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<document id="BA87E40D33025B8D7F387A52FC7E7895" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C787854628FFE7426C4121340FFDCD" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phialodonta Herbert 2020, gen. nov." docType="treatment" docUuid="39C2A680-FD1C-486C-AC3E-3F23EED1CD33" docUuidSource="ZooBank" docVersion="7" lastPageNumber="38" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="37" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A1FCFFF9A065063D02E2F09C4F1E2035">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
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<mods:namePart id="8A8E627025D28B73E30EF7CB55BEFCD3">Herbert, David G.</mods:namePart>
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<mods:affiliation id="66CCD47B3BC89F821042A4355D5759A3">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
<mods:nameIdentifier id="78D8B5A7742E0539ADE6B26FA78B9BCE" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:title id="5EE6AA3CBA342B87BABB880CA4B1A27E">European Journal of Taxonomy</mods:title>
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<mods:part id="5234D8A3CE1F9C9BC9EAEE2A53303D40">
<mods:date id="60FA8861498148B259EBC849D259688C">2020</mods:date>
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<mods:number id="D1E8D537988062AE50AA6C8CE4F01B84">2020-04-17</mods:number>
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<treatment id="03C787854628FFE7426C4121340FFDCD" ID-DOI="http://doi.org/10.5281/zenodo.3804752" ID-GBIF-Taxon="163732357" ID-Zenodo-Dep="3804752" LSID="urn:lsid:zoobank.org:act:39C2A680-FD1C-486C-AC3E-3F23EED1CD33" httpUri="http://treatment.plazi.org/id/03C787854628FFE7426C4121340FFDCD" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<subSubSection id="C37465184628FFE6426C412135F6FDE4" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="8BD136934628FFE6426C412132FEFDC2" blockId="36.[358,1230,512,573]" box="[622,966,512,539]" pageId="36" pageNumber="37">
<heading id="D09981FF4628FFE6426C412132FEFDC2" box="[622,966,512,539]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="37" reason="2">
Genus
<taxonomicName id="4C6E4D104628FFE642C24121326CFDC2" ID-CoL="7PCF3" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[704,852,512,539]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B91AEA814628FFE642C24121326CFDC2" bold="true" box="[704,852,512,539]" italics="true" pageId="36" pageNumber="37">Phialodonta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE64359412032FEFDC2" box="[859,966,513,539]" pageId="36" pageNumber="37" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C37465184628FFE640BF41443365FD77" pageId="36" pageNumber="37" type="type_taxon">
<paragraph id="8BD136934628FFE640BF41443060FDA7" blockId="36.[189,344,612,639]" box="[189,344,612,639]" pageId="36" pageNumber="37">
<heading id="D09981FF4628FFE640BF41443060FDA7" bold="true" box="[189,344,612,639]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B91AEA814628FFE640BF41443060FDA7" bold="true" box="[189,344,612,639]" pageId="36" pageNumber="37">
<typeStatus id="54D588314628FFE640BF414431C2FDA6" box="[189,250,613,639]" pageId="36" pageNumber="37">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934628FFE640BF41B23365FD77" blockId="36.[189,605,659,686]" box="[189,605,659,686]" pageId="36" pageNumber="37">
<taxonomicName id="4C6E4D104628FFE640BF41B2309DFD74" ID-CoL="7SP53" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[189,421,659,685]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="aviana" status="gen. et sp. nov.">
<emphasis id="B91AEA814628FFE640BF41B2309DFD74" box="[189,421,659,685]" italics="true" pageId="36" pageNumber="37">Phialodonta aviana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE641AF41B53365FD77" box="[429,605,660,686]" pageId="36" pageNumber="37" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C37465184628FFE640BF41EC3225FC30" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="8BD136934628FFE640BF41EC300EFD3E" blockId="36.[189,310,717,743]" box="[189,310,717,743]" pageId="36" pageNumber="37">
<heading id="D09981FF4628FFE640BF41EC300EFD3E" bold="true" box="[189,310,717,743]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B91AEA814628FFE640BF41EC300EFD3E" bold="true" box="[189,310,717,743]" pageId="36" pageNumber="37">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934628FFE640BF41DD3225FC30" blockId="36.[189,1399,764,1001]" pageId="36" pageNumber="37">Shell very small to relatively large (adult diameter ± 1.32.0 mm), spire slightly sunken to slightly elevated; umbilicus moderate to very wide; whorls strongly rounded, periphery at mid-whorl. Protoconch smooth or with low, weakly undulant axial sculpture. Teleoconch with close-set axial riblets; riblets compound, composed of several periostracal lamellae; intervals between riblets with several finer intermediary axial threads; spiral sculpture weak, at most comprising extremely fine, close-set spiral threads. Aperture lunate to broadly lunate, variously furnished with parietal and palatal dentition in the form of denticles and/or in-running lamellae/ridges.</paragraph>
</subSubSection>
<subSubSection id="C37465184628FFE640BF47283380FBAD" pageId="36" pageNumber="37" type="etymology">
<paragraph id="8BD136934628FFE640BF4728307CFBFA" blockId="36.[189,324,1033,1059]" box="[189,324,1033,1059]" pageId="36" pageNumber="37">
<heading id="D09981FF4628FFE640BF4728307CFBFA" bold="true" box="[189,324,1033,1059]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B91AEA814628FFE640BF4728307CFBFA" bold="true" box="[189,324,1033,1059]" pageId="36" pageNumber="37">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934628FFE640BF47193380FBAD" blockId="36.[189,1399,1079,1140]" pageId="36" pageNumber="37">
From the Latin
<emphasis id="B91AEA814628FFE6417B471630FCFB88" box="[377,452,1079,1105]" italics="true" pageId="36" pageNumber="37">phiala</emphasis>
: a saucer or bowl, and
<emphasis id="B91AEA814628FFE642E24716321BFB88" box="[736,803,1079,1105]" italics="true" pageId="36" pageNumber="37">donta</emphasis>
: a contraction of
<taxonomicName id="4C6E4D104628FFE643FA47163556FB88" authorityName="Melvill &amp; Ponsonby" authorityYear="1908" box="[1016,1134,1079,1105]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis id="B91AEA814628FFE643FA47163556FB88" box="[1016,1134,1079,1105]" italics="true" pageId="36" pageNumber="37">Afrodonta</emphasis>
</taxonomicName>
; with reference to the wide, shallow umbilicus. Gender feminine.
</paragraph>
</subSubSection>
<caption id="DF11661B4628FFE640BF44653539F832" ID-DOI="http://doi.org/10.5281/zenodo.3762308" ID-Zenodo-Dep="3762308" httpUri="https://zenodo.org/record/3762308/files/figure.png" pageId="36" pageNumber="37" startId="36.[189,232,1860,1886]" targetBox="[302,1285,1199,1820]" targetPageId="36">
<paragraph id="8BD136934628FFE640BF44653539F832" blockId="36.[189,1398,1860,2027]" pageId="36" pageNumber="37">
<emphasis id="B91AEA814628FFE640BF44653023F887" bold="true" box="[189,283,1860,1886]" pageId="36" pageNumber="37">Fig. 14.</emphasis>
Distribution of
<taxonomicName id="4C6E4D104628FFE641DE4465336FF887" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[476,599,1860,1886]" class="Gastropoda" family="Charopidae" genus="Iterodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B91AEA814628FFE641DE4465336FF887" box="[476,599,1860,1886]" italics="true" pageId="36" pageNumber="37">Iterodonta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE64262446533F5F887" box="[608,717,1860,1886]" pageId="36" pageNumber="37" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C6E4D104628FFE6430B446532A2F887" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[777,922,1860,1886]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B91AEA814628FFE6430B446532A2F887" box="[777,922,1860,1886]" italics="true" pageId="36" pageNumber="37">Phialodonta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE643A144653528F887" box="[931,1040,1860,1886]" pageId="36" pageNumber="37" rank="genus">gen. nov.</taxonomicNameLabel>
species.
<taxonomicName id="4C6E4D104628FFE6447F4465344EF887" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[1149,1398,1860,1886]" class="Gastropoda" family="Charopidae" genus="Iterodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="ammonita" status="gen. et sp. nov.">
<emphasis id="B91AEA814628FFE6447F4465344EF887" box="[1149,1398,1860,1886]" italics="true" pageId="36" pageNumber="37">Iterodonta ammonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE640BF4449305FF85B" box="[189,359,1896,1922]" pageId="36" pageNumber="37" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(brown triangles),
<taxonomicName id="4C6E4D104628FFE6424344463268F858" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[577,848,1895,1921]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="agulhasae" status="gen. et sp. nov.">
<emphasis id="B91AEA814628FFE6424344463268F858" box="[577,848,1895,1921]" italics="true" pageId="36" pageNumber="37">Phialodonta agulhasae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE64357444932C7F85B" box="[853,1023,1896,1922]" pageId="36" pageNumber="37" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(orange square),
<taxonomicName id="4C6E4D104628FFE644C74449344EF858" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[1221,1398,1896,1921]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="atromontana" status="gen. et sp. nov.">
<emphasis id="B91AEA814628FFE644C74449344EF858" box="[1221,1398,1896,1921]" italics="true" pageId="36" pageNumber="37">P. atromontana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE640BF44AA304CF87D" box="[189,372,1931,1957]" pageId="36" pageNumber="37" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(turquoise square),
<taxonomicName id="4C6E4D104628FFE6426644AA33EBF87D" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[612,723,1931,1956]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="aviana" status="gen. et sp. nov.">
<emphasis id="B91AEA814628FFE6426644AA33EBF87D" box="[612,723,1931,1956]" italics="true" pageId="36" pageNumber="37">P. aviana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE642DE44AA32ABF87D" box="[732,915,1931,1957]" pageId="36" pageNumber="37" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(royal blue squares),
<taxonomicName id="4C6E4D104628FFE6449844AA30BDF811" authority="(Connolly, 1933)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Connolly" baseAuthorityYear="1933" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="introtuberculata" status="gen. et comb. nov.">
<emphasis id="B91AEA814628FFE6449844AA344EF87D" box="[1178,1398,1930,1956]" italics="true" pageId="36" pageNumber="37">P. introtuberculata</emphasis>
(
<bibRefCitation id="EFFF4B624628FFE640C7448C3045F811" author="Connolly M." box="[197,381,1965,1992]" pageId="36" pageNumber="37" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly, 1933</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE6418C448F335EF811" box="[398,614,1965,1992]" pageId="36" pageNumber="37" rank="species">gen. et comb. nov.</taxonomicNameLabel>
(red squares),
<taxonomicName id="4C6E4D104628FFE64314448F357BF811" authority="(Burnup, 1912)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Burnup" baseAuthorityYear="1912" box="[790,1091,1966,1992]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="perfida" status="gen. et comb. nov.">
<emphasis id="B91AEA814628FFE64314448F32BEF811" box="[790,902,1966,1992]" italics="true" pageId="36" pageNumber="37">P. perfida</emphasis>
(
<bibRefCitation id="EFFF4B624628FFE64394448F3503F811" author="Burnup H. C." box="[918,1083,1966,1992]" pageId="36" pageNumber="37" pagination="333 - 343" refId="ref26463" refString="Burnup H. C. 1912. On Afrodonta Melv. and Pons., with descriptions of new species. Annals of the Natal Museum 2 (3): 333 - 343." type="journal article" year="1912">Burnup, 1912</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE64449448F341CF811" box="[1099,1316,1965,1992]" pageId="36" pageNumber="37" rank="species">gen. et comb. nov.</taxonomicNameLabel>
(green square),
<taxonomicName id="4C6E4D104628FFE6412044F33091F832" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[290,425,2001,2027]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="rivalalea" status="gen. et sp. nov.">
<emphasis id="B91AEA814628FFE6412044F33091F832" box="[290,425,2001,2027]" italics="true" pageId="36" pageNumber="37">P. rivalalea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE641B244F03358F832" box="[432,608,2001,2027]" pageId="36" pageNumber="37" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(pink squares). Contour at 1000 m.
</paragraph>
</caption>
<subSubSection id="C37465184629FFE740BF422A340FFDCD" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BD136934629FFE740BF422A3017FEFC" blockId="37.[189,303,267,293]" box="[189,303,267,293]" pageId="37" pageNumber="38">
<heading id="D09981FF4629FFE740BF422A3017FEFC" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B91AEA814629FFE740BF422A3017FEFC" bold="true" box="[189,303,267,293]" pageId="37" pageNumber="38">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934629FFE740BF421D340FFDCD" blockId="37.[189,1399,316,533]" pageId="37" pageNumber="38">
<taxonomicName id="4C6E4D104629FFE740BF421D3076FE8F" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[189,334,316,342]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B91AEA814629FFE740BF421D3076FE8F" box="[189,334,316,342]" italics="true" pageId="37" pageNumber="38">Phialodonta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4629FFE74155421C30FCFE8E" box="[343,452,317,343]" pageId="37" pageNumber="38" rank="genus">gen. nov.</taxonomicNameLabel>
is proposed for a group of species which share a smooth or weakly sculptured protoconch and a relatively coarse teleoconch sculpture of compound axial riblets with multiple fine intermediary axial threads. The genus ranges widely, from the Agulhas region, W.
<collectingRegion id="49AAF8714629FFE7447642A93588FE7B" box="[1140,1200,392,418]" country="South Africa" name="Western Cape" pageId="37" pageNumber="38">Cape</collectingRegion>
to the KwaZulu- Natal
<collectingRegion id="49AAF8714629FFE74106428F304FFE11" box="[260,375,430,456]" country="Zimbabwe" name="Midlands" pageId="37" pageNumber="38">Midlands</collectingRegion>
. In
<taxonomicName id="4C6E4D104629FFE741A0428F3378FE11" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[418,576,430,456]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B91AEA814629FFE741A0428F3378FE11" box="[418,576,430,456]" italics="true" pageId="37" pageNumber="38">Costulodonta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4629FFE7424A428E338BFE10" box="[584,691,431,457]" pageId="37" pageNumber="38" rank="genus">gen. nov.</taxonomicNameLabel>
the protoconch has distinct radial riblets and the teleoconch sculpture is generally somewhat finer. Members of
<taxonomicName id="4C6E4D104629FFE7431542F532B5FE37" authorityName="Melvill &amp; Ponsonby" authorityYear="1908" box="[791,909,468,494]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus">
<emphasis id="B91AEA814629FFE7431542F532B5FE37" box="[791,909,468,494]" italics="true" pageId="37" pageNumber="38">Afrodonta</emphasis>
</taxonomicName>
have a somewhat similar protoconch, but they possess a silky teleoconch sculpture comprised of simple axial riblets of alternating strength.
</paragraph>
</subSubSection>
</treatment>
</document>