<materialsCitation id="3B062B5BFFC7D72EFF207C5B227EF865" ID-GBIF-Occurrence="4029882499" collectionCode="MNHN" pageId="49" pageNumber="50" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="54D59FA4FFC7D72EFF207C5B2229F881" box="[199,303,1872,1898]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
|
||
(
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||
<collectionCode id="ED7FB9C3FFC7D72EFEA77C5B2291F881" box="[320,407,1872,1898]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="49" pageNumber="50">MNHN</collectionCode>
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||
IA-TYPE 80) complete, with slight histolysis; body subrectangular,
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||
<quantity id="4C968CE3FFC7D72EFB487C5B2606F881" box="[1199,1280,1872,1898]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="49" pageNumber="50" unit="mm" value="43.0">43 mm</quantity>
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||
long,
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||
<quantity id="4C968CE3FFC7D72EFAAC7C5B269BF881" box="[1355,1437,1872,1898]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="49" pageNumber="50" unit="mm" value="10.0">10 mm</quantity>
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||
wide, 27 chaetigers.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BD12106FFC7D72EFF207C932729F811" blockId="49.[151,1437,1655,2042]" pageId="49" pageNumber="50">
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||
<typeStatus id="54D59FA4FFC7D72EFF207C932229F859" box="[199,303,1944,1970]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
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||
pale, chaetae golden (
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||
<figureCitation id="13553D83FFC7D72EFDC17C93218BF859" box="[550,653,1944,1970]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="49" pageNumber="50">Fig. 21A</figureCitation>
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); prostomium, cephalic appendages, dorsal and ventral cirri pale, from chaetiger 14 a middorsal dark spot oval, longer than wide (
|
||
<figureCitation id="13553D83FFC7D72EFCC47CB7208FF83D" box="[803,905,1980,2006]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="49" pageNumber="50">Fig. 21D</figureCitation>
|
||
), better defined along last 5–6 chaetigers. Spots displaced towards posterior segmental margin, barely wider medially. Venter pale.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF207B9C27AEFEF6" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">
|
||
Prostomium anteriorly entire. Eyes blackish, small, anterior eyes slightly larger than posterior ones (
|
||
<figureCitation id="13553D83FFC4D72DFACE7B9C2689FF5A" box="[1321,1423,151,177]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="50" pageNumber="51">Fig. 21B</figureCitation>
|
||
). Median antenna inserted at anterior caruncular margin, contracted, about half as long as caruncle, 2× longer than lateral antennae. Lateral antennae bases very close to each other, about 2× longer than palps. Mouth ventral on chaetiger 2. Pharynx slightly exposed, smooth, with moderate histolysis, muscular wall very thin.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF207A2C27B7FE62" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">
|
||
Caruncle bent to the right, trilobed, tapered, reaching posterior margin of chaetiger 3 (
|
||
<figureCitation id="13553D83FFC4D72DFB6C7A2C27F5FEAA" box="[1163,1267,295,321]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="50" pageNumber="51">Fig. 21C</figureCitation>
|
||
). Median ridge plicate, with about 23 vertical folds, almost completely concealing lateral lobes at least along half its length. Lateral lobes narrow, with about 20 vertical folds, posterior ones indistinct, fused to median ridge tip.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF207A982201FDD6" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">
|
||
Bipinnate branchiae from chaetiger 3, continued throughout body. First pair of bipinnate branchiae small (
|
||
<figureCitation id="13553D83FFC4D72DFA887A9823CEFE3A" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="50" pageNumber="51">Fig. 21C</figureCitation>
|
||
, asterisks); cirriform branchiae present in the same segment. Bipinnate branchiae directed posteriorly, more or less convergent along posterior chaetigers; branchiae progressively larger to chaetiger 10, and then slightly larger posteriorly, becoming longer than segmental length. Bipinnate branchiae with up to 7–8 lateral branches in median segments.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF20794C22D1FD1A" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">Parapodia biramous, notopodia with cirriform branchiae along chaetigers 1–3, about as long as dorsal cirri in chaetiger 1, becoming half as long in chaetiger 3. Dorsal cirri slightly longer than bipinnate branchiae along median chaetigers, becoming up to 2× longer in posterior chaetigers. Second ventral cirri with cirrophores and cirrostyles slightly longer than those present in adjacent chaetigers, directed dorsally. Other ventral cirri directed ventrolaterally, as long as one segment.</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF2079F021E2FC4E" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">
|
||
Chaetae variably broken, distally covered by a mass of epizoic filamentous orgnisms. Complete chaetae with distal fragile hoods, usually eroded. Notochaetae in anterior chaetigers furcate, major tines 5–6× longer than minor ones (
|
||
<figureCitation id="13553D83FFC4D72DFF3E78482238FCB6" box="[217,318,835,861]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="50" pageNumber="51">Fig. 21E</figureCitation>
|
||
). Median chaetigers with
|
||
<specimenCount id="9D68EA8FFFC4D72DFDBC784821ACFCB6" box="[603,682,835,861]" pageId="50" pageNumber="51" type="generic" typeStatus="types">2 types</specimenCount>
|
||
of notochaetae: furcates with reduced minor tines, major ones 5–6× longer than shortest ones, and thicker harpoon-chaetae (
|
||
<figureCitation id="13553D83FFC4D72DFCE9786C2074FC6A" box="[782,882,871,897]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="50" pageNumber="51">Fig. 21F</figureCitation>
|
||
). Neurochaetae all furcates, in median chaetigers major tines 3–4× longer than minor ones (
|
||
<figureCitation id="13553D83FFC4D72DFD89788021D1FC4E" box="[622,727,907,933]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="50" pageNumber="51">Fig. 21G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF2078A42231FC06" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">
|
||
Posterior end tapered (
|
||
<figureCitation id="13553D83FFC4D72DFE2378A4212BFC22" box="[452,557,943,969]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="50" pageNumber="51">Fig. 21H</figureCitation>
|
||
); pygidium with anus terminal; anal cirri pale, subcylindrical, blunt, 7–8× longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF2078FD27B3FBFA" blockId="50.[151,1437,151,1977]" box="[199,1205,1014,1041]" pageId="50" pageNumber="51">
|
||
<emphasis id="B91AFD14FFC4D72DFF2078FD22A4FBFB" bold="true" box="[199,418,1014,1040]" pageId="50" pageNumber="51">Live pigmentation</emphasis>
|
||
. Unknown; dorsal cirri originally described as turned into intense red.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFC4D72DFF207F1021EFF87E" pageId="50" pageNumber="51" type="discussion">
|
||
<paragraph id="8BD12106FFC4D72DFF207F102167FB4A" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">
|
||
<emphasis id="B91AFD14FFC4D72DFF207F102232FBDE" bold="true" box="[199,308,1051,1077]" pageId="50" pageNumber="51">Remarks</emphasis>
|
||
. The specific epithet (Latin
|
||
<emphasis id="B91AFD14FFC4D72DFD957F102016FBDF" box="[626,784,1051,1077]" italics="true" pageId="50" pageNumber="51">fucatus, a, um</emphasis>
|
||
means to stain in red; French fardée, made-up) was given because “the only specimen had all body appendages with their purple pigmentation turned into intense red” (
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFA677F34229EFB96" author="De Quatrefages, A." pageId="50" pageNumber="51" pagination="578 - 600" refId="ref105083" refString="De Quatrefages, A. (1866 (1865 )) Histoire naturelle des Anneles marins et d'eau douce. Annelides et Gephyriens. Vol. 1. Librarie Encyclopedique de Roret, Paris, 588 pp. [publication date after Wright, E. P. (1866). Section 6 (Annelida, etc.). Zoological Record, 1866, 578 - 600] https: // doi. org / 10.5962 / bhl. title. 122818" type="book chapter" year="1866">de Quatrefages 1866: 390</bibRefCitation>
|
||
). In the Latin diagnosis, he indicated branchiae were medium-sized, and that the
|
||
<typeStatus id="54D59FA4FFC4D72DFAFF7F68264EFB96" box="[1304,1352,1123,1149]" pageId="50" pageNumber="51">type</typeStatus>
|
||
was 4o mm long,
|
||
<quantity id="4C968CE3FFC4D72DFEED7F8C224DFB4A" box="[266,331,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="50" pageNumber="51" unit="mm" value="9.0">9 mm</quantity>
|
||
wide, with 27 segments.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF207FA027BBFA76" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">
|
||
<taxonomicName id="4C6E5A85FFC4D72DFF207FA02169FB2E" authority="de Quatrefages, 1866" authorityName="de Quatrefages" authorityYear="1866" box="[199,623,1195,1221]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="fucata">
|
||
<emphasis id="B91AFD14FFC4D72DFF207FA02276FB2E" box="[199,368,1195,1221]" italics="true" pageId="50" pageNumber="51">Chloeia fucata</emphasis>
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFE9D7FA02169FB2E" author="De Quatrefages, A." box="[378,623,1195,1221]" pageId="50" pageNumber="51" pagination="578 - 600" refId="ref105083" refString="De Quatrefages, A. (1866 (1865 )) Histoire naturelle des Anneles marins et d'eau douce. Annelides et Gephyriens. Vol. 1. Librarie Encyclopedique de Roret, Paris, 588 pp. [publication date after Wright, E. P. (1866). Section 6 (Annelida, etc.). Zoological Record, 1866, 578 - 600] https: // doi. org / 10.5962 / bhl. title. 122818" type="book chapter" year="1866">de Quatrefages, 1866</bibRefCitation>
|
||
</taxonomicName>
|
||
is unique by having bipinnate branchiae from chaetiger 3, and by the presence of a discontinuous middorsal longitudinal band, slightly displaced towards the segmental posterior half, resembling middorsal oval spots. This pigmentation pattern is also present in
|
||
<taxonomicName id="4C6E5A85FFC4D72DFBE47FF8261BFAE6" authority="Baird, 1868" authorityName="Baird" authorityYear="1868" box="[1027,1309,1267,1293]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="pulchella">
|
||
<emphasis id="B91AFD14FFC4D72DFBE47FF8278BFAE6" box="[1027,1165,1267,1293]" italics="true" pageId="50" pageNumber="51">C. pulchella</emphasis>
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFB727FF8261BFAE6" author="Baird, W." box="[1173,1309,1267,1293]" pageId="50" pageNumber="51" pagination="215 - 250" refId="ref103011" refString="Baird, W. (1868) Contributions towards a monograph of the species of annelides belonging to the Amphinomacea, with a list of the known species, and a description of several new species (belonging to the group) contained in the National Collection of the British Museum. To which is appended a short account of two hitherto nondescript annulose animals of a larval character. The Journal of the Linnean Society of London, Zoology, 10 (44), 215 - 250, pls. 4 - 6. https: // doi. org / 10.1111 / j. 1096 - 3642.1868. tb 02233. x" type="journal article" year="1868">Baird, 1868</bibRefCitation>
|
||
</taxonomicName>
|
||
, reinstated, originally described from northeastern
|
||
<collectingCountry id="F3796196FFC4D72DFDB37E1C21B8FADA" box="[596,702,1303,1329]" name="Australia" pageId="50" pageNumber="51">Australia</collectingCountry>
|
||
, and its branchiae are rather short (after
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFB727E1C2613FADA" author="Horst, R." box="[1173,1301,1303,1329]" pageId="50" pageNumber="51" refId="ref108337" refString="Horst, R. (1912) Polychaeta errantia of the Siboga Expedition. Part 1. Amphinomidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 a, 1 - 43, 10 pls." type="journal volume" year="1912">Horst 1912</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFAB77E1C23C9FABE" author="Fauvel, P." pageId="50" pageNumber="51" refId="ref106154" refString="Fauvel, P. (1953) The Fauna of India, including Pakistan, Ceylon, Burma and Malaya. Annelida Polychaeta. Indian Press, Allahbad, 507 pp." type="book" year="1953">Fauvel 1953</bibRefCitation>
|
||
illustrations), not as long as one complete segment length, as in
|
||
<emphasis id="B91AFD14FFC4D72DFC517E302723FABE" box="[950,1061,1339,1365]" italics="true" pageId="50" pageNumber="51">
|
||
<taxonomicName id="4C6E5A85FFC4D72DFC517E302727FABE" authorityName="de Quatrefages" authorityYear="1866" box="[950,1057,1339,1365]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="fucata">C. fucata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
although this might result of the contraction of the body, more muscular than branchial stems. Thus, heavily contracted specimens might have apparently longer branchiae, in comparison to segment length, than those specimens less contracted.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF207EAC266AF972" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFF207EAC22A5FA2A" author="Fauvel, P." box="[199,419,1447,1473]" pageId="50" pageNumber="51" pagination="1 - 488" refId="ref105872" refString="Fauvel, P. (1923) Polychetes errantes. Faune de France, 5, 1 - 488." type="journal article" year="1923">Fauvel (1923: 134)</bibRefCitation>
|
||
, and later
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFDFA7EAC200BFA2A" author="Hartman, O." box="[541,781,1446,1473]" pageId="50" pageNumber="51" pagination="1 - 353" refId="ref107655" refString="Hartman, O. (1959) Catalogue of the polychaetous annelids of the World. Part 1. Allan Hancock Foundation Publications, Occasional Paper, 23, 1 - 353." type="journal article" year="1959">Hartman (1959: 131)</bibRefCitation>
|
||
regarded
|
||
<taxonomicName id="4C6E5A85FFC4D72DFC677EAC20EEFA2A" authorityName="de Quatrefages" authorityYear="1866" box="[896,1000,1447,1473]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="fucata">
|
||
<emphasis id="B91AFD14FFC4D72DFC677EAC20EEFA2A" box="[896,1000,1447,1473]" italics="true" pageId="50" pageNumber="51">C. fucata</emphasis>
|
||
</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName id="4C6E5A85FFC4D72DFAE67EAC226FFA0E" authority="de Quatrefages, 1866" authorityName="de Quatrefages" authorityYear="1866" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="venusta">
|
||
<emphasis id="B91AFD14FFC4D72DFAE67EAC267CFA2B" box="[1281,1402,1447,1473]" italics="true" pageId="50" pageNumber="51">C. venusta</emphasis>
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFA657EAC226FFA0E" author="De Quatrefages, A." pageId="50" pageNumber="51" pagination="578 - 600" refId="ref105083" refString="De Quatrefages, A. (1866 (1865 )) Histoire naturelle des Anneles marins et d'eau douce. Annelides et Gephyriens. Vol. 1. Librarie Encyclopedique de Roret, Paris, 588 pp. [publication date after Wright, E. P. (1866). Section 6 (Annelida, etc.). Zoological Record, 1866, 578 - 600] https: // doi. org / 10.5962 / bhl. title. 122818" type="book chapter" year="1866">de Quatrefages, 1866</bibRefCitation>
|
||
</taxonomicName>
|
||
, described from the Mediterranean Sea, probably after the confusion by
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFB477EC0269EFA0E" author="M'Intosh, W. C." box="[1184,1432,1482,1509]" pageId="50" pageNumber="51" pagination="395 - 416" refId="ref109853" refString="M'Intosh, W. C. (1876) On the Annelida of the ' Porcupine' expeditions of 1869 and 1870. Transactions of the Zoological Society of London, 9, 395 - 416, pls. 71 - 73." type="journal article" year="1876">M’Intosh (1876: 395)</bibRefCitation>
|
||
. However, M’Intosh hesitated about the species name and added a question mark after the name; his records include specimens collected in sediments from
|
||
<quantity id="4C968CE3FFC4D72DFDB47D1821C3F9C6" box="[595,709,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="1.555" metricValueMax="2.3" metricValueMin="0.8099999999999999" pageId="50" pageNumber="51" unit="m" value="155.5" valueMax="230.0" valueMin="81.0">81–230 m</quantity>
|
||
water depth in the Mediterranean Sea. The pigmentation pattern M’Intosh reported was a continuous brown middorsal band, and a similar band along the anterior part of each segment, thus rather approaching
|
||
<taxonomicName id="4C6E5A85FFC4D72DFE107D502176F99F" authorityName="de Quatrefages" authorityYear="1866" box="[503,624,1627,1653]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="venusta">
|
||
<emphasis id="B91AFD14FFC4D72DFE107D502176F99F" box="[503,624,1627,1653]" italics="true" pageId="50" pageNumber="51">C. venusta</emphasis>
|
||
</taxonomicName>
|
||
instead. This might hardly be regarded as conspecific with the Omani species, as the comparison of the specimens has shown, and hence they are herein regarded as distinct species.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFC4D72DFF207DA821EFF87E" blockId="50.[151,1437,151,1977]" pageId="50" pageNumber="51">
|
||
Although uncommon, cirriform branchiae might be present in the first segment where bipinnate branchiae start (
|
||
<bibRefCitation id="EFFF5CF7FFC4D72DFF307DCC218FF90A" author="Yanez-Rivera, B. & Salazar-Vallejo, S. I." box="[215,649,1735,1761]" pageId="50" pageNumber="51" pagination="503 - 537" refId="ref114234" refString="Yanez-Rivera, B. & Salazar-Vallejo, S. I. (2022) Revision of Chloeia Savigny in Lamarck, 1818 from tropical American seas (Annelida. Amphinomidae). Zootaxa, 5128 (4), 503 - 537. https: // doi. org / 10.11646 / zootaxa. 5128.4.3" type="journal article" year="2022">Yáñez-Rivera & Salazar-Vallejo 2022</bibRefCitation>
|
||
). The
|
||
<typeStatus id="54D59FA4FFC4D72DFD287DCC2037F90A" box="[719,817,1735,1761]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6E5A85FFC4D72DFCBE7DCC20C6F90A" authorityName="de Quatrefages" authorityYear="1866" box="[857,960,1735,1761]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="fucata">
|
||
<emphasis id="B91AFD14FFC4D72DFCBE7DCC20C6F90A" box="[857,960,1735,1761]" italics="true" pageId="50" pageNumber="51">C. fucata</emphasis>
|
||
</taxonomicName>
|
||
has bipinnate branchiae from chaetiger 3, together with cirriform branchiae. Solving the enigma about this early start whether this is an ontogenetic variant in a species having bipinnate branchiae from chaetiger 4, must wait for additional specimens. An ongoing project in
|
||
<collectingCountry id="F3796196FFC4D72DFF707C3823DDF8A6" box="[151,219,1843,1869]" name="Oman" pageId="50" pageNumber="51">Oman</collectingCountry>
|
||
has not found specimens of
|
||
<taxonomicName id="4C6E5A85FFC4D72DFDFB7C382173F8A6" authorityName="Savigny" authorityYear="1818" box="[540,629,1843,1869]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="genus">
|
||
<emphasis id="B91AFD14FFC4D72DFDFB7C382173F8A6" box="[540,629,1843,1869]" italics="true" pageId="50" pageNumber="51">Chloeia</emphasis>
|
||
</taxonomicName>
|
||
so far (G. Paulay, 2022, pers. comm.). If additional specimens show the
|
||
<typeStatus id="54D59FA4FFC4D72DFF707C5C23FFF89A" box="[151,249,1879,1905]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6E5A85FFC4D72DFEC77C5C228EF89A" authorityName="de Quatrefages" authorityYear="1866" box="[288,392,1879,1905]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="fucata">
|
||
<emphasis id="B91AFD14FFC4D72DFEC77C5C228EF89A" box="[288,392,1879,1905]" italics="true" pageId="50" pageNumber="51">C. fucata</emphasis>
|
||
</taxonomicName>
|
||
differs from typical specimens by having supernumerary bipinnate branchiae in chaetiger 3, then
|
||
<taxonomicName id="4C6E5A85FFC4D72DFF297C702233F87E" authorityName="de Quatrefages" authorityYear="1866" box="[206,309,1915,1941]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="fucata">
|
||
<emphasis id="B91AFD14FFC4D72DFF297C702233F87E" box="[206,309,1915,1941]" italics="true" pageId="50" pageNumber="51">C. fucata</emphasis>
|
||
</taxonomicName>
|
||
would have priority over
|
||
<emphasis id="B91AFD14FFC4D72DFDBE7C7021EFF87E" box="[601,745,1915,1941]" italics="true" pageId="50" pageNumber="51">
|
||
<taxonomicName id="4C6E5A85FFC4D72DFDBE7C7021E3F87E" authorityName="Baird" authorityYear="1868" box="[601,741,1915,1941]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="50" pageNumber="51" phylum="Annelida" rank="species" species="pulchella">C. pulchella</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFC4D72DFF207C952736F852" box="[199,1072,1950,1977]" pageId="50" pageNumber="51" type="distribution">
|
||
<paragraph id="8BD12106FFC4D72DFF207C952736F852" blockId="50.[151,1437,151,1977]" box="[199,1072,1950,1977]" pageId="50" pageNumber="51">
|
||
<emphasis id="B91AFD14FFC4D72DFF207C95225FF853" bold="true" box="[199,345,1950,1976]" pageId="50" pageNumber="51">Distribution</emphasis>
|
||
. Only known after the original description, from
|
||
<collectingCountry id="F3796196FFC4D72DFC647C9420DEF852" box="[899,984,1951,1977]" name="Oman" pageId="50" pageNumber="51">Muscat</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3796196FFC4D72DFC037C94272AF852" box="[996,1068,1951,1977]" name="Oman" pageId="50" pageNumber="51">Oman</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |