314 lines
38 KiB
XML
314 lines
38 KiB
XML
<document id="75D0F4B8E43E8CFCEA8D38F123FB6BE6" ID-DOI="10.5852/ejt.2024.932.2507" ID-ISSN="2118-9773" ID-Zenodo-Dep="11001504" ID-ZooBank="EA652637-A88D-41A7-A586-8D57770B9C0E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713609760163" checkinUser="plazi" docAuthor="Irwin, Michael E. & Winterton, Shaun L." docDate="2024" docId="EC6B87D89A43FFDCFD17C84DFE58FA75" docLanguage="en" docName="EJT.2024.932.1-33.pdf" docOrigin="European Journal of Taxonomy 932 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2507/11225" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.39:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="39" docTitle="Rinhatiana Irwin & Winterton 2024" docType="treatment" docVersion="3" lastPageNumber="4" masterDocId="1052FFA09A41FFDFFFB7CA08FFE0FF97" masterDocTitle="New genera of stiletto flies endemic to Madagascar (Therevidae: Therevinae)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="3" updateTime="1713798293351" updateUser="felipe" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="22F69FC5569F689AB8EB41588BD5F035">
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<mods:title id="4474980A1C1470DEFB023617D02E022A">New genera of stiletto flies endemic to Madagascar (Therevidae: Therevinae)</mods:title>
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<mods:name id="F8EB0EA874D59CF5B674A496BE410269" type="personal">
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<mods:namePart id="2CFDA378F1DFFBCA4D1DE8FADF6768C4">Irwin, Michael E.</mods:namePart>
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<mods:affiliation id="F019CFA49B3B056092225C2DBDCDA66D">Emeritus, University of Illinois, Urbana-Champaign, Illinois, USA. & University of Arizona, Tucson, USA.</mods:affiliation>
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<mods:nameIdentifier id="3BBA4069777A2758CDBFDC291D5049ED" type="email">meirwin@illinois.edu</mods:nameIdentifier>
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<mods:namePart id="818050ED68BBFADCE36387F0153565DB">Winterton, Shaun L.</mods:namePart>
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<mods:affiliation id="AC38F5603FDC149C05DB126F23548C25">California State Collection of Arthropods, California Department of Food & Agriculture, Sacramento, California, USA.</mods:affiliation>
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<mods:nameIdentifier id="6A5B7225EE8C111B36800F65E86E4BD6" type="email">wintertonshaun@gmail.com</mods:nameIdentifier>
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<mods:title id="E59285953EFE2488412F1E1312B87FE1">European Journal of Taxonomy</mods:title>
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<mods:part id="8E2E5C318164A8980D08E969A0649463">
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<mods:date id="928582F785B7A029D97F5101AB6856F0">2024</mods:date>
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<mods:number id="D17D9CFE5D2EB1A314FFAB7DEF5786FB">2024-04-19</mods:number>
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<mods:url id="D96D312D66CD5B108DA5D487314A4F13">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2507/11225</mods:url>
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<treatment id="EC6B87D89A43FFDCFD17C84DFE58FA75" LSID="urn:lsid:plazi:treatment:EC6B87D89A43FFDCFD17C84DFE58FA75" httpUri="http://treatment.plazi.org/id/EC6B87D89A43FFDCFD17C84DFE58FA75" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<subSubSection id="2CD865459A43FFDDFD17C84DFC74FDF7" box="[672,916,581,608]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="647D36CE9A43FFDDFD17C84DFC74FDF7" blockId="2.[373,1214,581,644]" box="[672,916,581,608]" pageId="2" pageNumber="3">
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<heading id="3F3581A29A43FFDDFD17C84DFC74FDF7" box="[672,916,581,608]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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<taxonomicName id="A3C24D4D9A43FFDDFD17C84DFCC9FDF7" authorityName="Irwin & Winterton" authorityYear="2024" box="[672,809,581,608]" class="Insecta" family="Therevidae" genus="Rinhatiana" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFD17C84DFCC9FDF7" bold="true" box="[672,809,581,608]" italics="true" pageId="2" pageNumber="3">Rinhatiana</emphasis>
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</taxonomicName>
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gen nov.
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="2CD865459A43FFDDFEC2C861FB5EFD13" box="[373,1214,617,644]" pageId="2" pageNumber="3" type="description">
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<paragraph id="647D36CE9A43FFDDFEC2C861FB5EFD13" blockId="2.[373,1214,581,644]" box="[373,1214,617,644]" pageId="2" pageNumber="3">
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urn:lsid:zoobank.org:act:
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<uuid id="10640C1B9A43FFDDFD2EC861FB5EFD13" box="[665,1214,617,644]" pageId="2" pageNumber="3">C41C01D5-81EE-46F3-B354-1161454B8A81</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="2CD865459A43FFDDFF0AC8A4FD98FD62" pageId="2" pageNumber="3" type="type_taxon">
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<paragraph id="647D36CE9A43FFDDFF0AC8A4FEB8FD52" blockId="2.[189,344,683,710]" box="[189,344,683,710]" pageId="2" pageNumber="3">
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<heading id="3F3581A29A43FFDDFF0AC8A4FEB8FD52" bold="true" box="[189,344,683,710]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="56B6EADC9A43FFDDFF0AC8A4FEB8FD52" bold="true" box="[189,344,683,710]" pageId="2" pageNumber="3">
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<typeStatus id="BB79886C9A43FFDDFF0AC8A4FF1AFD51" box="[189,250,684,710]" pageId="2" pageNumber="3">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="647D36CE9A43FFDDFF0AC8D2FD98FD62" blockId="2.[189,632,730,757]" box="[189,632,730,757]" pageId="2" pageNumber="3">
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<taxonomicName id="A3C24D4D9A43FFDDFF0AC8D2FE21FD63" authority="Irwin & Winterton, 2024" authorityName="Irwin & Winterton" authorityYear="2024" box="[189,449,730,756]" class="Insecta" family="Therevidae" genus="Rinhatiana" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="latifestuca" status="gen. et sp. nov.">
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<emphasis id="56B6EADC9A43FFDDFF0AC8D2FE21FD63" box="[189,449,730,756]" italics="true" pageId="2" pageNumber="3">Rinhatiana latifestuca</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4D8557A79A43FFDDFE7FC8D3FD98FD62" box="[456,632,731,757]" pageId="2" pageNumber="3" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="2CD865459A43FFDDFF0AC914FC72FA8F" pageId="2" pageNumber="3" type="diagnosis">
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||
<paragraph id="647D36CE9A43FFDDFF0AC914FED6FCA1" blockId="2.[189,310,796,822]" box="[189,310,796,822]" pageId="2" pageNumber="3">
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<heading id="3F3581A29A43FFDDFF0AC914FED6FCA1" bold="true" box="[189,310,796,822]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="56B6EADC9A43FFDDFF0AC914FED6FCA1" bold="true" box="[189,310,796,822]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="647D36CE9A43FFDDFF0AC943FC27FC3A" blockId="2.[189,1399,843,941]" pageId="2" pageNumber="3">Male and female with single row of postocular macrosetae (rarely more); prosternal setae absent medially; two pairs of scutellar macrosetae; setae absent on metanepisternum and posterior surface of mid coxa; wing vein R 1 with setulae along dorsal surface; wing cell m 3 open.</paragraph>
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<paragraph id="647D36CE9A43FFDDFF0AC9DDFE26FC78" blockId="2.[189,454,981,1007]" box="[189,454,981,1007]" pageId="2" pageNumber="3">
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<heading id="3F3581A29A43FFDDFF0AC9DDFE26FC78" bold="true" box="[189,454,981,1007]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="56B6EADC9A43FFDDFF0AC9DDFE26FC78" bold="true" box="[189,454,981,1007]" pageId="2" pageNumber="3">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="647D36CE9A43FFDDFF0ACE0CFC72FA8F" blockId="2.[188,1399,1028,1304]" pageId="2" pageNumber="3">
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<taxonomicName id="A3C24D4D9A43FFDDFF0ACE0CFEDFFB89" authority="Irwin & Winterton, 2024" authorityName="Irwin & Winterton" authorityYear="2024" box="[189,319,1028,1054]" class="Insecta" family="Therevidae" genus="Rinhatiana" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="56B6EADC9A43FFDDFF0ACE0CFEDFFB89" box="[189,319,1028,1054]" italics="true" pageId="2" pageNumber="3">Rinhatiana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4D8557A79A43FFDDFEF0CE0CFE52FB89" box="[327,434,1028,1054]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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belongs to the subfamily
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<taxonomicName id="A3C24D4D9A43FFDDFD54CE0CFC87FB89" box="[739,871,1028,1054]" class="Insecta" family="Therevidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Therevinae">Therevinae</taxonomicName>
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(characterized by the presence of adpressed, scale-like setae on the femora), but has setulae along wing vein R
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<subScript id="F846348B9A43FFDDFC73CE32FC2DFBDF" attach="left" box="[964,973,1082,1096]" fontSize="6" pageId="2" pageNumber="3">1</subScript>
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, a character shared with
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<taxonomicName id="A3C24D4D9A43FFDDFB43CE2FFA96FBD6" authority="Irwin & Winterton, 2024" authorityName="Irwin & Winterton" authorityYear="2024" box="[1268,1398,1063,1089]" class="Insecta" family="Therevidae" genus="Tianarinha" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="56B6EADC9A43FFDDFB43CE2FFA96FBD6" box="[1268,1398,1063,1089]" italics="true" pageId="2" pageNumber="3">Tianarinha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4D8557A79A43FFDDFF0ACE44FECBFBF1" box="[189,299,1100,1126]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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and the New World therevine genus
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<taxonomicName id="A3C24D4D9A43FFDDFD58CE43FBA4FBF2" authority="Malloch, 1932" authorityName="Malloch" authorityYear="1932" box="[751,1092,1099,1125]" class="Insecta" family="Therevidae" genus="Protothereva" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFD58CE43FC6AFBF2" box="[751,906,1099,1125]" italics="true" pageId="2" pageNumber="3">Protothereva</emphasis>
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<bibRefCitation id="00534B3F9A43FFDDFC25CE43FBA4FBF2" author="Malloch J. R." box="[914,1092,1099,1125]" pageId="2" pageNumber="3" pagination="199 - 293" refId="ref13152" refString="Malloch J. R. 1932. Rhagionidae, Therevidae. In: British Museum (Natural History), Dept. of Entomology (eds) Diptera of Patagonia and South Chile, Based Mainly on Material in the British Museum (Natural History). Part V. Fascicle 3. - Rhagionidae (Leptidae), Therevidae, Scenopinidae, Mydaidae, Asilidae, Lonchopteridae: 199 - 293. Trustees of the British Museum, London." type="book chapter" year="1932">Malloch, 1932</bibRefCitation>
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</taxonomicName>
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. It can be separated from
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<taxonomicName id="A3C24D4D9A43FFDDFF0ACE67FEDFFB1E" authorityName="Irwin & Winterton" authorityYear="2024" box="[189,319,1135,1161]" class="Insecta" family="Therevidae" genus="Tianarinha" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFF0ACE67FEDFFB1E" box="[189,319,1135,1161]" italics="true" pageId="2" pageNumber="3">Tianarinha</emphasis>
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</taxonomicName>
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by the absence of prosternal setae and two pairs of scutellar setae (in
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<taxonomicName id="A3C24D4D9A43FFDDFBC0CE67FB19FB1E" authorityName="Irwin & Winterton" authorityYear="2024" box="[1143,1273,1135,1161]" class="Insecta" family="Therevidae" genus="Tianarinha" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFBC0CE67FB19FB1E" box="[1143,1273,1135,1161]" italics="true" pageId="2" pageNumber="3">Tianarinha</emphasis>
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</taxonomicName>
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prosternal setae are present medially and there is only one pair of scutellar setae). In addition, it is separated from
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<taxonomicName id="A3C24D4D9A43FFDDFF49CEBEFE2DFB47" authorityName="Krober" authorityYear="1936" box="[254,461,1206,1232]" class="Insecta" family="Therevidae" genus="Schoutedenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFF49CEBEFE2DFB47" box="[254,461,1206,1232]" italics="true" pageId="2" pageNumber="3">Schoutedenomyia</emphasis>
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</taxonomicName>
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by the presence of regular sized macrosetae on the foretibia (relatively small in
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<taxonomicName id="A3C24D4D9A43FFDDFF0ACED2FE6CFB63" authorityName="Krober" authorityYear="1936" box="[189,396,1242,1268]" class="Insecta" family="Therevidae" genus="Schoutedenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFF0ACED2FE6CFB63" box="[189,396,1242,1268]" italics="true" pageId="2" pageNumber="3">Schoutedenomyia</emphasis>
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</taxonomicName>
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). It is distinguished from
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<taxonomicName id="A3C24D4D9A43FFDDFD03CED3FCACFB62" authorityName="Lyneborg" authorityYear="1976" box="[692,844,1243,1269]" class="Insecta" family="Therevidae" genus="Stenopomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFD03CED3FCACFB62" box="[692,844,1243,1269]" italics="true" pageId="2" pageNumber="3">Stenopomyia</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="A3C24D4D9A43FFDDFC35CED2FBE1FB63" authorityName="Lyneborg" authorityYear="1976" box="[898,1025,1242,1268]" class="Insecta" family="Therevidae" genus="Stenosathe" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFC35CED2FBE1FB63" box="[898,1025,1242,1268]" italics="true" pageId="2" pageNumber="3">Stenosathe</emphasis>
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</taxonomicName>
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by the wavy R
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<subScript id="F846348B9A43FFDDFB19CEE5FB57FB6C" attach="left" box="[1198,1207,1261,1275]" fontSize="6" pageId="2" pageNumber="3">4</subScript>
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vein in the wing (straight or gradually curved in
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<taxonomicName id="A3C24D4D9A43FFDDFD86CEF6FD29FA8F" authorityName="Lyneborg" authorityYear="1976" box="[561,713,1278,1304]" class="Insecta" family="Therevidae" genus="Stenopomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFD86CEF6FD29FA8F" box="[561,713,1278,1304]" italics="true" pageId="2" pageNumber="3">Stenopomyia</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="A3C24D4D9A43FFDDFCB5CEF6FC61FA8F" authorityName="Lyneborg" authorityYear="1976" box="[770,897,1278,1304]" class="Insecta" family="Therevidae" genus="Stenosathe" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="56B6EADC9A43FFDDFCB5CEF6FC61FA8F" box="[770,897,1278,1304]" italics="true" pageId="2" pageNumber="3">Stenosathe</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="2CD865459A43FFDDFF0ACF48FA8EF9AC" pageId="2" pageNumber="3" type="etymology">
|
||
<paragraph id="647D36CE9A43FFDDFF0ACF48FEA4FACD" blockId="2.[189,324,1344,1370]" box="[189,324,1344,1370]" pageId="2" pageNumber="3">
|
||
<heading id="3F3581A29A43FFDDFF0ACF48FEA4FACD" bold="true" box="[189,324,1344,1370]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="56B6EADC9A43FFDDFF0ACF48FEA4FACD" bold="true" box="[189,324,1344,1370]" pageId="2" pageNumber="3">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="647D36CE9A43FFDDFF0ACF67FA8EF9AC" blockId="2.[189,1399,1391,1596]" pageId="2" pageNumber="3">
|
||
ʻ
|
||
<taxonomicName id="A3C24D4D9A43FFDDFF70CF67FEA9FA1E" authorityName="Irwin & Winterton" authorityYear="2024" box="[199,329,1391,1417]" class="Insecta" family="Therevidae" genus="Rinhatiana" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A43FFDDFF70CF67FEA9FA1E" box="[199,329,1391,1417]" italics="true" pageId="2" pageNumber="3">Rinhatiana</emphasis>
|
||
</taxonomicName>
|
||
ʼ is a combination of the nicknames of two key personnel in
|
||
<collectingCountry id="1CD5765E9A43FFDDFBA8CF67FB4DFA1E" box="[1055,1197,1391,1417]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
|
||
who contributed significantly to the project on the
|
||
<taxonomicName id="A3C24D4D9A43FFDDFDE9CF9AFD58FA3B" authorityName="Linnaeus" authorityYear="1758" box="[606,696,1426,1452]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
|
||
of
|
||
<collectingCountry id="1CD5765E9A43FFDDFD53CF9AFC97FA3B" box="[740,887,1426,1452]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
|
||
. We have the honor of naming this genus after Rasolondalao Harin’Hala Hasinjaka (also known as Rin’ha) and Razafindratsita Vololontiana (also known as Tiana). Rin’ha managed and coordinated the Malaise trapping and sample sorting for the project throughout
|
||
<collectingCountry id="1CD5765E9A43FFDDFE15CFF6FDD2F98F" box="[418,562,1534,1560]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
|
||
, while Tiana managed the administrative aspects of the project within the
|
||
<collectingCountry id="1CD5765E9A43FFDDFF5FCC29FE96F9AC" box="[232,374,1569,1595]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
|
||
Institut pour la Conservation des Ecosystemes Tropicaux (MICET). Gender is female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2CD865459A43FFDCFF0ACC6BFB5AFC63" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="647D36CE9A43FFDDFF0ACC6BFEAEF9EA" blockId="2.[189,334,1635,1661]" box="[189,334,1635,1661]" pageId="2" pageNumber="3">
|
||
<heading id="3F3581A29A43FFDDFF0ACC6BFEAEF9EA" bold="true" box="[189,334,1635,1661]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="56B6EADC9A43FFDDFF0ACC6BFEAEF9EA" bold="true" box="[189,334,1635,1661]" pageId="2" pageNumber="3">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="647D36CE9A43FFDDFF0ACC9AFCE7F93B" blockId="2.[189,775,1682,1709]" box="[189,775,1682,1709]" pageId="2" pageNumber="3">Relatively small individuals with sparse vestiture.</paragraph>
|
||
<paragraph id="647D36CE9A43FFDDFF0ACCDDFA97F87F" blockId="2.[189,1399,1748,2025]" pageId="2" pageNumber="3">HEAD. Higher than long; frons flat, pubescent silver-tan dorsally, silver ventrally, matte black band medially; frons width strongly sexually dimorphic, male frons width at narrowest point narrower than ocellar tubercle width, nearly contiguous in some species, inner margin of female compound eyes subparallel; parafacial without setae; face flat; postocular ridge of male and female with single row of macrosetae along postocular ridge, rarely two distinct rows; occiput pubescence dull silver to gray or brown; gena rounded; antenna subequal to head length, positioned on lower half of head, directed anteriorly; scape relatively short, cylindrical, less than 3× pedicel length; flagellum tapered cone-shaped, subequal to combined length of scape and pedicel, style apical, elongate; palpus slender, mouthparts short.</paragraph>
|
||
<paragraph id="647D36CE9A42FFDCFF0ACB01FD91FDE9" blockId="3.[188,1399,265,638]" pageId="3" pageNumber="4">
|
||
THORAX. Central depression of prosternum bare, without setae; cervical sclerite lacking macrosetae; scutum covered with filiform setae, often of variable length; pleuron overlain with silver-tan pubescence; metanepisternum with postspiracular setae absent; metakatepisternum with setae absent; femora relatively short, hind femur and tibia distinctly longer and thicker than fore and mid legs; posterior surfaces of mid- and hind coxae without setae, hind coxal knob present; femoral vestiture includes filiform seta admixed with adpressed, lanceolate and scale-like setae; forefemur and midfemur macrosetae absent, hind femur with 0–3 subapical anteroventral macrosetae, posteroventral macrosetae present as subapical series; tibial macrosetae regular sized; scutal chaetotaxy (pairs of macrosetae): notopleural, 3; supra alar, 1 (rarely 2); post alar, 1; dorsocentral, 0–1; scutellar, 2; wing smoky (light infuscate), vein R
|
||
<subScript id="F846348B9A42FFDCFB37C83AFB69FDD7" attach="left" box="[1152,1161,562,576]" fontSize="6" pageId="3" pageNumber="4">1</subScript>
|
||
with setulae present on dorsal surface, R 4 slightly wavy, cell m 3 open, veins M 3 and M 4 separate to margin, wing membrane uniformly covered with microtrichia.
|
||
</paragraph>
|
||
<paragraph id="647D36CE9A42FFDCFF0AC8AEFC72FD28" blockId="3.[189,914,677,704]" box="[189,914,677,704]" pageId="3" pageNumber="4">ABDOMEN. Relatively slender, slightly narrowed along length.</paragraph>
|
||
<paragraph id="647D36CE9A42FFDCFF0AC8EFFB5AFC63" blockId="3.[189,1399,742,1012]" pageId="3" pageNumber="4">TERMINALIA. Epandrium slightly arched, posterolateral corners thickened and pointed; gonocoxites separate medially with posteromedial margins proximal; inner gonocoxal process (igp) present, not articulated; outer gonocoxal processes well-developed; ventral lobe large, elongate and narrow apically; phallus with dorsal apodeme of parameral sheath triangular, separate from gonocoxites laterally; ventral apodeme single lobe, narrow, single sub-triangular lobe; distiphallus narrow, short; female tergite 8 elongate with anteromedial process; sternite 8 posterior margin emarginate posteromedially; female acanthophorite setae as two sets (A1 & A2), A1 setae usually enlarged; spermathecal sac as single lobe with spermathecal ducts originating on main spermathecal sac duct; two spermathecae.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2CD865459A42FFDCFF0ACE13FE58FA75" pageId="3" pageNumber="4" type="discussion">
