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<document id="D73F3639CCF29AE34CB5E25677D9033C" ID-DOI="10.11646/zootaxa.3986.4.5" ID-GBIF-Dataset="b4a2993f-2308-4fdb-be95-a0a364f6a17e" ID-ISSN="1175-5326" ID-Zenodo-Dep="246080" ID-ZooBank="F0EF76A2-9584-4E3E-A94E-23B58365F313" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461264482319" checkinUser="plazi" docAuthor="Silvestro, Violeta A., Giraldo Mendoza, Alfredo E. &amp; Flores, Gustavo E." docDate="2015" docId="03C787C4131FFFB3D3AB8E37FEF5F945" docLanguage="en" docName="zt03986p471.pdf" docOrigin="Zootaxa 3986 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pumiliofossorum Silvestro &amp; Giraldo, gen. nov." docType="treatment" docVersion="7" lastPageNumber="464" masterDocId="FFFEFFBC131EFFB0D33C8A68FFD3FFBF" masterDocTitle="Pumiliofossorum: A new genus of Scotobiini (Coleoptera: Tenebrionidae) with two new species from Peru, and a revised key for the genera of the tribe" masterLastPageNumber="471" masterPageNumber="461" pageNumber="462" updateTime="1698610169863" updateUser="plazi">
<mods:mods id="2B20DBDE902FEC17D26FFC85A605CE89" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="97351098B4B7C3E27A001C6AD27AE7DA">Pumiliofossorum: A new genus of Scotobiini (Coleoptera: Tenebrionidae) with two new species from Peru, and a revised key for the genera of the tribe</mods:title>
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<mods:name id="99E3617197C07DF1BD01C994FA75824D" type="personal">
<mods:role id="21301610826A3ECBF8C86AC0B50A6D83">
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<mods:namePart id="F271F33038317B34C7F7504EDB05EC32">Silvestro, Violeta A.</mods:namePart>
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<mods:namePart id="B765B4B14BA81B90F20D038C303B49DC">Giraldo Mendoza, Alfredo E.</mods:namePart>
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<mods:namePart id="ACA88277C342B4C33E76C86C1935DBD3">Flores, Gustavo E.</mods:namePart>
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<mods:title id="296E899381B3E554C1CB82440ACAC6AF">Zootaxa</mods:title>
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<mods:part id="D931D16643B2C2CA1956D4C3BF331FC9">
<mods:date id="50CAC639408803114DFAEBC180C087A2">2015</mods:date>
<mods:detail id="1EDCCA9E3A2BDC35AC39AD829233187B" type="volume">
<mods:number id="8375FF74B12FB8F48197163736318C35">3986</mods:number>
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<mods:start id="B9EF23810167A7F7A3E0A4FE6C944FE7">461</mods:start>
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<mods:classification id="048FD9C6B32209A815692E2156FF7362">journal article</mods:classification>
<mods:identifier id="B8EF242ADEB066F9C531FF015A66D15F" type="DOI">10.11646/zootaxa.3986.4.5</mods:identifier>
<mods:identifier id="DD6CB9C06A8437BD2BB1CDA2D7B0CC99" type="GBIF-Dataset">b4a2993f-2308-4fdb-be95-a0a364f6a17e</mods:identifier>
<mods:identifier id="8D23CC6CF662E52EC2F09E7C857B4AF1" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C787C4131FFFB3D3AB8E37FEF5F945" ID-DOI="http://doi.org/10.5281/zenodo.6121375" ID-GBIF-Taxon="119637594" ID-Zenodo-Dep="6121375" LSID="urn:lsid:plazi:treatment:03C787C4131FFFB3D3AB8E37FEF5F945" httpUri="http://treatment.plazi.org/id/03C787C4131FFFB3D3AB8E37FEF5F945" lastPageId="3" lastPageNumber="464" pageId="1" pageNumber="462">
<subSubSection id="C3746559131FFFB1D3AB8E37FEF3FB24" pageId="1" pageNumber="462" type="nomenclature">
<paragraph id="8BD136D2131FFFB1D3AB8E37FD33FBC5" blockId="1.[151,736,1119,1179]" box="[151,736,1119,1146]" pageId="1" pageNumber="462">
<heading id="D09981BE131FFFB1D3AB8E37FD33FBC5" bold="true" box="[151,736,1119,1146]" fontSize="11" level="1" pageId="1" pageNumber="462" reason="1">
<emphasis id="B91AEAC0131FFFB1D3AB8E37FD33FBC5" bold="true" box="[151,736,1119,1146]" pageId="1" pageNumber="462">
<taxonomicName id="4C6E4D51131FFFB1D3AB8E37FDB6FBC5" authority="Silvestro &amp; Giraldo" authorityName="Silvestro &amp; Giraldo" box="[151,613,1119,1146]" class="Insecta" family="Tenebrionidae" genus="Pumiliofossorum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="462" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91AEAC0131FFFB1D3AB8E37FEB4FBC6" bold="true" box="[151,359,1119,1145]" italics="true" pageId="1" pageNumber="462">Pumiliofossorum</emphasis>
Silvestro &amp; Giraldo
</taxonomicName>
,
<taxonomicNameLabel id="A22957BB131FFFB1D14F8E08FD33FBC5" box="[627,736,1120,1146]" pageId="1" pageNumber="462" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136D2131FFFB1D3AB8EEAFEF3FB24" blockId="1.[151,736,1119,1179]" box="[151,288,1154,1179]" pageId="1" pageNumber="462">
(
<figureCitation id="13552A57131FFFB1D3A38EEAFEC4FB24" box="[159,279,1154,1179]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 7" captionStart-2="FIGURES 8 – 15" captionStart-3="FIGURES 16 – 19" captionStart-4="FIGURES 20 – 21" captionStartId-0="2.[151,264,1892,1915]" captionStartId-1="3.[151,264,1411,1434]" captionStartId-2="4.[151,264,1984,2007]" captionStartId-3="6.[151,264,1879,1902]" captionStartId-4="7.[151,264,1136,1159]" captionTargetBox-0="[175,1431,916,1868]" captionTargetBox-1="[154,1436,193,1389]" captionTargetBox-2="[156,1436,337,1923]" captionTargetBox-3="[190,1405,608,1836]" captionTargetBox-4="[157,1436,624,1109]" captionTargetId-0="figure@2.[155,1432,916,1869]" captionTargetId-1="figure@3.[154,1439,193,1390]" captionTargetId-2="figure@4.[156,1441,337,1932]" captionTargetId-3="figure@6.[169,1417,589,1859]" captionTargetId-4="figure@7.[157,1442,624,1109]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." captionText-1="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." captionText-2="FIGURES 8 – 15. Pumiliofossorum moche sp. nov. Protibia, dorsal and ventral view (8, 9), mesotibia, dorsal and ventral view (10, 11), metatibia, dorsal and ventral view (12, 13), last tarsomere (14) and meso- and metacoxae (15)." captionText-3="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." captionText-4="FIGURES 20 – 21. Sechura desert, habitat of Pumiliofossorum sechurae sp. nov. Sand dune (20), and pitfall trap (21)." httpUri-0="https://zenodo.org/record/246081/files/figure.png" httpUri-1="https://zenodo.org/record/246082/files/figure.png" httpUri-2="https://zenodo.org/record/246083/files/figure.png" httpUri-3="https://zenodo.org/record/246084/files/figure.png" httpUri-4="https://zenodo.org/record/246085/files/figure.png" pageId="1" pageNumber="462">Figs. 1–21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3746559131FFFB1D3AB8EA2FBA0FB5C" box="[151,1139,1226,1251]" pageId="1" pageNumber="462" type="materials_examined">
<paragraph id="8BD136D2131FFFB1D3AB8EA2FBA0FB5C" blockId="1.[151,1437,1226,2007]" box="[151,1139,1226,1251]" pageId="1" pageNumber="462">
<emphasis id="B91AEAC0131FFFB1D3AB8EA2FEE1FB5C" bold="true" box="[151,306,1226,1251]" pageId="1" pageNumber="462">
<typeStatus id="54D58870131FFFB1D3AB8EA2FF01FB5C" box="[151,210,1226,1251]" pageId="1" pageNumber="462">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6E4D51131FFFB1D2058EA3FD60FB5C" authority="Silvestro" authorityName="Silvestro" box="[313,691,1226,1251]" class="Insecta" family="Tenebrionidae" genus="Pumiliofossorum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="462" phylum="Arthropoda" rank="species" species="moche">
<emphasis id="B91AEAC0131FFFB1D2058EA3FD9BFB5C" box="[313,584,1227,1251]" italics="true" pageId="1" pageNumber="462">Pumiliofossorum moche</emphasis>
Silvestro
</taxonomicName>
&amp;
<collectingCountry id="F3797642131FFFB1D1EA8EA2FCCCFB5C" box="[726,799,1226,1251]" name="Indonesia" pageId="1" pageNumber="462">Flores</collectingCountry>
,
<emphasis id="B91AEAC0131FFFB1D0178EA2FC57FB5C" bold="true" box="[811,900,1226,1251]" pageId="1" pageNumber="462">sp. nov.</emphasis>
, present designation.
</paragraph>
</subSubSection>
<subSubSection id="C3746559131FFFB1D3FB8E86FF30FAF1" pageId="1" pageNumber="462" type="etymology">
<paragraph id="8BD136D2131FFFB1D3FB8E86FF30FAF1" blockId="1.[151,1437,1226,2007]" pageId="1" pageNumber="462">
<emphasis id="B91AEAC0131FFFB1D3FB8E86FE9DFAB8" bold="true" box="[199,334,1262,1287]" pageId="1" pageNumber="462">Etymology.</emphasis>
The generic epithet, which means “dwarf digger”, refers to the relatively small sizes of the two known species (the smallest ones currently known in Scotobiini) and the presence of fossorial tibiae. Gender neuter.
</paragraph>
</subSubSection>
<subSubSection id="C3746559131FFFB1D3FB8F32FC39F9F4" pageId="1" pageNumber="462" type="diagnosis">
<paragraph id="8BD136D2131FFFB1D3FB8F32FC39F9F4" blockId="1.[151,1437,1226,2007]" pageId="1" pageNumber="462">
<emphasis id="B91AEAC0131FFFB1D3FB8F32FE92FACC" bold="true" box="[199,321,1370,1395]" pageId="1" pageNumber="462">Diagnosis.</emphasis>
Within the tribe Scotobiini,
<emphasis id="B91AEAC0131FFFB1D1B98F33FC97FACC" box="[645,836,1371,1395]" italics="true" pageId="1" pageNumber="462">Pumiliofossorum</emphasis>
can be easily distinguished from other genera by the following combination of characters: antennae short, surpassing anterior margin of pronotum, capitate, antennomeres II–XI wider than long, the four last antennomeres bigger than the preceding ones; last antennomere truncated, slightly concave (
<figureCitation id="13552A57131FFFB1D2D98FAEFD88FA60" box="[485,603,1478,1503]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="1" pageNumber="462">Figs. 6, 7</figureCitation>
); pronotum tranverse, oval shape, margins not carinate, lateral margin rounded, slopping down, posterior angles not defined (
<figureCitation id="13552A57131FFFB1D1D28F82FCEEF9BC" box="[750,829,1514,1539]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="1" pageNumber="462">Figs. 2</figureCitation>
,
<figureCitation id="13552A57131FFFB1D0758F82FCBBF9BC" box="[841,872,1514,1539]" captionStart="FIGURES 16 – 19" captionStartId="6.[151,264,1879,1902]" captionTargetBox="[190,1405,608,1836]" captionTargetId="figure@6.[169,1417,589,1859]" captionTargetPageId="6" captionText="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." httpUri="https://zenodo.org/record/246084/files/figure.png" pageId="1" pageNumber="462">16</figureCitation>
); elytra acuminate, widest behind humeri (
<figureCitation id="13552A57131FFFB1D6748F82FA46F9BC" box="[1352,1429,1514,1539]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="1" pageNumber="462">Figs. 1</figureCitation>
,
<figureCitation id="13552A57131FFFB1D3AB8C66FF65F998" box="[151,182,1550,1575]" captionStart="FIGURES 16 – 19" captionStartId="6.[151,264,1879,1902]" captionTargetBox="[190,1405,608,1836]" captionTargetId="figure@6.[169,1417,589,1859]" captionTargetPageId="6" captionText="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." httpUri="https://zenodo.org/record/246084/files/figure.png" pageId="1" pageNumber="462">16</figureCitation>
); tibiae laterally compressed with outer margin expanded, convex and inner margin with long golden setae (
<figureCitation id="13552A57131FFFB1D3A38C5AFECAF9F4" box="[159,281,1586,1611]" captionStart="FIGURES 8 – 15" captionStartId="4.[151,264,1984,2007]" captionTargetBox="[156,1436,337,1923]" captionTargetId="figure@4.[156,1441,337,1932]" captionTargetPageId="4" captionText="FIGURES 8 – 15. Pumiliofossorum moche sp. nov. Protibia, dorsal and ventral view (8, 9), mesotibia, dorsal and ventral view (10, 11), metatibia, dorsal and ventral view (12, 13), last tarsomere (14) and meso- and metacoxae (15)." httpUri="https://zenodo.org/record/246083/files/figure.png" pageId="1" pageNumber="462">Figs. 8–13</figureCitation>
); protibiae flattened, markedly expanded outwards (
<figureCitation id="13552A57131FFFB1D05E8C5AFC63F9F4" box="[866,944,1586,1611]" captionStart="FIGURES 8 – 15" captionStartId="4.[151,264,1984,2007]" captionTargetBox="[156,1436,337,1923]" captionTargetId="figure@4.[156,1441,337,1932]" captionTargetPageId="4" captionText="FIGURES 8 – 15. Pumiliofossorum moche sp. nov. Protibia, dorsal and ventral view (8, 9), mesotibia, dorsal and ventral view (10, 11), metatibia, dorsal and ventral view (12, 13), last tarsomere (14) and meso- and metacoxae (15)." httpUri="https://zenodo.org/record/246083/files/figure.png" pageId="1" pageNumber="462">Figs. 8</figureCitation>
,
<figureCitation id="13552A57131FFFB1D0818C5BFC0FF9F4" box="[957,988,1587,1611]" captionStart="FIGURES 16 – 19" captionStartId="6.[151,264,1879,1902]" captionTargetBox="[190,1405,608,1836]" captionTargetId="figure@6.[169,1417,589,1859]" captionTargetPageId="6" captionText="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." httpUri="https://zenodo.org/record/246084/files/figure.png" pageId="1" pageNumber="462">18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3746559131FFFB3D3FB8C3EFF1BF9D5" lastPageId="3" lastPageNumber="464" pageId="1" pageNumber="462" type="description">
<paragraph id="8BD136D2131FFFB2D3FB8C3EFBFEFCE2" blockId="1.[151,1437,1226,2007]" lastBlockId="2.[151,1437,151,861]" lastPageId="2" lastPageNumber="463" pageId="1" pageNumber="462">
<emphasis id="B91AEAC0131FFFB1D3FB8C3EFE8AF9D0" bold="true" box="[199,345,1622,1647]" pageId="1" pageNumber="462">Description.</emphasis>
Length 4.0–6.0 mm, width 2.5 –3.0 mm. Apterous. Body outline oval, base of elytra constricted. Color reddish brown to dark brown, shiny; antennae and legs reddish brown (
<figureCitation id="13552A57131FFFB1D7228C12FBBDF92C" box="[1054,1134,1658,1683]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="1" pageNumber="462">Figs. 1</figureCitation>
,
<figureCitation id="13552A57131FFFB1D7438C13FB4DF92C" box="[1151,1182,1659,1683]" captionStart="FIGURES 16 – 19" captionStartId="6.[151,264,1879,1902]" captionTargetBox="[190,1405,608,1836]" captionTargetId="figure@6.[169,1417,589,1859]" captionTargetPageId="6" captionText="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." httpUri="https://zenodo.org/record/246084/files/figure.png" pageId="1" pageNumber="462">19</figureCitation>
).
<emphasis id="B91AEAC0131FFFB1D7898C12FB2FF92C" bold="true" box="[1205,1276,1658,1683]" pageId="1" pageNumber="462">Head.</emphasis>
Labrum light brown, visible, with round punctures and golden setae, anterior margin bisinuate with thick short golden setae (
<figureCitation id="13552A57131FFFB1D64C8CF6FF74F964" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="1" pageNumber="462">Fig. 4</figureCitation>
). Clypeus broad, width of anterior margin exceeding width of interocular distance, anterior margin concave (
<figureCitation id="13552A57131FFFB1D6708CAAFA5CF964" box="[1356,1423,1730,1755]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="1" pageNumber="462">Fig. 4</figureCitation>
). Clypeus and frons with strong and dense punctures, clypeal suture transverse, weakly depressed (
<figureCitation id="13552A57131FFFB1D7DF8C8EFA9FF940" box="[1251,1356,1766,1791]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="1" pageNumber="462">Figs. 4, 5</figureCitation>
). Head widest at epicanthus, before eyes. Epicanthus prominent, almost dividing the eyes into lower and upper portions (
<figureCitation id="13552A57131FFFB1D3A38D46FEDBF8F8" box="[159,264,1838,1863]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="1" pageNumber="462">Figs. 4, 5</figureCitation>
). Eyes narrow, sunken, groove-like in dorsal view, half-moon shaped in lateral view (
<figureCitation id="13552A57131FFFB1D78F8D46FAC8F8F8" box="[1203,1307,1838,1863]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="1" pageNumber="462">Figs. 4, 5</figureCitation>
). Antennae capitate, flattened dorsoventrally (
<figureCitation id="13552A57131FFFB1D1178D3AFD72F8D4" box="[555,673,1874,1899]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="1" pageNumber="462">Figs. 5, 6</figureCitation>
), short, surpassing anterior margin of pronotum. Antennomeres progressively increasing in width, II–XI wider than long, antennomeres IX and XI of same width, antennomere X wider than antennomeres IX and XI (
<figureCitation id="13552A57131FFFB1D1018DF2FD5CF80C" box="[573,655,1946,1971]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="1" pageNumber="462">Figs. 6</figureCitation>
,
<figureCitation id="13552A57131FFFB1D1A28DF2FD6EF80C" box="[670,701,1946,1971]" captionStart="FIGURES 16 – 19" captionStartId="6.[151,264,1879,1902]" captionTargetBox="[190,1405,608,1836]" captionTargetId="figure@6.[169,1417,589,1859]" captionTargetPageId="6" captionText="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." httpUri="https://zenodo.org/record/246084/files/figure.png" pageId="1" pageNumber="462">16</figureCitation>
); antennomeres VIII–X with dome-shaped placoid sensoriae on rims (
<figureCitation id="13552A57131FFFB1D3E48DD6FECEF868" box="[216,285,1982,2007]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="1" pageNumber="462">Fig. 6</figureCitation>
), antennomere XI apex truncate and slightly concave, dome-shaped placoid sensoriae on whole apical surface (
<figureCitation id="13552A57131CFFB2D3C48AFFFEEDFF0F" box="[248,318,151,176]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="2" pageNumber="463">Fig. 7</figureCitation>
). Mentum small, longer than wide, subtrapezoidal, subgenal process remote from submentum and mentum (
<figureCitation id="13552A57131CFFB2D2388AD4FE98FF6B" box="[260,331,188,213]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="2" pageNumber="463">Fig. 3</figureCitation>
).
<emphasis id="B91AEAC0131CFFB2D25E8AD4FE6CFF6A" bold="true" box="[354,447,188,213]" pageId="2" pageNumber="463">Thorax.</emphasis>
Pronotum transverse, 1.5 x wider than long, oval shaped, slightly domed, widest before midpoint, with round punctures (
<figureCitation id="13552A57131CFFB2D1368AB7FD81FF47" box="[522,594,223,248]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="2" pageNumber="463">Fig. 2</figureCitation>
). Margins not carinate, anterior margin slightly bisinuate, anterior angles rounded, lateral margin rounded, slopping towards venter, posterior angles not defined. Anterior and posterior margin with short golden setae projected outward (
<figureCitation id="13552A57131CFFB2D1D88B4FFCE5FEFF" box="[740,822,295,320]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="2" pageNumber="463">Figs. 2</figureCitation>
,
<figureCitation id="13552A57131CFFB2D0788B40FC86FEFF" box="[836,853,296,320]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1411,1434]" captionTargetBox="[154,1436,193,1389]" captionTargetId="figure@3.[154,1439,193,1390]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Pumiliofossorum moche sp. nov. Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7)." httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="2" pageNumber="463">4</figureCitation>
). Proepisternum not visible from above, with long golden setae (
<figureCitation id="13552A57131CFFB2D2008B24FE55FEDB" box="[316,390,332,357]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="2" pageNumber="463">Fig. 3</figureCitation>
). Hypomera, prosternum, mesosternum and metasternum with punctures less dense than pronotal punctures, clothed in long golden disperse setae. Intercoxal distances shorter than coxal diameters (
<figureCitation id="13552A57131CFFB2D6588B07FF74FE13" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="2" pageNumber="463">Figs. 3</figureCitation>
,
<figureCitation id="13552A57131CFFB2D38B8BFCFF05FE12" box="[183,214,404,429]" captionStart="FIGURES 8 – 15" captionStartId="4.[151,264,1984,2007]" captionTargetBox="[156,1436,337,1923]" captionTargetId="figure@4.[156,1441,337,1932]" captionTargetPageId="4" captionText="FIGURES 8 – 15. Pumiliofossorum moche sp. nov. Protibia, dorsal and ventral view (8, 9), mesotibia, dorsal and ventral view (10, 11), metatibia, dorsal and ventral view (12, 13), last tarsomere (14) and meso- and metacoxae (15)." httpUri="https://zenodo.org/record/246083/files/figure.png" pageId="2" pageNumber="463">15</figureCitation>
). Scutellum small, triangular, completely exposed (
<figureCitation id="13552A57131CFFB2D0118BFCFCADFE13" box="[813,894,404,429]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="2" pageNumber="463">Figs. 1</figureCitation>
,
<figureCitation id="13552A57131CFFB2D0AC8BFCFC7CFE12" box="[912,943,404,429]" captionStart="FIGURES 16 – 19" captionStartId="6.[151,264,1879,1902]" captionTargetBox="[190,1405,608,1836]" captionTargetId="figure@6.[169,1417,589,1859]" captionTargetPageId="6" captionText="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." httpUri="https://zenodo.org/record/246084/files/figure.png" pageId="2" pageNumber="463">16</figureCitation>
).
<emphasis id="B91AEAC0131CFFB2D0FB8BFCFBC8FE12" bold="true" box="[967,1051,404,429]" pageId="2" pageNumber="463">Elytral</emphasis>
length equal to 1.5–2.0 x of the pronotal length, acuminate, widest behind humeri, narrowing towards posterior apex, 1.4 x longer than wide; with four or five longitudinal grooves on internal half, with wrinkles, punctures, and sparse golden setae in lateral area; humeri curved, prominent (
<figureCitation id="13552A57131CFFB2D2F78B97FDCAFDA7" box="[459,537,511,536]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="2" pageNumber="463">Figs. 1</figureCitation>
,
<figureCitation id="13552A57131CFFB2D11B8B97FD95FDA7" box="[551,582,511,536]" captionStart="FIGURES 16 – 19" captionStartId="6.[151,264,1879,1902]" captionTargetBox="[190,1405,608,1836]" captionTargetId="figure@6.[169,1417,589,1859]" captionTargetPageId="6" captionText="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." httpUri="https://zenodo.org/record/246084/files/figure.png" pageId="2" pageNumber="463">16</figureCitation>
); suture flat; pseudopleuron with long golden setae; epipleuron not marked.
<emphasis id="B91AEAC0131CFFB2D3AB884CFEC0FD82" bold="true" box="[151,275,548,573]" pageId="2" pageNumber="463">Abdomen.</emphasis>
Ventrites with punctures separated by twice their diameter and sparse golden setae of moderate length.
<emphasis id="B91AEAC0131CFFB2D3AB882FFF05FDDF" bold="true" box="[151,214,583,608]" pageId="2" pageNumber="463">Legs.</emphasis>
Femora and tibiae with punctures separated by a distance twice or more the diameter of puncture, each puncture with a long, gold central seta. Tibiae laterally compressed, outer margin expanded, convex, inner margin with long golden setae (
<figureCitation id="13552A57131CFFB2D28988E7FDE6FD17" box="[437,565,655,680]" captionStart="FIGURES 8 – 15" captionStartId="4.[151,264,1984,2007]" captionTargetBox="[156,1436,337,1923]" captionTargetId="figure@4.[156,1441,337,1932]" captionTargetPageId="4" captionText="FIGURES 8 – 15. Pumiliofossorum moche sp. nov. Protibia, dorsal and ventral view (8, 9), mesotibia, dorsal and ventral view (10, 11), metatibia, dorsal and ventral view (12, 13), last tarsomere (14) and meso- and metacoxae (15)." httpUri="https://zenodo.org/record/246083/files/figure.png" pageId="2" pageNumber="463">Figs. 8–15</figureCitation>
). Protibiae median section flattened and expanded outwards, outer margin strongly convex, inner margin slightly concave (
<figureCitation id="13552A57131CFFB2D1FD88DCFCE3FD73" box="[705,816,692,717]" captionStart="FIGURES 8 – 15" captionStartId="4.[151,264,1984,2007]" captionTargetBox="[156,1436,337,1923]" captionTargetId="figure@4.[156,1441,337,1932]" captionTargetPageId="4" captionText="FIGURES 8 – 15. Pumiliofossorum moche sp. nov. Protibia, dorsal and ventral view (8, 9), mesotibia, dorsal and ventral view (10, 11), metatibia, dorsal and ventral view (12, 13), last tarsomere (14) and meso- and metacoxae (15)." httpUri="https://zenodo.org/record/246083/files/figure.png" pageId="2" pageNumber="463">Figs. 8, 9</figureCitation>
,
<figureCitation id="13552A57131CFFB2D00288DCFC8EFD73" box="[830,861,692,716]" captionStart="FIGURES 16 – 19" captionStartId="6.[151,264,1879,1902]" captionTargetBox="[190,1405,608,1836]" captionTargetId="figure@6.[169,1417,589,1859]" captionTargetPageId="6" captionText="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." httpUri="https://zenodo.org/record/246084/files/figure.png" pageId="2" pageNumber="463">18</figureCitation>
). Mesotibiae and metatibiae outer margins slightly convex, serrated, with row of contiguous spines, inner margin straight (
<figureCitation id="13552A57131CFFB2D0FB88BFFB87FD4F" box="[967,1108,727,752]" captionStart="FIGURES 8 – 15" captionStartId="4.[151,264,1984,2007]" captionTargetBox="[156,1436,337,1923]" captionTargetId="figure@4.[156,1441,337,1932]" captionTargetPageId="4" captionText="FIGURES 8 – 15. Pumiliofossorum moche sp. nov. Protibia, dorsal and ventral view (8, 9), mesotibia, dorsal and ventral view (10, 11), metatibia, dorsal and ventral view (12, 13), last tarsomere (14) and meso- and metacoxae (15)." httpUri="https://zenodo.org/record/246083/files/figure.png" pageId="2" pageNumber="463">Figs. 10–13</figureCitation>
). Mesotibiae median section triangular. Metatibiae median section flattened. Last tarsomere twice as long as each of the preceding ones (
<figureCitation id="13552A57131CFFB2D6748894FED8FC87" captionStart="FIGURES 8 – 15" captionStartId="4.[151,264,1984,2007]" captionTargetBox="[156,1436,337,1923]" captionTargetId="figure@4.[156,1441,337,1932]" captionTargetPageId="4" captionText="FIGURES 8 – 15. Pumiliofossorum moche sp. nov. Protibia, dorsal and ventral view (8, 9), mesotibia, dorsal and ventral view (10, 11), metatibia, dorsal and ventral view (12, 13), last tarsomere (14) and meso- and metacoxae (15)." httpUri="https://zenodo.org/record/246083/files/figure.png" pageId="2" pageNumber="463">Figs. 8, 10, 12, 14</figureCitation>
). Spurs of the protibiae short, approximately as long as the first tarsomere (forespurs of Mathews
<emphasis id="B91AEAC0131CFFB2D65F8949FA46FC87" box="[1379,1429,800,824]" italics="true" pageId="2" pageNumber="463">et al</emphasis>
. 2010: 582). Metafemoral-tibial articulation not reaching elytral apex (
<figureCitation id="13552A57131CFFB2D099892CFC21FCE3" box="[933,1010,836,861]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1892,1915]" captionTargetBox="[175,1431,916,1868]" captionTargetId="figure@2.[155,1432,916,1869]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Pumiliofossorum moche sp. nov. Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3)." httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="2" pageNumber="463">Figs. 1</figureCitation>
,
<figureCitation id="13552A57131CFFB2D73C892CFBCCFCE2" box="[1024,1055,836,861]" captionStart="FIGURES 16 – 19" captionStartId="6.[151,264,1879,1902]" captionTargetBox="[190,1405,608,1836]" captionTargetId="figure@6.[169,1417,589,1859]" captionTargetPageId="6" captionText="FIGURES 16 – 19. Pumiliofossorum sechurae sp. nov. Habitus of allotype in dorsal (16), lateral (17), frontal (18) and ventral view (19)." httpUri="https://zenodo.org/record/246084/files/figure.png" pageId="2" pageNumber="463">16</figureCitation>
).
</paragraph>
<caption id="DF11665A131CFFB2D3AB8D0CFAF4F8C5" box="[151,1319,1892,1915]" httpUri="https://zenodo.org/record/246081/files/figure.png" pageId="2" pageNumber="463" targetBox="[175,1431,916,1868]" targetPageId="2">
<paragraph id="8BD136D2131CFFB2D3AB8D0CFAF4F8C5" blockId="2.[151,1319,1892,1915]" box="[151,1319,1892,1915]" pageId="2" pageNumber="463">
<emphasis id="B91AEAC0131CFFB2D3AB8D0CFEE9F8C5" bold="true" box="[151,314,1892,1915]" pageId="2" pageNumber="463">FIGURES 1–3.</emphasis>
<taxonomicName id="4C6E4D51131CFFB2D27C8D0CFDE5F8C5" box="[320,566,1892,1914]" class="Insecta" family="Tenebrionidae" genus="Pumiliofossorum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="463" phylum="Arthropoda" rank="species" species="moche" status="sp. nov.">
<emphasis id="B91AEAC0131CFFB2D27C8D0CFDE5F8C5" box="[320,566,1892,1914]" italics="true" pageId="2" pageNumber="463">Pumiliofossorum moche</emphasis>
</taxonomicName>
<emphasis id="B91AEAC0131CFFB2D1008D0DFD5FF8C5" bold="true" box="[572,652,1893,1914]" pageId="2" pageNumber="463">
<taxonomicNameLabel id="A22957BB131CFFB2D1008D0DFD5FF8C5" box="[572,652,1893,1914]" pageId="2" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Habitus dorsal view (1), prothorax dorsal and ventral view (2, 3).
</paragraph>
</caption>
<caption id="DF11665A131DFFB3D3AB8FEBFE97FA07" httpUri="https://zenodo.org/record/246082/files/figure.png" pageId="3" pageNumber="464" targetBox="[154,1436,193,1389]" targetPageId="3">
<paragraph id="8BD136D2131DFFB3D3AB8FEBFE97FA07" blockId="3.[151,1435,1411,1464]" pageId="3" pageNumber="464">
<emphasis id="B91AEAC0131DFFB3D3AB8FEBFEECFA26" bold="true" box="[151,319,1411,1434]" pageId="3" pageNumber="464">FIGURES 4–7.</emphasis>
<taxonomicName id="4C6E4D51131DFFB3D2768FECFD97FA25" box="[330,580,1412,1434]" class="Insecta" family="Tenebrionidae" genus="Pumiliofossorum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="464" phylum="Arthropoda" rank="species" species="moche" status="sp. nov.">
<emphasis id="B91AEAC0131DFFB3D2768FECFD97FA25" box="[330,580,1412,1434]" italics="true" pageId="3" pageNumber="464">Pumiliofossorum moche</emphasis>
</taxonomicName>
<emphasis id="B91AEAC0131DFFB3D1738FECFD77FA26" bold="true" box="[591,676,1412,1433]" pageId="3" pageNumber="464">
<taxonomicNameLabel id="A22957BB131DFFB3D1738FECFD77FA26" box="[591,676,1412,1433]" pageId="3" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Head dorsal and latero-dorsal view (4, 5), antenna dorsal (6) and last antennomere (7).
</paragraph>
</caption>
<paragraph id="8BD136D2131DFFB3D3FB8F8CFC62F99D" blockId="3.[151,1436,1508,1786]" pageId="3" pageNumber="464">
<emphasis id="B91AEAC0131DFFB3D3FB8F8CFE79FA42" bold="true" box="[199,426,1508,1533]" pageId="3" pageNumber="464">Sexual dimorphism</emphasis>
: Female with abdominal sterna slightly convex; male with central concavity from first to third visible abdominal ventrites, more pronounced on second ventrite.
</paragraph>
<paragraph id="8BD136D2131DFFB3D3FB8C45FF1BF9D5" blockId="3.[151,1436,1508,1786]" pageId="3" pageNumber="464">
<emphasis id="B91AEAC0131DFFB3D3FB8C45FE40F9F9" bold="true" box="[199,403,1581,1606]" pageId="3" pageNumber="464">Species included.</emphasis>
<taxonomicName id="4C6E4D51131DFFB3D2A68C45FCC6F9F9" authority="Silvestro" authorityName="Silvestro" box="[410,789,1581,1606]" class="Insecta" family="Tenebrionidae" genus="Pumiliofossorum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="464" phylum="Arthropoda" rank="species" species="moche">
<emphasis id="B91AEAC0131DFFB3D2A68C45FD79F9FA" box="[410,682,1581,1605]" italics="true" pageId="3" pageNumber="464">Pumiliofossorum moche</emphasis>
Silvestro
</taxonomicName>
&amp;
<collectingCountry id="F3797642131DFFB3D0058C45FC51F9F9" box="[825,898,1581,1606]" name="Indonesia" pageId="3" pageNumber="464">Flores</collectingCountry>
,
<emphasis id="B91AEAC0131DFFB3D0B18C45FC35F9F9" bold="true" box="[909,998,1581,1606]" pageId="3" pageNumber="464">sp. nov.</emphasis>
, and
<taxonomicName id="4C6E4D51131DFFB3D7188C46FAD2F9F9" authority="Giraldo" authorityName="Giraldo" box="[1060,1281,1581,1606]" class="Insecta" family="Tenebrionidae" genus="Pumiliofossorum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="464" phylum="Arthropoda" rank="species" species="sechurae">
<emphasis id="B91AEAC0131DFFB3D7188C46FB76F9FA" box="[1060,1189,1581,1605]" italics="true" pageId="3" pageNumber="464">P. sechurae</emphasis>
Giraldo
</taxonomicName>
&amp;
<collectingCountry id="F3797642131DFFB3D61A8C45FABCF9F9" box="[1318,1391,1581,1606]" name="Indonesia" pageId="3" pageNumber="464">Flores</collectingCountry>
,
<emphasis id="B91AEAC0131DFFB3D6468C45FF1BF9D5" bold="true" pageId="3" pageNumber="464">sp. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3746559131DFFB3D3FB8C1DFEF5F945" pageId="3" pageNumber="464" type="distribution">
<paragraph id="8BD136D2131DFFB3D3FB8C1DFEF5F945" blockId="3.[151,1436,1508,1786]" pageId="3" pageNumber="464">
<emphasis id="B91AEAC0131DFFB3D3FB8C1DFE22F931" bold="true" box="[199,497,1653,1678]" pageId="3" pageNumber="464">Distribution and habitat.</emphasis>
Coastal desert of Northwestern
<collectingCountry id="F3797642131DFFB3D0678C1EFC5DF932" box="[859,910,1654,1677]" name="Peru" pageId="3" pageNumber="464">Peru</collectingCountry>
in sandy soils.
<emphasis id="B91AEAC0131DFFB3D7008C1DFB28F932" box="[1084,1275,1653,1677]" italics="true" pageId="3" pageNumber="464">Pumiliofossorum</emphasis>
species fit the morphological features of the sand swimmers (
<bibRefCitation id="EFFF4B23131DFFB3D19B8CF1FC70F90E" author="Matthews" box="[679,931,1689,1714]" pageId="3" pageNumber="464" refString="Matthews, E. G., Lawrence, J. F., Bouchard, P., Steiner, W. E. &amp; Slipinski, S. A. (2010) Tenebrionidae Latreille, 1802. In: Leschen, R. A. B., Beutel, R. G. &amp; Lawrence J. F. (Eds.), Handbook of Zoology. Coleoptera, Beetles. Vol. 2. Morphology and systematics (Elateroidea, Bostrichiformia partim). De Gruyter, Berlin, New York, pp. 574 - 659." type="book chapter" year="2010">
Matthews
<emphasis id="B91AEAC0131DFFB3D0238CF2FC84F90E" box="[799,855,1689,1713]" italics="true" pageId="3" pageNumber="464">et al.</emphasis>
, 2010
</bibRefCitation>
: 582): small size, shortened antenna, deeplyinserted eyes, body covered with long setae, and all tibiae bearing widened and flattened outer faces for digging and pushing.
</paragraph>
</subSubSection>
</treatment>
</document>
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