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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.205.82275" ID-Pensoft-Pub="1314-2003-205-239" ID-Pensoft-UUID="9A789A1610955E8581CC5A1796DAD919" ModsDocID="1314-2003-205-239" checkinTime="1661230140699" checkinUser="pensoft" docAuthor="Borges, Leonardo M., Inglis, Peter W., Simon, Marcelo F., Ribeiro, Petala Gomes &amp; de Queiroz, Luciano P." docDate="2022" docId="DE490BFFA4195D839D7046B7A724DD13" docLanguage="en" docName="PhytoKeys 205: 239-259" docOrigin="PhytoKeys 205" docPubDate="2022-08-22" docSource="http://dx.doi.org/10.3897/phytokeys.205.82275" docTitle="Pityrocarpa (Benth.) Britton &amp; Rose, N. Amer. Fl. 23 (3): 190. 1928." docType="treatment" docVersion="1" id="9A789A1610955E8581CC5A1796DAD919" lastPageNumber="239" masterDocId="9A789A1610955E8581CC5A1796DAD919" masterDocTitle="Misleading fruits: The non-monophyly of Pseudopiptadenia and Pityrocarpa supports generic re-circumscriptions and a new genus within mimosoid legumes" masterLastPageNumber="259" masterPageNumber="239" pageNumber="239" updateTime="1661230441930" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Misleading fruits: The non-monophyly of Pseudopiptadenia and Pityrocarpa supports generic re-circumscriptions and a new genus within mimosoid legumes</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Borges, Leonardo M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9269-7316</mods:nameIdentifier>
<mods:affiliation>Universidade Federal de Sao Carlos, Departamento de Botanica, Rodovia Washington Luis, Km 235, Sao Carlos, SP, 13565 - 905, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">aquitemcaqui@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Inglis, Peter W.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5513-8918</mods:nameIdentifier>
<mods:affiliation>Embrapa Recursos Geneticos e Biotecnologia, Parque Estacao Biologica, Caixa Postal 02372, Brasilia, DF, 70770 - 917, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Simon, Marcelo F.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5732-1716</mods:nameIdentifier>
<mods:affiliation>Embrapa Recursos Geneticos e Biotecnologia, Parque Estacao Biologica, Caixa Postal 02372, Brasilia, DF, 70770 - 917, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ribeiro, Petala Gomes</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0070-9971</mods:nameIdentifier>
<mods:affiliation>Universidade Estadual de Feira de Santana, Departamento de Ciencias Biologicas. Av. Transnordestina s. n., Novo Horizonte, Feira de Santana, BA, 44036 - 900, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Queiroz, Luciano P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7436-0939</mods:nameIdentifier>
<mods:affiliation>Universidade Estadual de Feira de Santana, Departamento de Ciencias Biologicas. Av. Transnordestina s. n., Novo Horizonte, Feira de Santana, BA, 44036 - 900, Brazil</mods:affiliation>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-22</mods:number>
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<mods:detail type="volume">
<mods:number>205</mods:number>
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<mods:start>239</mods:start>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.205.82275</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.205.82275</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-205-239</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:DE490BFFA4195D839D7046B7A724DD13" httpUri="http://treatment.plazi.org/id/DE490BFFA4195D839D7046B7A724DD13" lastPageNumber="239" pageId="0" pageNumber="239">
<subSubSection pageId="0" pageNumber="239" type="nomenclature">
<paragraph pageId="0" pageNumber="239">
1.
<taxonomicName LSID="DE490BFF-A419-5D83-9D70-46B7A724DD13" authority="(Benth.) Britton &amp; Rose, N. Amer. Fl. 23 (3): 190. 1928." authorityName="(Benth.) Britton &amp; Rose, N. Amer. Fl. 23 (3): 190. 1928." class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">Pityrocarpa (Benth.) Britton &amp; Rose, N. Amer. Fl. 23(3): 190. 1928.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="239" type="reference_group">
<paragraph pageId="0" pageNumber="239">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Monoschisma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Monoschisma" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">Monoschisma</taxonomicName>
Brenan,
<emphasis italics="true" pageId="0" pageNumber="239">Kew Bull.</emphasis>
10(2): 179. 1955,
<emphasis italics="true" pageId="0" pageNumber="239">nom. inval.</emphasis>
, non
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Monoschisma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Monoschisma" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">Monoschisma</taxonomicName>
Duby,
<normalizedToken originalValue="Mém">Mem</normalizedToken>
. Soc. Phys.
<normalizedToken originalValue="Genève">Geneve</normalizedToken>
19: 294. 1868. Type.
<taxonomicName authorityName="Borges &amp; Inglis &amp; Simon &amp; Ribeiro &amp; de Queiroz" authorityYear="2022" baseAuthorityName="Borges &amp; Inglis &amp; Simon &amp; Ribeiro &amp; de Queiroz" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Monoschisma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Monoschisma leptostachyum" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="species" species="leptostachyum">Monoschisma leptostachyum</taxonomicName>
(Benth.) Brenan, syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="239">
<taxonomicName authorityName="S.Rauschert" authorityYear="1982" class="Magnoliopsida" family="Fabaceae" genus="Pseudopiptadenia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudopiptadenia" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">Pseudopiptadenia</taxonomicName>
Rauschert, Taxon 31(3): 559. 1982. Type.
<taxonomicName authorityName="Borges &amp; Inglis &amp; Simon &amp; Ribeiro &amp; de Queiroz" authorityYear="2022" baseAuthorityName="Borges &amp; Inglis &amp; Simon &amp; Ribeiro &amp; de Queiroz" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Pseudopiptadenia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudopiptadenia leptostachya" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="species" species="leptostachya">Pseudopiptadenia leptostachya</taxonomicName>
(Benth.) Rauschert, syn. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="239" type="basionym">
<paragraph pageId="0" pageNumber="239">Basionym.</paragraph>
<paragraph pageId="0" pageNumber="239">
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Piptadenia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Piptadenia subsp. sect." order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="section" section="Pityrocarpa" species="leptostachya" subSpecies="sect.">Piptadenia sect. Pityrocarpa</taxonomicName>
Benth., J. Bot. (Hooker) 4: 339. 1842.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="239" type="materials_examined">
<paragraph pageId="0" pageNumber="239">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="239">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa moniliformis" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="species" species="moniliformis">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa moniliformis</emphasis>
</taxonomicName>
(Benth.) Luckow &amp; R.W. Jobson [≡
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Piptadenia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Piptadenia moniliformis" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="species" species="moniliformis">
<emphasis italics="true" pageId="0" pageNumber="239">Piptadenia moniliformis</emphasis>
</taxonomicName>
Benth., designated by
<bibRefCitation author="Britton, NL" journalOrPublisher="North American Flora" pageId="0" pageNumber="239" pagination="1 - 194" refId="B5" refString="Britton, NL, Rose, JN, 1928. Mimosaceae. North American Flora 23: 1 - 194" title="Mimosaceae." volume="23" year="1928">Britton and Rose 1928</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="239" type="description">
<paragraph pageId="0" pageNumber="239">Description.</paragraph>
<paragraph pageId="0" pageNumber="239">
Unarmed trees or shrubs.
<emphasis italics="true" pageId="0" pageNumber="239">Leaves</emphasis>
bipinnate; petiole with an extrafloral nectary between or shortly below the first pair of pinnae; pinnae 1-4 (5) pairs, exceptionally to 10 pairs in
<taxonomicName genus="Pi." lsidName="Pi. leptostachya" pageId="0" pageNumber="239" rank="species" species="leptostachya">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leptostachya</emphasis>
</taxonomicName>
; leaflets 1-10 pairs per pinna, rarely to 20 pairs (
<taxonomicName genus="Pi." lsidName="Pi. brenanii" pageId="0" pageNumber="239" rank="species" species="brenanii">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. brenanii</emphasis>
</taxonomicName>
and
<taxonomicName genus="Pi." lsidName="Pi. leptostachya" pageId="0" pageNumber="239" rank="species" species="leptostachya">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leptostachya</emphasis>
</taxonomicName>
), mostly rhomboid sometimes also asymmetrically elliptical or lanceolate.
<emphasis italics="true" pageId="0" pageNumber="239">Inflorescences</emphasis>
spikes, solitary in the axils of coeval leaves, commonly pendulous.
<emphasis italics="true" pageId="0" pageNumber="239">Flowers</emphasis>
pentamerous; petals free (except possibly
<taxonomicName genus="Pi." lsidName="Pi. leucoxylon" pageId="0" pageNumber="239" rank="species" species="leucoxylon">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leucoxylon</emphasis>
</taxonomicName>
), glabrous; stamens 10, anther gland present; ovary shortly stipitate and included within or exserted from the corolla.
<emphasis italics="true" pageId="0" pageNumber="239">Fruit</emphasis>
a follicle, dehiscing along the lower suture, flat compressed, mostly moniliform, the margins deeply and regularly constricted, rarely sinuous margins and shallowly constricted (
<taxonomicName genus="Pi." lsidName="Pi. brenanii" pageId="0" pageNumber="239" rank="species" species="brenanii">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. brenanii</emphasis>
</taxonomicName>
and occasionally in
<taxonomicName genus="Pi." lsidName="Pi. leucoxylon" pageId="0" pageNumber="239" rank="species" species="leucoxylon">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leucoxylon</emphasis>
</taxonomicName>
); valves stiffly coriaceous.
<emphasis italics="true" pageId="0" pageNumber="239">Seeds</emphasis>
mostly flat compressed with a coriaceous testa and a narrow marginal wing, lacking a pleurogram or, less frequently, ovoid or discoid with a hard, whitish testa, wingless and with a
<normalizedToken originalValue="U">'U'</normalizedToken>
-shaped pleurogram (
<taxonomicName genus="Pi." lsidName="Pi. leucoxylon" pageId="0" pageNumber="239" rank="species" species="leucoxylon">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leucoxylon</emphasis>
</taxonomicName>
,
<taxonomicName genus="Pi." lsidName="Pi. moniliformis" pageId="0" pageNumber="239" rank="species" species="moniliformis">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. moniliformis</emphasis>
</taxonomicName>
and
<taxonomicName genus="Pi." lsidName="Pi. obliqua" pageId="0" pageNumber="239" rank="species" species="obliqua">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. obliqua</emphasis>
</taxonomicName>
); embryo with a rudimentary plumule (except
<taxonomicName genus="Pi." lsidName="Pi. brenanii" pageId="0" pageNumber="239" rank="species" species="brenanii">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. brenanii</emphasis>
</taxonomicName>
).
<emphasis italics="true" pageId="0" pageNumber="239">Seedlings</emphasis>
with bifoliolate eophylls.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="239" type="distribution">
<paragraph pageId="0" pageNumber="239">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="239">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
is distributed in tropical America, from Mexico to southern Brazil and Paraguay. Most species occur in the Brazilian Atlantic rainforests (
<taxonomicName genus="Pi." lsidName="Pi. inaequalis" pageId="0" pageNumber="239" rank="species" species="inaequalis">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. inaequalis</emphasis>
</taxonomicName>
,
<taxonomicName genus="Pi." lsidName="Pi. leptostachya" pageId="0" pageNumber="239" rank="species" species="leptostachya">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leptostachya</emphasis>
</taxonomicName>
,
<taxonomicName genus="Pi." lsidName="Pi. schumanniana" pageId="0" pageNumber="239" rank="species" species="schumanniana">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. schumanniana</emphasis>
</taxonomicName>
), in the northern Amazonian rainforests (
<taxonomicName genus="Pi." lsidName="Pi. leucoxylon" pageId="0" pageNumber="239" rank="species" species="leucoxylon">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leucoxylon</emphasis>
</taxonomicName>
), in seasonally dry tropical forests and woodlands in the north-eastern Brazilian Caatinga (
<taxonomicName genus="Pi." lsidName="Pi. brenanii" pageId="0" pageNumber="239" rank="species" species="brenanii">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. brenanii</emphasis>
</taxonomicName>
,
<taxonomicName genus="Pi." lsidName="Pi. moniliformis" pageId="0" pageNumber="239" rank="species" species="moniliformis">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. moniliformis</emphasis>
</taxonomicName>
,
<taxonomicName genus="Pi." lsidName="Pi. obliqua subsp. brasiliensis" pageId="0" pageNumber="239" rank="subSpecies" species="obliqua" subSpecies="brasiliensis">Pi. obliqua subsp. brasiliensis</taxonomicName>
), western Mexico (
<taxonomicName authority="subsp. obliqua" genus="Pi." lsidName="Pi. obliqua subsp. obliqua" pageId="0" pageNumber="239" rank="subSpecies" species="obliqua" subSpecies="obliqua">Pi. obliqua subsp. obliqua</taxonomicName>
) or in Venezuelan savannas and Paraguayan Chaco (
<taxonomicName genus="Pi." lsidName="Pi. moniliformis" pageId="0" pageNumber="239" rank="species" species="moniliformis">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. moniliformis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="239" type="notes">
<paragraph pageId="0" pageNumber="239">Notes.</paragraph>
<paragraph pageId="0" pageNumber="239">
As circumscribed here,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
includes seven species, all with a moniliform fruit, with the margins deeply constricted between the seeds (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Fruits of Pityrocarpa species A Pi. brenanii (from Lewis et al. 1899, NY) B Pi. inaequalis (from Moreira et al. 3, F) C Pi. leptostachya (from Baez et al. 1174, NY) D Pi. leucoxylon (from de Bruijn 1750, NY) E Pi. moniliformis (from Nunes 597, HUEFS) F Pi. obliqua subsp. brasiliensis (from Mori 11837, NY) G Pi. schumanniana (from Lima 2994, RB)." figureDoi="10.3897/phytokeys.205.82275.figure4" httpUri="https://binary.pensoft.net/fig/732769" pageId="0" pageNumber="239">4</figureCitation>
). This trait is shared by species formerly included in
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
(sensu
<bibRefCitation DOI="https://doi.org/10.1600/036364407782250544" author="Jobson, RW" journalOrPublisher="Systematic Botany" pageId="0" pageNumber="239" pagination="569 - 575" refId="B11" refString="Jobson, RW, Luckow, M, 2007. Phylogenetic study of the genus Piptadenia (Mimosoideae: Leguminosae) using plastid trnL-F and trnK/matK sequence data. Systematic Botany 32 (3): 569 - 575, DOI: https://doi.org/10.1600/036364407782250544" title="Phylogenetic study of the genus Piptadenia (Mimosoideae: Leguminosae) using plastid trnL-F and trnK / matK sequence data." url="https://doi.org/10.1600/036364407782250544" volume="32" year="2007">Jobson and Luckow 2007</bibRefCitation>
) and some species previously placed in the genus
<taxonomicName authorityName="S.Rauschert" authorityYear="1982" class="Magnoliopsida" family="Leguminosae" genus="Pseudopiptadenia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudopiptadenia" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pseudopiptadenia</emphasis>
</taxonomicName>
(sensu
<bibRefCitation author="Lewis, GP" journalOrPublisher="Arquivos do Jardim Botanico do Rio de Janeiro" pageId="0" pageNumber="239" pagination="43 - 67" refId="B16" refString="Lewis, GP, Lima, MPM, 1991. Pseudopiptadenia Rauschert no Brasil (Leguminosae - Mimosoideae). Arquivos do Jardim Botanico do Rio de Janeiro 30: 43 - 67" title="Pseudopiptadenia Rauschert no Brasil (Leguminosae - Mimosoideae)." volume="30" year="1991">Lewis and Lima 1991</bibRefCitation>
). These two genera had been separated based on seed morphology,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
characterised by ovoid or discoid seeds with a hard, whitish seed coat and a
<normalizedToken originalValue="U">'U'</normalizedToken>
-shaped pleurogram, while
<taxonomicName authorityName="S.Rauschert" authorityYear="1982" class="Magnoliopsida" family="Leguminosae" genus="Pseudopiptadenia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudopiptadenia" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pseudopiptadenia</emphasis>
</taxonomicName>
included species with flat compressed and narrowly winged seeds with a coriaceous testa lacking a pleurogram.
<taxonomicName authorityName="L. P. Queiroz &amp; L. M. Borges" authorityYear="2022" baseAuthorityName="L. P. Queiroz &amp; L. M. Borges" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa brenanii" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="species" species="brenanii">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa brenanii</emphasis>
</taxonomicName>
and
<taxonomicName genus="Pi." lsidName="Pi. leucoxylon" pageId="0" pageNumber="239" rank="species" species="leucoxylon">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leucoxylon</emphasis>
</taxonomicName>
have fruits with only shallowly sinuous margins, more similar to species of the genus
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
.
</paragraph>
<caption doi="10.3897/phytokeys.205.82275.figure4" httpUri="https://binary.pensoft.net/fig/732769" pageId="0" pageNumber="239" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="239">
<emphasis bold="true" pageId="0" pageNumber="239">Figure 4.</emphasis>
Fruits of
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
species
<emphasis bold="true" pageId="0" pageNumber="239">A</emphasis>
<taxonomicName genus="Pi." lsidName="Pi. brenanii" pageId="0" pageNumber="239" rank="species" species="brenanii">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. brenanii</emphasis>
</taxonomicName>
(from
<emphasis italics="true" pageId="0" pageNumber="239">Lewis et al. 1899</emphasis>
, NY)
<emphasis bold="true" pageId="0" pageNumber="239">B</emphasis>
<taxonomicName genus="Pi." lsidName="Pi. inaequalis" pageId="0" pageNumber="239" rank="species" species="inaequalis">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. inaequalis</emphasis>
</taxonomicName>
(from
<emphasis italics="true" pageId="0" pageNumber="239">Moreira et al. 3</emphasis>
, F)
<emphasis bold="true" pageId="0" pageNumber="239">C</emphasis>
<taxonomicName genus="Pi." lsidName="Pi. leptostachya" pageId="0" pageNumber="239" rank="species" species="leptostachya">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leptostachya</emphasis>
</taxonomicName>
(from
<emphasis italics="true" pageId="0" pageNumber="239">Baez et al. 1174</emphasis>
, NY)
<emphasis bold="true" pageId="0" pageNumber="239">D</emphasis>
<taxonomicName genus="Pi." lsidName="Pi. leucoxylon" pageId="0" pageNumber="239" rank="species" species="leucoxylon">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leucoxylon</emphasis>
</taxonomicName>
(from
<emphasis italics="true" pageId="0" pageNumber="239">de Bruijn 1750</emphasis>
, NY)
<emphasis bold="true" pageId="0" pageNumber="239">E</emphasis>
<taxonomicName genus="Pi." lsidName="Pi. moniliformis" pageId="0" pageNumber="239" rank="species" species="moniliformis">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. moniliformis</emphasis>
</taxonomicName>
(from
<emphasis italics="true" pageId="0" pageNumber="239">Nunes 597</emphasis>
, HUEFS)
<emphasis bold="true" pageId="0" pageNumber="239">F</emphasis>
<taxonomicName genus="Pi." lsidName="Pi. obliqua subsp. brasiliensis" pageId="0" pageNumber="239" rank="subSpecies" species="obliqua" subSpecies="brasiliensis">Pi. obliqua subsp. brasiliensis</taxonomicName>
(from
<emphasis italics="true" pageId="0" pageNumber="239">Mori 11837</emphasis>
, NY)
<emphasis bold="true" pageId="0" pageNumber="239">G</emphasis>
<taxonomicName genus="Pi." lsidName="Pi. schumanniana" pageId="0" pageNumber="239" rank="species" species="schumanniana">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. schumanniana</emphasis>
</taxonomicName>
(from
<emphasis italics="true" pageId="0" pageNumber="239">Lima 2994</emphasis>
, RB).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="239">
Besides sharing these fruit traits,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
species also have leaves with few pinnae (1 to 4 [5] pairs, rarely up to 10 pairs in
<taxonomicName genus="Pi." lsidName="Pi. leptostachya" pageId="0" pageNumber="239" rank="species" species="leptostachya">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leptostachya</emphasis>
</taxonomicName>
) and relatively large rhomboid leaflets compared to species of
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
. One exception are the leaves of
<taxonomicName genus="Pi." lsidName="Pi. brenanii" pageId="0" pageNumber="239" rank="species" species="brenanii">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. brenanii</emphasis>
</taxonomicName>
, which are similar to those of
<taxonomicName genus="M." lsidName="M. bahiana" pageId="0" pageNumber="239" rank="species" species="bahiana">
<emphasis italics="true" pageId="0" pageNumber="239">M. bahiana</emphasis>
</taxonomicName>
. All species of
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
present an extrafloral nectary between or shortly below the first pair of pinnae, in contrast to species of
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
that have the nectary below mid-petiole, frequently close to the pulvinus.
</paragraph>
<paragraph pageId="0" pageNumber="239">
Floral traits, although previously disregarded as being generically diagnostic in the group, provide further evidence for the distinction between
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
. The solitary inflorescence spikes in the axils of coevally developing leaves in
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
contrast with the more complex synflorescences of
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Inflorescences of Parapiptadenia, Pityrocarpa and Marlimorimia A Parapiptadenia pterosperma (Benth.) Brenan showing reddish inflorescences in the axils of coeval leaves B Pa. rigida (Benth.) Brenan showing yellowish inflorescences and fruits with valves plicate above the seeds C Pityrocarpa brenanii showing whitish, solitary spikes in the axils of coeval leaves D Pi. moniliformis showing yellowish, solitary spikes in the axil of a coeval leaf E Marlimorimia bahiana (G. P. Lewis &amp; M. P. Lima) L. P. Queiroz &amp; L. M. Borges, showing whitish spikes clustered in efoliate terminal pseudoracemes F Marlimorimia sp. showing yellowish spikes clustered in efoliate terminal pseudoracemes (Photos: A PG Ribeiro; B RT de Queiroz C-E LP Queiroz; F G Siqueira)." figureDoi="10.3897/phytokeys.205.82275.figure3" httpUri="https://binary.pensoft.net/fig/732768" pageId="0" pageNumber="239">3</figureCitation>
; see notes under
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
). All species of
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
have free and glabrous petals, except for
<taxonomicName genus="Pi." lsidName="Pi. leucoxylon" pageId="0" pageNumber="239" rank="species" species="leucoxylon">
<emphasis italics="true" pageId="0" pageNumber="239">Pi. leucoxylon</emphasis>
</taxonomicName>
, in which the petals are connate for a little over 1 mm (
<bibRefCitation DOI="https://doi.org/10.2307/2806514" author="Barneby, RC" journalOrPublisher="Brittonia" pageId="0" pageNumber="239" pagination="236 - 240" refId="B1" refString="Barneby, RC, Grimes, JW, 1984. Two new mimosaceous trees from the American Tropics. Brittonia 36 (3): 236 - 240, DOI: https://doi.org/10.2307/2806514" title="Two new mimosaceous trees from the American Tropics." url="https://doi.org/10.2307/2806514" volume="36" year="1984">Barneby and Grimes 1984</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="239">
<bibRefCitation DOI="https://doi.org/10.1590/2175-78601985376206" author="Lima, MPM" journalOrPublisher="Rodriguesia" pageId="0" pageNumber="239" pagination="53 - 78" refId="B18" refString="Lima, MPM, 1985. Morfologia dos frutos e sementes dos generos da tribo Mimoseae (Leguminosae - Mimosoideae) aplicada a sistematica. Rodriguesia 37 (62): 53 - 78, DOI: https://doi.org/10.1590/2175-78601985376206" title="Morfologia dos frutos e sementes dos generos da tribo Mimoseae (Leguminosae - Mimosoideae) aplicada a sistematica." url="https://doi.org/10.1590/2175-78601985376206" volume="37" year="1985">Lima (1985)</bibRefCitation>
and
<bibRefCitation author="Lewis, GP" journalOrPublisher="Arquivos do Jardim Botanico do Rio de Janeiro" pageId="0" pageNumber="239" pagination="43 - 67" refId="B16" refString="Lewis, GP, Lima, MPM, 1991. Pseudopiptadenia Rauschert no Brasil (Leguminosae - Mimosoideae). Arquivos do Jardim Botanico do Rio de Janeiro 30: 43 - 67" title="Pseudopiptadenia Rauschert no Brasil (Leguminosae - Mimosoideae)." volume="30" year="1991">Lewis and Lima (1991)</bibRefCitation>
provided additional information on embryos and seedlings that are potentially useful for distinguishing
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
from
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
. Embryos of
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
species have a rudimentary plumule, while in
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
, the plumule is developed and multifid. This seems to be correlated with seedling morphology as the studied species of
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa</emphasis>
</taxonomicName>
have bifoliolate eophylls and those of
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
species have pinnate or bipinnate eophylls (
<bibRefCitation author="Lewis, GP" journalOrPublisher="Arquivos do Jardim Botanico do Rio de Janeiro" pageId="0" pageNumber="239" pagination="43 - 67" refId="B16" refString="Lewis, GP, Lima, MPM, 1991. Pseudopiptadenia Rauschert no Brasil (Leguminosae - Mimosoideae). Arquivos do Jardim Botanico do Rio de Janeiro 30: 43 - 67" title="Pseudopiptadenia Rauschert no Brasil (Leguminosae - Mimosoideae)." volume="30" year="1991">Lewis and Lima 1991</bibRefCitation>
).
<taxonomicName authorityName="L. P. Queiroz &amp; L. M. Borges" authorityYear="2022" baseAuthorityName="L. P. Queiroz &amp; L. M. Borges" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Pityrocarpa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pityrocarpa brenanii" order="Fabales" pageId="0" pageNumber="239" phylum="Tracheophyta" rank="species" species="brenanii">
<emphasis italics="true" pageId="0" pageNumber="239">Pityrocarpa brenanii</emphasis>
</taxonomicName>
, however, has embryo morphology more similar to that reported for species of
<taxonomicName authorityName="L. P. Queiroz, L. M. Borges, Marc. F. Simon &amp; P. G. Ribeiro" authorityYear="2022" family="Fabaceae" genus="Marlimorimia" kingdom="Plantae" lsidName="Marlimorimia" order="Fabales" pageId="0" pageNumber="239" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="239">Marlimorimia</emphasis>
</taxonomicName>
(
<bibRefCitation author="Lewis, GP" journalOrPublisher="Arquivos do Jardim Botanico do Rio de Janeiro" pageId="0" pageNumber="239" pagination="43 - 67" refId="B16" refString="Lewis, GP, Lima, MPM, 1991. Pseudopiptadenia Rauschert no Brasil (Leguminosae - Mimosoideae). Arquivos do Jardim Botanico do Rio de Janeiro 30: 43 - 67" title="Pseudopiptadenia Rauschert no Brasil (Leguminosae - Mimosoideae)." volume="30" year="1991">Lewis and Lima 1991</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>