|
||
<paragraph id="647D36CE9A42FFDCFF0ACE13FECFFBA2" blockId="3.[189,303,1051,1077]" box="[189,303,1051,1077]" pageId="3" pageNumber="4">
|
||
<heading id="3F3581A29A42FFDCFF0ACE13FECFFBA2" bold="true" box="[189,303,1051,1077]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="56B6EADC9A42FFDCFF0ACE13FECFFBA2" bold="true" box="[189,303,1051,1077]" pageId="3" pageNumber="4">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="647D36CE9A42FFDCFF0ACE41FE58FA75" blockId="3.[188,1400,1097,1506]" pageId="3" pageNumber="4">
|
||
<taxonomicName id="A3C24D4D9A42FFDCFF0ACE41FEA2FBF4" authority="Irwin & Winterton, 2024" authorityName="Irwin & Winterton" authorityYear="2024" box="[189,322,1097,1123]" class="Insecta" family="Therevidae" genus="Rinhatiana" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56B6EADC9A42FFDCFF0ACE41FEA2FBF4" box="[189,322,1097,1123]" italics="true" pageId="3" pageNumber="4">Rinhatiana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4D8557A79A42FFDCFEFBCE42FE5CFBF3" box="[332,444,1098,1124]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is a distinctive genus of diminutive flies likely the sister genus to
|
||
<taxonomicName id="A3C24D4D9A42FFDCFB46CE41FA97FBF4" authority="Irwin & Winterton, 2024" authorityName="Irwin & Winterton" authorityYear="2024" box="[1265,1399,1097,1123]" class="Insecta" family="Therevidae" genus="Tianarinha" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56B6EADC9A42FFDCFB46CE41FA97FBF4" box="[1265,1399,1097,1123]" italics="true" pageId="3" pageNumber="4">Tianarinha</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4D8557A79A42FFDCFF0ACE64FEC9FB11" box="[189,297,1132,1158]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
The presence of setulae along the dorsal surface of wing vein R 1 is known in all
|
||
<taxonomicName id="A3C24D4D9A42FFDCFB46CE64FA97FB11" authorityName="Lyneborg" authorityYear="1976" box="[1265,1399,1132,1158]" class="Insecta" family="Therevidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Phycusinae">Phycusinae</taxonomicName>
|
||
(except for
|
||
<taxonomicName id="A3C24D4D9A42FFDCFEFECE86FD84FB3E" authority="Irwin, 1977" authorityName="Irwin" authorityYear="1977" box="[329,612,1166,1193]" class="Insecta" family="Therevidae" genus="Schlingeria" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFEFECE86FE31FB3F" box="[329,465,1166,1192]" italics="true" pageId="3" pageNumber="4">Schlingeria</emphasis>
|
||
<bibRefCitation id="00534B3F9A42FFDCFE6FCE86FD84FB3E" author="Irwin M. E." box="[472,612,1166,1193]" pageId="3" pageNumber="4" pagination="422 - 451" refId="ref12595" refString="Irwin M. E. 1977. Two new genera and four new species of the Pherocera - group from western North America, with observations on habitats and behavior (Diptera: Therevidae: Phycinae). Proceedings of the Entomological Society of Washington 79 (3): 422 - 451." type="journal article" year="1977">Irwin, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
) and in only one genus of
|
||
<taxonomicName id="A3C24D4D9A42FFDCFC1FCE86FBB0FB3F" box="[936,1104,1166,1192]" class="Insecta" family="Therevidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Xestomyzinae">Xestomyzinae</taxonomicName>
|
||
(
|
||
<taxonomicName id="A3C24D4D9A42FFDCFBD5CE87FF1CFB5C" authority="Coquillett, 1898" authorityName="Coquillett" authorityYear="1898" class="Insecta" family="Therevidae" genus="Henicomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFBD5CE87FB10FB3F" box="[1122,1264,1167,1192]" italics="true" pageId="3" pageNumber="4">Henicomyia</emphasis>
|
||
<bibRefCitation id="00534B3F9A42FFDCFB4ECE86FF1CFB5C" author="Coquillett D. W." pageId="3" pageNumber="4" pagination="187 - 188" refId="ref12383" refString="Coquillett D. W. 1898. A new dipterous genus belonging to the Therevidae. Journal of the New York Entomological Society 6 (3): 187 - 188." type="journal article" year="1898">Coquillett, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
). They are absent in all known
|
||
<taxonomicName id="A3C24D4D9A42FFDCFDD3CEB9FCFAFB5C" authorityName="Winterton" authorityYear="2001" box="[612,794,1201,1227]" class="Insecta" family="Agapophytinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Agapophytinae</taxonomicName>
|
||
, while in
|
||
<taxonomicName id="A3C24D4D9A42FFDCFC3DCEB9FBEEFB5C" box="[906,1038,1201,1227]" class="Insecta" family="Therevidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Therevinae">Therevinae</taxonomicName>
|
||
setulate along R
|
||
<subScript id="F846348B9A42FFDCFB66CECCFB3AFB45" attach="left" box="[1233,1242,1220,1234]" fontSize="6" pageId="3" pageNumber="4">1</subScript>
|
||
is only found in the South American genus
|
||
<taxonomicName id="A3C24D4D9A42FFDCFD94CEDCFD5DFB79" authorityName="Malloch" authorityYear="1932" box="[547,701,1236,1262]" class="Insecta" family="Therevidae" genus="Protothereva" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFD94CEDCFD5DFB79" box="[547,701,1236,1262]" italics="true" pageId="3" pageNumber="4">Protothereva</emphasis>
|
||
</taxonomicName>
|
||
and some species of the Australian genus
|
||
<taxonomicName id="A3C24D4D9A42FFDCFB08CEDCFE99FA86" authority="Macquart, 1848" authorityName="Macquart" authorityYear="1848" class="Insecta" family="Therevidae" genus="Anabarhynchus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFB08CEDCFA98FB79" box="[1215,1400,1236,1262]" italics="true" pageId="3" pageNumber="4">Anabarhynchus</emphasis>
|
||
Macquart, 1848
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="A3C24D4D9A42FFDCFE33CEFFFDFDFA86" authorityName="Malloch" authorityYear="1932" box="[388,541,1271,1297]" class="Insecta" family="Therevidae" genus="Protothereva" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFE33CEFFFDFDFA86" box="[388,541,1271,1297]" italics="true" pageId="3" pageNumber="4">Protothereva</emphasis>
|
||
</taxonomicName>
|
||
is placed in the predominantly New World
|
||
<taxonomicName id="A3C24D4D9A42FFDCFBA0CEFFFB6FFA86" authorityName="Rondani" authorityYear="1848" box="[1047,1167,1271,1297]" class="Insecta" family="Therevidae" genus="Cerocatus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFBA0CEFFFB6FFA86" box="[1047,1167,1271,1297]" italics="true" pageId="3" pageNumber="4">Cerocatus</emphasis>
|
||
</taxonomicName>
|
||
genus group, likely close to
|
||
<taxonomicName id="A3C24D4D9A42FFDCFEACCF11FD2FFAA3" authority="Irwin & Lyneborg, 1981" authorityName="Irwin & Lyneborg" authorityYear="1981" box="[283,719,1305,1332]" class="Insecta" family="Therevidae" genus="Brachylinga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFEACCF11FE4CFAA4" box="[283,428,1305,1331]" italics="true" pageId="3" pageNumber="4">Brachylinga</emphasis>
|
||
<bibRefCitation id="00534B3F9A42FFDCFE04CF11FD2FFAA3" author="Irwin M. E. & Lyneborg L." box="[435,719,1305,1332]" pageId="3" pageNumber="4" pagination="193 - 277" refId="ref12692" refString="Irwin M. E. & Lyneborg L. 1981 [1980]. The genera of Nearctic Therevidae. Illinois Natural History Survey Bulletin 32: 193 - 277. https: // doi. org / 10.21900 / j. inhs. v 32.143" type="journal article" year="1981">Irwin & Lyneborg, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
, while
|
||
<taxonomicName id="A3C24D4D9A42FFDCFC95CF11FC44FAA4" authorityName="Irwin & Winterton" authorityYear="2024" box="[802,932,1305,1331]" class="Insecta" family="Therevidae" genus="Rinhatiana" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFC95CF11FC44FAA4" box="[802,932,1305,1331]" italics="true" pageId="3" pageNumber="4">Rinhatiana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A3C24D4D9A42FFDCFC6ECF11FBBBFAA4" authorityName="Irwin & Winterton" authorityYear="2024" box="[985,1115,1305,1331]" class="Insecta" family="Therevidae" genus="Tianarinha" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFC6ECF11FBBBFAA4" box="[985,1115,1305,1331]" italics="true" pageId="3" pageNumber="4">Tianarinha</emphasis>
|
||
</taxonomicName>
|
||
are placed in the largely Old World
|
||
<taxonomicName id="A3C24D4D9A42FFDCFE8CCF34FE7AFAC1" authorityName="Latreille" authorityYear="1796" box="[315,410,1340,1366]" class="Insecta" family="Therevidae" genus="Thereva" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFE8CCF34FE7AFAC1" box="[315,410,1340,1366]" italics="true" pageId="3" pageNumber="4">Thereva</emphasis>
|
||
</taxonomicName>
|
||
genus group, closely related to
|
||
<taxonomicName id="A3C24D4D9A42FFDCFCBECF35FC40FAC0" authorityName="Lyneborg" authorityYear="1976" box="[777,928,1341,1367]" class="Insecta" family="Therevidae" genus="Stenopomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFCBECF35FC40FAC0" box="[777,928,1341,1367]" italics="true" pageId="3" pageNumber="4">Stenopomyia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A3C24D4D9A42FFDCFC1ACF34FBCCFAC1" authorityName="Lyneborg" authorityYear="1976" box="[941,1068,1340,1366]" class="Insecta" family="Therevidae" genus="Stenosathe" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFC1ACF34FBCCFAC1" box="[941,1068,1340,1366]" italics="true" pageId="3" pageNumber="4">Stenosathe</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A3C24D4D9A42FFDCFB8DCF34FA97FAC1" authority="Lyneborg, 2001" authorityName="Lyneborg" authorityYear="2001" box="[1082,1399,1340,1366]" class="Insecta" family="Therevidae" genus="Megapalla" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFB8DCF34FB59FAC1" box="[1082,1209,1340,1366]" italics="true" pageId="3" pageNumber="4">Megapalla</emphasis>
|
||
Lyneborg, 2001
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A3C24D4D9A42FFDCFF58CF57FE5EFAEE" authorityName="Krober" authorityYear="1936" box="[239,446,1375,1401]" class="Insecta" family="Therevidae" genus="Schoutedenomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFF58CF57FE5EFAEE" box="[239,446,1375,1401]" italics="true" pageId="3" pageNumber="4">Schoutedenomyia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="00534B3F9A42FFDCFE7BCF57FD28FAEE" author="Webb D. W. & Metz M. A." box="[460,712,1375,1402]" pageId="3" pageNumber="4" pagination="1 - 12" refId="ref13319" refString="Webb D. W. & Metz M. A. 2003. The South American genus Protothereva Malloch (Diptera: Therevidae: Therevinae) with description of two new species. Zootaxa 234 (1): 1 - 12. https: // doi. org / 10.11646 / zootaxa. 234.1.1" type="journal article" year="2003">Webb & Metz (2003)</bibRefCitation>
|
||
mentioned that some undescribed species of
|
||
<taxonomicName id="A3C24D4D9A42FFDCFB68CF57FA97FAEE" authorityName="Lyneborg" authorityYear="1976" box="[1247,1399,1375,1401]" class="Insecta" family="Therevidae" genus="Stenopomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFB68CF57FA97FAEE" box="[1247,1399,1375,1401]" italics="true" pageId="3" pageNumber="4">Stenopomyia</emphasis>
|
||
</taxonomicName>
|
||
had R 1 setulae. Those species are described here in the genera
|
||
<taxonomicName id="A3C24D4D9A42FFDCFC16CF8AFBC3FA0B" authorityName="Irwin & Winterton" authorityYear="2024" box="[929,1059,1410,1436]" class="Insecta" family="Therevidae" genus="Rinhatiana" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFC16CF8AFBC3FA0B" box="[929,1059,1410,1436]" italics="true" pageId="3" pageNumber="4">Rinhatiana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A3C24D4D9A42FFDCFBEACF8AFB3FFA0B" authorityName="Irwin & Winterton" authorityYear="2024" box="[1117,1247,1410,1436]" class="Insecta" family="Therevidae" genus="Tianarinha" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFBEACF8AFB3FFA0B" box="[1117,1247,1410,1436]" italics="true" pageId="3" pageNumber="4">Tianarinha</emphasis>
|
||
</taxonomicName>
|
||
. In addition, one previously described species,
|
||
<taxonomicName id="A3C24D4D9A42FFDCFDEACFADFBD3FA29" authority="Lyneborg, 1976" authorityName="Lyneborg" authorityYear="1976" box="[605,1075,1444,1471]" class="Insecta" family="Therevidae" genus="Stenopomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="distincta">
|
||
<emphasis id="56B6EADC9A42FFDCFDEACFADFC88FA29" box="[605,872,1444,1471]" italics="true" pageId="3" pageNumber="4">Stenopomyia distincta</emphasis>
|
||
<bibRefCitation id="00534B3F9A42FFDCFCC6CFACFBD3FA29" author="Lyneborg L." box="[881,1075,1444,1470]" pageId="3" pageNumber="4" pagination="189 - 346" refId="ref12921" refString="Lyneborg L. 1976. A revision of the therevine stiletto-flies (Diptera: Therevidae) of the Ethiopian Region. Bulletin of the British Museum (Natural History), Entomology 33 (3): 189 - 346." type="journal article" year="1976">Lyneborg, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
, also has R
|
||
<subScript id="F846348B9A42FFDCFB77CFBFFB29FA52" attach="left" box="[1216,1225,1463,1477]" fontSize="6" pageId="3" pageNumber="4">1</subScript>
|
||
setulae and is placed in
|
||
<taxonomicName id="A3C24D4D9A42FFDCFE98CFCFFE51FA76" authorityName="Irwin & Winterton" authorityYear="2024" box="[303,433,1479,1505]" class="Insecta" family="Therevidae" genus="Rinhatiana" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56B6EADC9A42FFDCFE98CFCFFE51FA76" box="[303,433,1479,1505]" italics="true" pageId="3" pageNumber="4">Rinhatiana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |