227 lines
18 KiB
XML
227 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/nl.45.78574" ID-GBIF-Dataset="396526d5-15c6-4ccf-9aa7-71d65c160998" ID-Pensoft-Pub="2367-5365-45-169" ID-Pensoft-UUID="AFF4A590A57550959B0E0010A2248D23" ID-ZooBank="23C83728C2BD47A7BEF2519AE8B1B42E" ModsDocID="2367-5365-45-169" checkinTime="1650371355456" checkinUser="pensoft" docAuthor="Mey, Wolfram" docDate="2022" docId="1B9D08768ED65CCEB5F351DE7ED74251" docLanguage="en" docName="NotaLepidopt 45: 169-190" docOrigin="Nota Lepidopterologica 45" docPubDate="2022-04-18" docSource="http://dx.doi.org/10.3897/nl.45.78574" docTitle="Homadaula schaeuffelei Mey 2022, comb. nov." docType="treatment" docVersion="2" id="AFF4A590A57550959B0E0010A2248D23" lastPageNumber="169" masterDocId="AFF4A590A57550959B0E0010A2248D23" masterDocTitle="Taxonomic notes on Palearctic taxa of Galacticidae, a little-known family of Lepidoptera (Galacticoidea)" masterLastPageNumber="190" masterPageNumber="169" pageId="0" pageNumber="169" updateTime="1650372000879" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic notes on Palearctic taxa of Galacticidae, a little-known family of Lepidoptera (Galacticoidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mey, Wolfram</mods:namePart>
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<mods:affiliation>Museum fuer Naturkunde, Leibniz Institute of Evolution and Biodiversity Research, Invalidenstr. 43, 10115 Berlin, Germany; wolfram. mey @ mfn. de</mods:affiliation>
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<mods:nameIdentifier type="email">wolfram.mey@gmx.de</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Nota Lepidopterologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-04-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>45</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>169</mods:start>
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<mods:end>190</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/nl.45.78574</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/nl.45.78574</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2367-5365-45-169</mods:identifier>
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<mods:identifier type="ZooBank">23C83728C2BD47A7BEF2519AE8B1B42E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">AFF4A590A57550959B0E0010A2248D23</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:1B9D08768ED65CCEB5F351DE7ED74251" httpUri="http://treatment.plazi.org/id/1B9D08768ED65CCEB5F351DE7ED74251" lastPageNumber="169" pageId="0" pageNumber="169">
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<subSubSection pageId="0" pageNumber="169" type="nomenclature">
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<paragraph pageId="0" pageNumber="169">
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<taxonomicName LSID="1B9D0876-8ED6-5CCE-B5F3-51DE7ED74251" authority="(Amsel, 1959)" authorityName="Mey" authorityYear="2022" baseAuthorityName="Amsel" baseAuthorityYear="1959" class="Insecta" family="Galacticidae" genus="Homadaula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homadaula schaeuffelei" order="Lepidoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="schaeuffelei" status="comb. nov.">Homadaula schaeuffelei (Amsel, 1959)</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="169">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="description">
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<paragraph pageId="0" pageNumber="169">
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<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Adults of Homadaula spp. 1 - 2. H. caradjae (Walsingham), male paralectotype and label; 3 - 4. H. punctigera (Rebel), male and label; 5 - 6. H. schaeuffelei (Amsel), female paratype and label; 7. H. schaeuffelei (Amsel), female, Iran; 8. H. schaeuffelei (Amsel), male, Pakistan (photos by W. Mey)." figureDoi="10.3897/nl.45.78574.figures1-8" httpUri="https://binary.pensoft.net/fig/673999" pageId="0" pageNumber="169">Figs 5-8</figureCitation>
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<figureCitation captionStart="Figures 18–26" captionStartId="F4" captionText="Figures 18 - 26. Male genitalia of Homadaula punctigera (Rebel), 18. Lateral; 19. Dorsal; 20. Ventral; 21 - 23. Male genitalia of H. schaeuffelei (Amsel); 21. Lateral; 22. Ventral; 23. Dorsal; 24 - 26. Female genitalia of H. schaeuffelei (Amsel); 24. Ventral; 25. Segment VII dorsal; 26. Lateral." figureDoi="10.3897/nl.45.78574.figures18-26" httpUri="https://binary.pensoft.net/fig/674002" pageId="0" pageNumber="169">, 21-26</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="reference_group">
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<paragraph pageId="0" pageNumber="169">
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<taxonomicName authorityName="Mey" authorityYear="2022" class="Insecta" family="Galacticidae" genus="Bahrlutia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bahrlutia schaeuffelei" order="Lepidoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="schaeuffelei" status="comb. nov.">Bahrlutia schaeuffelei</taxonomicName>
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Amsel, 1959: 35-36, type locality: Iran, Iranshar, Holotype and Paratype deposited in SMNS [Paratype examined].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="materials_examined">
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<paragraph pageId="0" pageNumber="169">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="169">
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type material,
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<materialsCitation collectingDate="1954-03-01" collectingDateMax="1954-03-10" collectingDateMin="1954-03-01" collectorName="H. Amsel, Mey" country="IRAN" elevation="800" location="Beluchistan" municipality="Jranshar" specimenCount="1" specimenCount-female="1" typeStatus="Paratype">
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<specimenCount type="female">1 ♀</specimenCount>
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<typeStatus>Paratype</typeStatus>
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, "
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<collectingCountry name="Iran">IRAN</collectingCountry>
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<location LSID="urn:lsid:plazi:treatment:1B9D08768ED65CCEB5F351DE7ED74251:392B65971E249C6B028470A5D1A7894E" country="IRAN" municipality="Jranshar" name="Beluchistan">Beluchistan</location>
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/
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<collectingMunicipality>Jranshar</collectingMunicipality>
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,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
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</quantity>
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/
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<collectingDate value="1954-03-01" valueMax="1954-03-10" valueMin="1954-03-01">1.-10.iii.1954</collectingDate>
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/Richter u.
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<normalizedToken originalValue="Schäuffele”">Schaeuffele"</normalizedToken>
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[printed on grey card], "
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<typeStatus>Paratypus</typeStatus>
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/leg.
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<collectorName>H. Amsel</collectorName>
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" [printed on red card], genitalia slide
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<collectorName>Mey</collectorName>
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(SMNS)
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</materialsCitation>
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.
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<materialsCitation collectingDate="1965-06-13" collectorName="Kasy, Vartian, Mey" country="Iran" elevation="900" location="Pakistan" municipality="Quetta" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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,
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<emphasis bold="true" pageId="0" pageNumber="169">
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<location LSID="urn:lsid:plazi:treatment:1B9D08768ED65CCEB5F351DE7ED74251:7FD56A39B02408636B654745C41F482B" country="Iran" municipality="Quetta" name="Pakistan">Pakistan</location>
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</emphasis>
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, [Province Baluchistan],
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<quantity metricMagnitude="5" metricUnit="m" metricValue="1.5" unit="km" value="150.0">150 km</quantity>
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south-west of
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<collectingMunicipality>Quetta</collectingMunicipality>
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,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
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</quantity>
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,
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<collectingDate value="1965-06-13">13.vi.1965</collectingDate>
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, leg.
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<collectorName>Kasy</collectorName>
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&
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<collectorName>Vartian</collectorName>
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, genitalia slide
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<collectorName>Mey</collectorName>
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23/21 (NHMW)
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</materialsCitation>
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;
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<materialsCitation collectingDate="1972-03-31" collectorName="Exped. Mus. Vind., Mey" country="Iran" location="South Iran" municipality="Bandar-Abbas" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<specimenCount type="female">1 ♀</specimenCount>
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,
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<location LSID="urn:lsid:plazi:treatment:1B9D08768ED65CCEB5F351DE7ED74251:5A8BB59E0F31449C581F0663B31C940E" country="Iran" municipality="Bandar-Abbas" name="South Iran">
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South
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<emphasis bold="true" pageId="0" pageNumber="169">Iran</emphasis>
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</location>
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,
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<quantity metricMagnitude="4" metricUnit="m" metricValue="4.0" unit="km" value="40.0">40 km</quantity>
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north of
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<collectingMunicipality>Bandar-Abbas</collectingMunicipality>
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,
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<collectingDate value="1972-03-31">31.iii.1972</collectingDate>
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,
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<collectorName>Exped. Mus. Vind.</collectorName>
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, genitalia slide
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<collectorName>Mey</collectorName>
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24/21 (NHMW)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="diagnosis">
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<paragraph pageId="0" pageNumber="169">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="169">Male without uncus and gnathos, valvae long, dorsal process broad, elongate, directed dorsad and laterad. Female without bulla seminalis, bursa copuatrix with a sclerotised antrum of ductus bursae, dorsal and ventral side of bursa with thin medial line.</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<taxonomicName family="Galacticidae" genus="H." kingdom="Animalia" lsidName="H. schaeuffelei" order="Lepidoptera" pageId="0" pageNumber="169" rank="species" species="schaeuffelei">
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<emphasis italics="true" pageId="0" pageNumber="169">H. schaeuffelei</emphasis>
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</taxonomicName>
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is an unusual species of the genus by showing a completely cream-white, external appearance of the adults, resembling externally
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<taxonomicName family="Galacticidae" genus="H." kingdom="Animalia" lsidName="H. caradjae" order="Lepidoptera" pageId="0" pageNumber="169" rank="species" species="caradjae">
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<emphasis italics="true" pageId="0" pageNumber="169">H. caradjae</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="redescription">
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<paragraph pageId="0" pageNumber="169">Redescription.</paragraph>
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<paragraph pageId="0" pageNumber="169">
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Adult (Figs
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<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Adults of Homadaula spp. 1 - 2. H. caradjae (Walsingham), male paralectotype and label; 3 - 4. H. punctigera (Rebel), male and label; 5 - 6. H. schaeuffelei (Amsel), female paratype and label; 7. H. schaeuffelei (Amsel), female, Iran; 8. H. schaeuffelei (Amsel), male, Pakistan (photos by W. Mey)." figureDoi="10.3897/nl.45.78574.figures1-8" httpUri="https://binary.pensoft.net/fig/673999" pageId="0" pageNumber="169">5-8</figureCitation>
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). Length of forewing 5-7 mm, wingspan 11.5-14.8 mm (n = 3); head white, frons with appressed scales, vertex and scapes with semi-erected scales; labial palpi as long as eye-diameter, straight, white and with brown patch on second segment; unscaled proboscis yellow-brown; antennae filiform, 0.5 of forewing length, finely ciliated on ventral side; fore-tibia with epiphysis, tarsomeres of fore and mid-legs pale brown on dorsal side, hind legs white, spurs 0.2.4.; forewing white, with small, black scales on termen and in cell forming short, indistinct or distinct patches in basal half; hindwings white, fringes white; wing venation with accessory cell in forewing and M present in cell, R4 and R5 with a short stalk, originating from distal end of accessory cell; hindwing with M3 and Cu1a confluent over short distance; one frenular bristle in male, two in females; retinaculum with white scales on underside of Sc.
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</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Male genitalia</emphasis>
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(Figs
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<figureCitation captionStart="Figures 18–26" captionStartId="F4" captionText="Figures 18 - 26. Male genitalia of Homadaula punctigera (Rebel), 18. Lateral; 19. Dorsal; 20. Ventral; 21 - 23. Male genitalia of H. schaeuffelei (Amsel); 21. Lateral; 22. Ventral; 23. Dorsal; 24 - 26. Female genitalia of H. schaeuffelei (Amsel); 24. Ventral; 25. Segment VII dorsal; 26. Lateral." figureDoi="10.3897/nl.45.78574.figures18-26" httpUri="https://binary.pensoft.net/fig/674002" pageId="0" pageNumber="169">21-23</figureCitation>
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). Tergal plate of segment VIII tapering from broad base towards an acute apex in dorsal view; pleurae membranous, simple, not lobe-like; segment IX vestigial, uncus and gnathos absent, valvae long, dorsal process broad, elongate, directed dorsad and laterad, length of apical part longer, somewhat curved dorsad, rounded apically; phallus with large bulbus ejaculatorius and short, sickle-shaped phallus, connected sub-basally with anellus.
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</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Female genitalia</emphasis>
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(Figs
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<figureCitation captionStart="Figures 18–26" captionStartId="F4" captionText="Figures 18 - 26. Male genitalia of Homadaula punctigera (Rebel), 18. Lateral; 19. Dorsal; 20. Ventral; 21 - 23. Male genitalia of H. schaeuffelei (Amsel); 21. Lateral; 22. Ventral; 23. Dorsal; 24 - 26. Female genitalia of H. schaeuffelei (Amsel); 24. Ventral; 25. Segment VII dorsal; 26. Lateral." figureDoi="10.3897/nl.45.78574.figures18-26" httpUri="https://binary.pensoft.net/fig/674002" pageId="0" pageNumber="169">24-26</figureCitation>
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). Tergum VII with median, shallow groove extending to caudal margin; sternum VIII slightly overlapping with tergum laterally; distal margin of sternum VII in ventral view slightly asymmetrical, ostium bursae on tip of henia, originating in central position of inter-segmental membrane, ductus bursae shorter than bursa, small bulla seminalis present, signa in oval bursa copulatrix absent.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="biology_ecology">
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<paragraph pageId="0" pageNumber="169">Biology.</paragraph>
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<paragraph pageId="0" pageNumber="169">Host plant unknown. Adults have been collected at the lights in March and June at elevations of 800-900 m and at lower elevations.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="distribution">
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<paragraph pageId="0" pageNumber="169">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="169">Iran and Pakistan.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="remarks">
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<paragraph pageId="0" pageNumber="169">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="169">
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The male specimen from Pakistan, widely separated geographically from the localities in south Iran, is associated with
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<taxonomicName family="Galacticidae" genus="H." kingdom="Animalia" lsidName="H. schaeuffelei" order="Lepidoptera" pageId="0" pageNumber="169" rank="species" species="schaeuffelei">
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<emphasis italics="true" pageId="0" pageNumber="169">H. schaeuffelei</emphasis>
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</taxonomicName>
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by observing the presence of a forked R4 and R5 in the forewing, a character, which is unique and unknown from other species of
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<taxonomicName authorityName="Minet" authorityYear="1986" class="Insecta" family="Galacticidae" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="family">Galacticidae</taxonomicName>
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in the region.
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</paragraph>
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<caption doi="10.3897/nl.45.78574.figures1-8" httpUri="https://binary.pensoft.net/fig/673999" pageId="0" pageNumber="169" start="Figures 1–8" startId="F1">
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Figures 1-8.</emphasis>
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Adults of
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<taxonomicName authorityName="Lower" authorityYear="1899" class="Insecta" family="Galacticidae" genus="Homadaula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homadaula" order="Lepidoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="169">Homadaula</emphasis>
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</taxonomicName>
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spp.
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<emphasis bold="true" pageId="0" pageNumber="169">1-2.</emphasis>
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<taxonomicName family="Galacticidae" genus="H." kingdom="Animalia" lsidName="H. caradjae" order="Lepidoptera" pageId="0" pageNumber="169" rank="species" species="caradjae">
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<emphasis italics="true" pageId="0" pageNumber="169">H. caradjae</emphasis>
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</taxonomicName>
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(Walsingham), male paralectotype and label;
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<emphasis bold="true" pageId="0" pageNumber="169">3-4.</emphasis>
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<taxonomicName family="Galacticidae" genus="H." kingdom="Animalia" lsidName="H. punctigera" order="Lepidoptera" pageId="0" pageNumber="169" rank="species" species="punctigera">
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<emphasis italics="true" pageId="0" pageNumber="169">H. punctigera</emphasis>
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</taxonomicName>
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(Rebel), male and label;
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<emphasis bold="true" pageId="0" pageNumber="169">5-6.</emphasis>
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<taxonomicName family="Galacticidae" genus="H." kingdom="Animalia" lsidName="H. schaeuffelei" order="Lepidoptera" pageId="0" pageNumber="169" rank="species" species="schaeuffelei">
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<emphasis italics="true" pageId="0" pageNumber="169">H. schaeuffelei</emphasis>
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</taxonomicName>
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(Amsel), female paratype and label;
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<emphasis bold="true" pageId="0" pageNumber="169">7.</emphasis>
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<taxonomicName family="Galacticidae" genus="H." kingdom="Animalia" lsidName="H. schaeuffelei" order="Lepidoptera" pageId="0" pageNumber="169" rank="species" species="schaeuffelei">
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<emphasis italics="true" pageId="0" pageNumber="169">H. schaeuffelei</emphasis>
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</taxonomicName>
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(Amsel), female, Iran;
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<emphasis bold="true" pageId="0" pageNumber="169">8.</emphasis>
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<taxonomicName family="Galacticidae" genus="H." kingdom="Animalia" lsidName="H. schaeuffelei" order="Lepidoptera" pageId="0" pageNumber="169" rank="species" species="schaeuffelei">
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<emphasis italics="true" pageId="0" pageNumber="169">H. schaeuffelei</emphasis>
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</taxonomicName>
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(Amsel), male, Pakistan (photos by W. Mey).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |