692 lines
83 KiB
XML
692 lines
83 KiB
XML
<document id="01E652A65DB987CEE51943C7BD4183B5" ID-CLB-Dataset="282032" ID-DOI="10.11646/phytotaxa.629.2.6" ID-GBIF-Dataset="3e117228-19db-4094-9aad-90a026cf8456" ID-ISSN="1179-3163" ID-Zenodo-Dep="10275901" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1701851201194" checkinUser="plazi" docAuthor="Ocupa-Horna, Luis, Wilson, Mark & Jiménez, Marco M." docDate="2023" docId="03C78782FFA8FFA76085E94EFBD8FDDB" docLanguage="en" docName="phytotaxa.629.2.6.pdf" docOrigin="Phytotaxa 629 (2)" docSource="https://phytotaxa.mapress.com/pt/article/download/phytotaxa.629.2.6/51338" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Pleurothallis carpishensis Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="174" masterDocId="FFFEFFFAFFAAFFAC600DEB5AFF98FFB3" masterDocTitle="A new species of Pleurothallis (Orchidaceae: Pleurothallidinae) in subsection Macrophyllae-Fasciculatae found in Carpish Montane Forest Regional Conservation Area, Peru" masterLastPageNumber="175" masterPageNumber="163" pageNumber="165" updateTime="1702641665498" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="799B66E9AFD9B4874A66DBC1BDC00D39">
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<mods:title id="A486EBE5A7C30BEAA296CC9906E32352">A new species of Pleurothallis (Orchidaceae: Pleurothallidinae) in subsection Macrophyllae-Fasciculatae found in Carpish Montane Forest Regional Conservation Area, Peru</mods:title>
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<mods:roleTerm id="55A18298F79058CD2CD7D86BFFD7B84B">Author</mods:roleTerm>
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<mods:namePart id="5A662373D3121328C39EA0A99AFF11E6">Ocupa-Horna, Luis</mods:namePart>
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<mods:nameIdentifier id="663D0BBF4423562A092A943DB261861D" type="ORCID">0000-0002-4904-1060</mods:nameIdentifier>
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<mods:affiliation id="C142D0D3625ED0B7A98EF053B5B05E23">Departamento de Orquideología, Centro de Investigación en Biología Tropical y Conservación - CINBIOTYC, Cal. Nestor Martos Mza. e Lote. 16 A. H. Almirante Miguel Grau II, Piura, Peru. & Departamento de Gimnospermas y Monocotiledóneas, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Av. Arenales 1256, Jesús María, Lima, Peru. & luis. ocupa @ unmsm. edu. pe; https: // orcid. org / 0000 - 0002 - 4904 - 1060</mods:affiliation>
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<mods:nameIdentifier id="CCB6506047F3F9A9302F51574FEF5742" type="email">luis.ocupa@unmsm.edu.pe</mods:nameIdentifier>
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</mods:name>
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<mods:name id="9B420A06065C1DD55A46CE02A3A50306" type="personal">
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<mods:roleTerm id="E1F310BCEC167E7B582C4E03805736EC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="10F139E683C3397484CCFC25F94BEDDB">Wilson, Mark</mods:namePart>
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<mods:nameIdentifier id="30DA07F358EADC59E4F7316F33DC8901" type="ORCID">0000-0003-3044-7471</mods:nameIdentifier>
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<mods:affiliation id="5F4BCDCEE0E1C7B8F4E3599887092A99">Department of Organismal Biology and Ecology, Colorado College, Colorado Springs, CO 80903, USA. & mwilson @ coloradocollege. edu; https: // orcid. org / 0000 - 0003 - 3044 - 7471</mods:affiliation>
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<mods:nameIdentifier id="9FBAE2E2CEA9D081230AC4290D8EB94B" type="email">mwilson@coloradocollege.edu</mods:nameIdentifier>
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<mods:name id="A9784451B1FEF0A14BB03307C7431A7C" type="personal">
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<mods:roleTerm id="7A725986447EA87803A39C1A470FAE01">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A8808A159CCC3660A16ABC204CA2F2D5">Jiménez, Marco M.</mods:namePart>
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<mods:nameIdentifier id="CEEB71FDB61B870417017816CE9D7A22" type="ORCID">0000-0002-9502-5651</mods:nameIdentifier>
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<mods:affiliation id="91F94566C3765104AC72C77057CFD092">Grupo de Investigación en Medio Ambiente y Salud (BIOMAS), Ingeniería en Agroindustria, Facultad de Ingeniería y Ciencias Agropecuarias. Universidad de Las Américas, Vía a Nayón, Quito 170124, Pichincha, Ecuador. & marco. jimenez. leon @ udla. edu. ec; https: // orcid. org / 0000 - 0002 - 9502 - 5651</mods:affiliation>
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<mods:nameIdentifier id="C014834C88B7F252BBAE85DAF872ACCC" type="email">marco.jimenez.leon@udla.edu.ec</mods:nameIdentifier>
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<mods:typeOfResource id="185416F11976FE9041FB5D75497187A9">text</mods:typeOfResource>
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<mods:title id="EDE6D47F523CB9098280AE0817C6919F">Phytotaxa</mods:title>
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<mods:part id="65907E5865D05276971B5208077F251F">
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<mods:date id="B39FC008F43650679B3983D6FA9CC1D9">2023</mods:date>
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<mods:number id="476699E78B0ED3DCD3622C0064B4825A">2023-12-06</mods:number>
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<mods:detail id="05F574EB1028FF6EEC179F1F4E049171" type="volume">
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<mods:number id="78D6CCC6CB233996F0AC9A75B9CAB60B">629</mods:number>
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<mods:detail id="662A2FC9B363E29A23E5D33BB89EEEC1" type="issue">
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<mods:number id="99C096F0CE2FA44E5F62042516D4898B">2</mods:number>
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<mods:start id="FEAADBF7A9309BEEE3A8D8D37114E855">163</mods:start>
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<mods:end id="8B640A884D8B016F11BCF1F26274D3E1">175</mods:end>
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<mods:location id="D8AEDDE2945909F3565E09B4E377BCCF">
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<mods:url id="022A28B75A741F05DE20CA9EC34914ED">https://phytotaxa.mapress.com/pt/article/download/phytotaxa.629.2.6/51338</mods:url>
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</mods:location>
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<mods:classification id="89C0EAF92E034E859EA379E2432AA058">journal article</mods:classification>
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<mods:identifier id="B227145E2AD284ACFE89E002D263873E" type="CLB-Dataset">282032</mods:identifier>
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<mods:identifier id="D4584D1EA1A903385901B5A75480E041" type="DOI">10.11646/phytotaxa.629.2.6</mods:identifier>
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<mods:identifier id="32C78B3C0D42A783EBD1448BADA657E9" type="GBIF-Dataset">3e117228-19db-4094-9aad-90a026cf8456</mods:identifier>
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<mods:identifier id="877E07BEBBE04090DD3D4D973914FCC4" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="4C875C9BC0AD8D67AD3E8E807E1CC32F" type="Zenodo-Dep">10275901</mods:identifier>
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<treatment id="03C78782FFA8FFA76085E94EFBD8FDDB" ID-DOI="http://doi.org/10.5281/zenodo.10390966" ID-Zenodo-Dep="10390966" LSID="urn:lsid:plazi:treatment:03C78782FFA8FFA76085E94EFBD8FDDB" httpUri="http://treatment.plazi.org/id/03C78782FFA8FFA76085E94EFBD8FDDB" lastPageId="11" lastPageNumber="174" pageId="2" pageNumber="165">
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<subSubSection id="C374651FFFA8FFAE6085E94EFBC5FD9D" box="[136,1117,532,558]" pageId="2" pageNumber="165" type="nomenclature">
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<paragraph id="8BD13694FFA8FFAE6085E94EFBC5FD9D" blockId="2.[136,1451,532,630]" box="[136,1117,532,558]" pageId="2" pageNumber="165">
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<taxonomicName id="4C6E4D17FFA8FFAE6085E94EFCF8FD9D" authority="Ocupa, M.M.Jimenez & Mark Wilson" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[136,864,532,558]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="165" phylum="Tracheophyta" rank="species" species="carpishensis" status="sp. nov.">
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<emphasis id="B91AEA86FFA8FFAE6085E94EFE34FD9D" box="[136,428,532,558]" italics="true" pageId="2" pageNumber="165">Pleurothallis carpishensis</emphasis>
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Ocupa, M.M.Jiménez & Mark Wilson
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</taxonomicName>
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||
,
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||
<emphasis id="B91AEA86FFA8FFAE6366E94FFC27FD9D" box="[875,959,533,558]" italics="true" pageId="2" pageNumber="165">
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||
<taxonomicNameLabel id="A22957FDFFA8FFAE6366E94FFC27FD9D" box="[875,959,533,558]" pageId="2" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
(
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<figureCitation id="13552A11FFA8FFAE63C0E94EFBCEFD9D" box="[973,1110,532,558]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[136,229,1877,1899]" captionStartId-1="4.[136,229,1877,1899]" captionTargetBox-0="[151,1436,190,1847]" captionTargetBox-1="[151,1436,190,1853]" captionTargetId-0="figure-21@3.[151,1436,190,1853]" captionTargetId-1="figure-25@4.[151,1436,190,1853]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Illustration of Pleurothallis carpishensis Ocupa, M.M.Jiménez & Mark Wilson. A. Habit. B. Flower. C. Dissected perianth. D. Lip, column and ovary, lateral view. E. Column. F. Lip, adaxial view. G. Anther cap and pollinarium. Illustrated by S. Moreno from the holotype." captionText-1="FIGURE 3. Composite plate of Pleurothallis carpishensis Ocupa, M.M.Jiménez & Mark Wilson. A. Plant. B. Flower. C. Dissected perianth. D. Lip, column and ovary, lateral view.E. Column.F. Lip, adaxial (right) and abaxial (left) views. G.Anther cap and pollinarium. Prepared by L. Ocupa-Horna." figureDoi-0="http://doi.org/10.5281/zenodo.10275907" figureDoi-1="http://doi.org/10.5281/zenodo.10275909" httpUri-0="https://zenodo.org/record/10275907/files/figure.png" httpUri-1="https://zenodo.org/record/10275909/files/figure.png" pageId="2" pageNumber="165">Figures 2–3</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C374651FFFA8FFAE6085E966FB8DFDC6" pageId="2" pageNumber="165" type="materials_examined">
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<paragraph id="8BD13694FFA8FFAE6085E966FB8DFDC6" blockId="2.[136,1451,532,630]" pageId="2" pageNumber="165">
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<materialsCitation id="3B063CC9FFA8FFAE6085E966FB89FDC6" collectingDate="2022-03-22" collectionCode="USM" collectorName="L. Ocupa" country="Peru" elevation="1722" location="Distrito Chinchao" municipality="Prov" pageId="2" pageNumber="165" specimenCount="1" stateProvince="Huanuco" typeStatus="holotype">
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||
<emphasis id="B91AEA86FFA8FFAE6085E966FF5AFDE2" bold="true" box="[136,194,572,593]" pageId="2" pageNumber="165">Type:</emphasis>
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||
—
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||
<collectingCountry id="F3797604FFA8FFAE60D6E966FE83FDE2" box="[219,283,572,593]" name="Peru" pageId="2" pageNumber="165" value="Peru">PERU</collectingCountry>
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||
.
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||
<collectingRegion id="49AAF876FFA8FFAE612BE966FE18FDE2" box="[294,384,572,593]" country="Peru" name="Huanuco" pageId="2" pageNumber="165">Huánuco</collectingRegion>
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||
:
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||
<collectingMunicipality id="6BB5ACEEFFA8FFAE6181E966FE23FDE2" box="[396,443,572,593]" pageId="2" pageNumber="165">Prov</collectingMunicipality>
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||
. Huánuco,
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||
<location id="8EB1604FFFA8FFAE6221E966FD44FDE1" LSID="urn:lsid:plazi:treatment:03C78782FFA8FFA76085E94EFBD8FDDB:8EB1604FFFA8FFAE6221E966FD44FDE1" box="[556,732,572,594]" country="Peru" municipality="Prov" name="Distrito Chinchao" pageId="2" pageNumber="165" stateProvince="Huanuco">Distrito Chinchao</location>
|
||
,
|
||
<location id="8EB1604FFFA8FFAE62E5E966FE43FDC5" LSID="urn:lsid:plazi:treatment:03C78782FFA8FFA76085E94EFBD8FDDB:8EB1604FFFA8FFAE62E5E966FE43FDC5" country="Peru" municipality="Prov" name="Area de Conservacion Regional Bosque Montano de Carpish, cerca del caserio Santa Rosa de Quives" pageId="2" pageNumber="165" stateProvince="Huanuco">Área de Conservación Regional Bosque Montano de Carpish, cerca del caserío Santa Rosa de Quives</location>
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||
,
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||
<quantity id="4C969B71FFA8FFAE61E8E93AFDA8FDC6" box="[485,560,608,629]" metricMagnitude="3" metricUnit="m" metricValue="1.722" pageId="2" pageNumber="165" unit="m" value="1722.0">
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<elevation id="0043D1A7FFA8FFAE61E8E93AFDA8FDC6" box="[485,560,608,629]" metricMagnitude="3" metricUnit="m" metricValue="1.722" pageId="2" pageNumber="165" unit="m" value="1722.0">1722 m</elevation>
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||
</quantity>
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||
,
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||
<date id="FFD01054FFA8FFAE6237E93AFD57FDC6" box="[570,719,608,630]" pageId="2" pageNumber="165" value="2022-03-22">
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||
<collectingDate id="EF94E9BCFFA8FFAE6237E93AFD57FDC6" box="[570,719,608,630]" pageId="2" pageNumber="165" value="2022-03-22">22 March 2022</collectingDate>
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</date>
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,
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||
<emphasis id="B91AEA86FFA8FFAE62D4E93AFCC4FDC6" box="[729,860,608,629]" italics="true" pageId="2" pageNumber="165">
|
||
<collectorName id="269B5342FFA8FFAE62D4E93AFCAAFDC6" box="[729,818,608,629]" pageId="2" pageNumber="165">L. Ocupa</collectorName>
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||
301
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||
</emphasis>
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||
(
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||
<typeStatus id="54D58836FFA8FFAE6364E93AFC5AFDC5" box="[873,962,608,630]" pageId="2" pageNumber="165" type="holotype">holotype</typeStatus>
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||
:
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||
<collectionCode id="ED7FAE51FFA8FFAE63C6E93AFB9FFDC6" box="[971,1031,608,629]" collectionName="USM" pageId="2" pageNumber="165">USM</collectionCode>
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||
!)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C374651FFFA8FFAE6085E9FEFF61FCC1" pageId="2" pageNumber="165" type="diagnosis">
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<paragraph id="8BD13694FFA8FFAE6085E9FEFF61FCC1" blockId="2.[136,1452,676,1999]" pageId="2" pageNumber="165">
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||
Morphologically similar to
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<taxonomicName id="4C6E4D17FFA8FFAE61CFE9FFFDB3FD0D" authorityName="Luer" authorityYear="1979" box="[450,555,676,702]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="165" phylum="Tracheophyta" rank="species" species="radula">
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||
<emphasis id="B91AEA86FFA8FFAE61CFE9FFFDB3FD0D" box="[450,555,676,702]" italics="true" pageId="2" pageNumber="165">P. radula</emphasis>
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||
</taxonomicName>
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||
but is distinguished by the inflorescence that arises from a depressed, oblong, obtuse spathaceous bract (
|
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<emphasis id="B91AEA86FFA8FFAE61A0E993FE53FD51" box="[429,459,713,738]" italics="true" pageId="2" pageNumber="165">vs.</emphasis>
|
||
erect, oblanceolate and sub-truncate), dorsal sepal lanceolate (
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||
<emphasis id="B91AEA86FFA8FFAE6489E993FB3AFD51" box="[1156,1186,713,738]" italics="true" pageId="2" pageNumber="165">vs.</emphasis>
|
||
ovate to ovate-elliptic), petals oblong-lanceolate, apiculate, with large papillae on the margins (
|
||
<emphasis id="B91AEA86FFA8FFAE63CDE9B7FC47FCB5" box="[960,991,749,774]" italics="true" pageId="2" pageNumber="165">vs.</emphasis>
|
||
narrowly lanceolate, acute and serrate), a triangular, sub-acute lip (
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||
<emphasis id="B91AEA86FFA8FFAE61B0E84BFE44FC99" box="[445,476,785,810]" italics="true" pageId="2" pageNumber="165">vs.</emphasis>
|
||
ovate to oblong-ovate, acute), with a callus broadly ovate and emarginate, papillose-verrucose (
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||
<emphasis id="B91AEA86FFA8FFAE6107E86FFEB1FCFD" box="[266,297,821,846]" italics="true" pageId="2" pageNumber="165">vs.</emphasis>
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||
callus glabrous,
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||
<quantity id="4C969B71FFA8FFAE61FBE86EFDA4FCFC" box="[502,572,820,847]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="165" unit="mm" value="1.0">1 mm</quantity>
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||
long) and column edge surrounding stigmatic surfaces ciliate (
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<emphasis id="B91AEA86FFA8FFAE6520E86FFAD3FCFD" box="[1325,1355,821,846]" italics="true" pageId="2" pageNumber="165">vs.</emphasis>
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||
slightly papillate).
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C374651FFFA8FFAE60B0E826FBD4F961" pageId="2" pageNumber="165" type="description">
|
||
<paragraph id="8BD13694FFA8FFAE60B0E826FBD4F961" blockId="2.[136,1452,676,1999]" pageId="2" pageNumber="165">
|
||
<emphasis id="B91AEA86FFA8FFAE60B0E826FEF5FC25" bold="true" box="[189,365,892,918]" pageId="2" pageNumber="165">Description:—</emphasis>
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||
Epiphytic
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||
<emphasis id="B91AEA86FFA8FFAE61E8E826FD80FC25" box="[485,536,892,918]" italics="true" pageId="2" pageNumber="165">herb</emphasis>
|
||
, caespitose, up to
|
||
<quantity id="4C969B71FFA8FFAE62F8E826FCA6FC24" box="[757,830,892,919]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="165" unit="cm" value="23.0">23 cm</quantity>
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||
tall,
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||
<emphasis id="B91AEA86FFA8FFAE6373E827FC2FFC25" box="[894,951,893,918]" italics="true" pageId="2" pageNumber="165">roots</emphasis>
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||
ca. 1.0 mm in diameter, slender, flexuous.
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<emphasis id="B91AEA86FFA8FFAE6085E8FAFF66FC09" box="[136,254,928,954]" italics="true" pageId="2" pageNumber="165">Ramicauls</emphasis>
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||
suberect to erect, terete, ca. 8.0–20.0 cm long,
|
||
<quantity id="4C969B71FFA8FFAE632BE8FAFC37FC08" box="[806,943,928,955]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="2" pageNumber="165" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
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in diameter, curved at the apex, forming an angle of about 90° in the abscission layer, enclosed by two papyraceous, sulcate, pale brown, tubular sheaths, upper sheath ca. 20.0–23.0 × 3.0 mm, basal sheath ca. 8.0–12.0 × 3.0 mm, unifoliate.
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<emphasis id="B91AEA86FFA8FFAE6435E8B3FB1EFBB1" box="[1080,1158,1001,1026]" italics="true" pageId="2" pageNumber="165">Leaves</emphasis>
|
||
borne at the apex of the ramicaul, pale green, thinly coriaceous, deflexed toward the stem, concave, narrowly ovate, apically somewhat falcate, with a navicular concavity at the base of the spathe, margins involute, acuminate, tridentate sometimes with a longer central tooth, ca. 5.2–9.3 ×
|
||
<quantity id="4C969B71FFA8FFAE61CCEF0EFDDAFBDC" box="[449,578,1108,1135]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.7" metricValueMin="2.9" pageId="2" pageNumber="165" unit="cm" value="3.3" valueMax="3.7" valueMin="2.9">2.9–3.7 cm</quantity>
|
||
, base sessile, cordate, lobes unequal, sometimes overlapping.
|
||
<emphasis id="B91AEA86FFA8FFAE650CEF0EFA0FFBDD" box="[1281,1431,1108,1134]" italics="true" pageId="2" pageNumber="165">Inflorescence</emphasis>
|
||
a solitary flower, resting on a navicular concavity at the base of the bract and borne from a depressed, conduplicate, oblong, obtuse spathaceous bract at the base of the leaf, ca.
|
||
<quantity id="4C969B71FFA8FFAE632FEFC6FC31FB04" box="[802,937,1180,1207]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="2" pageNumber="165" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.5–9.5 mm</quantity>
|
||
long, striate, yellowish-green, becoming light brown, dry-papyraceous when mature, dotted with black, concealing peduncle, pedicel and floral bract;
|
||
<emphasis id="B91AEA86FFA8FFAE654BEF9AFA33FB69" box="[1350,1451,1216,1242]" italics="true" pageId="2" pageNumber="165">peduncle</emphasis>
|
||
terete, up to
|
||
<quantity id="4C969B71FFA8FFAE611BEFBEFEC0FB4C" box="[278,344,1252,1279]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="165" unit="mm" value="3.0">3 mm</quantity>
|
||
long;
|
||
<emphasis id="B91AEA86FFA8FFAE61ADEFBEFDB4FB4D" box="[416,556,1252,1278]" italics="true" pageId="2" pageNumber="165">floral bracts</emphasis>
|
||
infundibuliform, membranaceous, 3.5–5.0 mm long;
|
||
<emphasis id="B91AEA86FFA8FFAE6487EFBEFB43FB4D" box="[1162,1243,1252,1278]" italics="true" pageId="2" pageNumber="165">pedicel</emphasis>
|
||
terete, ca. 4.0–5.0 mm long;
|
||
<emphasis id="B91AEA86FFA8FFAE60F6EE53FEA3FA91" box="[251,315,1289,1314]" italics="true" pageId="2" pageNumber="165">ovary</emphasis>
|
||
terete, pale green, somewhat incurved, 6-furrowed, ca. 2.7–3.0 mm long, with black dots all over the surface.
|
||
<emphasis id="B91AEA86FFA8FFAE60EAEE76FEDBFAF5" box="[231,323,1324,1350]" italics="true" pageId="2" pageNumber="165">Flowers</emphasis>
|
||
resupinate, spreading, yellow-hyaline, column white, anther cap cream, pollinia yellow;
|
||
<emphasis id="B91AEA86FFA8FFAE652FEE76FA33FAF5" box="[1314,1451,1324,1350]" italics="true" pageId="2" pageNumber="165">dorsal sepal</emphasis>
|
||
erect, slightly concave, lanceolate, sub-acute, 8.0–10.3 ×
|
||
<quantity id="4C969B71FFA8FFAE6308EE0AFC16FAD8" box="[773,910,1360,1387]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="2" pageNumber="165" unit="mm" value="3.5" valueMax="3.8" valueMin="3.2">3.2–3.8 mm</quantity>
|
||
, 3-veined;
|
||
<emphasis id="B91AEA86FFA8FFAE6404EE0AFB38FAD9" box="[1033,1184,1360,1386]" italics="true" pageId="2" pageNumber="165">lateral sepals</emphasis>
|
||
connate into a reflexed, ovate-lanceolate synsepal, centrally concave-channeled, obtuse, minutely emarginate, 8.0–10.0 ×
|
||
<quantity id="4C969B71FFA8FFAE64EDEE2EFAF3FA3C" box="[1248,1387,1396,1423]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="2" pageNumber="165" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5–4.8 mm</quantity>
|
||
, each half 2-veined;
|
||
<emphasis id="B91AEA86FFA8FFAE6120EEC2FEE8FA01" box="[301,368,1432,1458]" italics="true" pageId="2" pageNumber="165">petals</emphasis>
|
||
porrect, shallowly concave, oblong-lanceolate, apiculate, thickened apex, with large papillae on the margins, 4.5–6.0 × 1.5–2.0 mm, single-veined, abaxially 1-keeled;
|
||
<emphasis id="B91AEA86FFA8FFAE63BEEEE6FC49FA65" box="[947,977,1468,1494]" italics="true" pageId="2" pageNumber="165">lip</emphasis>
|
||
thick, fleshy, triangular, channeled in the midline, sub-acute, apex sometimes slightly recurved, 4.5–5.0 × 2.0–
|
||
<quantity id="4C969B71FFA8FFAE6391EEBAFC6CFA48" box="[924,1012,1504,1531]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="165" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, with three incomplete nerves, abaxial side foveolate, adaxial side cellular-papillose, without glenion; callus basal, widely ovate, emarginate, occupying 1/3 of the lip length, waxy, papillose-verrucose, the papillae irregularly coarse, especially at the margins, 1.7 ×
|
||
<quantity id="4C969B71FFA8FFAE6585ED72FF2EF9D4" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="165" unit="mm" value="2.3">2.3 mm</quantity>
|
||
, with two oblique, lateral, marginal and elevated keels;
|
||
<emphasis id="B91AEA86FFA8FFAE633BED16FC10F9D5" box="[822,904,1612,1638]" italics="true" pageId="2" pageNumber="165">column</emphasis>
|
||
short, stout, terete, ca. 1.0 × 2.0 mm, periphery of stigmatic surfaces ciliate, rostellum apical, rostellar flap long, linear, obtuse;
|
||
<emphasis id="B91AEA86FFA8FFAE6408ED2AFBD6F939" box="[1029,1102,1648,1674]" italics="true" pageId="2" pageNumber="165">anther</emphasis>
|
||
apical, incumbent, papillose at the margins, anther cap cucullate, long-ovate, sub-cordate, 2-celled, 1.2 ×
|
||
<quantity id="4C969B71FFA8FFAE63CEEDCEFB83F91C" box="[963,1051,1684,1711]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="165" unit="mm" value="0.5">0.5 mm</quantity>
|
||
;
|
||
<emphasis id="B91AEA86FFA8FFAE642BEDCFFBE9F91D" box="[1062,1137,1685,1710]" italics="true" pageId="2" pageNumber="165">stigma</emphasis>
|
||
apical, reniform;
|
||
<emphasis id="B91AEA86FFA8FFAE6534EDCEFA08F91D" box="[1337,1424,1684,1710]" italics="true" pageId="2" pageNumber="165">pollinia</emphasis>
|
||
2, narrowly obovoid, complanate,
|
||
<quantity id="4C969B71FFA8FFAE61E0EDE2FDDBF960" box="[493,579,1720,1747]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="165" unit="mm" value="0.8">0.8 mm</quantity>
|
||
long, brought together by a minute viscidium.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374651FFFA8FFAE60B0ED86FD76F87D" pageId="2" pageNumber="165" type="distribution">
|
||
<paragraph id="8BD13694FFA8FFAE60B0ED86FD76F87D" blockId="2.[136,1452,676,1999]" pageId="2" pageNumber="165">
|
||
<emphasis id="B91AEA86FFA8FFAE60B0ED86FE6BF945" bold="true" box="[189,499,1756,1782]" pageId="2" pageNumber="165">Distribution and ecology:</emphasis>
|
||
—
|
||
<taxonomicName id="4C6E4D17FFA8FFAE6202ED86FCAEF945" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[527,822,1756,1782]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="165" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA8FFAE6202ED86FCAEF945" box="[527,822,1756,1782]" italics="true" pageId="2" pageNumber="165">Pleurothallis carpishensis</emphasis>
|
||
</taxonomicName>
|
||
is known only from a small area on the eastern slope side of the central branch of the Peruvian Andes, forming part of the ecosystem known as the Yunga montane forest (
|
||
<bibRefCitation id="EFFF4B65FFA8FFAE6082EC7EFEAFF88D" author="MINAM" box="[143,311,1828,1854]" pageId="2" pageNumber="165" refId="ref4843" refString="MINAM. (2018) Definiciones conceptuales de los ecosistemas del Peru. Ministerio del Ambiente - MINAM, Lima, 112 pp." type="book" year="2018">
|
||
<collectionCode id="ED7FAE51FFA8FFAE6082EC7EFF6CF88D" box="[143,244,1828,1854]" pageId="2" pageNumber="165">MINAM</collectionCode>
|
||
2018
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFF4B65FFA8FFAE614CEC7EFEE5F88D" author="MINAM" box="[321,381,1828,1854]" pageId="2" pageNumber="165" refId="ref4867" refString="MINAM. (2019) Mapa Nacional de Ecosistemas: Memoria descriptiva. NEGRAPATA S. A. C., Lima, 124 pp." type="book" year="2019">2019</bibRefCitation>
|
||
), between 1720 and
|
||
<quantity id="4C969B71FFA8FFAE626FEC7EFD2EF88C" box="[610,694,1828,1855]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="2" pageNumber="165" unit="m" value="1730.0">1730 m</quantity>
|
||
elevation. This locality corresponds to the Carpish Montane Forest, located in the district of Chinchao, in the department of Huánuco (
|
||
<figureCitation id="13552A11FFA8FFAE6395EC12FC66F8D1" box="[920,1022,1864,1890]" captionStart="FIGURE 4" captionStartId="5.[136,229,1056,1078]" captionTargetBox="[190,1391,188,1035]" captionTargetId="figure-21@5.[177,1411,175,1047]" captionTargetPageId="5" captionText="FIGURE 4. Location map of Pleurothallis carpishensis Ocupa, M.M.Jiménez & Mark Wilson in the Carpish Montane Forest Regional Conservation Area. Prepared by L. Ocupa-Horna." figureDoi="http://doi.org/10.5281/zenodo.10275911" httpUri="https://zenodo.org/record/10275911/files/figure.png" pageId="2" pageNumber="165">Figure 4</figureCitation>
|
||
). Individuals of this species grow as epiphytes on branches located in the tree canopy, especially in the zone IV (
|
||
<bibRefCitation id="EFFF4B65FFA8FFAE63C4EC36FB18F835" author="Johansson, D. R." box="[969,1152,1900,1926]" pageId="2" pageNumber="165" pagination="1 - 129" refId="ref4462" refString="Johansson, D. R. (1974) Ecology of the vascular epiphytes in West African rainforest. Acta Phytogeographica Suecia 59: 1 - 129." type="journal article" year="1974">Johansson 1974</bibRefCitation>
|
||
), at ~
|
||
<quantity id="4C969B71FFA8FFAE64B1EC36FB7EF834" box="[1212,1254,1900,1927]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="165" unit="m" value="8.0">8 m</quantity>
|
||
above the ground (
|
||
<figureCitation id="13552A11FFA8FFAE609DECCAFF6AF818" box="[144,242,1936,1963]" captionStart="FIGURE 5" captionStartId="6.[136,229,2028,2050]" captionTargetBox="[163,1424,190,2004]" captionTargetId="figure-25@6.[163,1424,190,2004]" captionTargetPageId="6" captionText="FIGURE 5. Habitat of Pleurothallis carpishensis Ocupa, M.M.Jiménez & Mark Wilson. A. Close-up to tree canopy branches. B. P. carpishensis growing in situ. Photos by L. Ocupa-Horna." figureDoi="http://doi.org/10.5281/zenodo.10275913" httpUri="https://zenodo.org/record/10275913/files/figure.png" pageId="2" pageNumber="165">Figure 5</figureCitation>
|
||
). The flowering of this species occurs, at the beginning of the rainy season, between March and April, and during the dry season, between August and September.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF11661CFFA9FFAF6085EC0FFE9FF800" ID-DOI="http://doi.org/10.5281/zenodo.10275907" ID-Zenodo-Dep="10275907" httpUri="https://zenodo.org/record/10275907/files/figure.png" pageId="3" pageNumber="166" startId="3.[136,229,1877,1899]" targetBox="[151,1436,190,1847]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BD13694FFA9FFAF6085EC0FFE9FF800" blockId="3.[136,1453,1877,1971]" pageId="3" pageNumber="166">
|
||
<emphasis id="B91AEA86FFA9FFAF6085EC0FFF66F8D9" bold="true" box="[136,254,1877,1899]" pageId="3" pageNumber="166">FIGURE 2.</emphasis>
|
||
Illustration of
|
||
<taxonomicName id="4C6E4D17FFA9FFAF6183EC0FFAC8F8D8" authority="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[398,1360,1877,1899]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="166" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA9FFAF6183EC0FFD11F8D9" box="[398,649,1877,1898]" italics="true" pageId="3" pageNumber="166">Pleurothallis carpishensis</emphasis>
|
||
Ocupa, M.M.Jiménez & Mark Wilson.
|
||
<emphasis id="B91AEA86FFA9FFAF6407EC0FFBBAF8D9" bold="true" box="[1034,1058,1877,1898]" pageId="3" pageNumber="166">A.</emphasis>
|
||
Habit.
|
||
<emphasis id="B91AEA86FFA9FFAF6467EC0CFB18F8D8" bold="true" box="[1130,1152,1878,1899]" pageId="3" pageNumber="166">B.</emphasis>
|
||
Flower.
|
||
<emphasis id="B91AEA86FFA9FFAF64D8EC0FFB75F8D8" bold="true" box="[1237,1261,1877,1899]" pageId="3" pageNumber="166">C.</emphasis>
|
||
Dissected
|
||
</taxonomicName>
|
||
perianth.
|
||
<emphasis id="B91AEA86FFA9FFAF6085EC20FF38F83C" bold="true" box="[136,160,1914,1935]" pageId="3" pageNumber="166">D.</emphasis>
|
||
Lip, column and ovary, lateral view.
|
||
<emphasis id="B91AEA86FFA9FFAF6201EC20FDBBF83C" bold="true" box="[524,547,1914,1935]" pageId="3" pageNumber="166">E.</emphasis>
|
||
Column.
|
||
<emphasis id="B91AEA86FFA9FFAF628EEC20FD0EF83C" bold="true" box="[643,662,1914,1935]" pageId="3" pageNumber="166">F.</emphasis>
|
||
Lip, adaxial view.
|
||
<emphasis id="B91AEA86FFA9FFAF635CEC23FCF2F83C" bold="true" box="[849,874,1913,1935]" pageId="3" pageNumber="166">G.</emphasis>
|
||
Anther cap and pollinarium. Illustrated by S. Moreno from the holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF11661CFFAEFFA86085EC0FFE3EF800" ID-DOI="http://doi.org/10.5281/zenodo.10275909" ID-Zenodo-Dep="10275909" httpUri="https://zenodo.org/record/10275909/files/figure.png" pageId="4" pageNumber="167" startId="4.[136,229,1877,1899]" targetBox="[151,1436,190,1853]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BD13694FFAEFFA86085EC0FFE3EF800" blockId="4.[136,1453,1877,1971]" pageId="4" pageNumber="167">
|
||
<emphasis id="B91AEA86FFAEFFA86085EC0FFE98F8D9" bold="true" box="[136,256,1877,1899]" pageId="4" pageNumber="167">FIGURE 3.</emphasis>
|
||
Composite plate of
|
||
<taxonomicName id="4C6E4D17FFAEFFA861C2EC0FFA33F8D8" authority="Ocupa, M. M. Jimenez & Mark Wilson. A. Plant. B. Flower. C. Dissected" box="[463,1451,1877,1899]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="167" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFAEFFA861C2EC0FFD54F8D9" box="[463,716,1877,1898]" italics="true" pageId="4" pageNumber="167">Pleurothallis carpishensis</emphasis>
|
||
Ocupa, M.M.Jiménez & Mark Wilson.
|
||
<emphasis id="B91AEA86FFAEFFA86450EC0FFBECF8D9" bold="true" box="[1117,1140,1877,1898]" pageId="4" pageNumber="167">A.</emphasis>
|
||
Plant.
|
||
<emphasis id="B91AEA86FFAEFFA864B0EC0CFB4BF8D8" bold="true" box="[1213,1235,1878,1899]" pageId="4" pageNumber="167">B.</emphasis>
|
||
Flower.
|
||
<emphasis id="B91AEA86FFAEFFA86523EC0FFADDF8D8" bold="true" box="[1326,1349,1877,1899]" pageId="4" pageNumber="167">C.</emphasis>
|
||
Dissected
|
||
</taxonomicName>
|
||
perianth.
|
||
<emphasis id="B91AEA86FFAEFFA860EFEC20FF61F83C" bold="true" box="[226,249,1914,1935]" pageId="4" pageNumber="167">D.</emphasis>
|
||
Lip, column and ovary, lateral view.
|
||
<emphasis id="B91AEA86FFAEFFA8625AEC20FDF6F83C" bold="true" box="[599,622,1914,1935]" pageId="4" pageNumber="167">E.</emphasis>
|
||
Column.
|
||
<emphasis id="B91AEA86FFAEFFA862C7EC20FD45F83C" bold="true" box="[714,733,1914,1935]" pageId="4" pageNumber="167">F.</emphasis>
|
||
Lip, adaxial (right) and abaxial (left) views.
|
||
<emphasis id="B91AEA86FFAEFFA86489EC23FB05F83C" bold="true" box="[1156,1181,1913,1935]" pageId="4" pageNumber="167">G.</emphasis>
|
||
Anther cap and pollinarium. Prepared by L. Ocupa-Horna.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF11661CFFAFFFA96085EF7AFDFDFBEA" ID-DOI="http://doi.org/10.5281/zenodo.10275911" ID-Zenodo-Dep="10275911" httpUri="https://zenodo.org/record/10275911/files/figure.png" pageId="5" pageNumber="168" startId="5.[136,229,1056,1078]" targetBox="[190,1391,188,1035]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BD13694FFAFFFA96085EF7AFDFDFBEA" blockId="5.[136,1452,1056,1114]" pageId="5" pageNumber="168">
|
||
<emphasis id="B91AEA86FFAFFFA96085EF7AFF66FB86" bold="true" box="[136,254,1056,1078]" pageId="5" pageNumber="168">FIGURE 4.</emphasis>
|
||
Location map of
|
||
<taxonomicName id="4C6E4D17FFAFFFA961A0EF7AFB87FB85" authority="Ocupa, M. M. Jimenez & Mark Wilson" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[429,1055,1056,1078]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFAFFFA961A0EF7AFD30FB86" box="[429,680,1056,1077]" italics="true" pageId="5" pageNumber="168">Pleurothallis carpishensis</emphasis>
|
||
Ocupa, M.M.Jiménez & Mark Wilson
|
||
</taxonomicName>
|
||
in the Carpish Montane Forest Regional Conservation Area. Prepared by L. Ocupa-Horna.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C374651FFFAFFFA960B0EFD8FDC8FB73" pageId="5" pageNumber="168" type="etymology">
|
||
<paragraph id="8BD13694FFAFFFA960B0EFD8FDC8FB73" blockId="5.[136,1452,1154,2080]" pageId="5" pageNumber="168">
|
||
<emphasis id="B91AEA86FFAFFFA960B0EFD8FED0FB2F" bold="true" box="[189,328,1154,1180]" pageId="5" pageNumber="168">Etymology:</emphasis>
|
||
—In reference to the
|
||
<typeStatus id="54D58836FFAFFFA96222EFD8FDC7FB2F" box="[559,607,1154,1180]" pageId="5" pageNumber="168">type</typeStatus>
|
||
locality, the Carpish Montane Forest Regional Conservation Area, Huánuco, Peru, where this species was discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374651FFFAFFFA960B0EF93FE5FFAC7" pageId="5" pageNumber="168" type="conservation">
|
||
<paragraph id="8BD13694FFAFFFA960B0EF93FE5FFAC7" blockId="5.[136,1452,1154,2080]" pageId="5" pageNumber="168">
|
||
<emphasis id="B91AEA86FFAFFFA960B0EF93FE2EFB57" bold="true" box="[189,438,1225,1252]" pageId="5" pageNumber="168">Conservation status:</emphasis>
|
||
—This species is only known from the
|
||
<typeStatus id="54D58836FFAFFFA9638DEF90FC28FB57" box="[896,944,1226,1252]" pageId="5" pageNumber="168">type</typeStatus>
|
||
collection in central Peru.
|
||
<location id="8EB1604FFFAFFFA964E7EF90FA8EFB57" LSID="urn:lsid:plazi:treatment:03C78782FFA8FFA76085E94EFBD8FDDB:8EB1604FFFAFFFA964E7EF90FA8EFB57" box="[1258,1302,1226,1252]" country="Peru" municipality="Carpish Montane Forest" name="The" pageId="5" pageNumber="168">The</location>
|
||
status of the population present in the
|
||
<collectingMunicipality id="6BB5ACEEFFAFFFA961A9EFB4FD2AFABB" box="[420,690,1262,1288]" pageId="5" pageNumber="168">Carpish Montane Forest</collectingMunicipality>
|
||
is unknown; however, the habitat of this species may be affected by anthropogenic activities, such as deforestation and burning of trees, which occur in the area (
|
||
<figureCitation id="13552A11FFAFFFA964AEEE48FA9EFA9F" box="[1187,1286,1298,1324]" captionStart="FIGURE 6" captionStartId="7.[136,229,1011,1033]" captionTargetBox="[227,1360,175,987]" captionTargetId="figure-21@7.[227,1360,175,987]" captionTargetPageId="7" captionText="FIGURE 6. Habitat of Pleurothallis carpishensis Ocupa, M.M.Jiménez & Mark Wilson impacted by deforestation and tree burning.Photo by L. Ocupa-Horna." figureDoi="http://doi.org/10.5281/zenodo.10275915" httpUri="https://zenodo.org/record/10275915/files/figure.png" pageId="5" pageNumber="168">Figure 6</figureCitation>
|
||
). Until a more comprehensive assessment can be made, the species should be considered "Data Deficient" (
|
||
<collectionCode id="ED7FAE51FFAFFFA964B8EE6CFB78FAE3" box="[1205,1248,1334,1360]" pageId="5" pageNumber="168">DD</collectionCode>
|
||
) according to the IUCN criteria (
|
||
<bibRefCitation id="EFFF4B65FFAFFFA9613CEE00FE24FAC7" author="IUCN" box="[305,444,1370,1396]" pageId="5" pageNumber="168" refId="ref4351" refString="IUCN (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. 110 pp. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 20 June 2022)" type="book" year="2019">
|
||
<collectionCode id="ED7FAE51FFAFFFA9613CEE00FEEFFAC7" box="[305,375,1370,1396]" pageId="5" pageNumber="168">IUCN</collectionCode>
|
||
2019
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374651FFFAFFFA760B0EE27FBD8FDDB" lastPageId="11" lastPageNumber="174" pageId="5" pageNumber="168" type="discussion">
|
||
<paragraph id="8BD13694FFAFFFA960B0EE27FB1AF9B7" blockId="5.[136,1452,1154,2080]" pageId="5" pageNumber="168">
|
||
<emphasis id="B91AEA86FFAFFFA960B0EE27FE57FA24" bold="true" box="[189,463,1405,1431]" pageId="5" pageNumber="168">Taxonomic Discussion:</emphasis>
|
||
—
|
||
<taxonomicName id="4C6E4D17FFAFFFA961E6EE24FC97FA2B" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[491,783,1406,1432]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFAFFFA961E6EE24FC97FA2B" box="[491,783,1406,1432]" italics="true" pageId="5" pageNumber="168">Pleurothallis carpishensis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by the pale yellow-to-hyaline flowers, that are borne from a depressed spathaceous bract that lies over a navicular concavity on the leaf, by petals oblong-lanceolate, apiculate, with large papillae on the margins and a triangular, sub-acute lip, with a callus basal, widely ovate, emarginate, waxy, papillose-verrucose, with two oblique, lateral and marginal keels.
|
||
</paragraph>
|
||
<paragraph id="8BD13694FFAFFFA960B0ED54FDE5F8FB" blockId="5.[136,1452,1154,2080]" pageId="5" pageNumber="168">
|
||
Among all the species belonging to
|
||
<taxonomicName id="4C6E4D17FFAFFFA9625FED54FD7BF99B" box="[594,739,1550,1576]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B91AEA86FFAFFFA9625FED54FD7BF99B" box="[594,739,1550,1576]" italics="true" pageId="5" pageNumber="168">Pleurothallis</emphasis>
|
||
</taxonomicName>
|
||
subsection
|
||
<emphasis id="B91AEA86FFAFFFA96365ED54FB06F99B" box="[872,1182,1550,1576]" italics="true" pageId="5" pageNumber="168">Macrophyllae-Fasciculatae</emphasis>
|
||
,
|
||
<taxonomicName id="4C6E4D17FFAFFFA964A1ED54FACCF99B" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[1196,1364,1550,1576]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFAFFFA964A1ED54FACCF99B" box="[1196,1364,1550,1576]" italics="true" pageId="5" pageNumber="168">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
is most similar morphologically to
|
||
<taxonomicName id="4C6E4D17FFAFFFA961BAED68FD85F9FF" authorityName="Luer" authorityYear="1979" box="[439,541,1586,1612]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="species" species="radula">
|
||
<emphasis id="B91AEA86FFAFFFA961BAED68FD85F9FF" box="[439,541,1586,1612]" italics="true" pageId="5" pageNumber="168">P. radula</emphasis>
|
||
</taxonomicName>
|
||
from Costa Rica (
|
||
<figureCitation id="13552A11FFAFFFA962EAED68FCD0F9FF" box="[743,840,1586,1612]" captionStart="FIGURE 7" captionStartId="7.[136,229,2024,2046]" captionTargetBox="[234,1353,1087,2008]" captionTargetId="figure-45@7.[227,1360,1080,2015]" captionTargetPageId="7" captionText="FIGURE 7. Comparison of a morphologically similar species. A–C. Pleurothallis radula. D–F. P. carpishensis. Photos by P.Aratoun (A), R. Parsons (B–C), L. Ocupa-Horna (D–F)." figureDoi="http://doi.org/10.5281/zenodo.10275917" httpUri="https://zenodo.org/record/10275917/files/figure.png" pageId="5" pageNumber="168">Figure 7</figureCitation>
|
||
), sharing characteristics such as the ovate to lanceolate leaves, deeply cordate at the base with the basal lobes sometimes overlapping, the solitary flowers, erect dorsal sepal, reflexed, single-veined petals and the thick and fleshy lip. However,
|
||
<taxonomicName id="4C6E4D17FFAFFFA96392ED20FBD0F927" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[927,1096,1658,1684]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFAFFFA96392ED20FBD0F927" box="[927,1096,1658,1684]" italics="true" pageId="5" pageNumber="168">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
is easily differentiated from
|
||
<taxonomicName id="4C6E4D17FFAFFFA9659AED20FF4BF90B" authorityName="Luer" authorityYear="1979" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="species" species="radula">
|
||
<emphasis id="B91AEA86FFAFFFA9659AED20FF4BF90B" italics="true" pageId="5" pageNumber="168">P. radula</emphasis>
|
||
</taxonomicName>
|
||
by the smaller plants, up to
|
||
<quantity id="4C969B71FFAFFFA96216EDC4FDF9F90B" box="[539,609,1694,1720]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="168" unit="cm" value="23.0">23 cm</quantity>
|
||
tall (
|
||
<emphasis id="B91AEA86FFAFFFA962ADEDC4FD2FF904" box="[672,695,1694,1719]" italics="true" pageId="5" pageNumber="168">vs</emphasis>
|
||
. up to
|
||
<quantity id="4C969B71FFAFFFA96304EDC7FCC8F90B" box="[777,848,1693,1720]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="168" unit="cm" value="50.0">50 cm</quantity>
|
||
tall), ovary 2.7–3.0 mm long (
|
||
<emphasis id="B91AEA86FFAFFFA964BFEDC4FB48F904" box="[1202,1232,1694,1719]" italics="true" pageId="5" pageNumber="168">vs.</emphasis>
|
||
4.0–6.0 mm long), dorsal sepal 8.0–
|
||
<quantity id="4C969B71FFAFFFA9614AED98FE31F96F" box="[327,425,1730,1756]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="5" pageNumber="168" unit="mm" value="10.3">10.3 mm</quantity>
|
||
long (
|
||
<emphasis id="B91AEA86FFAFFFA961FFED98FD88F968" box="[498,528,1730,1755]" italics="true" pageId="5" pageNumber="168">vs.</emphasis>
|
||
<quantity id="4C969B71FFAFFFA9621AED98FD08F96F" box="[535,656,1730,1756]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="5" pageNumber="168" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
|
||
long), synsepal 8.0–10 ×
|
||
<quantity id="4C969B71FFAFFFA963BEED9BFBA2F96F" box="[947,1082,1729,1756]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="5" pageNumber="168" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5–4.8 mm</quantity>
|
||
(
|
||
<emphasis id="B91AEA86FFAFFFA96447ED98FBF0F968" box="[1098,1128,1730,1755]" italics="true" pageId="5" pageNumber="168">vs.</emphasis>
|
||
12.0–14.0 × 7.0 mm), petals 4.5–6.0 mm long (
|
||
<emphasis id="B91AEA86FFAFFFA9615BEDBCFEF5F94C" box="[342,365,1766,1791]" italics="true" pageId="5" pageNumber="168">vs</emphasis>
|
||
. 8.0–
|
||
<quantity id="4C969B71FFAFFFA961A0EDBFFD9AF8B3" box="[429,514,1765,1792]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="168" unit="mm" value="9.5">9.5 mm</quantity>
|
||
long), lip with a callus widely ovate, emarginate, papillose-verrucose, 1.7 ×
|
||
<quantity id="4C969B71FFAFFFA9655BEDBCFA34F8B3" box="[1366,1452,1766,1792]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="168" unit="mm" value="2.3">2.3 mm</quantity>
|
||
(
|
||
<emphasis id="B91AEA86FFAFFFA9609CEC50FF28F890" box="[145,176,1802,1827]" italics="true" pageId="5" pageNumber="168">vs.</emphasis>
|
||
glabrous, callus
|
||
<quantity id="4C969B71FFAFFFA96160EC50FE36F897" box="[365,430,1802,1828]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="168" unit="mm" value="1.0">1 mm</quantity>
|
||
long), without glenion (
|
||
<emphasis id="B91AEA86FFAFFFA962B3EC50FD45F890" box="[702,733,1802,1827]" italics="true" pageId="5" pageNumber="168">vs.</emphasis>
|
||
glenion slightly raised above callus) and the ciliate periphery of the stigmatic surfaces (
|
||
<emphasis id="B91AEA86FFAFFFA96186EC74FE31F8F4" box="[395,425,1838,1863]" italics="true" pageId="5" pageNumber="168">vs.</emphasis>
|
||
slightly papillate).
|
||
</paragraph>
|
||
<paragraph id="8BD13694FFAFFFA960B0EC08FC7FF793" blockId="5.[136,1452,1154,2080]" pageId="5" pageNumber="168">
|
||
As in many species of
|
||
<taxonomicName id="4C6E4D17FFAFFFA961C1EC08FD16F8DF" authorityName="Lindl. ex G.Don" authorityYear="1839" box="[460,654,1874,1900]" class="Liliopsida" family="Orchidaceae" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="subTribe" subTribe="Pleurothallidinae">Pleurothallidinae</taxonomicName>
|
||
, the leaves of
|
||
<taxonomicName id="4C6E4D17FFAFFFA96331EC08FC7FF8DF" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[828,999,1874,1900]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFAFFFA96331EC08FC7FF8DF" box="[828,999,1874,1900]" italics="true" pageId="5" pageNumber="168">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
show vegetative variability. Therefore, some individuals have leaves ranging from ovate to ovate-lanceolate, and occasionally falcate near the apex, with the apex acute to shortly acuminate, with the basal lobes cordate and sometimes overlapping (
|
||
<figureCitation id="13552A11FFAFFFA96482ECC0FAB2F807" box="[1167,1322,1946,1972]" captionStart="FIGURE 8" captionStartId="8.[136,229,1266,1288]" captionTargetBox="[151,1436,190,1241]" captionTargetId="figure-25@8.[151,1436,190,1241]" captionTargetPageId="8" captionText="FIGURE 8. A. Leaf narrowly ovate, with overlapping basal lobes and detail of the concavity (red arrow). B. Ovate leaf, with nonoverlapping basal lobes. C. Leaf, abaxial view and detail of the apex of the ramicaul. D. Detail of the flower above the concavity. Prepared by L. Ocupa-Horna." figureDoi="http://doi.org/10.5281/zenodo.10275919" httpUri="https://zenodo.org/record/10275919/files/figure.png" pageId="5" pageNumber="168">
|
||
Figure 8
|
||
<collectionCode id="ED7FAE51FFAFFFA964FDECC0FA9FF807" box="[1264,1287,1946,1972]" pageId="5" pageNumber="168">A</collectionCode>
|
||
–
|
||
<collectionCode id="ED7FAE51FFAFFFA9651AECC0FAB2F807" box="[1303,1322,1946,1972]" pageId="5" pageNumber="168">B</collectionCode>
|
||
</figureCitation>
|
||
). However, a peculiar characteristic in
|
||
<taxonomicName id="4C6E4D17FFAFFFA961CDECE4FDF2F86B" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[448,618,1982,2008]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="168" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFAFFFA961CDECE4FDF2F86B" box="[448,618,1982,2008]" italics="true" pageId="5" pageNumber="168">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
are the apices of the ramicaul, since these are curved, forming an angle of approximately 90°, before the leaf abscission layer (
|
||
<figureCitation id="13552A11FFAFFFA96305ECB8FC18F84F" box="[776,896,2018,2044]" captionStart="FIGURE 8" captionStartId="8.[136,229,1266,1288]" captionTargetBox="[151,1436,190,1241]" captionTargetId="figure-25@8.[151,1436,190,1241]" captionTargetPageId="8" captionText="FIGURE 8. A. Leaf narrowly ovate, with overlapping basal lobes and detail of the concavity (red arrow). B. Ovate leaf, with nonoverlapping basal lobes. C. Leaf, abaxial view and detail of the apex of the ramicaul. D. Detail of the flower above the concavity. Prepared by L. Ocupa-Horna." figureDoi="http://doi.org/10.5281/zenodo.10275919" httpUri="https://zenodo.org/record/10275919/files/figure.png" pageId="5" pageNumber="168">
|
||
Figure 8
|
||
<collectionCode id="ED7FAE51FFAFFFA96366ECB8FC18F84F" box="[875,896,2018,2044]" pageId="5" pageNumber="168">C</collectionCode>
|
||
</figureCitation>
|
||
), unlike other
|
||
<taxonomicName id="4C6E4D17FFAFFFA96420ECB8FA81F84F" box="[1069,1305,2018,2044]" pageId="5" pageNumber="168">
|
||
<emphasis id="B91AEA86FFAFFFA96420ECB8FB26F84F" box="[1069,1214,2018,2044]" italics="true" pageId="5" pageNumber="168">Pleurothallis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
of the same subsection, which generally have leaves attached to a non-angulate ramicaul.
|
||
</paragraph>
|
||
<caption id="DF11661CFFACFFAA6085ECB6FD35F795" ID-DOI="http://doi.org/10.5281/zenodo.10275913" ID-Zenodo-Dep="10275913" httpUri="https://zenodo.org/record/10275913/files/figure.png" pageId="6" pageNumber="169" startId="6.[136,229,2028,2050]" targetBox="[163,1424,190,2004]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8BD13694FFACFFAA6085ECB6FD35F795" blockId="6.[136,1453,2028,2087]" pageId="6" pageNumber="169">
|
||
<emphasis id="B91AEA86FFACFFAA6085ECB6FE98F7B1" bold="true" box="[136,256,2028,2050]" pageId="6" pageNumber="169">FIGURE 5.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="4C6E4D17FFACFFAA6178ECB7FB81F7B2" authority="Ocupa, M. M. Jimenez & Mark Wilson. A." authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[373,1049,2028,2051]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="169" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFACFFAA6178ECB7FDEAF7B1" box="[373,626,2029,2050]" italics="true" pageId="6" pageNumber="169">Pleurothallis carpishensis</emphasis>
|
||
Ocupa, M.M.Jiménez & Mark Wilson.
|
||
<emphasis id="B91AEA86FFACFFAA640CECB6FB81F7B2" bold="true" box="[1025,1049,2028,2049]" pageId="6" pageNumber="169">A.</emphasis>
|
||
</taxonomicName>
|
||
Close-up to tree canopy branches.
|
||
<emphasis id="B91AEA86FFACFFAA6576ECB7FA09F7B1" bold="true" box="[1403,1425,2029,2050]" pageId="6" pageNumber="169">B.</emphasis>
|
||
<taxonomicName id="4C6E4D17FFACFFAA6597ECB7FE98F795" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="169" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFACFFAA6597ECB7FE98F795" italics="true" pageId="6" pageNumber="169">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
growing
|
||
<emphasis id="B91AEA86FFACFFAA6151E34BFE00F795" box="[348,408,2065,2086]" italics="true" pageId="6" pageNumber="169">in situ</emphasis>
|
||
. Photos by L. Ocupa-Horna.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF11661CFFADFFAB6085E8A9FED3FB9E" ID-DOI="http://doi.org/10.5281/zenodo.10275915" ID-Zenodo-Dep="10275915" httpUri="https://zenodo.org/record/10275915/files/figure.png" pageId="7" pageNumber="170" startId="7.[136,229,1011,1033]" targetBox="[227,1360,175,987]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8BD13694FFADFFAB6085E8A9FED3FB9E" blockId="7.[136,1451,1011,1070]" pageId="7" pageNumber="170">
|
||
<emphasis id="B91AEA86FFADFFAB6085E8A9FF64FBBB" bold="true" box="[136,252,1011,1033]" pageId="7" pageNumber="170">FIGURE 6.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="4C6E4D17FFADFFAB616BE8AEFC54FBB9" authority="Ocupa, M. M. Jimenez & Mark Wilson" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[358,972,1012,1034]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="170" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFADFFAB616BE8AEFDC7FBBA" box="[358,607,1012,1033]" italics="true" pageId="7" pageNumber="170">Pleurothallis carpishensis</emphasis>
|
||
Ocupa, M.M.Jiménez & Mark Wilson
|
||
</taxonomicName>
|
||
impacted by deforestation and tree burning. Photo by L. Ocupa-Horna.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF11661CFFADFFAB6085ECB2FDB9F791" ID-DOI="http://doi.org/10.5281/zenodo.10275917" ID-Zenodo-Dep="10275917" httpUri="https://zenodo.org/record/10275917/files/figure.png" pageId="7" pageNumber="170" startId="7.[136,229,2024,2046]" targetBox="[234,1353,1087,2008]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8BD13694FFADFFAB6085ECB2FDB9F791" blockId="7.[136,1451,2024,2082]" pageId="7" pageNumber="170">
|
||
<emphasis id="B91AEA86FFADFFAB6085ECB2FF65F84D" bold="true" box="[136,253,2024,2046]" pageId="7" pageNumber="170">FIGURE 7.</emphasis>
|
||
Comparison of a morphologically similar species.
|
||
<emphasis id="B91AEA86FFADFFAB62ECECB2FC97F84D" bold="true" box="[737,783,2024,2046]" pageId="7" pageNumber="170">A–C</emphasis>
|
||
.
|
||
<taxonomicName id="4C6E4D17FFADFFAB6316ECB3FC44F84D" authorityName="Luer" authorityYear="1979" box="[795,988,2025,2046]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="170" phylum="Tracheophyta" rank="species" species="radula">
|
||
<emphasis id="B91AEA86FFADFFAB6316ECB3FC44F84D" box="[795,988,2025,2046]" italics="true" pageId="7" pageNumber="170">Pleurothallis radula</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B91AEA86FFADFFAB63EAECB3FB81F84D" bold="true" box="[999,1049,2025,2046]" pageId="7" pageNumber="170">D–F.</emphasis>
|
||
<taxonomicName id="4C6E4D17FFADFFAB6413ECB3FB35F84D" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[1054,1197,2025,2046]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="170" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFADFFAB6413ECB3FB35F84D" box="[1054,1197,2025,2046]" italics="true" pageId="7" pageNumber="170">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
. Photos by P.Aratoun (A), R. Parsons (B
|
||
<emphasis id="B91AEA86FFADFFAB6100E357FE81F791" bold="true" box="[269,281,2061,2082]" pageId="7" pageNumber="170">–</emphasis>
|
||
C), L. Ocupa-Horna (D
|
||
<emphasis id="B91AEA86FFADFFAB61F4E357FD9DF791" bold="true" box="[505,517,2061,2082]" pageId="7" pageNumber="170">–</emphasis>
|
||
F).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF11661CFFA2FFA46085EFA8FED3FAFC" ID-DOI="http://doi.org/10.5281/zenodo.10275919" ID-Zenodo-Dep="10275919" httpUri="https://zenodo.org/record/10275919/files/figure.png" pageId="8" pageNumber="171" startId="8.[136,229,1266,1288]" targetBox="[151,1436,190,1241]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BD13694FFA2FFA46085EFA8FED3FAFC" blockId="8.[136,1451,1266,1360]" pageId="8" pageNumber="171">
|
||
<emphasis id="B91AEA86FFA2FFA46085EFA8FEB8FAB4" bold="true" box="[136,288,1266,1288]" pageId="8" pageNumber="171">FIGURE 8. A.</emphasis>
|
||
Leaf narrowly ovate, with overlapping basal lobes and detail of the concavity (red arrow).
|
||
<emphasis id="B91AEA86FFA2FFA464BAEFA8FB56FAB4" bold="true" box="[1207,1230,1266,1287]" pageId="8" pageNumber="171">B.</emphasis>
|
||
Ovate leaf, with nonoverlapping basal lobes.
|
||
<emphasis id="B91AEA86FFA2FFA46179EE4CFE14FA9F" bold="true" box="[372,396,1302,1324]" pageId="8" pageNumber="171">C.</emphasis>
|
||
Leaf, abaxial view and detail of the apex of the ramicaul.
|
||
<emphasis id="B91AEA86FFA2FFA463BFEE4CFC5BFA98" bold="true" box="[946,963,1302,1323]" pageId="8" pageNumber="171">D</emphasis>
|
||
. Detail of the flower above the concavity. Prepared by L. Ocupa-Horna.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD13694FFA2FFA460B0EE24FD17F9C2" blockId="8.[136,1452,1406,2009]" pageId="8" pageNumber="171">
|
||
Interestingly, in some species of subsection
|
||
<emphasis id="B91AEA86FFA2FFA462BBEE24FC74FA2B" box="[694,1004,1406,1432]" italics="true" pageId="8" pageNumber="171">Macrophyllae-Fasciculatae</emphasis>
|
||
, the single-flowered inflorescences are borne from an erect spathaceous bract (
|
||
<bibRefCitation id="EFFF4B65FFA2FFA4625DEEF8FCB7FA0F" author="Pupulin, F. & Aguilar, J. & Belfort-Oconitrillo, N. & Diaz-Morales, M. & Bogarin, D." box="[592,815,1442,1468]" pageId="8" pageNumber="171" pagination="203 - 295" refId="ref4961" refString="Pupulin, F., Aguilar, J., Belfort-Oconitrillo, N., Diaz-Morales, M. & Bogarin, D. (2021) Florae Costaricensis Subtribui Pleurothallidinis (orchidaceae) Prodromus II. Systematics of the Pleurothallis cardiothallis and P. phyllocardia groups, and other related groups of Pleurothallis with large vegetative habit. Harvard Papers in Botany 26: 203 - 295. https: // doi. org / 10.3100 / hpib. v 26 iss 1.2021. n 14" type="journal article" year="2021">
|
||
Pupulin
|
||
<emphasis id="B91AEA86FFA2FFA462BFEEF9FD75FA0F" box="[690,749,1442,1468]" italics="true" pageId="8" pageNumber="171">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
) [e.g.
|
||
<taxonomicName id="4C6E4D17FFA2FFA46370EEF9FA10FA0E" authority="Bogarin & Belfort (2021: 265)" authorityName="Bogarin & Belfort" authorityPageNumber="265" authorityYear="2021" box="[893,1416,1442,1469]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="longipetala">
|
||
<emphasis id="B91AEA86FFA2FFA46370EEF9FB83FA0F" box="[893,1051,1442,1468]" italics="true" pageId="8" pageNumber="171">P. longipetala</emphasis>
|
||
Bogarín & Belfort (2021: 265)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6E4D17FFA2FFA4659AEEF9FDA8FA53" authority="Reichenbach (1866: 97)" authorityName="Reichenbach" authorityPageNumber="97" authorityYear="1866" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="phyllocardia">
|
||
<emphasis id="B91AEA86FFA2FFA4659AEEF9FE8EFA53" italics="true" pageId="8" pageNumber="171">P. phyllocardia</emphasis>
|
||
<bibRefCitation id="EFFF4B65FFA2FFA46113EE9CFDA8FA53" author="Reichenbach, H. G." box="[286,560,1478,1504]" pageId="8" pageNumber="171" refId="ref5048" refString="Reichenbach, H. G. (1866) Beitrage zu einer Orchideenkunde Central-Amerika's. Druck von T. G. Meissner, Hamburg. 117 pp. https: // doi. org / 10.5962 / bhl. title. 60976" type="book" year="1866">Reichenbach (1866: 97)</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6E4D17FFA2FFA46231EE9DFD3BFA53" authorityName="Luer" authorityYear="1979" box="[572,675,1478,1504]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="radula">
|
||
<emphasis id="B91AEA86FFA2FFA46231EE9DFD3BFA53" box="[572,675,1478,1504]" italics="true" pageId="8" pageNumber="171">P. radula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6E4D17FFA2FFA462BFEE9DFB59FA53" authority="Ames & C. Schweinf. (1925: 32)" authorityName="Ames & C. Schweinf." authorityPageNumber="32" authorityYear="1925" box="[690,1217,1478,1505]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="rectipetala">
|
||
<emphasis id="B91AEA86FFA2FFA462BFEE9DFCDEFA53" box="[690,838,1478,1504]" italics="true" pageId="8" pageNumber="171">P. rectipetala</emphasis>
|
||
Ames & C.Schweinf. (1925: 32)
|
||
</taxonomicName>
|
||
], that emerge from a horizontal to erect leaf or leaf deflexed toward the ramicaul. Nevertheless, in
|
||
<taxonomicName id="4C6E4D17FFA2FFA463E1EEB1FB0CF9B7" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[1004,1172,1514,1540]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA2FFA463E1EEB1FB0CF9B7" box="[1004,1172,1514,1540]" italics="true" pageId="8" pageNumber="171">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
the flower is borne from a depressed spathaceous bract that lies over a navicular concavity on the leaf (
|
||
<figureCitation id="13552A11FFA2FFA46406ED54FBF6F99B" box="[1035,1134,1550,1576]" captionStart="FIGURE 8" captionStartId="8.[136,229,1266,1288]" captionTargetBox="[151,1436,190,1241]" captionTargetId="figure-25@8.[151,1436,190,1241]" captionTargetPageId="8" captionText="FIGURE 8. A. Leaf narrowly ovate, with overlapping basal lobes and detail of the concavity (red arrow). B. Ovate leaf, with nonoverlapping basal lobes. C. Leaf, abaxial view and detail of the apex of the ramicaul. D. Detail of the flower above the concavity. Prepared by L. Ocupa-Horna." figureDoi="http://doi.org/10.5281/zenodo.10275919" httpUri="https://zenodo.org/record/10275919/files/figure.png" pageId="8" pageNumber="171">Figure 8</figureCitation>
|
||
, red arrow). This concavity is as long as the flower which overlies it (
|
||
<figureCitation id="13552A11FFA2FFA46261ED68FD7DF9FF" box="[620,741,1586,1612]" captionStart="FIGURE 8" captionStartId="8.[136,229,1266,1288]" captionTargetBox="[151,1436,190,1241]" captionTargetId="figure-25@8.[151,1436,190,1241]" captionTargetPageId="8" captionText="FIGURE 8. A. Leaf narrowly ovate, with overlapping basal lobes and detail of the concavity (red arrow). B. Ovate leaf, with nonoverlapping basal lobes. C. Leaf, abaxial view and detail of the apex of the ramicaul. D. Detail of the flower above the concavity. Prepared by L. Ocupa-Horna." figureDoi="http://doi.org/10.5281/zenodo.10275919" httpUri="https://zenodo.org/record/10275919/files/figure.png" pageId="8" pageNumber="171">
|
||
Figure 8
|
||
<collectionCode id="ED7FAE51FFA2FFA462C0ED68FD7DF9FF" box="[717,741,1586,1612]" pageId="8" pageNumber="171">D</collectionCode>
|
||
</figureCitation>
|
||
). This makes the species unmistakable, compared to the other
|
||
<emphasis id="B91AEA86FFA2FFA46085ED0CFE26F9C3" box="[136,446,1622,1648]" italics="true" pageId="8" pageNumber="171">Macrophyllae-Fasciculatae</emphasis>
|
||
occurring in Peru.
|
||
</paragraph>
|
||
<paragraph id="8BD13694FFA2FFA460B0ED20FD4EF86B" blockId="8.[136,1452,1406,2009]" pageId="8" pageNumber="171">
|
||
Since
|
||
<taxonomicName id="4C6E4D17FFA2FFA4610BED21FEE8F927" authorityName="Luer" authorityYear="1979" box="[262,368,1658,1684]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="radula">
|
||
<emphasis id="B91AEA86FFA2FFA4610BED21FEE8F927" box="[262,368,1658,1684]" italics="true" pageId="8" pageNumber="171">P. radula</emphasis>
|
||
</taxonomicName>
|
||
is not found in Peru, field botanists in that country need to distinguish
|
||
<taxonomicName id="4C6E4D17FFA2FFA464B2ED21FAF2F927" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[1215,1386,1658,1684]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA2FFA464B2ED21FAF2F927" box="[1215,1386,1658,1684]" italics="true" pageId="8" pageNumber="171">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
from the most morphologically similar species to be encountered in that country, in this case
|
||
<taxonomicName id="4C6E4D17FFA2FFA46489EDC5FA92F90B" box="[1156,1290,1694,1720]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="crucifera">
|
||
<emphasis id="B91AEA86FFA2FFA46489EDC5FA92F90B" box="[1156,1290,1694,1720]" italics="true" pageId="8" pageNumber="171">P. crucifera</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C6E4D17FFA2FFA46517EDC4FE8CF96F" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA2FFA46517EDC4FE8CF96F" italics="true" pageId="8" pageNumber="171">Pleurothallis carpishensis</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C6E4D17FFA2FFA461B8ED99FDA4F96F" box="[437,572,1730,1756]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="crucifera">
|
||
<emphasis id="B91AEA86FFA2FFA461B8ED99FDA4F96F" box="[437,572,1730,1756]" italics="true" pageId="8" pageNumber="171">P. crucifera</emphasis>
|
||
</taxonomicName>
|
||
in the coriaceous, ovate, acute leaves, the yellow-brown, glabrous flowers, that rest on the leaf and the long, acute petals (
|
||
<figureCitation id="13552A11FFA2FFA462AEEDBCFC9FF8B3" box="[675,775,1766,1792]" captionStart="FIGURE 9" captionStartId="9.[136,229,1771,1793]" captionTargetBox="[161,1427,198,1737]" captionTargetId="figure-21@9.[151,1436,190,1747]" captionTargetPageId="9" captionText="FIGURE 9. Comparison of morphologically similar species. A–B. P. carpishensis Ocupa, M.M.Jiménez & Mark Wilson. C–D. Pleurothallis crucifera Luer & Hirtz. Photos by L. Ocupa-Horna (A–B), H. Rice (C–D)." figureDoi="http://doi.org/10.5281/zenodo.10275921" httpUri="https://zenodo.org/record/10275921/files/figure.png" pageId="8" pageNumber="171">Figure 9</figureCitation>
|
||
). However,
|
||
<taxonomicName id="4C6E4D17FFA2FFA46399EDBDFBA5F8B3" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[916,1085,1766,1792]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA2FFA46399EDBDFBA5F8B3" box="[916,1085,1766,1792]" italics="true" pageId="8" pageNumber="171">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C6E4D17FFA2FFA464DAEDBDFAC4F8B3" box="[1239,1372,1766,1792]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="171" phylum="Tracheophyta" rank="species" species="crucifera">
|
||
<emphasis id="B91AEA86FFA2FFA464DAEDBDFAC4F8B3" box="[1239,1372,1766,1792]" italics="true" pageId="8" pageNumber="171">P. crucifera</emphasis>
|
||
</taxonomicName>
|
||
by the length of the ramicauls (8.0–
|
||
<quantity id="4C969B71FFA2FFA461DCEC50FDB6F896" box="[465,558,1802,1829]" metricMagnitude="-1" metricUnit="m" metricValue="2.06" pageId="8" pageNumber="171" unit="cm" value="20.6">20.6 cm</quantity>
|
||
long
|
||
<emphasis id="B91AEA86FFA2FFA4627FEC51FD08F897" box="[626,656,1803,1828]" italics="true" pageId="8" pageNumber="171">vs.</emphasis>
|
||
5.0–10.0 cm long), arrangement and direction of the leaves (reflexed
|
||
<emphasis id="B91AEA86FFA2FFA46085EC75FF3FF8FB" box="[136,167,1839,1864]" italics="true" pageId="8" pageNumber="171">vs.</emphasis>
|
||
spreading), with attenuate, tridentate (
|
||
<emphasis id="B91AEA86FFA2FFA4625AEC75FDEDF8FB" box="[599,629,1839,1864]" italics="true" pageId="8" pageNumber="171">vs.</emphasis>
|
||
acute) apex, the floral bracts long-conical in shape, 3.5–5.0 mm long (
|
||
<emphasis id="B91AEA86FFA2FFA46580EC75FA33F8FB" box="[1421,1451,1839,1864]" italics="true" pageId="8" pageNumber="171">vs.</emphasis>
|
||
tubular, 5.0–6.0 mm long), the pedicels 4.0–5.0 mm long (
|
||
<emphasis id="B91AEA86FFA2FFA4631BEC09FCACF8DF" box="[790,820,1875,1900]" italics="true" pageId="8" pageNumber="171">vs.</emphasis>
|
||
6.0–8.0 mm long), the flowers hyaline, yellow concolor (
|
||
<emphasis id="B91AEA86FFA2FFA4609CEC2DFF30F823" box="[145,168,1911,1936]" italics="true" pageId="8" pageNumber="171">vs</emphasis>
|
||
. light yellowish or brown flowers), arrangement and direction of the petals (porrect
|
||
<emphasis id="B91AEA86FFA2FFA46450EC2DFBE4F823" box="[1117,1148,1911,1936]" italics="true" pageId="8" pageNumber="171">vs.</emphasis>
|
||
spreading), the lip without glenion (
|
||
<emphasis id="B91AEA86FFA2FFA460E3ECC1FE95F807" box="[238,269,1947,1972]" italics="true" pageId="8" pageNumber="171">vs.</emphasis>
|
||
large glenion), and the stigma ventral, reniform (
|
||
<emphasis id="B91AEA86FFA2FFA46348ECC1FCFCF807" box="[837,868,1947,1972]" italics="true" pageId="8" pageNumber="171">vs.</emphasis>
|
||
apical, sub-rectangular), with margins of stigmatic surface ciliate margins (
|
||
<emphasis id="B91AEA86FFA2FFA46198ECE5FE34F86B" box="[405,428,1983,2008]" italics="true" pageId="8" pageNumber="171">vs</emphasis>
|
||
. papillose-erose margins).
|
||
</paragraph>
|
||
<caption id="DF11661CFFA3FFA56085EDB1FC4FF896" ID-DOI="http://doi.org/10.5281/zenodo.10275921" ID-Zenodo-Dep="10275921" httpUri="https://zenodo.org/record/10275921/files/figure.png" pageId="9" pageNumber="172" startId="9.[136,229,1771,1793]" targetBox="[161,1427,198,1737]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8BD13694FFA3FFA56085EDB1FC4FF896" blockId="9.[136,1451,1771,1830]" pageId="9" pageNumber="172">
|
||
<emphasis id="B91AEA86FFA3FFA56085EDB1FE9BF8B2" bold="true" box="[136,259,1771,1793]" pageId="9" pageNumber="172">FIGURE 9.</emphasis>
|
||
Comparison of morphologically similar species.
|
||
<emphasis id="B91AEA86FFA3FFA5630DEDB1FCABF8B3" bold="true" box="[768,819,1771,1792]" pageId="9" pageNumber="172">A–B.</emphasis>
|
||
<taxonomicName id="4C6E4D17FFA3FFA5634DEDB6FA10F8B2" authority="Ocupa, M. M. Jimenez & Mark Wilson. C" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[832,1416,1771,1794]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="172" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA3FFA5634DEDB6FC4DF8B2" box="[832,981,1772,1793]" italics="true" pageId="9" pageNumber="172">P. carpishensis</emphasis>
|
||
Ocupa, M.M.Jiménez & Mark Wilson.
|
||
<emphasis id="B91AEA86FFA3FFA5657AEDB1FA10F8B2" bold="true" box="[1399,1416,1771,1793]" pageId="9" pageNumber="172">C</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91AEA86FFA3FFA56585EDB1FA33F8B2" bold="true" box="[1416,1451,1771,1793]" pageId="9" pageNumber="172">–D.</emphasis>
|
||
<taxonomicName id="4C6E4D17FFA3FFA56085EC4AFDABF895" authority="Luer & Hirtz. Photos" authorityName="Luer & Hirtz. Photos" box="[136,563,1808,1830]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="172" phylum="Tracheophyta" rank="species" species="crucifera">
|
||
<emphasis id="B91AEA86FFA3FFA56085EC4AFEFBF896" box="[136,355,1808,1829]" italics="true" pageId="9" pageNumber="172">Pleurothallis crucifera</emphasis>
|
||
Luer & Hirtz. Photos
|
||
</taxonomicName>
|
||
by L. Ocupa-Horna (A
|
||
<emphasis id="B91AEA86FFA3FFA56319EC4AFCB8F896" bold="true" box="[788,800,1808,1829]" pageId="9" pageNumber="172">–</emphasis>
|
||
B), H. Rice (C
|
||
<emphasis id="B91AEA86FFA3FFA563A6EC4AFC2FF896" bold="true" box="[939,951,1808,1829]" pageId="9" pageNumber="172">–</emphasis>
|
||
D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF11661CFFA0FFA66085EDB1FB39F896" ID-DOI="http://doi.org/10.5281/zenodo.10275923" ID-Zenodo-Dep="10275923" httpUri="https://zenodo.org/record/10275923/files/figure.png" pageId="10" pageNumber="173" startId="10.[136,229,1771,1793]" targetBox="[161,1427,198,1737]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8BD13694FFA0FFA66085EDB1FB39F896" blockId="10.[136,1451,1771,1830]" pageId="10" pageNumber="173">
|
||
<emphasis id="B91AEA86FFA0FFA66085EDB1FEDDF8B2" bold="true" box="[136,325,1771,1793]" pageId="10" pageNumber="173">FIGURE 10. A–C.</emphasis>
|
||
Flowers of
|
||
<taxonomicName id="4C6E4D17FFA0FFA661B3EDB6FCB2F8B2" authority="Schltr. D." authorityName="Schltr. D." box="[446,810,1772,1794]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="10" pageNumber="173" phylum="Tracheophyta" rank="species" species="rhopalocarpa">
|
||
<emphasis id="B91AEA86FFA0FFA661B3EDB6FD5DF8B2" box="[446,709,1772,1793]" italics="true" pageId="10" pageNumber="173">Pleurothallis rhopalocarpa</emphasis>
|
||
Schltr.
|
||
<emphasis id="B91AEA86FFA0FFA6631FEDB6FCB2F8B2" bold="true" box="[786,810,1772,1793]" pageId="10" pageNumber="173">D.</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C6E4D17FFA0FFA6633CEDB6FAA3F8B1" authority="Ocupa, M. M. Jimenez & Mark Wilson" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[817,1339,1772,1794]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="10" pageNumber="173" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA0FFA6633CEDB6FC5AF8B2" box="[817,962,1772,1793]" italics="true" pageId="10" pageNumber="173">P. carpishensis</emphasis>
|
||
Ocupa, M.M.Jiménez & Mark Wilson
|
||
</taxonomicName>
|
||
illustrating absence of glenion. Photos by A. Kay † (A), F. Lopez-Machado (B), J. Varigos (C), and L. Ocupa-Horna (D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD13694FFA1FFA760B0EBC4FBD8FDDB" blockId="11.[136,1452,158,616]" pageId="11" pageNumber="174">
|
||
Within
|
||
<taxonomicName id="4C6E4D17FFA1FFA7611CEBC4FE3AFF0B" box="[273,418,158,184]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="174" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B91AEA86FFA1FFA7611CEBC4FE3AFF0B" box="[273,418,158,184]" italics="true" pageId="11" pageNumber="174">Pleurothallis</emphasis>
|
||
</taxonomicName>
|
||
subsection
|
||
<emphasis id="B91AEA86FFA1FFA7622BEBC4FCC4FF0B" box="[550,860,158,184]" italics="true" pageId="11" pageNumber="174">Macrophyllae-Fasciculatae</emphasis>
|
||
, most species exhibit a glenion at the base of the lip, located below the apically located viscidium of the pollinarium. Morphologically, the glenion appears to be a small secretory structure (
|
||
<bibRefCitation id="EFFF4B65FFA1FFA76161EBBCFDDFFEB3" author="Wilson, M. & Baquero, L. & Dupree, K. & Jimenez, M. & LeBlanc, C. & Merino, G. & Portilla, J. & Salas Guerrero, M. & Tobar Suarez, F. & Werner, J." box="[364,583,230,257]" pageId="11" pageNumber="174" pagination="349 - 366" refId="ref5542" refString="Wilson, M., Baquero, L., Dupree, K., Jimenez, M., LeBlanc, C., Merino, G., Portilla, J., Salas Guerrero, M., Tobar Suarez, F. & Werner, J. (2016) Three new species of Pleurothallis (Orchidaceae: Pleurothallidinae) in subsection Macrophyllae-Fasciculatae from northern South America. Lankesteriana 16: 349 - 366. https: // doi. org / 10.15517 / lank. v 16 i 3.27314" type="journal article" year="2016">
|
||
Wilson
|
||
<emphasis id="B91AEA86FFA1FFA761CAEBBDFE63FEB3" box="[455,507,230,256]" italics="true" pageId="11" pageNumber="174">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFF4B65FFA1FFA76259EBBCFCB5FEB3" author="Wilson, M. & Larsen, B. & Moreno, J. S. & Ward, R. & Riksen, J. A. G. & Pina, L. & Sierra-Ariza, M. A. & Jimenez, M. M. & Rincon-Gonzalez, M. & Galindo-Tarazona, R. & Garzon Suarez, H. & Haelterman, D." box="[596,813,230,257]" pageId="11" pageNumber="174" pagination="187 - 220" refId="ref5636" refString="Wilson, M., Larsen, B., Moreno, J. S., Ward, R., Riksen, J. A. G., Pina, L., Sierra-Ariza, M. A., Jimenez, M. M., Rincon-Gonzalez, M., Galindo-Tarazona, R., Garzon Suarez, H. & Haelterman, D. (2022) New species of Pleurothallis (Orchidaceae: Pleurothallidinae), a new country record, and labellar morphology in the P. cardiostola-P. lilijae complex of subsection Macrophyllae-Fasciculatae. Harvard Papers in Botany 27: 187 - 220. https: // doi. org / 10.3100 / hpib. v 27 iss 2.2022. n 10" type="journal article" year="2022">
|
||
Wilson
|
||
<emphasis id="B91AEA86FFA1FFA762A3EBBDFD7AFEB3" box="[686,738,230,256]" italics="true" pageId="11" pageNumber="174">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
) and we hypothesize that the glenion aids in pollinator positioning for optimal pollinarium acquisition and subsequent deposition on another flower. Yet surprisingly some species, such as
|
||
<taxonomicName id="4C6E4D17FFA1FFA76132EA75FE7EFEFB" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[319,486,302,328]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="174" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA1FFA76132EA75FE7EFEFB" box="[319,486,302,328]" italics="true" pageId="11" pageNumber="174">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6E4D17FFA1FFA761F9EA75FD08FEFB" authorityName="Mark Wilson, Tobar & A. J. Perez" authorityYear="2018" box="[500,656,302,328]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="174" phylum="Tracheophyta" rank="species" species="minutilabia">
|
||
<emphasis id="B91AEA86FFA1FFA761F9EA75FD08FEFB" box="[500,656,302,328]" italics="true" pageId="11" pageNumber="174">P. minutilabia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6E4D17FFA1FFA76293EA75FCCAFEFB" box="[670,850,302,328]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="174" phylum="Tracheophyta" rank="species" species="rhopalocarpa">
|
||
<emphasis id="B91AEA86FFA1FFA76293EA75FCCAFEFB" box="[670,850,302,328]" italics="true" pageId="11" pageNumber="174">P. rhopalocarpa</emphasis>
|
||
</taxonomicName>
|
||
, and the
|
||
<taxonomicName id="4C6E4D17FFA1FFA763B5EA75FBFFFEFB" authorityName="C. Schweinfurth" authorityYear="1951" box="[952,1127,302,328]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="174" phylum="Tracheophyta" rank="species" species="crateriformis">
|
||
<emphasis id="B91AEA86FFA1FFA763B5EA75FBFFFEFB" box="[952,1127,302,328]" italics="true" pageId="11" pageNumber="174">P. crateriformis</emphasis>
|
||
</taxonomicName>
|
||
complex appear to lack such structure. In
|
||
<taxonomicName id="4C6E4D17FFA1FFA76110EA09FE4DFEDF" box="[285,469,338,364]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="174" phylum="Tracheophyta" rank="species" species="rhopalocarpa">
|
||
<emphasis id="B91AEA86FFA1FFA76110EA09FE4DFEDF" box="[285,469,338,364]" italics="true" pageId="11" pageNumber="174">P. rhopalocarpa</emphasis>
|
||
</taxonomicName>
|
||
, a very widely distributed species, the lateral margins produce copious quantities of liquid (
|
||
<figureCitation id="13552A11FFA1FFA760D5EA2CFEE5FE23" box="[216,381,374,400]" captionStart="FIGURE 10" captionStartId="10.[136,229,1771,1793]" captionTargetBox="[161,1427,198,1737]" captionTargetId="figure-25@10.[151,1436,190,1747]" captionTargetPageId="10" captionText="FIGURE 10. A–C. Flowers of Pleurothallis rhopalocarpa Schltr. D. P. carpishensis Ocupa, M.M.Jiménez & Mark Wilson illustrating absence of glenion. Photos by A. Kay † (A), F. Lopez-Machado (B), J. Varigos (C), and L. Ocupa-Horna (D)." figureDoi="http://doi.org/10.5281/zenodo.10275923" httpUri="https://zenodo.org/record/10275923/files/figure.png" pageId="11" pageNumber="174">
|
||
Figure 10
|
||
<collectionCode id="ED7FAE51FFA1FFA76148EA2CFEC2FE23" box="[325,346,374,400]" pageId="11" pageNumber="174">A</collectionCode>
|
||
–
|
||
<collectionCode id="ED7FAE51FFA1FFA76167EA2CFEE5FE23" box="[362,381,374,400]" pageId="11" pageNumber="174">C</collectionCode>
|
||
</figureCitation>
|
||
) that is presumably a nectar reward, and this species may have evolved to permit more flexible pollinator positioning during pollinarium acquisition than occurs in species with a glenion. On the other hand,
|
||
<taxonomicName id="4C6E4D17FFA1FFA7659AEAC1FE8CFE6B" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="174" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA1FFA7659AEAC1FE8CFE6B" italics="true" pageId="11" pageNumber="174">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
neither possesses a glenion nor secretes nectar-like liquid from the medial and apical regions of the lip (
|
||
<figureCitation id="13552A11FFA1FFA7609DEAB8FE8BFE4F" box="[144,275,482,508]" captionStart="FIGURE 10" captionStartId="10.[136,229,1771,1793]" captionTargetBox="[161,1427,198,1737]" captionTargetId="figure-25@10.[151,1436,190,1747]" captionTargetPageId="10" captionText="FIGURE 10. A–C. Flowers of Pleurothallis rhopalocarpa Schltr. D. P. carpishensis Ocupa, M.M.Jiménez & Mark Wilson illustrating absence of glenion. Photos by A. Kay † (A), F. Lopez-Machado (B), J. Varigos (C), and L. Ocupa-Horna (D)." figureDoi="http://doi.org/10.5281/zenodo.10275923" httpUri="https://zenodo.org/record/10275923/files/figure.png" pageId="11" pageNumber="174">
|
||
Figure 10
|
||
<collectionCode id="ED7FAE51FFA1FFA760F0EAB8FE8BFE4F" box="[253,275,482,508]" pageId="11" pageNumber="174">D</collectionCode>
|
||
</figureCitation>
|
||
), suggesting a unique pollination strategy yet to be elucidated. In addition,
|
||
<taxonomicName id="4C6E4D17FFA1FFA7645FEAB9FB61FE4F" authorityName="Ocupa, M. M. Jimenez & Mark Wilson. A. Habit. B. Flower. C. Dissected" authorityYear="2023" box="[1106,1273,482,508]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="174" phylum="Tracheophyta" rank="species" species="carpishensis">
|
||
<emphasis id="B91AEA86FFA1FFA7645FEAB9FB61FE4F" box="[1106,1273,482,508]" italics="true" pageId="11" pageNumber="174">P. carpishensis</emphasis>
|
||
</taxonomicName>
|
||
exhibits ciliatepapillate margins around the stigmatic surfaces of the column (
|
||
<figureCitation id="13552A11FFA1FFA7635FE95CFC5DFD93" box="[850,965,518,544]" captionStart="FIGURE 3" captionStartId="4.[136,229,1877,1899]" captionTargetBox="[151,1436,190,1853]" captionTargetId="figure-25@4.[151,1436,190,1853]" captionTargetPageId="4" captionText="FIGURE 3. Composite plate of Pleurothallis carpishensis Ocupa, M.M.Jiménez & Mark Wilson. A. Plant. B. Flower. C. Dissected perianth. D. Lip, column and ovary, lateral view.E. Column.F. Lip, adaxial (right) and abaxial (left) views. G.Anther cap and pollinarium. Prepared by L. Ocupa-Horna." figureDoi="http://doi.org/10.5281/zenodo.10275909" httpUri="https://zenodo.org/record/10275909/files/figure.png" pageId="11" pageNumber="174">
|
||
Figure 3
|
||
<collectionCode id="ED7FAE51FFA1FFA763BFE95CFC5DFD93" box="[946,965,518,544]" pageId="11" pageNumber="174">E</collectionCode>
|
||
</figureCitation>
|
||
) and the petals (
|
||
<figureCitation id="13552A11FFA1FFA7648FE95CFB6FFD93" box="[1154,1271,518,544]" captionStart="FIGURE 3" captionStartId="4.[136,229,1877,1899]" captionTargetBox="[151,1436,190,1853]" captionTargetId="figure-25@4.[151,1436,190,1853]" captionTargetPageId="4" captionText="FIGURE 3. Composite plate of Pleurothallis carpishensis Ocupa, M.M.Jiménez & Mark Wilson. A. Plant. B. Flower. C. Dissected perianth. D. Lip, column and ovary, lateral view.E. Column.F. Lip, adaxial (right) and abaxial (left) views. G.Anther cap and pollinarium. Prepared by L. Ocupa-Horna." figureDoi="http://doi.org/10.5281/zenodo.10275909" httpUri="https://zenodo.org/record/10275909/files/figure.png" pageId="11" pageNumber="174">
|
||
Figure 3
|
||
<collectionCode id="ED7FAE51FFA1FFA764EFE95CFB6FFD93" box="[1250,1271,518,544]" pageId="11" pageNumber="174">C</collectionCode>
|
||
</figureCitation>
|
||
), both of which are rare in
|
||
<emphasis id="B91AEA86FFA1FFA76107E970FDD8FDF7" box="[266,576,554,580]" italics="true" pageId="11" pageNumber="174">Macrophyllae-Fasciculatae</emphasis>
|
||
. The ciliate papillae could be involved in secretion of volatiles involved in pollinator attraction, but determination of their function will require additional study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |