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<document ID-DOI="http://doi.org/10.5281/zenodo.4766390" ID-GBIF-Dataset="53754ae5-8913-4e45-b546-b9edae8f726b" ID-GBIF-Taxon="182135164" ID-ISSN="1854-0392" ID-Zenodo-Dep="4766390" ID-ZooBank="150B2555-91E3-4110-8BBC-25F1AAC0E6B2" checkinTime="1620964466541" checkinUser="felipe" docAuthor="Sandberg, John B. &amp; Kondratieff, Boris C." docDate="2013" docId="1B32387A8314B27DF676A7E5FA3FA3DC" docLanguage="en" docName="Illiesia.9.05.34-64.pdf" docOrigin="Illiesia 9 (5)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Isoperla OF" docType="key" docVersion="9" lastPageId="5" lastPageNumber="39" masterDocId="E70B40028317B278F535A704FF9CA45D" masterDocTitle="The Isoperla Of California (Plecoptera: Perlodidae); Updated Male Descriptions And Adult Keys For 18 Western Nearctic Species" masterLastPageNumber="64" masterPageNumber="34" pageId="3" pageNumber="37" updateTime="1632021822729" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Isoperla Of California (Plecoptera: Perlodidae); Updated Male Descriptions And Adult Keys For 18 Western Nearctic Species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sandberg, John B.</mods:namePart>
<mods:affiliation>, &amp; Boris C. Kondratieff &amp; California Dept. of Fish &amp; Wildlife - A. B. L. and CSUC Research Foundation, California State University, Chico, CA 95929 - 0555 U. S. A. E-mail: jsandberg @ csuchico. edu</mods:affiliation>
<mods:nameIdentifier type="email">jsandberg@csuchico.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kondratieff, Boris C.</mods:namePart>
<mods:affiliation>Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, Colorado 80523, U. S. A. E-mail: Boris. Kondratieff @ Colostate. edu</mods:affiliation>
<mods:nameIdentifier type="email">Boris.Kondratieff@Colostate.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>34</mods:start>
<mods:end>64</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760870</mods:identifier>
<mods:identifier type="GBIF-Dataset">53754ae5-8913-4e45-b546-b9edae8f726b</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4760870</mods:identifier>
<mods:identifier type="ZooBank">150B2555-91E3-4110-8BBC-25F1AAC0E6B2</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4766390" ID-GBIF-Taxon="182135164" ID-Zenodo-Dep="4766390" LSID="urn:lsid:plazi:treatment:1B32387A8314B27DF676A7E5FA3FA3DC" httpUri="http://treatment.plazi.org/id/1B32387A8314B27DF676A7E5FA3FA3DC" lastPageId="5" lastPageNumber="39" pageId="3" pageNumber="37">
<subSubSection pageId="3" pageNumber="37" type="nomenclature">
<paragraph blockId="3.[835,1443,224,285]" pageId="3" pageNumber="37">
<heading bold="true" centered="true" fontSize="10" level="2" pageId="3" pageNumber="37" reason="2">
<emphasis bold="true" pageId="3" pageNumber="37">
Keys to the
<taxonomicName authority="of" authorityName="OF" box="[998,1095,224,248]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[998,1095,224,248]" italics="true" pageId="3" pageNumber="37">Isoperla</emphasis>
</taxonomicName>
of California (including
<taxonomicName authorityName="Jewett" authorityYear="1954" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="37" phylum="Arthropoda" rank="species" species="rainiera">
<emphasis bold="true" italics="true" pageId="3" pageNumber="37">I. rainiera</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="1979" box="[943,1065,260,284]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="37" phylum="Arthropoda" rank="species" species="tilasqua">
<emphasis bold="true" box="[943,1065,260,284]" italics="true" pageId="3" pageNumber="37">I. tilasqua</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="2013" box="[1125,1243,261,284]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="37" phylum="Arthropoda" rank="species" species="umpqua">
<emphasis bold="true" box="[1125,1243,261,284]" italics="true" pageId="3" pageNumber="37">I. umpqua</emphasis>
</taxonomicName>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1123,1193,332,356]" pageId="3" pageNumber="37" type="description">
<paragraph blockId="3.[1123,1193,332,356]" box="[1123,1193,332,356]" pageId="3" pageNumber="37">Males</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="39" pageId="3" pageNumber="37" type="key">
<key lastPageId="5" lastPageNumber="39" pageId="3" pageNumber="37">
<keyStep pageId="3" pageNumber="37">
<paragraph blockId="3.[835,1443,403,640]" pageId="3" pageNumber="37">
<keyLead pageId="3" pageNumber="37">
1 Pronotum checkered black on yellow; aedeagus without sclerotized process; paraprocts abruptly tapering to blunt apical tips, usually secondarily barbed ventrally; vesicle a broadly rounded lobe, wider at base then apex (
<figureCitation box="[1157,1245,545,569]" captionStart="Figs" captionStartId="18.[189,236,1752,1774]" captionTargetBox="[337,1297,251,1707]" captionTargetId="figure-54@18.[334,1297,251,1707]" captionText="Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA &amp; Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764060" httpUri="https://zenodo.org/record/4764060/files/figure.png" pageId="3" pageNumber="37">Figs. 11</figureCitation>
a-d) ..…...
<taxonomicName authorityName="Frison" authorityYear="1937" box="[1360,1443,545,569]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="37" phylum="Arthropoda" rank="species" species="pinta">
<emphasis bold="true" box="[1360,1443,545,569]" italics="true" pageId="3" pageNumber="37">I. pinta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[835,1443,403,640]" pageId="3" pageNumber="37">
<keyLead pageId="3" pageNumber="37">
1’ Pronotum without checkered pigment pattern, aedeagus, paraprocts and vesicle variable ….....
<emphasis bold="true" box="[1430,1443,616,640]" pageId="3" pageNumber="37">2</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="37">
<paragraph blockId="3.[835,1443,652,783]" pageId="3" pageNumber="37">
<keyLead pageId="3" pageNumber="37">
2 Aedeagus without posterior sclerotized process (
<figureCitation box="[881,956,687,712]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="3" pageNumber="37">Figs. 1</figureCitation>
,
<figureCitation box="[969,984,688,712]" captionStart="Figs" captionStartId="8.[189,236,1752,1774]" captionTargetBox="[336,1296,246,1699]" captionTargetId="figure-54@8.[336,1296,246,1699]" captionText="Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch &amp; thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb &amp; apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle" figureDoi="http://doi.org/10.5281/zenodo.4764052" httpUri="https://zenodo.org/record/4764052/files/figure.png" pageId="3" pageNumber="37">3</figureCitation>
,
<figureCitation box="[995,1044,688,712]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="16.[189,236,1717,1739]" captionStartId-1="18.[189,236,1752,1774]" captionTargetBox-0="[334,1297,215,1671]" captionTargetBox-1="[337,1297,251,1707]" captionTargetId-0="figure-54@16.[334,1297,215,1671]" captionTargetId-1="figure-54@18.[334,1297,251,1707]" captionTargetPageId-0="16" captionText-0="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." captionText-1="Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA &amp; Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle." figureDoi-0="http://doi.org/10.5281/zenodo.4764058" figureDoi-1="http://doi.org/10.5281/zenodo.4764060" httpUri-0="https://zenodo.org/record/4764058/files/figure.png" httpUri-1="https://zenodo.org/record/4764060/files/figure.png" pageId="3" pageNumber="37">9-12</figureCitation>
,
<figureCitation box="[1057,1084,687,711]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="3" pageNumber="37">15</figureCitation>
,
<figureCitation box="[1096,1123,688,712]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="3" pageNumber="37">17</figureCitation>
) ………..……....………….
<emphasis bold="true" box="[1430,1443,687,711]" pageId="3" pageNumber="37">3</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="3.[835,1443,652,783]" pageId="3" pageNumber="37">
<keyLead pageId="3" pageNumber="37">
2’ Aedeagus with variable posterior sclerotized process (
<figureCitation box="[975,1051,759,783]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="3" pageNumber="37">Figs. 2</figureCitation>
,
<figureCitation box="[1062,1099,759,783]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStartId-0="8.[189,236,1752,1774]" captionStartId-1="11.[189,236,1717,1739]" captionStartId-2="13.[189,236,1681,1703]" captionTargetBox-0="[336,1296,246,1699]" captionTargetBox-1="[335,1297,215,1671]" captionTargetBox-2="[334,1297,195,1650]" captionTargetId-0="figure-54@8.[336,1296,246,1699]" captionTargetId-1="figure-54@11.[335,1297,215,1671]" captionTargetId-2="figure-54@13.[334,1297,195,1650]" captionTargetPageId-0="8" captionTargetPageId-1="11" captionText-0="Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch &amp; thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb &amp; apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle" captionText-1="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." captionText-2="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi-0="http://doi.org/10.5281/zenodo.4764052" figureDoi-1="http://doi.org/10.5281/zenodo.4764054" figureDoi-2="http://doi.org/10.5281/zenodo.4764056" httpUri-0="https://zenodo.org/record/4764052/files/figure.png" httpUri-1="https://zenodo.org/record/4764054/files/figure.png" httpUri-2="https://zenodo.org/record/4764056/files/figure.png" pageId="3" pageNumber="37">4-8</figureCitation>
,
<figureCitation box="[1110,1173,759,783]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="3" pageNumber="37">13-14</figureCitation>
,
<figureCitation box="[1185,1212,759,783]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="3" pageNumber="37">16</figureCitation>
,
<figureCitation box="[1224,1251,759,783]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="3" pageNumber="37">18</figureCitation>
) ………...…….
<emphasis bold="true" box="[1430,1443,759,783]" pageId="3" pageNumber="37">9</emphasis>
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3(2) Vesicle absent or weakly produced (
<figureCitation box="[1342,1437,795,819]" captionStart="Figs" captionStartId="16.[189,236,1717,1739]" captionTargetBox="[334,1297,215,1671]" captionTargetId="figure-54@16.[334,1297,215,1671]" captionText="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764058" httpUri="https://zenodo.org/record/4764058/files/figure.png" pageId="3" pageNumber="37">Figs. 9d</figureCitation>
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<figureCitation box="[873,916,830,854]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="3" pageNumber="37">15d</figureCitation>
).………………...….………….………………
<emphasis bold="true" box="[1430,1443,830,854]" pageId="3" pageNumber="37">4</emphasis>
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3’ Vesicle present and variably shaped (
<figureCitation box="[1304,1396,866,890]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="3" pageNumber="37">Figs. 1d</figureCitation>
,
<figureCitation box="[1406,1438,866,890]" captionStart="Figs" captionStartId="8.[189,236,1752,1774]" captionTargetBox="[336,1296,246,1699]" captionTargetId="figure-54@8.[336,1296,246,1699]" captionText="Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch &amp; thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb &amp; apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle" figureDoi="http://doi.org/10.5281/zenodo.4764052" httpUri="https://zenodo.org/record/4764052/files/figure.png" pageId="3" pageNumber="37">3d</figureCitation>
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<figureCitation box="[873,918,902,926]" captionStart="Figs" captionStartId="16.[189,236,1717,1739]" captionTargetBox="[334,1297,215,1671]" captionTargetId="figure-54@16.[334,1297,215,1671]" captionText="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764058" httpUri="https://zenodo.org/record/4764058/files/figure.png" pageId="3" pageNumber="37">10d</figureCitation>
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<figureCitation box="[928,972,902,926]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="3" pageNumber="37">13d</figureCitation>
,
<figureCitation box="[983,1027,902,926]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="3" pageNumber="37">18d</figureCitation>
) …………..……..…………………..
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4(3) Aedeagal posterior margin with a median patch of spinulae (
<figureCitation box="[1089,1179,973,997]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="3" pageNumber="37">Figs. 15</figureCitation>
a-c); apex usually with two small, pointed lobes (
<figureCitation box="[1284,1400,1008,1032]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="3" pageNumber="37">Figs. 15</figureCitation>
a-c) ……………………………………………..
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4’ Aedeagal posterobasal margin with a median patch of spinulae concentrated into two distinct longitudinal bands (
<figureCitation box="[1126,1210,1151,1175]" captionStart="Figs" captionStartId="16.[189,236,1717,1739]" captionTargetBox="[334,1297,215,1671]" captionTargetId="figure-54@16.[334,1297,215,1671]" captionText="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764058" httpUri="https://zenodo.org/record/4764058/files/figure.png" pageId="3" pageNumber="37">Figs. 9</figureCitation>
a-b); and a second posteroapical patch of spinulae above posterior lobe (
<figureCitation box="[938,1053,1222,1246]" captionStart="Figs" captionStartId="16.[189,236,1717,1739]" captionTargetBox="[334,1297,215,1671]" captionTargetId="figure-54@16.[334,1297,215,1671]" captionText="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764058" httpUri="https://zenodo.org/record/4764058/files/figure.png" pageId="3" pageNumber="37">Figs. 9a, c</figureCitation>
) …………………….…..
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5(3) Ninth tergum with short stout spinulae (sometimes arranged in a bipartite patch) or ninth and tenth terga with distinct patches of long stout setae near posterior margins (
<figureCitation box="[1233,1325,1364,1388]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="3" pageNumber="37">Figs. 1c</figureCitation>
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<figureCitation box="[1340,1381,1364,1388]" captionStart="Figs" captionStartId="16.[189,236,1717,1739]" captionTargetBox="[334,1297,215,1671]" captionTargetId="figure-54@16.[334,1297,215,1671]" captionText="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764058" httpUri="https://zenodo.org/record/4764058/files/figure.png" pageId="3" pageNumber="37">10c</figureCitation>
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<figureCitation box="[1395,1434,1364,1388]" captionStart="Figs" captionStartId="18.[189,236,1752,1774]" captionTargetBox="[337,1297,251,1707]" captionTargetId="figure-54@18.[334,1297,251,1707]" captionText="Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA &amp; Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764060" httpUri="https://zenodo.org/record/4764060/files/figure.png" pageId="3" pageNumber="37">12c</figureCitation>
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5’ Ninth and tenth terga without spinulae, long setae usually scattered ……………...………..…..
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6(5) Vesicle rounded apically, base broader than apex, occasionally appearing constricted at middle; paraproct length if straightened, subequal to combined length of first two cercal segments, tapering gradually to blunt apices; posterolateral margins of ninth and tenth terga with patches of long stout setae, fewer and smaller on tergum ten (
<figureCitation box="[881,969,1755,1779]" captionStart="Figs" captionStartId="18.[189,236,1752,1774]" captionTargetBox="[337,1297,251,1707]" captionTargetId="figure-54@18.[334,1297,251,1707]" captionText="Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA &amp; Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764060" httpUri="https://zenodo.org/record/4764060/files/figure.png" pageId="3" pageNumber="37">Figs. 12</figureCitation>
a-d) ……………...…...
<taxonomicName authorityName=", Yuba R., Yuba Co., CA &amp; Nehalem R., Clatsop Co., OR" baseAuthorityName="Banks" baseAuthorityYear="1902" box="[1227,1443,1755,1779]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="37" phylum="Arthropoda" rank="species" species="quinquepunctata">
<emphasis bold="true" box="[1227,1443,1755,1779]" italics="true" pageId="3" pageNumber="37">I. quinquepunctata</emphasis>
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</paragraph>
<paragraph blockId="3.[835,1443,1506,1921]" lastBlockId="4.[227,797,225,249]" lastPageId="4" lastPageNumber="38" pageId="3" pageNumber="37">
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6’ Vesicle truncated or rectangular apically; paraproct length and shape variable; ninth and tenth terga with scattered fine setae, ninth tergum with mesal patches of short stout spinulae (
<figureCitation captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" lastPageId="4" lastPageNumber="38" pageId="3" pageNumber="37">Figs. 1</figureCitation>
,
<figureCitation box="[252,280,225,249]" captionStart="Figs" captionStartId="16.[189,236,1717,1739]" captionTargetBox="[334,1297,215,1671]" captionTargetId="figure-54@16.[334,1297,215,1671]" captionText="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764058" httpUri="https://zenodo.org/record/4764058/files/figure.png" pageId="4" pageNumber="38">10</figureCitation>
) ………………………………………...…......
<emphasis bold="true" box="[784,797,225,249]" pageId="4" pageNumber="38">7</emphasis>
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<keyStep pageId="4" pageNumber="38">
<paragraph blockId="4.[189,797,261,853]" pageId="4" pageNumber="38">
<keyLead pageId="4" pageNumber="38">
7(6) Vesicle rectangular, length half as long as wide (
<figureCitation box="[235,330,296,320]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="4" pageNumber="38">Fig. 1d</figureCitation>
); apical 1/3 of aedeagus tube-like, directed ventrally, covered with minute spinulae and a posterior pair of finer spinule patches divided or joined apically, apex a complex nipplelike tip (
<figureCitation box="[365,458,438,462]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="4" pageNumber="38">Figs. 1</figureCitation>
a-c); paraproct length if straightened greater than combined length of first and second cercal segments, tapering gradually to sharp apices (
<figureCitation box="[386,462,545,569]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="4" pageNumber="38">Fig. 1c</figureCitation>
) …………………..…
<taxonomicName authorityName="Jewett" authorityYear="1962" box="[710,797,544,568]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="acula">
<emphasis bold="true" box="[710,797,544,568]" italics="true" pageId="4" pageNumber="38">I. acula</emphasis>
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</paragraph>
<paragraph blockId="4.[189,797,261,853]" pageId="4" pageNumber="38">
<keyLead pageId="4" pageNumber="38">
7’ Vesicle trapezoid shaped, about as long as wide (
<figureCitation box="[235,344,616,640]" captionStart="Figs" captionStartId="16.[189,236,1717,1739]" captionTargetBox="[334,1297,215,1671]" captionTargetId="figure-54@16.[334,1297,215,1671]" captionText="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764058" httpUri="https://zenodo.org/record/4764058/files/figure.png" pageId="4" pageNumber="38">Fig. 10d</figureCitation>
); apical 1/2 of aedeagus tube-like, directed posteriorly, covered with minute spinulae and concentrated somewhat above bilobed tip (
<figureCitation box="[376,480,722,747]" captionStart="Figs" captionStartId="16.[189,236,1717,1739]" captionTargetBox="[334,1297,215,1671]" captionTargetId="figure-54@16.[334,1297,215,1671]" captionText="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764058" httpUri="https://zenodo.org/record/4764058/files/figure.png" pageId="4" pageNumber="38">Figs. 10</figureCitation>
a-c); paraproct length if straightened subequal to combined length of first two cercal segments, tapering gradually to blunt apices (
<figureCitation box="[313,402,829,853]" captionStart="Figs" captionStartId="16.[189,236,1717,1739]" captionTargetBox="[334,1297,215,1671]" captionTargetId="figure-54@16.[334,1297,215,1671]" captionText="Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764058" httpUri="https://zenodo.org/record/4764058/files/figure.png" pageId="4" pageNumber="38">Fig. 10c</figureCitation>
) .....................................
<taxonomicName authorityName="Banks" authorityYear="1920" box="[665,797,829,852]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="mormona">
<emphasis bold="true" box="[665,797,829,852]" italics="true" pageId="4" pageNumber="38">I. mormona</emphasis>
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<keyStep pageId="4" pageNumber="38">
<paragraph blockId="4.[189,798,864,1527]" pageId="4" pageNumber="38">
<keyLead pageId="4" pageNumber="38">
8(5) Abdomen yellow; when everted live and fixed in hot water, aedeagus with one large patch of spinulae concentrated below posterobasal lobe (
<figureCitation box="[235,320,971,995]" captionStart="Figs" captionStartId="8.[189,236,1752,1774]" captionTargetBox="[336,1296,246,1699]" captionTargetId="figure-54@8.[336,1296,246,1699]" captionText="Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch &amp; thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb &amp; apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle" figureDoi="http://doi.org/10.5281/zenodo.4764052" httpUri="https://zenodo.org/record/4764052/files/figure.png" pageId="4" pageNumber="38">Fig. 3b</figureCitation>
), a long thin patch along posteroapical margin (
<figureCitation box="[328,409,1007,1031]" captionStart="Figs" captionStartId="8.[189,236,1752,1774]" captionTargetBox="[336,1296,246,1699]" captionTargetId="figure-54@8.[336,1296,246,1699]" captionText="Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch &amp; thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb &amp; apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle" figureDoi="http://doi.org/10.5281/zenodo.4764052" httpUri="https://zenodo.org/record/4764052/files/figure.png" pageId="4" pageNumber="38">Fig. 3c</figureCitation>
), apex bulbous, deeply inverted, and without a pair of short apicolateral lobes; when cleared in KOH, apex with a single inverted tube-like process (
<figureCitation box="[440,534,1113,1137]" captionStart="Figs" captionStartId="27.[189,236,1503,1525]" captionTargetBox="[294,1339,251,1437]" captionTargetId="figure-54@27.[294,1339,251,1437]" captionText="Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d &amp; e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4764597" httpUri="https://zenodo.org/record/4764597/files/figure.png" pageId="4" pageNumber="38">Fig. 20a</figureCitation>
), CA-Domingo Spring …………………………...……...…......
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="1984" box="[658,797,1148,1172]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="baumanni">
<emphasis bold="true" box="[658,797,1148,1172]" italics="true" pageId="4" pageNumber="38">I. baumanni</emphasis>
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<paragraph blockId="4.[189,798,864,1527]" pageId="4" pageNumber="38">
<keyLead pageId="4" pageNumber="38">
8’ Abdomen dark; when everted live and fixed in hot water, aedeagus with one large patch of spinulae concentrated below posterobasal lobe (
<figureCitation box="[235,331,1291,1315]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="4" pageNumber="38">Fig. 17b</figureCitation>
), a long thin patch along posteroapical margin (
<figureCitation box="[326,418,1326,1350]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="4" pageNumber="38">Fig. 17c</figureCitation>
), apex bulbous, deeply inverted, and usually with one pair of short sharp apicolateral lobes near apex (similar to
<taxonomicName authorityName=": Szczytko &amp; Stewart" authorityYear="1979" baseAuthorityName="Hagen" baseAuthorityYear="1874" box="[695,782,1397,1421]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="sobria">
<emphasis box="[695,782,1397,1421]" italics="true" pageId="4" pageNumber="38">I. sobria</emphasis>
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); when cleared in KOH, apex with two inverted tube-like processes (
<figureCitation box="[514,630,1468,1492]" captionStart="Figs" captionStartId="27.[189,236,1503,1525]" captionTargetBox="[294,1339,251,1437]" captionTargetId="figure-54@27.[294,1339,251,1437]" captionText="Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d &amp; e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4764597" httpUri="https://zenodo.org/record/4764597/files/figure.png" pageId="4" pageNumber="38">Fig. 20f</figureCitation>
), OR, WA …………………………………………...
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="1979" box="[677,797,1503,1527]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="tilasqua">
<emphasis bold="true" box="[677,797,1503,1527]" italics="true" pageId="4" pageNumber="38">I. tilasqua</emphasis>
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<keyStep pageId="4" pageNumber="38">
<paragraph blockId="4.[189,797,1539,1741]" pageId="4" pageNumber="38">
<keyLead pageId="4" pageNumber="38">
9(2) Vesicle a broadly rounded lobe, wider at base then apex (
<figureCitation box="[353,445,1575,1599]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 6d</figureCitation>
,
<figureCitation box="[456,487,1575,1599]" captionStart="Figs" captionStartId="13.[189,236,1681,1703]" captionTargetBox="[334,1297,195,1650]" captionTargetId="figure-54@13.[334,1297,195,1650]" captionText="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764056" httpUri="https://zenodo.org/record/4764056/files/figure.png" pageId="4" pageNumber="38">8d</figureCitation>
,
<figureCitation box="[497,542,1575,1599]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">14d</figureCitation>
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<figureCitation box="[552,595,1575,1599]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="4" pageNumber="38">16d</figureCitation>
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9’ Vesicle pedunculate, width subequal to length, constricted near base with curved lateral margins, and apex variable (
<figureCitation box="[451,545,1682,1706]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="4" pageNumber="38">Figs. 2d</figureCitation>
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<figureCitation box="[557,589,1682,1706]" captionStart="Figs" captionStartId="8.[189,236,1752,1774]" captionTargetBox="[336,1296,246,1699]" captionTargetId="figure-54@8.[336,1296,246,1699]" captionText="Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch &amp; thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb &amp; apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle" figureDoi="http://doi.org/10.5281/zenodo.4764052" httpUri="https://zenodo.org/record/4764052/files/figure.png" pageId="4" pageNumber="38">4d</figureCitation>
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<figureCitation box="[601,632,1682,1706]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">5d</figureCitation>
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<figureCitation box="[644,675,1682,1706]" captionStart="Figs" captionStartId="13.[189,236,1681,1703]" captionTargetBox="[334,1297,195,1650]" captionTargetId="figure-54@13.[334,1297,195,1650]" captionText="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764056" httpUri="https://zenodo.org/record/4764056/files/figure.png" pageId="4" pageNumber="38">7d</figureCitation>
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<figureCitation box="[688,733,1682,1706]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">13d</figureCitation>
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<figureCitation box="[745,788,1682,1706]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="4" pageNumber="38">18d</figureCitation>
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10(9) Aedeagus with at least one patch of stout spinules above sclerotized process (
<figureCitation box="[655,749,1789,1813]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 6a</figureCitation>
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<figureCitation box="[762,791,1789,1813]" captionStart="Figs" captionStartId="13.[189,236,1681,1703]" captionTargetBox="[334,1297,195,1650]" captionTargetId="figure-54@13.[334,1297,195,1650]" captionText="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764056" httpUri="https://zenodo.org/record/4764056/files/figure.png" pageId="4" pageNumber="38">8b</figureCitation>
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<figureCitation box="[227,269,1824,1848]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="4" pageNumber="38">15a</figureCitation>
); apex of sclerotized process rounded in lateral view (
<figureCitation box="[377,467,1860,1884]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 6a</figureCitation>
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<figureCitation box="[477,504,1860,1884]" captionStart="Figs" captionStartId="13.[189,236,1681,1703]" captionTargetBox="[334,1297,195,1650]" captionTargetId="figure-54@13.[334,1297,195,1650]" captionText="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764056" httpUri="https://zenodo.org/record/4764056/files/figure.png" pageId="4" pageNumber="38">8a</figureCitation>
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<figureCitation box="[515,557,1860,1884]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="4" pageNumber="38">15a</figureCitation>
), distal margin either thin or expanded in posterior view (
<figureCitation box="[657,750,1895,1919]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 6b</figureCitation>
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<figureCitation box="[762,791,1895,1919]" captionStart="Figs" captionStartId="13.[189,236,1681,1703]" captionTargetBox="[334,1297,195,1650]" captionTargetId="figure-54@13.[334,1297,195,1650]" captionText="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764056" httpUri="https://zenodo.org/record/4764056/files/figure.png" pageId="4" pageNumber="38">8b</figureCitation>
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<figureCitation box="[873,914,225,249]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="4" pageNumber="38">15b</figureCitation>
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10’ Aedeagus without a spinule patch above sclerotized process (
<figureCitation box="[1110,1203,296,320]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="4" pageNumber="38">Fig. 16a</figureCitation>
); apex of sclerotized process expanded, flattened and projecting dorsally in lateral view (
<figureCitation box="[1202,1307,367,391]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="4" pageNumber="38">Fig. 16b</figureCitation>
), bifurcate subapically with two ventrally directed rounded lobes in posterior view (
<figureCitation box="[1161,1255,438,462]" captionStart="Figs" captionStartId="22.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1669]" captionTargetId="figure-54@22.[336,1296,215,1671]" captionText="Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow &amp; inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764064" httpUri="https://zenodo.org/record/4764064/files/figure.png" pageId="4" pageNumber="38">Fig. 16b</figureCitation>
inset), width of lobes
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………………….....
<taxonomicName authorityName="Banks" authorityYear="1906" box="[1330,1443,473,497]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="sordida">
<emphasis bold="true" box="[1330,1443,473,497]" italics="true" pageId="4" pageNumber="38">I. sordida</emphasis>
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11(10) A single patch above sclerotized process composed of fine pale spinules; apex of sclerotized process rounded in lateral view (
<figureCitation captionStart="Figs" captionStartId="13.[189,236,1681,1703]" captionTargetBox="[334,1297,195,1650]" captionTargetId="figure-54@13.[334,1297,195,1650]" captionText="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764056" httpUri="https://zenodo.org/record/4764056/files/figure.png" pageId="4" pageNumber="38">Figs. 8</figureCitation>
a-b), distal margin thin and blade-like in posterior view (
<figureCitation box="[1056,1134,651,675]" captionStart="Figs" captionStartId="13.[189,236,1681,1703]" captionTargetBox="[334,1297,195,1650]" captionTargetId="figure-54@13.[334,1297,195,1650]" captionText="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764056" httpUri="https://zenodo.org/record/4764056/files/figure.png" pageId="4" pageNumber="38">Fig. 8b</figureCitation>
) ….………….
<taxonomicName authorityName="Needham &amp; Claassen" authorityYear="1925" box="[1291,1443,652,675]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis bold="true" box="[1291,1443,652,675]" italics="true" pageId="4" pageNumber="38">I. marmorata</emphasis>
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11’ Spinule patch either single or triple and usually dark (
<figureCitation box="[942,1031,722,746]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 6c</figureCitation>
,
<figureCitation box="[1042,1081,723,747]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">14c</figureCitation>
); apex of expanded sclerotized process rounded in lateral view (
<figureCitation box="[1277,1372,758,782]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 6a</figureCitation>
,
<figureCitation box="[1387,1429,758,782]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">14a</figureCitation>
), distal margin expanded slightly in posterior view (
<figureCitation box="[881,970,829,853]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 6c</figureCitation>
,
<figureCitation box="[979,1018,829,853]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">14c</figureCitation>
) ………………………………….....
<emphasis bold="true" box="[1416,1443,829,853]" pageId="4" pageNumber="38">12</emphasis>
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<paragraph blockId="4.[835,1443,865,1031]" pageId="4" pageNumber="38">
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12(11) A single spinule patch above sclerotized process (
<figureCitation box="[976,1110,900,924]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 6a &amp; c</figureCitation>
) ..……………………
<taxonomicName authorityName="Claassen" authorityYear="1937" box="[1358,1443,900,924]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="fulva">
<emphasis bold="true" box="[1358,1443,900,924]" italics="true" pageId="4" pageNumber="38">I. fulva</emphasis>
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12’ One bi-hemispherical shaped spinule patch above sclerotized process and a pair of smaller rounded patches laterally (
<figureCitation box="[1080,1170,1007,1031]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">Figs. 14</figureCitation>
a-c) ………
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="1984" box="[1306,1442,1007,1031]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="roguensis">
<emphasis bold="true" box="[1306,1442,1007,1031]" italics="true" pageId="4" pageNumber="38">I. roguensis</emphasis>
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13(9) Aedeagal sclerotized process bifurcate in posterior view (
<figureCitation box="[1058,1157,1078,1102]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="8.[189,236,1752,1774]" captionStartId-1="11.[189,236,1717,1739]" captionTargetBox-0="[336,1296,246,1699]" captionTargetBox-1="[335,1297,215,1671]" captionTargetId-0="figure-54@8.[336,1296,246,1699]" captionTargetId-1="figure-54@11.[335,1297,215,1671]" captionTargetPageId-0="8" captionText-0="Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch &amp; thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb &amp; apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle" captionText-1="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi-0="http://doi.org/10.5281/zenodo.4764052" figureDoi-1="http://doi.org/10.5281/zenodo.4764054" httpUri-0="https://zenodo.org/record/4764052/files/figure.png" httpUri-1="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 4-5</figureCitation>
) …………...………….
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13’ Aedeagal sclerotized process not bifurcate in posterior view (
<figureCitation box="[1056,1131,1149,1173]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="4" pageNumber="38">Figs. 2</figureCitation>
,
<figureCitation box="[1140,1155,1150,1173]" captionStart="Figs" captionStartId="13.[189,236,1681,1703]" captionTargetBox="[334,1297,195,1650]" captionTargetId="figure-54@13.[334,1297,195,1650]" captionText="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764056" httpUri="https://zenodo.org/record/4764056/files/figure.png" pageId="4" pageNumber="38">7</figureCitation>
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<figureCitation box="[1164,1191,1149,1173]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">13</figureCitation>
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<figureCitation box="[1202,1228,1149,1173]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="4" pageNumber="38">18</figureCitation>
) ………………..
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14(13) Sclerotized process with long apically pointed bifurcate arms; length of projecting process greater than
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<figureCitation box="[1136,1220,1256,1280]" captionStart="Figs" captionStartId="8.[189,236,1752,1774]" captionTargetBox="[336,1296,246,1699]" captionTargetId="figure-54@8.[336,1296,246,1699]" captionText="Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch &amp; thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb &amp; apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle" figureDoi="http://doi.org/10.5281/zenodo.4764052" httpUri="https://zenodo.org/record/4764052/files/figure.png" pageId="4" pageNumber="38">Fig. 4a</figureCitation>
); abdomen mostly brown ……………………..…………...
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<emphasis bold="true" box="[1313,1444,1291,1315]" italics="true" pageId="4" pageNumber="38">I. bifurcata</emphasis>
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14’ Sclerotized process with short rounded bifurcate lobes; length of projecting process less than
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<figureCitation box="[937,1016,1398,1422]" captionStart="Figs" captionStartId="11.[189,236,1717,1739]" captionTargetBox="[335,1297,215,1671]" captionTargetId="figure-54@11.[335,1297,215,1671]" captionText="Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO &amp; Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow &amp; inset-lateral view); spinule patch (right white arrow &amp; inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764054" httpUri="https://zenodo.org/record/4764054/files/figure.png" pageId="4" pageNumber="38">Figs. 5</figureCitation>
a-b); width of bifurcate apicolateral lobes
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; abdomen mostly yellow ……………………………………….......
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<emphasis bold="true" box="[1323,1444,1469,1493]" italics="true" pageId="4" pageNumber="38">I. denningi</emphasis>
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15(13) Posterior margin of aedeagus with elliptical shaped patch of long stout spinulae separated from and above sclerotized process; sclerotized process digitate, with nearly parallel dorsal and ventral margins in lateral view, apex narrowly rounded (
<figureCitation box="[990,1065,1683,1707]" captionStart="Figs" captionStartId="13.[189,236,1681,1703]" captionTargetBox="[334,1297,195,1650]" captionTargetId="figure-54@13.[334,1297,195,1650]" captionText="Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow &amp; inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow &amp; inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch &amp; sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764056" httpUri="https://zenodo.org/record/4764056/files/figure.png" pageId="4" pageNumber="38">Figs. 7</figureCitation>
a-c) ..……………….…..
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<emphasis bold="true" box="[1342,1440,1682,1706]" italics="true" pageId="4" pageNumber="38">I. laucki</emphasis>
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15’ Posterior margin of aedeagus without a patch of long stout spinulae; sclerotized process otherwise (
<figureCitation box="[881,957,1789,1813]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="4" pageNumber="38">Figs. 2</figureCitation>
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<figureCitation box="[967,994,1789,1813]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">13</figureCitation>
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<figureCitation box="[1006,1032,1789,1813]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="4" pageNumber="38">18</figureCitation>
) ………….………………...……....
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16(15) Sclerotized process crescent shaped in dorsal and ventral views (
<figureCitation box="[1141,1245,1860,1885]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">Figs. 13</figureCitation>
b-c); paraprocts gradually tapering to sharp apices (
<figureCitation box="[1292,1384,1896,1920]" captionStart="Figs" captionStartId="20.[189,236,1717,1739]" captionTargetBox="[336,1296,215,1668]" captionTargetId="figure-54@20.[336,1296,215,1671]" captionText="Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA &amp; Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764062" httpUri="https://zenodo.org/record/4764062/files/figure.png" pageId="4" pageNumber="38">Fig. 13c</figureCitation>
) OR, WA.………………………….………...….
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16’ Sclerotized process clavate; paraprocts tapering gradually to blunt apices; OR, CA ...….………
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17(16) Wings hyaline; aedeagus with one distinct lobe (
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); sclerotized process length approximately
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<figureCitation box="[624,747,403,427]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="5" pageNumber="39">Figs. 18</figureCitation>
a-c); mesoposterior area of 8th and 9th terga with bipartite patches of stout spinulae and long fine scattered setae (
<figureCitation box="[410,500,509,533]" captionStart="Figs" captionStartId="24.[189,236,1717,1739]" captionTargetBox="[336,1296,216,1671]" captionTargetId="figure-54@24.[336,1296,215,1671]" captionText="Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch &amp; thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow &amp; insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764066" httpUri="https://zenodo.org/record/4764066/files/figure.png" pageId="5" pageNumber="39">Fig. 18c</figureCitation>
), OR ...………...
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17’ Wings tinted; aedeagus with four distinct lobes in lateral view (
<figureCitation box="[384,467,580,604]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="5" pageNumber="39">Fig. 2b</figureCitation>
); sclerotized process length approximately
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(
<figureCitation box="[517,597,616,640]" captionStart="Figs" captionStartId="6.[189,236,1752,1774]" captionTargetBox="[335,1297,243,1696]" captionTargetId="figure-54@6.[335,1297,243,1696]" captionText="Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. &amp; Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow &amp; inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow &amp; inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow &amp; inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764050" httpUri="https://zenodo.org/record/4764050/files/figure.png" pageId="5" pageNumber="39">Fig. 2b</figureCitation>
), spinule patches absent on terga 8-9, CA ….……………..
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<emphasis bold="true" box="[689,797,651,675]" italics="true" pageId="5" pageNumber="39">I. adunca</emphasis>
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<subSubSection box="[445,541,722,746]" pageId="5" pageNumber="39" type="description">
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<taxonomicName authority="of" authorityName="OF" box="[445,541,722,746]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="genus">
<keyLead box="[445,541,722,746]" pageId="5" pageNumber="39">
<keyStep box="[445,541,722,746]" pageId="5" pageNumber="39">Females</keyStep>
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</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="39" type="key">
<keyStep pageId="5" pageNumber="39">
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1 Subgenital plate usually with invagination along apical margin (
<figureCitation box="[401,488,829,853]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19</figureCitation>
b-c, 19l-m, 19q-s) .........….
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1’ Subgenital plate usually not invaginated (
<figureCitation captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19a, 19</figureCitation>
d-k, 19n-p) ..…………………………….…
<emphasis bold="true" box="[784,797,900,924]" pageId="5" pageNumber="39">9</emphasis>
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<paragraph blockId="5.[189,797,936,1138]" pageId="5" pageNumber="39">
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2(1) Subgenital plate with deep median invagination, the apex 3–4 times wider than depth (
<figureCitation box="[235,390,1007,1031]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19b, 19r</figureCitation>
) ………………………………........
<emphasis bold="true" box="[784,797,1007,1031]" pageId="5" pageNumber="39">3</emphasis>
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</paragraph>
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2’ Subgenital plate with shallow median invagination, the apex 8–10 times wider than depth (
<figureCitation box="[309,447,1114,1138]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19c, 19</figureCitation>
l-m, 19q, 19s) ..……………....
<emphasis bold="true" box="[784,797,1114,1138]" pageId="5" pageNumber="39">4</emphasis>
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<paragraph blockId="5.[189,798,1150,1281]" pageId="5" pageNumber="39">
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3(2) Subgenital plate length less than half the width at base (
<figureCitation box="[318,407,1185,1209]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19b</figureCitation>
) …………...…………..…
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<emphasis bold="true" box="[693,798,1185,1209]" italics="true" pageId="5" pageNumber="39">I. adunca</emphasis>
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3’ Subgenital plate length approximately half the width at base (
<figureCitation box="[394,482,1257,1281]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19r</figureCitation>
) ………..…….…
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="1979" box="[680,797,1256,1280]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="tilasqua">
<emphasis bold="true" box="[680,797,1256,1280]" italics="true" pageId="5" pageNumber="39">I. tilasqua</emphasis>
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<paragraph blockId="5.[189,797,1292,1529]" pageId="5" pageNumber="39">
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4(2) Subgenital plate apex truncated, with nearly parallel lateral margins (
<figureCitation box="[552,657,1328,1352]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19c</figureCitation>
); deflected ventrally in lateral view, CA-Domingo Spring ……………………………………...….
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="1984" box="[657,797,1398,1422]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="baumanni">
<emphasis bold="true" box="[657,797,1398,1422]" italics="true" pageId="5" pageNumber="39">I. baumanni</emphasis>
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4’ Subgenital plate apex rounded and not deflected ventrally in lateral view (
<figureCitation box="[540,632,1470,1494]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19</figureCitation>
l-m, 19q, 19s) ………………………………………………...........
<emphasis bold="true" box="[784,797,1505,1529]" pageId="5" pageNumber="39">5</emphasis>
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<paragraph blockId="5.[189,798,1541,1743]" pageId="5" pageNumber="39">
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5(4) Subgenital plate width at base approximately half the width of segment 8 and nearly twice that of the subgenital plate length (
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Fig.
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) ………………………………......
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5’ Subgenital plate width at base greater than half the width of segment 8 (
<figureCitation box="[509,710,1719,1743]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19l, 19q, 19s</figureCitation>
) ……
<emphasis bold="true" box="[784,797,1719,1743]" pageId="5" pageNumber="39">6</emphasis>
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6(5) Subgenital plate lateral margins broadly rounded (
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) ………….…….……
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6’ Subgenital plate with short, nearly parallel lateral margins (
<figureCitation box="[336,479,1861,1885]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19l, 19s</figureCitation>
) ………………….…..…….
<emphasis bold="true" box="[784,797,1861,1885]" pageId="5" pageNumber="39">7</emphasis>
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7(6) Pronotum with checkerboard pattern (
<figureCitation box="[707,797,1897,1921]" captionStart="Figs" captionStartId="18.[189,236,1752,1774]" captionTargetBox="[337,1297,251,1707]" captionTargetId="figure-54@18.[334,1297,251,1707]" captionText="Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA &amp; Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle." figureDoi="http://doi.org/10.5281/zenodo.4764060" httpUri="https://zenodo.org/record/4764060/files/figure.png" pageId="5" pageNumber="39">Fig. 12d</figureCitation>
inset) …………………………...…………….
<taxonomicName authorityName="Frison" authorityYear="1937" box="[1363,1444,225,249]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="pinta">
<emphasis bold="true" box="[1363,1444,225,249]" italics="true" pageId="5" pageNumber="39">I. pinta</emphasis>
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</paragraph>
<paragraph blockId="5.[835,1444,225,284]" box="[835,1443,260,284]" pageId="5" pageNumber="39">
<keyLead box="[835,1443,260,284]" pageId="5" pageNumber="39">
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7’ Pronotum without checkerboard pattern ……....
<emphasis bold="true" box="[1430,1443,260,284]" pageId="5" pageNumber="39">8</emphasis>
</keyStep>
</keyLead>
</paragraph>
<keyStep pageId="5" pageNumber="39">
<paragraph blockId="5.[835,1443,296,533]" pageId="5" pageNumber="39">
<keyLead pageId="5" pageNumber="39">
8(7) Apical margin of subgenital plate sometimes invaginated (broadly rounded in
<figureCitation box="[1276,1363,332,356]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19j</figureCitation>
), apex reaches approximately mid-length of ninth sternum (CA) …………………
<taxonomicName authorityName="Bottorff &amp; Szczytko" authorityYear="1990" box="[1235,1341,402,426]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="miwok">
<emphasis bold="true" box="[1235,1341,402,426]" italics="true" pageId="5" pageNumber="39">I. miwok</emphasis>
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(in part)
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<paragraph blockId="5.[835,1443,296,533]" pageId="5" pageNumber="39">
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8’ Apical margin of subgenital plate typically invaginated (
<figureCitation box="[1033,1128,474,498]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19s</figureCitation>
), apex not reaching midlength of ninth sternum (OR) ………...
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="2013" box="[1325,1443,510,533]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="umpqua">
<emphasis bold="true" box="[1325,1443,510,533]" italics="true" pageId="5" pageNumber="39">I. umpqua</emphasis>
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<paragraph blockId="5.[835,1443,545,676]" pageId="5" pageNumber="39">
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9(1) Subgenital plate with lateral margins nearly parallel (
<figureCitation box="[976,1127,580,604]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19j, 19p</figureCitation>
) ........................................
<emphasis bold="true" box="[1416,1443,580,604]" pageId="5" pageNumber="39">10</emphasis>
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<paragraph blockId="5.[835,1443,545,676]" pageId="5" pageNumber="39">
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9’ Subgenital plate with lateral margins not nearly parallel (
<figureCitation box="[976,1117,652,676]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19a, 19</figureCitation>
d-i, 19k, 19n-o) ...................
<emphasis bold="true" box="[1417,1443,651,675]" pageId="5" pageNumber="39">11</emphasis>
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<paragraph blockId="5.[835,1443,687,819]" pageId="5" pageNumber="39">
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10(9) Subgenital plate width nearly as wide as segment 8 (
<figureCitation box="[1004,1088,723,747]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19j</figureCitation>
) …………...………….
<taxonomicName authorityName="Bottorff &amp; Szczytko" authorityYear="1990" box="[1341,1443,723,747]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="miwok">
<emphasis bold="true" box="[1341,1443,723,747]" italics="true" pageId="5" pageNumber="39">I. miwok</emphasis>
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10’ Subgenital plate width distinctly less than width of segment 8 (
<figureCitation box="[1036,1130,794,818]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19p</figureCitation>
) ……..……...……
<taxonomicName authorityName=": Szczytko &amp; Stewart" authorityYear="1979" baseAuthorityName="Hagen" baseAuthorityYear="1874" box="[1345,1443,794,818]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="sobria">
<emphasis bold="true" box="[1345,1443,794,818]" italics="true" pageId="5" pageNumber="39">I. sobria</emphasis>
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<paragraph blockId="5.[835,1443,830,960]" pageId="5" pageNumber="39">
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11(9) Subgenital plate triangular (
<figureCitation box="[1261,1434,830,854]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">
Figs.
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, 19o
</figureCitation>
) …..…………………..
<emphasis bold="true" box="[1117,1346,865,889]" italics="true" pageId="5" pageNumber="39">
<taxonomicName authorityName="Claassen" authorityYear="1937" box="[1117,1202,865,889]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="fulva">I. fulva</taxonomicName>
,
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="1984" box="[1213,1346,865,889]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="roguensis">I. roguensis</taxonomicName>
</emphasis>
(in part)
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<paragraph blockId="5.[835,1443,830,960]" pageId="5" pageNumber="39">
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11’ Subgenital plate rounded (
<figureCitation box="[1184,1325,901,925]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19a, 19</figureCitation>
d-g, 19h-i, 19k, 19n) ………….…………………..……….….
<emphasis bold="true" box="[1417,1443,936,960]" pageId="5" pageNumber="39">12</emphasis>
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<paragraph blockId="5.[835,1443,972,1174]" pageId="5" pageNumber="39">
<keyLead pageId="5" pageNumber="39">
12(11) Subgenital plate width at base approximately 4-5 times that of the length (
<figureCitation box="[1205,1295,1008,1032]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19</figureCitation>
d-f, 19i, 19n) ………………………………………….………....
<emphasis bold="true" box="[1416,1443,1043,1067]" pageId="5" pageNumber="39">13</emphasis>
</keyLead>
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<paragraph blockId="5.[835,1443,972,1174]" pageId="5" pageNumber="39">
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12’ Subgenital plate width at base approximately 2-3 times that of the length (
<figureCitation box="[1198,1435,1114,1139]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19a, 19h, 19k</figureCitation>
) …………………………………………………......
<emphasis bold="true" box="[1417,1443,1150,1174]" pageId="5" pageNumber="39">16</emphasis>
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13(12) Subgenital plate completely suffused with dark pigment (
<figureCitation box="[1046,1140,1221,1246]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19n</figureCitation>
) ……………….
<taxonomicName authorityName="Jewett" authorityYear="1954" box="[1326,1443,1221,1245]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="rainiera">
<emphasis bold="true" box="[1326,1443,1221,1245]" italics="true" pageId="5" pageNumber="39">I. rainiera</emphasis>
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13’ Subgenital plate partially patterned with pigment or completely light (
<figureCitation box="[1108,1197,1292,1317]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19</figureCitation>
d-f, 19i) ……….….
<emphasis bold="true" box="[1416,1443,1292,1316]" pageId="5" pageNumber="39">14</emphasis>
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<paragraph blockId="5.[835,1444,1328,1459]" pageId="5" pageNumber="39">
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14(13) Subgenital plate partially patterned with pigment (
<figureCitation box="[985,1126,1364,1388]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19d, 19</figureCitation>
f-g).….
<emphasis bold="true" box="[1220,1444,1363,1387]" italics="true" pageId="5" pageNumber="39">
<taxonomicName authorityName="Szczytko &amp; Stewart" authorityYear="1979" box="[1220,1351,1363,1387]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="bifurcata">I. bifurcata</taxonomicName>
,
<taxonomicName authorityName="Claassen" authorityYear="1937" box="[1361,1444,1363,1387]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="fulva">I. fulva</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="5.[835,1444,1328,1459]" pageId="5" pageNumber="39">
<keyLead pageId="5" pageNumber="39">
14’ Subgenital plate completely light (
<figureCitation box="[1281,1434,1399,1423]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19e, 19i</figureCitation>
) …………………………………………………......
<emphasis bold="true" box="[1417,1443,1435,1459]" pageId="5" pageNumber="39">15</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="5" pageNumber="39">
<paragraph blockId="5.[835,1444,1470,1601]" pageId="5" pageNumber="39">
<keyLead pageId="5" pageNumber="39">
15(14) Subgenital plate width at base approximately 3-4 times the length (
<figureCitation box="[1112,1200,1506,1530]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19e</figureCitation>
) …………
<taxonomicName authorityName="Jewett" authorityYear="1955" box="[1323,1444,1506,1530]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="denningi">
<emphasis bold="true" box="[1323,1444,1506,1530]" italics="true" pageId="5" pageNumber="39">I. denningi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="5.[835,1444,1470,1601]" pageId="5" pageNumber="39">
<keyLead pageId="5" pageNumber="39">
15’ Subgenital plate width at base 6-7 times the length (
<figureCitation box="[881,967,1577,1601]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19i</figureCitation>
) …………………….………
<taxonomicName authorityName="Needham &amp; Claassen" authorityYear="1925" box="[1287,1443,1578,1601]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis bold="true" box="[1287,1443,1578,1601]" italics="true" pageId="5" pageNumber="39">I. marmorata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="5" pageNumber="39">
<paragraph blockId="5.[835,1443,1613,1779]" pageId="5" pageNumber="39">
<keyLead pageId="5" pageNumber="39">
16(12) Subgenital plate width at base approximately 3 times as wide than length; apex slightly truncated or broadly rounded (
<figureCitation box="[1119,1213,1684,1708]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19k</figureCitation>
) ...……
<taxonomicName authorityName="Banks" authorityYear="1920" box="[1307,1443,1684,1707]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="mormona">
<emphasis bold="true" box="[1307,1443,1684,1707]" italics="true" pageId="5" pageNumber="39">I. mormona</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="5.[835,1443,1613,1779]" pageId="5" pageNumber="39">
<keyLead pageId="5" pageNumber="39">
16’ Subgenital plate width at base approximately twice that of the length (
<figureCitation box="[1159,1318,1755,1779]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Figs. 19a, 19h</figureCitation>
) …..….
<emphasis bold="true" box="[1413,1440,1755,1779]" pageId="5" pageNumber="39">17</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="5" pageNumber="39">
<paragraph blockId="5.[835,1444,1791,1921]" pageId="5" pageNumber="39">
<keyLead pageId="5" pageNumber="39">
17(16) Subgenital plate darkly pigmented (
<figureCitation box="[1339,1434,1791,1815]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19h</figureCitation>
) ……………………………………..…………….
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[1353,1444,1826,1850]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="laucki">
<emphasis bold="true" box="[1353,1444,1826,1850]" italics="true" pageId="5" pageNumber="39">I. laucki</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="5.[835,1444,1791,1921]" pageId="5" pageNumber="39">
<keyLead pageId="5" pageNumber="39">
17’ Subgenital plate not darkly pigmented (
<figureCitation box="[1342,1434,1862,1886]" captionStart="Figs" captionStartId="26.[189,236,1752,1774]" captionTargetBox="[328,1296,255,1686]" captionTargetId="figure-54@26.[327,1304,251,1687]" captionText="Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua." figureDoi="http://doi.org/10.5281/zenodo.4764593" httpUri="https://zenodo.org/record/4764593/files/figure.png" pageId="5" pageNumber="39">Fig. 19a</figureCitation>
) ……………………………………………...
<taxonomicName authorityName="Jewett" authorityYear="1962" box="[1353,1443,1897,1921]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="acula">
<emphasis bold="true" box="[1353,1443,1897,1921]" italics="true" pageId="5" pageNumber="39">I. acula</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</key>
</treatment>
</document>
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