1126 lines
103 KiB
XML
1126 lines
103 KiB
XML
<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3893.4.2" ID-GBIF-Dataset="c50b4017-beb6-4783-820e-8a1da797f2d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="252022" ID-ZooBank="65D00761-21AC-4B5D-ACB9-7BFFC69A75FC" checkinTime="1460995836878" checkinUser="plazi" docAuthor="Sparks, Kathryn S., Andersen, Alan N. & Austin, Andrew D." docDate="2014" docId="03A509737E362F41EBF2FEEF4B9275C5" docLanguage="en" docName="zt03893p529.pdf" docOrigin="Zootaxa 3893 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monomorium rothsteini" docType="treatment" docVersion="1" lastPageNumber="497" masterDocId="FF9C710B7E352F49EB65FF804E0E707D" masterDocTitle="Systematics of the Monomorium rothsteini Forel species complex (Hymenoptera: Formicidae), a problematic ant group in Australia" masterLastPageNumber="529" masterPageNumber="489" pageNumber="492" updateTime="1678126142864" updateUser="tatiana">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematics of the Monomorium rothsteini Forel species complex (Hymenoptera: Formicidae), a problematic ant group in Australia</mods:title>
|
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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||
<mods:namePart>Sparks, Kathryn S.</mods:namePart>
|
||
<mods:affiliation>South Australian Museum, North Terrace, Adelaide SA 5000, and Australian Centre for Evolutionary Biology and Biodiversity, and School of Earth & Environmental Sciences, The University of Adelaide, 5005, Australia</mods:affiliation>
|
||
<mods:nameIdentifier type="email">kate.sparks@samuseum.sa.gov.au</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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||
<mods:namePart>Andersen, Alan N.</mods:namePart>
|
||
<mods:affiliation>CSIRO Land and Water Flagship, Tropical Ecosystems Research Centre, PMB 44 Winnellie, NT 0822, Australia</mods:affiliation>
|
||
<mods:nameIdentifier type="email">Alan.Andersen@csiro.au</mods:nameIdentifier>
|
||
</mods:name>
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||
<mods:name type="personal">
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||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Austin, Andrew D.</mods:namePart>
|
||
<mods:affiliation>Australian Centre for Evolutionary Biology and Biodiversity, and School of Earth & Environmental Sciences, The University of Adelaide, 5005, Australia</mods:affiliation>
|
||
<mods:nameIdentifier type="email">andy.austin@adelaide.edu.au</mods:nameIdentifier>
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||
</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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||
<mods:part>
|
||
<mods:date>2014</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2014-12-10</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>3893</mods:number>
|
||
</mods:detail>
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||
<mods:detail type="issue">
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||
<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>489</mods:start>
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<mods:end>529</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3893.4.2</mods:identifier>
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<mods:identifier type="GBIF-Dataset">c50b4017-beb6-4783-820e-8a1da797f2d9</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">252022</mods:identifier>
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<mods:identifier type="ZooBank">65D00761-21AC-4B5D-ACB9-7BFFC69A75FC</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5683106" ID-GBIF-Taxon="119597204" ID-Zenodo-Dep="5683106" LSID="urn:lsid:plazi:treatment:03A509737E362F41EBF2FEEF4B9275C5" httpUri="http://treatment.plazi.org/id/03A509737E362F41EBF2FEEF4B9275C5" lastPageId="8" lastPageNumber="497" pageId="3" pageNumber="492">
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<subSection lastPageId="5" lastPageNumber="494" pageId="3" pageNumber="492" type="multiple">
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<subSubSection box="[151,668,367,392]" pageId="3" pageNumber="492" type="nomenclature">
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<paragraph blockId="3.[151,668,367,392]" box="[151,668,367,392]" pageId="3" pageNumber="492">
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<heading bold="true" box="[151,668,367,392]" fontSize="10" level="2" pageId="3" pageNumber="492" reason="2">
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<emphasis bold="true" box="[151,668,367,392]" pageId="3" pageNumber="492">
|
||
Biology of the
|
||
<taxonomicName ID-CoL="73P7H" ID-ENA="612456" box="[323,472,367,392]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis bold="true" box="[323,472,367,392]" italics="true" pageId="3" pageNumber="492">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
species complex
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="5" lastPageNumber="494" pageId="3" pageNumber="492" type="biology_ecology">
|
||
<paragraph blockId="3.[151,1436,439,933]" pageId="3" pageNumber="492">
|
||
Species in the
|
||
<taxonomicName box="[317,463,440,464]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis box="[317,463,440,464]" italics="true" pageId="3" pageNumber="492">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
species complex are a common and abundant component of the ant fauna throughout most of the Australian mainland outside mesic forests. They are predominantly seed harvesters and the nests of many of the species described here, particular the larger species that occur in the arid and northern monsoonal zones, are easily recognised by the dense middens of mostly grass seed husks that have been discarded to one side of the nest entrance (
|
||
<figureCitation box="[398,470,583,608]" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[151,1436,1009,1867]" captionTargetId="figure@3.[151,1436,1009,1868]" captionTargetPageId="3" captionText="FIGURE 2. Field images of the M. rothsteini (Forel) species complex. Fig. 2 a – c nest entrances showing (a) single entrance with discarded seed husks, (b) multiple entrances without seed husks and (c) single nest entrance without sees husks. Fig. 2 d M. subapterum sp. nov. feeding on nectar from Melaleuca sp. flowers." httpUri="https://zenodo.org/record/252024/files/figure.png" pageId="3" pageNumber="492">
|
||
Fig.
|
||
<number box="[454,470,584,608]" pageId="3" pageNumber="492" value="2.0">2</number>
|
||
</figureCitation>
|
||
a). Worker ants carrying a variety of seeds and arthropod material can be observed foraging in long columns that radiate out from these nest entrances. The nests of smaller species that occur both in the northern monsoonal zone and the central and southern arid zones are less distinctive, with smaller nest middens or no obvious midden at all and with fewer workers (
|
||
<figureCitation box="[739,806,692,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[151,1436,1009,1867]" captionTargetId="figure@3.[151,1436,1009,1868]" captionTargetPageId="3" captionText="FIGURE 2. Field images of the M. rothsteini (Forel) species complex. Fig. 2 a – c nest entrances showing (a) single entrance with discarded seed husks, (b) multiple entrances without seed husks and (c) single nest entrance without sees husks. Fig. 2 d M. subapterum sp. nov. feeding on nectar from Melaleuca sp. flowers." httpUri="https://zenodo.org/record/252024/files/figure.png" pageId="3" pageNumber="492">
|
||
Fig.
|
||
<number box="[791,806,692,717]" pageId="3" pageNumber="492" value="2.0">2</number>
|
||
</figureCitation>
|
||
b,
|
||
<number box="[832,848,692,716]" pageId="3" pageNumber="492" value="2.0">2</number>
|
||
c). In many areas, the nest of one species with a large midden occurs within metres of another species with no nest midden. Although seeds appear to make up the bulk of the diet of the complex generally, species are opportunistic feeders on other food sources, and it is possible that seeds are only a minor component of the diets of some species. For example, one of us (KSS) has observed several colonies of
|
||
<taxonomicName box="[281,424,836,860]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="speculum" status="sp. nov.">
|
||
<emphasis box="[281,424,836,860]" italics="true" pageId="3" pageNumber="492">M. speculum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[430,517,837,860]" pageId="3" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in a dry lake bed. There was very little vegetation within foraging distance and all workers moving around the nest entrance were carrying minute gastropod shells. Workers of
|
||
<taxonomicName box="[1205,1375,872,896]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="subapterum">
|
||
<emphasis box="[1205,1375,872,896]" italics="true" pageId="3" pageNumber="492">M. subapterum</emphasis>
|
||
</taxonomicName>
|
||
have been observed feeding on the nectar of
|
||
<taxonomicName box="[592,710,908,932]" class="Agaricomycetes" family="Tricholomataceae" genus="Melaleuca" higherTaxonomySource="CoL" kingdom="Fungi" order="Agaricales" pageId="3" pageNumber="492" phylum="Basidiomycota" rank="genus">
|
||
<emphasis box="[592,710,908,932]" italics="true" pageId="3" pageNumber="492">Melaleuca</emphasis>
|
||
</taxonomicName>
|
||
flowers (
|
||
<figureCitation box="[816,883,908,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[151,1436,1009,1867]" captionTargetId="figure@3.[151,1436,1009,1868]" captionTargetPageId="3" captionText="FIGURE 2. Field images of the M. rothsteini (Forel) species complex. Fig. 2 a – c nest entrances showing (a) single entrance with discarded seed husks, (b) multiple entrances without seed husks and (c) single nest entrance without sees husks. Fig. 2 d M. subapterum sp. nov. feeding on nectar from Melaleuca sp. flowers." httpUri="https://zenodo.org/record/252024/files/figure.png" pageId="3" pageNumber="492">
|
||
Fig.
|
||
<number box="[868,883,908,933]" pageId="3" pageNumber="492" value="2.0">2</number>
|
||
</figureCitation>
|
||
d).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1594,2015]" pageId="4" pageNumber="493">
|
||
Queens and males are rarely seen. Both are rare in collections and during two extensive field trips and several shorter excursions queens and males were each observed on only one occasion. There are published accounts of species in the
|
||
<taxonomicName box="[312,457,1666,1690]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis box="[312,457,1666,1690]" italics="true" pageId="4" pageNumber="493">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
complex having brachypterous queens.
|
||
<bibRefCitation author="Briese" box="[912,1066,1666,1691]" firstAuthor="Briese" pageId="4" pageNumber="493" refString="Briese, D. T. (1983) Different modes of reproductive-behaviour (including a description of colony fission) in a species of Chelaner (Hymenoptera, Formicidae). Insectes Sociaux, 30, 308 - 316." type="journal article" year="1983">
|
||
Briese (
|
||
<number box="[1000,1058,1666,1690]" pageId="4" pageNumber="493" value="1983.0">1983</number>
|
||
)
|
||
</bibRefCitation>
|
||
observed
|
||
<number box="[1183,1211,1666,1691]" pageId="4" pageNumber="493" value="63.0">63</number>
|
||
queens with poorly developed wings and orange gasters emerging from a colony that subsequently underwent colony fission. The colony was within
|
||
<quantity box="[369,415,1738,1763]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="493" unit="m" value="5.0">5 m</quantity>
|
||
of another colony that later produced queens described as fully winged with dark brown gasters.
|
||
<bibRefCitation author="Wheeler, W. M." box="[247,447,1774,1799]" journalOrPublisher="Proceedings of the National Academy of Sciences of the United States of America" pageId="4" pageNumber="493" pagination="109 - 117" part="3" refId="ref30518" refString="Wheeler, W. M. (1917) The development of subapterous and apterous castes in the Formicidae. Proceedings of the National Academy of Sciences of the United States of America, 3, 109 - 117." title="The development of subapterous and apterous castes in the Formicidae" type="journal article" year="1917">
|
||
Wheeler’s (
|
||
<number box="[381,439,1775,1799]" pageId="4" pageNumber="493" value="1917.0">1917</number>
|
||
)
|
||
</bibRefCitation>
|
||
erection of
|
||
<taxonomicName box="[595,768,1774,1798]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="subapterum">
|
||
<emphasis box="[595,768,1774,1798]" italics="true" pageId="4" pageNumber="493">M. subapterum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[832,1006,1774,1798]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="subapterum">
|
||
<emphasis box="[832,1006,1774,1798]" italics="true" pageId="4" pageNumber="493">M. subapterum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName box="[1017,1165,1774,1798]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="variety" species="subapterum" variety="bogischi">
|
||
var.
|
||
<emphasis box="[1070,1165,1774,1798]" italics="true" pageId="4" pageNumber="493">bogischi</emphasis>
|
||
</taxonomicName>
|
||
is based on queens he received from the South Australian Museum that had been collected from Harding River in northwest Western
|
||
<collectingCountry box="[151,254,1846,1871]" name="Australia" pageId="4" pageNumber="493">Australia</collectingCountry>
|
||
and Port Wakefield in southern South
|
||
<collectingCountry box="[693,799,1846,1871]" name="Australia" pageId="4" pageNumber="493">Australia</collectingCountry>
|
||
. He described these specimens as being smaller than the queens of
|
||
<taxonomicName box="[267,410,1882,1906]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis box="[267,410,1882,1906]" italics="true" pageId="4" pageNumber="493">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName box="[418,563,1882,1906]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="variety" species="rothsteini" variety="humilior">
|
||
var.
|
||
<emphasis box="[466,563,1882,1906]" italics="true" pageId="4" pageNumber="493">humilior</emphasis>
|
||
</taxonomicName>
|
||
and having very small wings. He alluded to other specific differences but did not discuss them. The lead author has examined queens from both these series in the South Australian Museum that were dealate, having just the remains of both pairs of wings and resembling fully developed queens in all other aspects. Queens that were excavated from a nest in Birdsville in southwest Queensland and here attributed to
|
||
<emphasis box="[1405,1436,1991,2014]" italics="true" pageId="4" pageNumber="493">M.</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,151,536]" pageId="5" pageNumber="494">
|
||
<taxonomicName box="[151,246,152,176]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="bogischi">
|
||
<emphasis box="[151,246,152,176]" italics="true" pageId="5" pageNumber="494">bogischi</emphasis>
|
||
</taxonomicName>
|
||
were all dealate with obvious wing remnants (
|
||
<figureCitation box="[774,842,151,176]" captionStart="FIGURE 9" captionStartId="33.[151,250,1913,1935]" captionTargetBox="[204,1383,193,1893]" captionTargetId="figure@33.[204,1384,193,1894]" captionTargetPageId="33" captionText="FIGURE 9. Queens and males of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 9 a – c: M. bogischi queen (light form), paralectotype (Port Wakefield. SA), d – f: M. bogischi queen (dark form) (Birdsville, QLD, KSS 46), g – i: M. humilior queen, paralectotype (Tennant Ck, NT), j – l: M. humilior male (Tennant Creek, NT), m – o: M. subapterum queen, paralectotype (Harding River, WA). Scale bars are 1 mm for all views." httpUri="https://zenodo.org/record/252031/files/figure.png" pageId="5" pageNumber="494">
|
||
Fig.
|
||
<number box="[827,842,151,176]" pageId="5" pageNumber="494" value="9.0">9</number>
|
||
</figureCitation>
|
||
d). Wheeler may have had access to other specimens than those currently available at the South Australian Museum, but based on our observations of specimens collected in the field and examined in collections, we cannot confirm brachyptery exists in this group. Larger workers that appeared to bear some aberrant characteristics (overall larger size, well defined promesonotal suture and coarser sculpture) were found foraging with other workers from a colony of
|
||
<taxonomicName box="[1083,1205,296,320]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="pilbara" status="sp. nov.">
|
||
<emphasis box="[1083,1205,296,320]" italics="true" pageId="5" pageNumber="494">M. pilbara</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1214,1305,296,319]" pageId="5" pageNumber="494" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and small, black, heavily sculptured workers have been observed foraging alongside workers of
|
||
<taxonomicName box="[1129,1244,332,356]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="topend" status="sp. nov.">
|
||
<emphasis box="[1129,1244,332,356]" italics="true" pageId="5" pageNumber="494">M. topend</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1253,1342,333,356]" pageId="5" pageNumber="494" rank="species">sp. nov.</taxonomicNameLabel>
|
||
that are typically large and amber orange in colour. Whether these aberrant forms represent an intercaste, a developmental anomaly or an apterous female is unknown.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,151,536]" pageId="5" pageNumber="494">
|
||
A number of morphospecies within the
|
||
<taxonomicName box="[667,815,440,464]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis box="[667,815,440,464]" italics="true" pageId="5" pageNumber="494">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
species complex have been the subject of ecological studies.
|
||
<tableCitation box="[242,324,476,501]" captionStart="TABLE 1" captionStartId="5.[151,239,584,607]" captionTargetBox="[159,1294,668,1702]" captionTargetPageId="5" captionText="TABLE 1. This table provides a list of the species within the M. rothsteini species complex that have been referred to in previous publications and their respective species codes used in those publications." httpUri="http://table.plazi.org/id/DF73E8ED7E302F4CEBF2FDC84A0272FD" pageId="5" pageNumber="494" tableUuid="DF73E8ED7E302F4CEBF2FDC84A0272FD">
|
||
Table
|
||
<number box="[310,324,476,500]" pageId="5" pageNumber="494" value="1.0">1</number>
|
||
</tableCitation>
|
||
provides a list of these publications, the morphospecies codes that were used and the corresponding species name as per this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
<subSection pageId="6" pageNumber="495" type="multiple">
|
||
<subSubSection pageId="6" pageNumber="495" type="diagnosis">
|
||
<paragraph blockId="6.[151,693,151,176]" box="[151,693,151,176]" pageId="6" pageNumber="495">
|
||
<heading bold="true" box="[151,693,151,176]" fontSize="10" level="2" pageId="6" pageNumber="495" reason="2">
|
||
<emphasis bold="true" box="[151,693,151,176]" pageId="6" pageNumber="495">
|
||
Diagnosis of the
|
||
<taxonomicName box="[348,497,151,176]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis bold="true" box="[348,497,151,176]" italics="true" pageId="6" pageNumber="495">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
species complex
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,223,392]" pageId="6" pageNumber="495">
|
||
Members of the
|
||
<taxonomicName box="[346,493,224,248]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis box="[346,493,224,248]" italics="true" pageId="6" pageNumber="495">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
species complex can be distinguished from all other species in the genus by the presence of a
|
||
<number box="[316,344,260,284]" pageId="6" pageNumber="495" value="12.0">12</number>
|
||
segmented antenna, mandible with three teeth, mildly to deeply concave anterodorsal clypeal margin and alveolate sculpture on the meso- and metapleuron. Members of the
|
||
<taxonomicName box="[1044,1188,296,320]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis box="[1044,1188,296,320]" italics="true" pageId="6" pageNumber="495">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
species complex bear a close morphological relationship to
|
||
<taxonomicName box="[590,735,332,356]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="sordidum">
|
||
<emphasis box="[590,735,332,356]" italics="true" pageId="6" pageNumber="495">M. sordidum</emphasis>
|
||
</taxonomicName>
|
||
but can be easily distinguished by the shape of the anterior clypeal margin, which is rounded in the latter (
|
||
<taxonomicName box="[675,940,368,392]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="sordidum">
|
||
<emphasis box="[675,940,368,392]" italics="true" pageId="6" pageNumber="495">Monomorium sordidum</emphasis>
|
||
</taxonomicName>
|
||
is not granivorous).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
<subSection pageId="6" pageNumber="495" type="multiple">
|
||
<subSubSection pageId="6" pageNumber="495" type="description">
|
||
<paragraph blockId="6.[151,716,439,464]" box="[151,716,439,464]" pageId="6" pageNumber="495">
|
||
<heading bold="true" box="[151,716,439,464]" fontSize="10" level="2" pageId="6" pageNumber="495" reason="2">
|
||
<emphasis bold="true" box="[151,716,439,464]" pageId="6" pageNumber="495">
|
||
Description of the
|
||
<taxonomicName box="[371,520,439,464]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis bold="true" box="[371,520,439,464]" italics="true" pageId="6" pageNumber="495">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
species complex
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,511,1400]" pageId="6" pageNumber="495">
|
||
<emphasis bold="true" box="[151,533,511,536]" pageId="6" pageNumber="495">
|
||
Worker measurements (n=
|
||
<number box="[476,520,511,536]" pageId="6" pageNumber="495" value="201.0">201</number>
|
||
).
|
||
</emphasis>
|
||
HW
|
||
<number box="[599,714,511,536]" pageId="6" pageNumber="495" value="0.835" valueMax="1.05" valueMin="0.62">0.62–1.05</number>
|
||
, HL
|
||
<number box="[774,888,511,536]" pageId="6" pageNumber="495" value="0.855" valueMax="1.03" valueMin="0.68">0.68–1.03</number>
|
||
, EL
|
||
<number box="[946,1060,512,536]" pageId="6" pageNumber="495" value="0.18" valueMax="0.22" valueMin="0.14">0.14–0.22</number>
|
||
, PMH
|
||
<number box="[1144,1258,512,536]" pageId="6" pageNumber="495" value="0.36" valueMax="0.49" valueMin="0.23">0.23–0.49</number>
|
||
,
|
||
<collectingCountry box="[1272,1308,512,535]" name="Philippines" pageId="6" pageNumber="495">PH</collectingCountry>
|
||
<number box="[1317,1432,512,536]" pageId="6" pageNumber="495" value="0.31" valueMax="0.41" valueMin="0.21">0.21–0.41</number>
|
||
, PNH
|
||
<number box="[214,329,548,573]" pageId="6" pageNumber="495" value="0.245" valueMax="0.35" valueMin="0.14">0.14–0.35</number>
|
||
, LHW
|
||
<number box="[411,525,548,573]" pageId="6" pageNumber="495" value="0.505" valueMax="0.62" valueMin="0.39">0.39–0.62</number>
|
||
, EW
|
||
<number box="[587,688,548,572]" pageId="6" pageNumber="495" value="0.12000000000000001" valueMax="0.14" valueMin="0.1">0.1–0.14</number>
|
||
, PML
|
||
<number box="[764,864,548,573]" pageId="6" pageNumber="495" value="0.6799999999999999" valueMax="0.7" valueMin="0.66">0.66–0.7</number>
|
||
, ML
|
||
<number box="[925,1039,548,573]" pageId="6" pageNumber="495" value="0.905" valueMax="1.17" valueMin="0.64">0.64–1.17</number>
|
||
, PL
|
||
<number box="[1090,1206,548,573]" pageId="6" pageNumber="495" value="0.41000000000000003" valueMax="0.55" valueMin="0.27">0.27–0.55</number>
|
||
, PNWdv
|
||
<number box="[1314,1429,548,573]" pageId="6" pageNumber="495" value="0.255" valueMax="0.35" valueMin="0.16">0.16–0.35</number>
|
||
. Head: Shape appearing square or rectangular but always longer than wide; posterior cephalic margin in full face view straight to mildly concave or broadly v-shaped, antennae
|
||
<number box="[923,949,620,645]" pageId="6" pageNumber="495" value="12.0">12</number>
|
||
-segmented with a well differentiated
|
||
<number box="[1413,1427,620,644]" pageId="6" pageNumber="495" value="3.0">3</number>
|
||
- segmented club; scape reaching approximately three quarters of the way to vertexal margin of head; mandible with three teeth; anterior margin of clypeus mildly to deeply concave; coarse strigae present laterally between mandibular articulation and eye; strigae may be present on frons above antennal lobes or restricted to antennal lobes or area in between; eye oval in shape, ventral margin flattened.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,511,1400]" pageId="6" pageNumber="495">Mesosoma: Pronotum rounded in lateral profile, promesonotal suture clearly present, faint, or absent, posterior mesonotum slightly raised or flat; metanotal groove shallow or deeply impressed, porcate; mesonotum coarsely strigate, punctuate, both, or smooth. Propodeum with or without a raised anterodorsal transverse carina, dorsal margin in lateral view rounded, flat or sloping posteriorly, dorsal surface concave or flat to slightly rounded, with or without longitudinal carinae and transverse strigae, posterior surface near vertical or sloping. Petiole node broad to narrow and rounded dorsally, postpetiole rounded, approximately three quarters the height of the petiole. Mesopleuron and propodeum coarsely alveolate with fine to coarse striations; metanotal groove always with numerous transverse ridges; petiole without sculpture or with fine reticulation, postpetiole almost always finely reticulate; very fine reticulation present or absent on first metasomal segment.</paragraph>
|
||
<paragraph blockId="6.[151,1437,511,1400]" pageId="6" pageNumber="495">Pilosity sparse with long and short erect or appressed white or yellow setae on head; always with one pair of elongate, inwardly curved setae arising from centre of clypeal lateral carinae, and one pair of elongate erect setae arising from antennal lobes; erect setae on vertex curved anteriorly, setae on dorsal surface of pronotum inwardly curved, elongate erect setae on propodeum curved anteriorly, erect setae on petiole and postpetiole curved posteriorly.</paragraph>
|
||
<paragraph blockId="6.[151,1437,511,1400]" pageId="6" pageNumber="495">Colour variable; head and mesosoma amber orange, dark orange brown, dark brown or almost black; metasoma almost always dark brown to black, rarely light brown or amber, cuticle with a high gloss on nonsculptured areas.</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="496" pageId="6" pageNumber="495" type="discussion">
|
||
<subSection lastPageId="7" lastPageNumber="496" pageId="6" pageNumber="495" type="multiple">
|
||
<paragraph blockId="6.[151,389,1447,1472]" box="[151,389,1447,1472]" pageId="6" pageNumber="495">
|
||
<heading bold="true" box="[151,389,1447,1472]" fontSize="10" level="2" pageId="6" pageNumber="495" reason="2">
|
||
<emphasis bold="true" box="[151,389,1447,1472]" pageId="6" pageNumber="495">Taxonomic synopsis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,805,1519,1544]" pageId="6" pageNumber="495">
|
||
<taxonomicName authority="Wheeler, 1917" authorityName="Wheeler" authorityYear="1917" box="[151,576,1519,1544]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="bogischi" status="stat. rev.">
|
||
<emphasis box="[151,405,1520,1544]" italics="true" pageId="6" pageNumber="495">Monomorium bogischi</emphasis>
|
||
<bibRefCitation author="Wheeler" box="[412,576,1519,1544]" firstAuthor="Wheeler" pageId="6" pageNumber="495" refString="Wheeler, W. M. (1917) The development of subapterous and apterous castes in the Formicidae. Proceedings of the National Academy of Sciences of the United States of America, 3, 109 - 117." type="journal article" year="1917">
|
||
Wheeler,
|
||
<number box="[520,576,1520,1544]" pageId="6" pageNumber="495" value="1917.0">1917</number>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[583,805,1519,1544]" pageId="6" pageNumber="495">
|
||
<taxonomicNameLabel box="[583,686,1519,1544]" pageId="6" pageNumber="495">stat. rev.</taxonomicNameLabel>
|
||
, stat. nov.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,541,1556,1581]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,444,1556,1580]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="broschorum" status="sp. nov.">
|
||
<emphasis box="[151,444,1556,1580]" italics="true" pageId="6" pageNumber="495">Monomorium broschorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[452,541,1556,1581]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[452,541,1556,1581]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,509,1591,1616]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,413,1592,1616]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="capeyork" status="sp. nov.">
|
||
<emphasis box="[151,413,1592,1616]" italics="true" pageId="6" pageNumber="495">Monomorium capeyork</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[420,509,1591,1616]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[420,509,1591,1616]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,520,1628,1653]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,424,1628,1652]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="eremoides" status="sp. nov.">
|
||
<emphasis box="[151,424,1628,1652]" italics="true" pageId="6" pageNumber="495">Monomorium eremoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[431,520,1628,1653]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[431,520,1628,1653]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,496,1663,1688]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,399,1664,1688]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="eremum" status="sp. nov.">
|
||
<emphasis box="[151,399,1664,1688]" italics="true" pageId="6" pageNumber="495">Monomorium eremum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[406,496,1663,1688]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[406,496,1663,1688]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,509,1700,1725]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,413,1701,1724]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="geminum" status="sp. nov.">
|
||
<emphasis box="[151,413,1701,1724]" italics="true" pageId="6" pageNumber="495">Monomorium geminum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[420,509,1700,1725]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[420,509,1700,1725]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,488,1735,1760]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,391,1736,1760]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="hertogi" status="sp. nov.">
|
||
<emphasis box="[151,391,1736,1760]" italics="true" pageId="6" pageNumber="495">Monomorium hertogi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[398,488,1735,1760]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[398,488,1735,1760]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,520,1772,1797]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,424,1772,1796]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="hoffmanni" status="sp. nov.">
|
||
<emphasis box="[151,424,1772,1796]" italics="true" pageId="6" pageNumber="495">Monomorium hoffmanni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[431,520,1772,1797]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[431,520,1772,1797]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,765,1807,1832]" pageId="6" pageNumber="495">
|
||
<taxonomicName authority="Forel 1910" authorityName="Forel" authorityYear="1910" box="[151,536,1807,1832]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="humilior" status="stat. rev.">
|
||
<emphasis box="[151,407,1808,1832]" italics="true" pageId="6" pageNumber="495">Monomorium humilior</emphasis>
|
||
<bibRefCitation author="Forel" box="[414,536,1807,1832]" firstAuthor="Forel" pageId="6" pageNumber="495" refString="Forel, A. (1910) Formicides australiens recs de M. M. Frogatt et Rowland Turner. Revue Suisse de Zoologie, 18, 1 - 94." type="journal article" year="1910">
|
||
Forel
|
||
<number box="[480,536,1808,1832]" pageId="6" pageNumber="495" value="1910.0">1910</number>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[543,765,1807,1832]" pageId="6" pageNumber="495">
|
||
<taxonomicNameLabel box="[543,646,1807,1832]" pageId="6" pageNumber="495">stat. rev.</taxonomicNameLabel>
|
||
, stat. nov.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,489,1844,1869]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,393,1844,1868]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="kidman" status="sp. nov.">
|
||
<emphasis box="[151,393,1844,1868]" italics="true" pageId="6" pageNumber="495">Monomorium kidman</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[400,489,1844,1869]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[400,489,1844,1869]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,717,1879,1904]" pageId="6" pageNumber="495">
|
||
<taxonomicName authority="Forel 1915" authorityName="Forel" authorityYear="1915" box="[151,487,1879,1904]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="leda" status="stat. rev.">
|
||
<emphasis box="[151,358,1880,1904]" italics="true" pageId="6" pageNumber="495">Monomorium leda</emphasis>
|
||
<bibRefCitation author="Forel" box="[365,487,1879,1904]" firstAuthor="Forel" pageId="6" pageNumber="495" refString="Forel, A. (1915) Results of Dr Mj oberg's Swedish Scientific Expeditions to Australia 1910 - 1913. 2. Ameisen. Arkiv fur Zoologi, 9 (16), 1 - 119." type="journal article" year="1915">
|
||
Forel
|
||
<number box="[431,487,1879,1904]" pageId="6" pageNumber="495" value="1915.0">1915</number>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[494,717,1879,1904]" pageId="6" pageNumber="495">
|
||
<taxonomicNameLabel box="[494,597,1879,1904]" pageId="6" pageNumber="495">stat. rev.</taxonomicNameLabel>
|
||
, stat. nov.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,533,1916,1941]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,436,1917,1940]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="maryannae" status="sp. nov.">
|
||
<emphasis box="[151,436,1917,1940]" italics="true" pageId="6" pageNumber="495">Monomorium maryannae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[443,533,1916,1941]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[443,533,1916,1941]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,504,1952,1977]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,407,1952,1976]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="merepah" status="sp. nov.">
|
||
<emphasis box="[151,407,1952,1976]" italics="true" pageId="6" pageNumber="495">Monomorium merepah</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[414,504,1952,1977]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[414,504,1952,1977]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,805,1519,2013]" box="[151,497,1988,2013]" pageId="6" pageNumber="495">
|
||
<taxonomicName box="[151,400,1988,2012]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="mitchell" status="sp. nov.">
|
||
<emphasis box="[151,400,1988,2012]" italics="true" pageId="6" pageNumber="495">Monomorium mitchell</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[407,497,1988,2013]" pageId="6" pageNumber="495" rank="species">
|
||
<emphasis bold="true" box="[407,497,1988,2013]" pageId="6" pageNumber="495">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[151,534,151,176]" pageId="7" pageNumber="496">
|
||
<taxonomicName box="[151,438,152,176]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="oodnadatta" status="sp. nov.">
|
||
<emphasis box="[151,438,152,176]" italics="true" pageId="7" pageNumber="496">Monomorium oodnadatta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[445,534,151,176]" pageId="7" pageNumber="496" rank="species">
|
||
<emphasis bold="true" box="[445,534,151,176]" pageId="7" pageNumber="496">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[151,489,188,213]" pageId="7" pageNumber="496">
|
||
<taxonomicName box="[151,393,188,212]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="pilbara" status="sp. nov.">
|
||
<emphasis box="[151,393,188,212]" italics="true" pageId="7" pageNumber="496">Monomorium pilbara</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[400,489,188,213]" pageId="7" pageNumber="496" rank="species">
|
||
<emphasis bold="true" box="[400,489,188,213]" pageId="7" pageNumber="496">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[151,553,223,248]" pageId="7" pageNumber="496">
|
||
<taxonomicName authority="Forel 1902" authorityName="Forel" authorityYear="1902" box="[151,549,223,248]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis box="[151,416,224,248]" italics="true" pageId="7" pageNumber="496">Monomorium rothsteini</emphasis>
|
||
<bibRefCitation author="Forel" box="[423,549,223,248]" firstAuthor="Forel" pageId="7" pageNumber="496" refString="Forel, A. (1902) Fourmis nouvelles d'Australie. Revue Suisse de Zoologie, 10, 405 - 408." type="journal article" year="1902">
|
||
Forel
|
||
<number box="[489,549,224,248]" pageId="7" pageNumber="496" value="1902.0">1902</number>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[199,737,260,285]" pageId="7" pageNumber="496">
|
||
=
|
||
<taxonomicName authority="Santschi, 1919" authorityName="Santschi" authorityYear="1919" box="[221,733,260,285]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="subSpecies" species="rothsteini" subSpecies="doddi">
|
||
<emphasis box="[221,558,260,284]" italics="true" pageId="7" pageNumber="496">Monomorium rothsteini doddi</emphasis>
|
||
<bibRefCitation author="Santschi" box="[565,733,260,285]" firstAuthor="Santschi" pageId="7" pageNumber="496" refString="Santschi, F. (1919) Cinq notes myrmecologiques. Bulletin de la Societe Vaudoise des Sciences Naturelles, 52, 325 - 350." type="journal article" year="1919">
|
||
Santschi,
|
||
<number box="[674,733,260,284]" pageId="7" pageNumber="496" value="1919.0">1919</number>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[151,512,295,320]" pageId="7" pageNumber="496">
|
||
<taxonomicName box="[151,416,296,320]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="speculum" status="sp. nov.">
|
||
<emphasis box="[151,416,296,320]" italics="true" pageId="7" pageNumber="496">Monomorium speculum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[423,512,295,320]" pageId="7" pageNumber="496" rank="species">
|
||
<emphasis bold="true" box="[423,512,295,320]" pageId="7" pageNumber="496">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[151,502,332,357]" pageId="7" pageNumber="496">
|
||
<taxonomicName box="[151,405,333,356]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="stagnum" status="sp. nov.">
|
||
<emphasis box="[151,405,333,356]" italics="true" pageId="7" pageNumber="496">Monomorium stagnum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[412,502,332,357]" pageId="7" pageNumber="496" rank="species">
|
||
<emphasis bold="true" box="[412,502,332,357]" pageId="7" pageNumber="496">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[151,722,367,392]" pageId="7" pageNumber="496">
|
||
<taxonomicName authority="Wheeler, 1917" authorityName="Wheeler" authorityYear="1917" box="[151,613,367,392]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="subapterum" status="stat. rev.">
|
||
<emphasis box="[151,442,368,392]" italics="true" pageId="7" pageNumber="496">Monomorium subapterum</emphasis>
|
||
<bibRefCitation author="Wheeler" box="[449,613,367,392]" firstAuthor="Wheeler" pageId="7" pageNumber="496" refString="Wheeler, W. M. (1917) The development of subapterous and apterous castes in the Formicidae. Proceedings of the National Academy of Sciences of the United States of America, 3, 109 - 117." type="journal article" year="1917">
|
||
Wheeler,
|
||
<number box="[557,613,368,392]" pageId="7" pageNumber="496" value="1917.0">1917</number>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[620,722,367,392]" pageId="7" pageNumber="496">
|
||
<emphasis bold="true" box="[620,722,367,392]" pageId="7" pageNumber="496">stat. rev.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[151,536,404,429]" pageId="7" pageNumber="496">
|
||
<taxonomicName box="[151,440,404,428]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="tenebrosum" status="sp. nov.">
|
||
<emphasis box="[151,440,404,428]" italics="true" pageId="7" pageNumber="496">Monomorium tenebrosum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[447,536,404,429]" pageId="7" pageNumber="496" rank="species">
|
||
<emphasis bold="true" box="[447,536,404,429]" pageId="7" pageNumber="496">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[151,483,439,464]" pageId="7" pageNumber="496">
|
||
<taxonomicName box="[151,387,440,464]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="topend" status="sp. nov.">
|
||
<emphasis box="[151,387,440,464]" italics="true" pageId="7" pageNumber="496">Monomorium topend</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[393,483,439,464]" pageId="7" pageNumber="496" rank="species">
|
||
<emphasis bold="true" box="[393,483,439,464]" pageId="7" pageNumber="496">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,737,151,501]" box="[151,486,476,501]" pageId="7" pageNumber="496">
|
||
<taxonomicName box="[151,390,477,500]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="torrens" status="sp. nov.">
|
||
<emphasis box="[151,390,477,500]" italics="true" pageId="7" pageNumber="496">Monomorium torrens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[397,486,476,501]" pageId="7" pageNumber="496" rank="species">
|
||
<emphasis bold="true" box="[397,486,476,501]" pageId="7" pageNumber="496">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSection>
|
||
<paragraph blockId="7.[151,753,547,573]" box="[151,753,547,573]" pageId="7" pageNumber="496">
|
||
<heading bold="true" box="[151,753,547,573]" fontSize="10" level="2" pageId="7" pageNumber="496" reason="2">
|
||
<emphasis bold="true" box="[151,753,547,573]" pageId="7" pageNumber="496">
|
||
Key to species of the
|
||
<taxonomicName box="[400,549,547,572]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis bold="true" box="[400,549,547,572]" italics="true" pageId="7" pageNumber="496">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
species complex.
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1436,620,968]" pageId="7" pageNumber="496">
|
||
This key is designed for the identification of the majority of the most commonly encountered species in the
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="rothsteini">
|
||
<emphasis italics="true" pageId="7" pageNumber="496">M. rothsteini</emphasis>
|
||
</taxonomicName>
|
||
complex. All of these species, with the exception of
|
||
<emphasis box="[880,975,656,680]" italics="true" pageId="7" pageNumber="496">
|
||
<taxonomicName box="[880,972,656,680]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="leda">M. leda</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
were recognised as separately evolving lineages in the mitochondrial
|
||
<collectionCode LSID="urn:lsid:biocol.org:col:15536" box="[502,563,692,716]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="7" pageNumber="496">DNA</collectionCode>
|
||
analysis of
|
||
<bibRefCitation author="Sparks, K. S. & Andersen, A. N. & Donnellan, S. C. & Austin, A. D." box="[710,946,692,717]" etAl="et al." firstAuthor="Sparks" journalOrPublisher="Systematic Entomology" pageId="7" pageNumber="496" pagination="264 - 278" part="39" refId="ref30365" refString="Sparks, K. S., Andersen, A. N., Donnellan, S. C. & Austin, A. D. (2014) Navigating the mtDNA road map out of the morphological maze: interpreting morphological variation in the diverse Monomorium rothsteini (Forel) complex (Hymenoptera: Formicidae). Systematic Entomology, 39, 264 - 278. http: // dx. doi. org / 10.1111 / syen. 12051" title="Navigating the mtDNA road map out of the morphological maze: interpreting morphological variation in the diverse Monomorium rothsteini (Forel) complex (Hymenoptera: Formicidae)" type="journal article" year="2014">
|
||
Sparks et al. (
|
||
<number box="[876,938,692,716]" pageId="7" pageNumber="496" value="2014.0">2014</number>
|
||
)
|
||
</bibRefCitation>
|
||
. The phylogenetic tree from that study is reproduced in
|
||
<figureCitation box="[327,400,727,752]" captionStart="FIGURE 3" captionStartId="4.[151,250,1441,1463]" captionTargetBox="[359,1253,231,1418]" captionTargetId="figure@4.[332,1254,193,1419]" captionTargetPageId="4" captionText="FIGURE 3. Bayesian phylogeny based on 934 bp region of cytochrome oxidase I (COI) (Sparks et al. 2014). Nodes with> 0.95 Bayesian posterior probability support and> 75 % maximum likelihood bootstrap values are indicated with a star. Well supported morphologically cohesive terminal clades have been compressed for ease of interpretation. The molecular clades are labelled with the corresponding species names." httpUri="https://zenodo.org/record/252025/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<number box="[386,400,728,752]" pageId="7" pageNumber="496" value="3.0">3</number>
|
||
</figureCitation>
|
||
with the molecular clades labelled with the corresponding species names. Additional morphotypes are known to exist within the complex, but there is insufficient evidence available at this time to formally describe them. For species identifications that are particularly important it is strongly recommended that the CO
|
||
<number box="[235,244,836,860]" pageId="7" pageNumber="496" value="1.0">1</number>
|
||
sequence data are obtained and compared with those deposited in Genbank. All species exhibit some level of intraspecific variation that can in some cases lead to incorrect identification when using this key. It is recommended that a range of specimens from a colony are examined to determine the character state that best represents the majority of specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="8" lastPageNumber="497" pageId="7" pageNumber="496" type="key">
|
||
<keyStep pageId="7" pageNumber="496">
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
<number box="[151,164,1015,1036]" pageId="7" pageNumber="496" value="1.0">1</number>
|
||
. Anterodorsal margin of the clypeus deeply concave, anterolateral carinae produced as acute angles, often with a wavy margin or minute preapical teeth medial to the acute angles (
|
||
<figureCitation box="[732,789,1045,1066]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[204,1391,181,1899]" captionTargetId="figure@9.[204,1392,181,1903]" captionTargetPageId="9" captionText="FIGURE 4. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 4 a – c: M. bogischi, (Port Wakefield, SA, KSS 241), d – f: M. broschorum holotype (Dajarra, QLD, KSS 60), g – i: M. capeyork, holotype (69 km N Archer River Roadhouse, QLD, TERC 4), j – l: M. eremoides, holotype (9 km E 80 mile Beach, WA, KSS 89), m – o: M. eremum, holotype (33 km N Barkley Homestead, NT, KSS 77). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252026/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<number box="[776,789,1045,1066]" pageId="7" pageNumber="496" value="4.0">4</number>
|
||
</figureCitation>
|
||
h,
|
||
<number box="[812,825,1045,1066]" pageId="7" pageNumber="496" value="6.0">6</number>
|
||
e,
|
||
<number box="[847,859,1045,1066]" pageId="7" pageNumber="496" value="6.0">6</number>
|
||
h,
|
||
<number box="[883,895,1045,1066]" pageId="7" pageNumber="496" value="7.0">7</number>
|
||
k)..........................................
|
||
<number box="[1423,1435,1045,1066]" pageId="7" pageNumber="496" value="2.0">2</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1436,1015,2034]" box="[151,1435,1074,1095]" pageId="7" pageNumber="496">
|
||
<keyLead box="[151,1435,1074,1095]" pageId="7" pageNumber="496">
|
||
- Anterodorsal margin of the clypeus not as above, may be concave but without acute anterolateral angles or preapical teeth...
|
||
<number box="[1423,1435,1074,1095]" pageId="7" pageNumber="496" value="5.0">5</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="496">
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
<number box="[151,166,1103,1124]" pageId="7" pageNumber="496" value="2.0">2</number>
|
||
. Petiole node, postpetiole and anterior half of T
|
||
<number box="[673,682,1103,1124]" pageId="7" pageNumber="496" value="1.0">1</number>
|
||
completely covered in a fine reticulate pattern; very large, robust species (HW and ML>
|
||
<quantity box="[327,376,1133,1154]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="496" unit="mm" value="1.0">1mm</quantity>
|
||
for a majority of specimens)..........................................................
|
||
<taxonomicName box="[1352,1435,1133,1154]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="merepah">
|
||
<emphasis box="[1352,1435,1133,1154]" italics="true" pageId="7" pageNumber="496">merepah</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
- Petiole and postpetiole with a fine reticulate pattern that does not extend onto dorsal surface; smaller species (HW and ML <
|
||
<quantity box="[225,274,1191,1212]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="496" unit="mm" value="1.0">1mm</quantity>
|
||
for a majority of specimens)........................................................................
|
||
<number box="[1423,1435,1191,1212]" pageId="7" pageNumber="496" value="3.0">3</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="496">
|
||
<paragraph blockId="7.[151,1436,1015,2034]" box="[151,1435,1221,1242]" pageId="7" pageNumber="496">
|
||
<keyLead box="[151,1435,1221,1242]" pageId="7" pageNumber="496">
|
||
<number box="[151,165,1221,1242]" pageId="7" pageNumber="496" value="3.0">3</number>
|
||
. First metasomal tergite (T
|
||
<number box="[477,486,1221,1242]" pageId="7" pageNumber="496" value="1.0">1</number>
|
||
) with fine reticulate sculpture restricted to anterior third.........................
|
||
<taxonomicName box="[1327,1435,1221,1242]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="maryannae">
|
||
<emphasis box="[1327,1435,1221,1242]" italics="true" pageId="7" pageNumber="496">maryannae</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1436,1015,2034]" box="[151,1435,1250,1271]" pageId="7" pageNumber="496">
|
||
<keyLead box="[151,1435,1250,1271]" pageId="7" pageNumber="496">
|
||
- T
|
||
<number box="[243,252,1250,1271]" pageId="7" pageNumber="496" value="1.0">1</number>
|
||
with fine reticulate sculpture covering at least anterior half..................................................
|
||
<number box="[1423,1435,1250,1271]" pageId="7" pageNumber="496" value="4.0">4</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="496">
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
<number box="[151,166,1279,1300]" pageId="7" pageNumber="496" value="4.0">4</number>
|
||
. A majority of specimens with mesonotum sculpture present over entire area posteriad of promesonotal suture, sculpture consisting of longitudinal rows of shallow alveolae, rarely with a small central area without sculpture; known only from the Kimberley region of Western
|
||
<collectingCountry box="[468,557,1338,1359]" name="Australia" pageId="7" pageNumber="496">Australia</collectingCountry>
|
||
..................................................................
|
||
<taxonomicName box="[1354,1435,1338,1359]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="stagnum">
|
||
<emphasis box="[1354,1435,1338,1359]" italics="true" pageId="7" pageNumber="496">stagnum</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
- Mesonotum sculpture restricted to promesonotal suture, posterior mesonotum and lateral curvature; central area always smooth; known only from the Cape York area of Queensland.............................................
|
||
<taxonomicName box="[1344,1435,1397,1418]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="capeyork">
|
||
<emphasis box="[1344,1435,1397,1418]" italics="true" pageId="7" pageNumber="496">capeyork</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="496">
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
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<keyLead pageId="7" pageNumber="496">
|
||
<number box="[151,165,1426,1447]" pageId="7" pageNumber="496" value="5.0">5</number>
|
||
. Anterodorsal margin of the clypeus distinctly concave, with frontolateral carinae forming angular ridges that extend beyond anteroventral clypeal margin, forming obtuse angles (
|
||
<figureCitation box="[738,795,1455,1476]" captionStart="FIGURE 5" captionStartId="15.[151,250,1924,1946]" captionTargetBox="[201,1388,181,1900]" captionTargetId="figure@15.[201,1389,181,1903]" captionTargetPageId="15" captionText="FIGURE 5. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 5 a – c: M. geminum, paratype (Litchfield NP, NT, ANA 10 – 22), d – f: M. hertogi, holotype (Lakefield NP, QLD, TERC 2), g – i: M. hoffmani, holotype (Mt Stanford Station, NT, TERC 70), j – l: M. humilior, (Alice Springs, NT, ANA 11 – 13), m – o: M. kidman, holotype (Top Springs, NT, TERC 71). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252027/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<number box="[783,795,1455,1476]" pageId="7" pageNumber="496" value="5.0">5</number>
|
||
</figureCitation>
|
||
h,
|
||
<number box="[819,831,1455,1476]" pageId="7" pageNumber="496" value="5.0">5</number>
|
||
n,
|
||
<number box="[854,865,1455,1476]" pageId="7" pageNumber="496" value="8.0">8</number>
|
||
b)............................................
|
||
<number box="[1423,1435,1455,1476]" pageId="7" pageNumber="496" value="6.0">6</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
- Anterodorsal margin of the clypeus sinuous to almost straight, frontolateral carinae forming smooth ridges that do not extend beyond anteroventral clypeal margin (
|
||
<figureCitation box="[589,647,1514,1535]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[204,1391,181,1899]" captionTargetId="figure@9.[204,1392,181,1903]" captionTargetPageId="9" captionText="FIGURE 4. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 4 a – c: M. bogischi, (Port Wakefield, SA, KSS 241), d – f: M. broschorum holotype (Dajarra, QLD, KSS 60), g – i: M. capeyork, holotype (69 km N Archer River Roadhouse, QLD, TERC 4), j – l: M. eremoides, holotype (9 km E 80 mile Beach, WA, KSS 89), m – o: M. eremum, holotype (33 km N Barkley Homestead, NT, KSS 77). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252026/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<number box="[634,647,1514,1535]" pageId="7" pageNumber="496" value="4.0">4</number>
|
||
</figureCitation>
|
||
a,
|
||
<number box="[669,681,1514,1535]" pageId="7" pageNumber="496" value="4.0">4</number>
|
||
k,
|
||
<number box="[705,717,1514,1535]" pageId="7" pageNumber="496" value="8.0">8</number>
|
||
e)........................................................
|
||
<number box="[1411,1435,1514,1535]" pageId="7" pageNumber="496" value="15.0">15</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="496">
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
<number box="[151,166,1543,1564]" pageId="7" pageNumber="496" value="6.0">6</number>
|
||
. Mesonotum sculptured over most of area posteriad of promesonotal suture (
|
||
<figureCitation box="[957,1015,1543,1564]" captionStart="FIGURE 6" captionStartId="22.[151,250,1912,1934]" captionTargetBox="[208,1379,193,1888]" captionTargetId="figure@22.[208,1379,193,1891]" captionTargetPageId="22" captionText="FIGURE 6. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 6 a – c: M. leda, lectotype (Kimberley District, WA), d – f: M. maryannae holotype (Berry Springs, NT, ANA 09 – 15), g – i: M. merepah, paratype (Lakefield NP, QLD, TERC 1), j – l: M. mitchell, holotype (Mitchell Falls, WA, TERC 17), m – o: M. oodnadatta, holotype (160 km E Mala, SA, KSS 114). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252028/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<number box="[1002,1015,1543,1564]" pageId="7" pageNumber="496" value="6.0">6</number>
|
||
</figureCitation>
|
||
l), propodeum in lateral view with dorsolateral angles almost forming a right angle, such that propodeum appears cuboid (
|
||
<figureCitation box="[983,1040,1573,1594]" captionStart="FIGURE 6" captionStartId="22.[151,250,1912,1934]" captionTargetBox="[208,1379,193,1888]" captionTargetId="figure@22.[208,1379,193,1891]" captionTargetPageId="22" captionText="FIGURE 6. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 6 a – c: M. leda, lectotype (Kimberley District, WA), d – f: M. maryannae holotype (Berry Springs, NT, ANA 09 – 15), g – i: M. merepah, paratype (Lakefield NP, QLD, TERC 1), j – l: M. mitchell, holotype (Mitchell Falls, WA, TERC 17), m – o: M. oodnadatta, holotype (160 km E Mala, SA, KSS 114). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252028/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<number box="[1028,1040,1573,1594]" pageId="7" pageNumber="496" value="6.0">6</number>
|
||
</figureCitation>
|
||
j)........................
|
||
<taxonomicName box="[1358,1435,1573,1594]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="mitchell">
|
||
<emphasis box="[1358,1435,1573,1594]" italics="true" pageId="7" pageNumber="496">mitchell</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
- Mesonotum with sculpture restricted to at most posterior half of area posteriad of promesonotal suture; propodeum in lateral view with dorsolateral angles forming an obtuse angle or rounded (
|
||
<figureCitation box="[868,925,1631,1652]" captionStart="FIGURE 5" captionStartId="15.[151,250,1924,1946]" captionTargetBox="[201,1388,181,1900]" captionTargetId="figure@15.[201,1389,181,1903]" captionTargetPageId="15" captionText="FIGURE 5. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 5 a – c: M. geminum, paratype (Litchfield NP, NT, ANA 10 – 22), d – f: M. hertogi, holotype (Lakefield NP, QLD, TERC 2), g – i: M. hoffmani, holotype (Mt Stanford Station, NT, TERC 70), j – l: M. humilior, (Alice Springs, NT, ANA 11 – 13), m – o: M. kidman, holotype (Top Springs, NT, TERC 71). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252027/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<quantity box="[913,925,1631,1652]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="7" pageNumber="496" unit="g" value="5.0">5</quantity>
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||
</figureCitation>
|
||
g,
|
||
<number box="[949,961,1631,1652]" pageId="7" pageNumber="496" value="7.0">7</number>
|
||
a).....................................
|
||
<number box="[1423,1435,1631,1652]" pageId="7" pageNumber="496" value="7.0">7</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="7" pageNumber="496">
|
||
<paragraph blockId="7.[151,1436,1015,2034]" box="[151,1436,1661,1682]" pageId="7" pageNumber="496">
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<keyLead box="[151,1436,1661,1682]" pageId="7" pageNumber="496">
|
||
<number box="[151,165,1661,1682]" pageId="7" pageNumber="496" value="7.0">7</number>
|
||
. T
|
||
<number box="[243,252,1661,1682]" pageId="7" pageNumber="496" value="1.0">1</number>
|
||
with fine reticulate sculpture over at least anterior quarter; coarse lateral cephalic strigae reaching anterior margin of eye.
|
||
<number box="[1424,1436,1661,1682]" pageId="7" pageNumber="496" value="8.0">8</number>
|
||
</keyLead>
|
||
</paragraph>
|
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<paragraph blockId="7.[151,1436,1015,2034]" box="[151,1436,1690,1711]" pageId="7" pageNumber="496">
|
||
<keyLead box="[151,1436,1690,1711]" pageId="7" pageNumber="496">
|
||
- T
|
||
<number box="[243,252,1690,1711]" pageId="7" pageNumber="496" value="1.0">1</number>
|
||
without sculpture, smooth and shining; coarse lateral cephalic strigae may or may not reach anterior margin of eye....
|
||
<number box="[1411,1436,1690,1711]" pageId="7" pageNumber="496" value="10.0">10</number>
|
||
</keyLead>
|
||
</paragraph>
|
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</keyStep>
|
||
<keyStep pageId="7" pageNumber="496">
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
<number box="[151,165,1719,1740]" pageId="7" pageNumber="496" value="8.0">8</number>
|
||
. Anteroventral margin of the clypeus without an anterior median projection, petiole node broadly rounded in posterior view...........................................................................................
|
||
<taxonomicName box="[1293,1376,1749,1770]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="humilior">
|
||
<emphasis box="[1293,1376,1749,1770]" italics="true" pageId="7" pageNumber="496">humilior</emphasis>
|
||
</taxonomicName>
|
||
(part)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
- A majority of specimens with a median projection on the anteroventral margin of the clypeus; petiole node tapering to a rounded point in posterior view..........................................................................
|
||
<number box="[1423,1435,1807,1828]" pageId="7" pageNumber="496" value="9.0">9</number>
|
||
</keyLead>
|
||
</paragraph>
|
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</keyStep>
|
||
<keyStep pageId="7" pageNumber="496">
|
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<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
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<keyLead pageId="7" pageNumber="496">
|
||
<number box="[151,165,1837,1858]" pageId="7" pageNumber="496" value="9.0">9</number>
|
||
. Mesonotum strigulate on lateral curvature, in metanotal groove and extending onto posterior mesonotum (
|
||
<figureCitation box="[1277,1337,1837,1858]" captionStart="FIGURE 7" captionStartId="27.[151,250,1912,1934]" captionTargetBox="[208,1379,193,1888]" captionTargetId="figure@27.[208,1379,193,1891]" captionTargetPageId="27" captionText="FIGURE 7. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 7 a – c: M. pilbara, holotype (4 km S Newman, WA, KSS 188), d – f: M. rothsteini, (Alice River, QLD, ES 38), g – i: M. speculum paratype (Niagara Dam, WA, KSS 218), j – l: M. stagnum, holotype (Mitchell Falls, WA, TERC 11), m – o: M. subapterum, (160 km E Mala, SA, KSS 116). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252029/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<number box="[1324,1337,1837,1858]" pageId="7" pageNumber="496" value="7.0">7</number>
|
||
</figureCitation>
|
||
c); petiole node narrow, width less than
|
||
<date box="[505,535,1866,1887]" pageId="7" pageNumber="496">
|
||
<number box="[505,517,1866,1887]" pageId="7" pageNumber="496" value="2.0">2</number>
|
||
x
|
||
</date>
|
||
eye width when viewed from above; longitudinal strigae absent from dorsal surface of propodeum (
|
||
<figureCitation box="[291,349,1895,1916]" captionStart="FIGURE 7" captionStartId="27.[151,250,1912,1934]" captionTargetBox="[208,1379,193,1888]" captionTargetId="figure@27.[208,1379,193,1891]" captionTargetPageId="27" captionText="FIGURE 7. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 7 a – c: M. pilbara, holotype (4 km S Newman, WA, KSS 188), d – f: M. rothsteini, (Alice River, QLD, ES 38), g – i: M. speculum paratype (Niagara Dam, WA, KSS 218), j – l: M. stagnum, holotype (Mitchell Falls, WA, TERC 11), m – o: M. subapterum, (160 km E Mala, SA, KSS 116). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252029/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<number box="[336,349,1895,1916]" pageId="7" pageNumber="496" value="7.0">7</number>
|
||
</figureCitation>
|
||
c); known only from the
|
||
<taxonomicName box="[587,655,1895,1916]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="526" phylum="Arthropoda" rank="species" species="pilbara">Pilbara</taxonomicName>
|
||
region of Western
|
||
<collectingCountry box="[838,926,1895,1916]" name="Australia" pageId="7" pageNumber="496">Australia</collectingCountry>
|
||
....................................
|
||
<taxonomicName box="[1364,1435,1895,1916]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="pilbara">
|
||
<emphasis box="[1364,1435,1895,1916]" italics="true" pageId="7" pageNumber="496">pilbara</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1436,1015,2034]" pageId="7" pageNumber="496">
|
||
<keyLead pageId="7" pageNumber="496">
|
||
- Mesonotum almost entirely smooth, a few strigulae on posterior and lateral portions of posterior mesonotum (
|
||
<figureCitation box="[1285,1343,1925,1946]" captionStart="FIGURE 5" captionStartId="15.[151,250,1924,1946]" captionTargetBox="[201,1388,181,1900]" captionTargetId="figure@15.[201,1389,181,1903]" captionTargetPageId="15" captionText="FIGURE 5. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 5 a – c: M. geminum, paratype (Litchfield NP, NT, ANA 10 – 22), d – f: M. hertogi, holotype (Lakefield NP, QLD, TERC 2), g – i: M. hoffmani, holotype (Mt Stanford Station, NT, TERC 70), j – l: M. humilior, (Alice Springs, NT, ANA 11 – 13), m – o: M. kidman, holotype (Top Springs, NT, TERC 71). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252027/files/figure.png" pageId="7" pageNumber="496">
|
||
Fig.
|
||
<number box="[1330,1343,1925,1946]" pageId="7" pageNumber="496" value="5.0">
|
||
<date box="[1330,1343,1925,1946]" pageId="7" pageNumber="496">5</date>
|
||
</number>
|
||
</figureCitation>
|
||
i); petiole node of medium breadth, between
|
||
<number box="[557,569,1954,1975]" pageId="7" pageNumber="496" value="2.0">2</number>
|
||
and
|
||
<number box="[616,646,1954,1975]" pageId="7" pageNumber="496" value="2.5">2.5</number>
|
||
x eye width when viewed from above; propodeal longitudinal strigae very weakly present as broken lines on posterior half only; known from the Victoria River district of the Northern Territory....
|
||
<taxonomicName box="[1338,1435,1983,2004]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="hoffmanni">
|
||
<emphasis box="[1338,1435,1983,2004]" italics="true" pageId="7" pageNumber="496">hoffmanni</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
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|
||
<keyStep lastPageId="8" lastPageNumber="497" pageId="7" pageNumber="496">
|
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<keyLead lastPageId="8" lastPageNumber="497" pageId="7" pageNumber="496">
|
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<paragraph blockId="7.[151,1436,1015,2034]" box="[151,1436,2013,2034]" pageId="7" pageNumber="496">
|
||
<number box="[151,177,2013,2034]" pageId="7" pageNumber="496" value="10.0">10</number>
|
||
. Coarse lateral cephalic strigae not reaching anterior eye margin and transverse strigae absent from dorsal surface of propodeum
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
(
|
||
<figureCitation box="[232,289,151,172]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[204,1391,181,1899]" captionTargetId="figure@9.[204,1392,181,1903]" captionTargetPageId="9" captionText="FIGURE 4. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 4 a – c: M. bogischi, (Port Wakefield, SA, KSS 241), d – f: M. broschorum holotype (Dajarra, QLD, KSS 60), g – i: M. capeyork, holotype (69 km N Archer River Roadhouse, QLD, TERC 4), j – l: M. eremoides, holotype (9 km E 80 mile Beach, WA, KSS 89), m – o: M. eremum, holotype (33 km N Barkley Homestead, NT, KSS 77). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252026/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[277,289,151,172]" pageId="8" pageNumber="497" value="4.0">4</number>
|
||
</figureCitation>
|
||
f,
|
||
<number box="[309,321,151,172]" pageId="8" pageNumber="497" value="5.0">5</number>
|
||
o).........................................................................................
|
||
<number box="[1412,1435,151,172]" pageId="8" pageNumber="497" value="11.0">11</number>
|
||
<keyLead pageId="8" pageNumber="497">
|
||
- At least some coarse lateral cephalic strigae reaching anterior eye margin; propodeal transverse strigae present on dorsal surface, although may be very weak (
|
||
<figureCitation box="[555,612,210,231]" captionStart="FIGURE 7" captionStartId="27.[151,250,1912,1934]" captionTargetBox="[208,1379,193,1888]" captionTargetId="figure@27.[208,1379,193,1891]" captionTargetPageId="27" captionText="FIGURE 7. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 7 a – c: M. pilbara, holotype (4 km S Newman, WA, KSS 188), d – f: M. rothsteini, (Alice River, QLD, ES 38), g – i: M. speculum paratype (Niagara Dam, WA, KSS 218), j – l: M. stagnum, holotype (Mitchell Falls, WA, TERC 11), m – o: M. subapterum, (160 km E Mala, SA, KSS 116). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252029/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[600,612,210,231]" pageId="8" pageNumber="497" value="7.0">7</number>
|
||
</figureCitation>
|
||
f,
|
||
<number box="[631,644,210,231]" pageId="8" pageNumber="497" value="6.0">6</number>
|
||
o,
|
||
<number box="[667,678,210,231]" pageId="8" pageNumber="497" value="8.0">8</number>
|
||
c)...........................................................
|
||
<number box="[1411,1435,210,231]" pageId="8" pageNumber="497" value="13.0">13</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyLead>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="497">
|
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<paragraph blockId="8.[151,1436,151,1464]" box="[151,1436,239,260]" pageId="8" pageNumber="497">
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<keyLead box="[151,1436,239,260]" pageId="8" pageNumber="497">
|
||
<number box="[151,176,239,260]" pageId="8" pageNumber="497" value="11.0">11</number>
|
||
. Medium to large species (ML>
|
||
<quantity box="[535,584,239,260]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="497" unit="mm" value="1.0">1mm</quantity>
|
||
) with medium to small eyes (EW <
|
||
<number box="[933,975,239,260]" pageId="8" pageNumber="497" value="0.23">0.23</number>
|
||
x LHW)..........................
|
||
<taxonomicName box="[1394,1436,239,260]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="leda">
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<emphasis box="[1394,1436,239,260]" italics="true" pageId="8" pageNumber="497">leda</emphasis>
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</taxonomicName>
|
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|
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</paragraph>
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||
<paragraph blockId="8.[151,1436,151,1464]" box="[151,1435,269,290]" pageId="8" pageNumber="497">
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<keyLead box="[151,1435,269,290]" pageId="8" pageNumber="497">
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- Relatively small species (ML <
|
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<quantity box="[532,589,269,290]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="497" unit="mm" value="1.0">1 mm</quantity>
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) with large eyes (EW>
|
||
<number box="[822,864,269,290]" pageId="8" pageNumber="497" value="0.23">0.23</number>
|
||
x LHW)......................................
|
||
<number box="[1411,1435,269,290]" pageId="8" pageNumber="497" value="12.0">12</number>
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||
</keyLead>
|
||
</paragraph>
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</keyStep>
|
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<keyStep pageId="8" pageNumber="497">
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<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
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||
<keyLead pageId="8" pageNumber="497">
|
||
<number box="[151,178,298,319]" pageId="8" pageNumber="497" value="12.0">12</number>
|
||
. Petiole node without sculpture and narrow (<
|
||
<date box="[690,722,298,319]" pageId="8" pageNumber="497">
|
||
<number box="[690,702,298,319]" pageId="8" pageNumber="497" value="2.0">2</number>
|
||
x
|
||
</date>
|
||
eye width when viewed from above); head and mesosoma light to dark brown with a reddish-orange tinge; anteroventral margin of the clypeus with a small median projection........
|
||
<taxonomicName box="[1320,1435,327,348]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="broschorum">
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||
<emphasis box="[1320,1435,327,348]" italics="true" pageId="8" pageNumber="497">broschorum</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
- Petiole node with fine reticulate sculpture laterally and a long posterobasal margin, of medium breadth (
|
||
<number box="[1255,1310,357,378]" pageId="8" pageNumber="497" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5</number>
|
||
x eye width when viewed from above); head, mesosoma and legs amber orange to orange brown; anteroventral margin of the clypeus rarely with a small median projection.................................................................
|
||
<taxonomicName box="[1365,1436,415,436]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="kidman">
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||
<emphasis box="[1365,1436,415,436]" italics="true" pageId="8" pageNumber="497">kidman</emphasis>
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||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="497">
|
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<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
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||
<keyLead pageId="8" pageNumber="497">
|
||
<number box="[151,177,445,466]" pageId="8" pageNumber="497" value="13.0">13</number>
|
||
. Longitudinal strigae absent from dorsal surface of propodeum and longitudinal depression very shallow or absent; propodeum appears dorsally rounded (
|
||
<figureCitation box="[483,542,474,495]" captionStart="FIGURE 7" captionStartId="27.[151,250,1912,1934]" captionTargetBox="[208,1379,193,1888]" captionTargetId="figure@27.[208,1379,193,1891]" captionTargetPageId="27" captionText="FIGURE 7. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 7 a – c: M. pilbara, holotype (4 km S Newman, WA, KSS 188), d – f: M. rothsteini, (Alice River, QLD, ES 38), g – i: M. speculum paratype (Niagara Dam, WA, KSS 218), j – l: M. stagnum, holotype (Mitchell Falls, WA, TERC 11), m – o: M. subapterum, (160 km E Mala, SA, KSS 116). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252029/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[530,542,474,495]" pageId="8" pageNumber="497" value="7.0">7</number>
|
||
</figureCitation>
|
||
f), dorsolateral angles of the propodeum rounded; petiole node of medium breadth (
|
||
<number box="[1361,1416,474,495]" pageId="8" pageNumber="497" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5</number>
|
||
x eye width when viewed from above)...............................................................
|
||
<taxonomicName box="[1345,1435,503,524]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="rothsteini">
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||
<emphasis box="[1345,1435,503,524]" italics="true" pageId="8" pageNumber="497">rothsteini</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
- Longitudinal strigae present on dorsal surface of propodeum either as raised convergent ridges or irregular convergent lines; longitudinal depression present; petiole node narrow (<
|
||
<date box="[762,792,562,583]" pageId="8" pageNumber="497">
|
||
<number box="[762,774,562,583]" pageId="8" pageNumber="497" value="2.0">2</number>
|
||
x
|
||
</date>
|
||
eye width when viewed from above)......................
|
||
<number box="[1411,1435,562,583]" pageId="8" pageNumber="497" value="14.0">14</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="497">
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
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||
<keyLead pageId="8" pageNumber="497">
|
||
<number box="[151,178,591,612]" pageId="8" pageNumber="497" value="14.0">14</number>
|
||
. Longitudinal strigae on the dorsal surface of the propodeum converging anteriorly, forming a triangle with a carina that extends between the dorsolateral angles (
|
||
<figureCitation box="[633,693,621,642]" captionStart="FIGURE 8" captionStartId="31.[151,250,1660,1682]" captionTargetBox="[151,1436,519,1638]" captionTargetId="figure@31.[151,1436,519,1639]" captionTargetPageId="31" captionText="FIGURE 8. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 8 a – c: M. tenebrosum, holotype (10 km N Tom Price, WA, KSS 177), d – f: M. topend, holotype (Darwin, NT, TERC 75), g – i: M. torrens, paratype (Flinders Rangers, SA, KSS 4). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252030/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[681,693,621,642]" pageId="8" pageNumber="497" value="8.0">8</number>
|
||
</figureCitation>
|
||
c); head, mesosoma and legs dark chestnut brown, except trochanters light brown; known only from the
|
||
<taxonomicName box="[506,574,650,671]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="526" phylum="Arthropoda" rank="species" species="pilbara">Pilbara</taxonomicName>
|
||
region of Western
|
||
<collectingCountry box="[757,844,650,671]" name="Australia" pageId="8" pageNumber="497">Australia</collectingCountry>
|
||
........................................
|
||
<taxonomicName box="[1320,1435,650,671]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="tenebrosum">
|
||
<emphasis box="[1320,1435,650,671]" italics="true" pageId="8" pageNumber="497">tenebrosum</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
- Longitudinal strigae on the dorsal surface of the propodeum not forming a triangle, carina absent between the dorsolateral angles; head and mesosoma dark brown with a reddish-orange tinge; known only from the stony plains region of northern South
|
||
<collectingCountry box="[287,374,738,759]" name="Australia" pageId="8" pageNumber="497">Australia</collectingCountry>
|
||
...............................................................................
|
||
<taxonomicName box="[1326,1435,738,759]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="oodnadatta">
|
||
<emphasis box="[1326,1435,738,759]" italics="true" pageId="8" pageNumber="497">oodnadatta</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="497">
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
<number box="[151,177,767,788]" pageId="8" pageNumber="497" value="15.0">15</number>
|
||
. Dorsolateral angles of the propodeum almost forming a right angle, such that the propodeum appears cuboid (
|
||
<figureCitation box="[1318,1378,767,788]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[204,1391,181,1899]" captionTargetId="figure@9.[204,1392,181,1903]" captionTargetPageId="9" captionText="FIGURE 4. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 4 a – c: M. bogischi, (Port Wakefield, SA, KSS 241), d – f: M. broschorum holotype (Dajarra, QLD, KSS 60), g – i: M. capeyork, holotype (69 km N Archer River Roadhouse, QLD, TERC 4), j – l: M. eremoides, holotype (9 km E 80 mile Beach, WA, KSS 89), m – o: M. eremum, holotype (33 km N Barkley Homestead, NT, KSS 77). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252026/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[1365,1378,767,788]" pageId="8" pageNumber="497" value="4.0">4</number>
|
||
</figureCitation>
|
||
j,
|
||
<number box="[1398,1410,767,788]" pageId="8" pageNumber="497" value="5.0">5</number>
|
||
a); propodeum laterally alveolate with irregular strigae extending onto dorsal surface or on posterior half of lateral surface; mesopleuron with at least a few strigae extending anteriorly from metapleural groove..............................
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||
<number box="[1411,1435,826,847]" pageId="8" pageNumber="497" value="16.0">16</number>
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||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
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||
<keyLead pageId="8" pageNumber="497">
|
||
- Dorsolateral angles of the propodeum clearly forming an obtuse angle (
|
||
<figureCitation box="[907,964,855,876]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[204,1391,181,1899]" captionTargetId="figure@9.[204,1392,181,1903]" captionTargetPageId="9" captionText="FIGURE 4. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 4 a – c: M. bogischi, (Port Wakefield, SA, KSS 241), d – f: M. broschorum holotype (Dajarra, QLD, KSS 60), g – i: M. capeyork, holotype (69 km N Archer River Roadhouse, QLD, TERC 4), j – l: M. eremoides, holotype (9 km E 80 mile Beach, WA, KSS 89), m – o: M. eremum, holotype (33 km N Barkley Homestead, NT, KSS 77). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252026/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[951,964,855,876]" pageId="8" pageNumber="497" value="4.0">4</number>
|
||
</figureCitation>
|
||
a,
|
||
<number box="[986,998,855,876]" pageId="8" pageNumber="497" value="5.0">5</number>
|
||
a), laterally alveolate without irregular strigae extending onto dorsal surface or on posterior half of lateral surface; mesopleuron with or without strigae...............
|
||
<number box="[1411,1435,885,906]" pageId="8" pageNumber="497" value="17.0">17</number>
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||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="497">
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
<number box="[151,177,914,935]" pageId="8" pageNumber="497" value="16.0">16</number>
|
||
. Mesonotum strigate-rugose over most of surface posteriad of promesonotal suture (
|
||
<figureCitation box="[1041,1101,914,935]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[204,1391,181,1899]" captionTargetId="figure@9.[204,1392,181,1903]" captionTargetPageId="9" captionText="FIGURE 4. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 4 a – c: M. bogischi, (Port Wakefield, SA, KSS 241), d – f: M. broschorum holotype (Dajarra, QLD, KSS 60), g – i: M. capeyork, holotype (69 km N Archer River Roadhouse, QLD, TERC 4), j – l: M. eremoides, holotype (9 km E 80 mile Beach, WA, KSS 89), m – o: M. eremum, holotype (33 km N Barkley Homestead, NT, KSS 77). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252026/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[1087,1101,914,935]" pageId="8" pageNumber="497" value="4.0">4</number>
|
||
</figureCitation>
|
||
l,
|
||
<number box="[1120,1134,914,935]" pageId="8" pageNumber="497" value="4.0">4</number>
|
||
o); sculpture on frons absent, a few strigae present only on antennal lobes (
|
||
<figureCitation box="[640,697,943,964]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[204,1391,181,1899]" captionTargetId="figure@9.[204,1392,181,1903]" captionTargetPageId="9" captionText="FIGURE 4. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 4 a – c: M. bogischi, (Port Wakefield, SA, KSS 241), d – f: M. broschorum holotype (Dajarra, QLD, KSS 60), g – i: M. capeyork, holotype (69 km N Archer River Roadhouse, QLD, TERC 4), j – l: M. eremoides, holotype (9 km E 80 mile Beach, WA, KSS 89), m – o: M. eremum, holotype (33 km N Barkley Homestead, NT, KSS 77). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252026/files/figure.png" pageId="8" pageNumber="497">
|
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Fig.
|
||
<number box="[684,697,943,964]" pageId="8" pageNumber="497" value="4.0">4</number>
|
||
</figureCitation>
|
||
k,
|
||
<number box="[720,733,943,964]" pageId="8" pageNumber="497" value="4.0">4</number>
|
||
n); petiole node narrow, <
|
||
<date box="[985,1015,943,964]" pageId="8" pageNumber="497">
|
||
<number box="[985,997,943,964]" pageId="8" pageNumber="497" value="2.0">2</number>
|
||
x
|
||
</date>
|
||
eye width when viewed from above; colour of head and mesosoma predominantly dark chestnut brown......................................
|
||
<emphasis box="[1254,1434,973,994]" italics="true" pageId="8" pageNumber="497">
|
||
<taxonomicName box="[1254,1329,973,994]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="eremum">eremum</taxonomicName>
|
||
/
|
||
<taxonomicName box="[1340,1434,973,994]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="eremoides">eremoides</taxonomicName>
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</emphasis>
|
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</keyLead>
|
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</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
- Mesonotum rugulose-strigate on lateral curvature and in metanotal groove, smooth medially (
|
||
<figureCitation box="[1140,1198,1002,1023]" captionStart="FIGURE 5" captionStartId="15.[151,250,1924,1946]" captionTargetBox="[201,1388,181,1900]" captionTargetId="figure@15.[201,1389,181,1903]" captionTargetPageId="15" captionText="FIGURE 5. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 5 a – c: M. geminum, paratype (Litchfield NP, NT, ANA 10 – 22), d – f: M. hertogi, holotype (Lakefield NP, QLD, TERC 2), g – i: M. hoffmani, holotype (Mt Stanford Station, NT, TERC 70), j – l: M. humilior, (Alice Springs, NT, ANA 11 – 13), m – o: M. kidman, holotype (Top Springs, NT, TERC 71). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252027/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[1186,1198,1002,1023]" pageId="8" pageNumber="497" value="5.0">5</number>
|
||
</figureCitation>
|
||
c,
|
||
<number box="[1222,1234,1002,1023]" pageId="8" pageNumber="497" value="8.0">8</number>
|
||
f); frons with strigae that extend well above antennal lobes (
|
||
<figureCitation box="[611,671,1031,1052]" captionStart="FIGURE 8" captionStartId="31.[151,250,1660,1682]" captionTargetBox="[151,1436,519,1638]" captionTargetId="figure@31.[151,1436,519,1639]" captionTargetPageId="31" captionText="FIGURE 8. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 8 a – c: M. tenebrosum, holotype (10 km N Tom Price, WA, KSS 177), d – f: M. topend, holotype (Darwin, NT, TERC 75), g – i: M. torrens, paratype (Flinders Rangers, SA, KSS 4). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252030/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[659,671,1031,1052]" pageId="8" pageNumber="497" value="8.0">8</number>
|
||
</figureCitation>
|
||
e); petiole node broad,>
|
||
<number box="[923,953,1031,1052]" pageId="8" pageNumber="497" value="2.5">2.5</number>
|
||
x eye width when viewed from above (
|
||
<figureCitation box="[1351,1410,1031,1052]" captionStart="FIGURE 5" captionStartId="15.[151,250,1924,1946]" captionTargetBox="[201,1388,181,1900]" captionTargetId="figure@15.[201,1389,181,1903]" captionTargetPageId="15" captionText="FIGURE 5. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 5 a – c: M. geminum, paratype (Litchfield NP, NT, ANA 10 – 22), d – f: M. hertogi, holotype (Lakefield NP, QLD, TERC 2), g – i: M. hoffmani, holotype (Mt Stanford Station, NT, TERC 70), j – l: M. humilior, (Alice Springs, NT, ANA 11 – 13), m – o: M. kidman, holotype (Top Springs, NT, TERC 71). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252027/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[1398,1410,1031,1052]" pageId="8" pageNumber="497" value="5.0">5</number>
|
||
</figureCitation>
|
||
a); colour of head and mesosoma predominantly dark amber orange.................................
|
||
<emphasis box="[1245,1435,1061,1082]" italics="true" pageId="8" pageNumber="497">
|
||
<taxonomicName box="[1245,1332,1061,1082]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="geminum">geminum</taxonomicName>
|
||
/topendese
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="497">
|
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<paragraph blockId="8.[151,1436,151,1464]" box="[151,1436,1090,1111]" pageId="8" pageNumber="497">
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<keyLead box="[151,1436,1090,1111]" pageId="8" pageNumber="497">
|
||
<number box="[151,178,1090,1111]" pageId="8" pageNumber="497" value="17.0">17</number>
|
||
. T
|
||
<number box="[243,252,1090,1111]" pageId="8" pageNumber="497" value="1.0">1</number>
|
||
with a fine reticulate pattern over at least anterior third; colour of head and mesosoma predominantly dark amber orange
|
||
<number box="[1414,1436,1090,1111]" pageId="8" pageNumber="497" value="18.0">18</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
- T
|
||
<number box="[243,252,1119,1140]" pageId="8" pageNumber="497" value="1.0">1</number>
|
||
without sculpture, smooth and shining; colour of head and mesosoma predominantly light to dark brown, sometimes with a reddish tinge........................................................................................
|
||
<number box="[1411,1435,1149,1170]" pageId="8" pageNumber="497" value="19.0">19</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="497">
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
<number box="[151,178,1178,1199]" pageId="8" pageNumber="497" value="18.0">18</number>
|
||
. Petiole node of medium breadth,
|
||
<number box="[555,609,1178,1199]" pageId="8" pageNumber="497" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5</number>
|
||
x eye width when viewed from above; a majority of specimens with fine reticulate sculpture restricted to anterior half or less; known only from the central desert region of the Northern Territory
|
||
<taxonomicName box="[1294,1377,1207,1228]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="humilior">
|
||
<emphasis box="[1294,1377,1207,1228]" italics="true" pageId="8" pageNumber="497">humilior</emphasis>
|
||
</taxonomicName>
|
||
(part)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
- Petiole node width broad,>
|
||
<number box="[506,536,1237,1258]" pageId="8" pageNumber="497" value="2.5">2.5</number>
|
||
x eye width when viewed from above; all specimens with T
|
||
<number box="[1138,1147,1237,1258]" pageId="8" pageNumber="497" value="1.0">1</number>
|
||
completely covered in a fine reticulate pattern; known only from Cape York Peninsula.................................................
|
||
<taxonomicName box="[1365,1434,1266,1287]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="hertogi">
|
||
<emphasis box="[1365,1434,1266,1287]" italics="true" pageId="8" pageNumber="497">hertogi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="497">
|
||
<paragraph blockId="8.[151,1436,151,1464]" box="[151,1434,1295,1316]" pageId="8" pageNumber="497">
|
||
<keyLead box="[151,1434,1295,1316]" pageId="8" pageNumber="497">
|
||
<number box="[151,177,1295,1316]" pageId="8" pageNumber="497" value="19.0">19</number>
|
||
. Anterior transverse carina prominent on propodeum....................................................
|
||
<taxonomicName box="[1353,1434,1295,1316]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="bogischi">
|
||
<emphasis box="[1353,1434,1295,1316]" italics="true" pageId="8" pageNumber="497">bogischi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" box="[151,1435,1325,1346]" pageId="8" pageNumber="497">
|
||
<keyLead box="[151,1435,1325,1346]" pageId="8" pageNumber="497">
|
||
- Anterior transverse carina on propodeum very weak or absent.................................................
|
||
<number box="[1411,1435,1325,1346]" pageId="8" pageNumber="497" value="20.0">20</number>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="497">
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
<number box="[151,178,1354,1375]" pageId="8" pageNumber="497" value="20.0">20</number>
|
||
. Dorsal surface of propodeum with a shallow longitudinal depression that runs most of its length; a majority of specimens without sculpture on the posterior mesonotum (
|
||
<figureCitation box="[636,693,1383,1404]" captionStart="FIGURE 8" captionStartId="31.[151,250,1660,1682]" captionTargetBox="[151,1436,519,1638]" captionTargetId="figure@31.[151,1436,519,1639]" captionTargetPageId="31" captionText="FIGURE 8. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 8 a – c: M. tenebrosum, holotype (10 km N Tom Price, WA, KSS 177), d – f: M. topend, holotype (Darwin, NT, TERC 75), g – i: M. torrens, paratype (Flinders Rangers, SA, KSS 4). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252030/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[681,693,1383,1404]" pageId="8" pageNumber="497" value="8.0">
|
||
<date box="[681,693,1383,1404]" pageId="8" pageNumber="497">8</date>
|
||
</number>
|
||
</figureCitation>
|
||
i)......................................................
|
||
<taxonomicName box="[1366,1434,1383,1404]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="torrens">
|
||
<emphasis box="[1366,1434,1383,1404]" italics="true" pageId="8" pageNumber="497">torrens</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1436,151,1464]" pageId="8" pageNumber="497">
|
||
<keyLead pageId="8" pageNumber="497">
|
||
- Dorsal surface of the propodeum appears flat or slightly rounded along most of its length, a depression present only between posterodorsal angles; a majority of specimens finely strigulate on posterior mesonotum (
|
||
<figureCitation box="[1072,1130,1443,1464]" captionStart="FIGURE 7" captionStartId="27.[151,250,1912,1934]" captionTargetBox="[208,1379,193,1888]" captionTargetId="figure@27.[208,1379,193,1891]" captionTargetPageId="27" captionText="FIGURE 7. Workers of the M. rothsteini (Forel) species complex in lateral, head and dorsal view of the mesosoma. Fig. 7 a – c: M. pilbara, holotype (4 km S Newman, WA, KSS 188), d – f: M. rothsteini, (Alice River, QLD, ES 38), g – i: M. speculum paratype (Niagara Dam, WA, KSS 218), j – l: M. stagnum, holotype (Mitchell Falls, WA, TERC 11), m – o: M. subapterum, (160 km E Mala, SA, KSS 116). Scale bars are 0.5 mm for all views." httpUri="https://zenodo.org/record/252029/files/figure.png" pageId="8" pageNumber="497">
|
||
Fig.
|
||
<number box="[1117,1130,1443,1464]" pageId="8" pageNumber="497" value="7.0">
|
||
<date box="[1117,1130,1443,1464]" pageId="8" pageNumber="497">7</date>
|
||
</number>
|
||
</figureCitation>
|
||
i,
|
||
<number box="[1148,1160,1443,1464]" pageId="8" pageNumber="497" value="7.0">7</number>
|
||
o)...
|
||
<emphasis box="[1225,1436,1443,1464]" italics="true" pageId="8" pageNumber="497">
|
||
<taxonomicName box="[1225,1313,1443,1464]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="speculum">speculum</taxonomicName>
|
||
/
|
||
<taxonomicName box="[1325,1436,1443,1464]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="subapterum">subapterum</taxonomicName>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |