189 lines
21 KiB
XML
189 lines
21 KiB
XML
<document id="214FD40D7E009045802A0E0A4D13BB75" ID-CLB-Dataset="7262" ID-DOI="10.11646/zootaxa.4954.1.1" ID-GBIF-Dataset="64eab05b-4539-4be2-bb06-7af4369949e0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4690775" ID-ZooBank="A8D2AA60-562C-4F98-8000-D792F1E40C87" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618451905616" checkinUser="plazi" docAuthor="Zahniser, James N." docDate="2021" docId="DE47C351FFD2C171FF67D966FBC57B81" docLanguage="en" docName="zootaxa.4954.1.1.pdf" docOrigin="Zootaxa 4954 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hecullus mexicanus Zahniser 2021, n. sp." docType="treatment" docUuid="7420FE24-2D06-48EA-AB5E-EAE1C6BBA0CB" docUuidSource="ZooBank" docVersion="7" lastPageNumber="75" masterDocId="227EBB29FF9AC13AFFF0DE5DFFF67B54" masterDocTitle="Revision of the New World leafhopper tribe Faltalini (Hemiptera: Cicadellidae: Deltocephalinae) and the evolution of brachyptery" masterLastPageNumber="160" masterPageNumber="1" pageNumber="73" updateTime="1698936753110" updateUser="ExternalLinkService">
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<mods:titleInfo id="960EF875D2303E3B52A0031A7EA8380D">
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<mods:title id="615B422C10B6370DE60148F604BB13ED">Revision of the New World leafhopper tribe Faltalini (Hemiptera: Cicadellidae: Deltocephalinae) and the evolution of brachyptery</mods:title>
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<mods:name id="46D592BAA3F5E3AE41828913F1CBD680" type="personal">
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<mods:namePart id="95BEBF05F6B7A04AC9E6FE42E0E33796">Zahniser, James N.</mods:namePart>
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<mods:title id="A024FE87B10DDE61DB655849172AFD59">Zootaxa</mods:title>
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<mods:part id="C4D2145B17F6285AD2A4D05E27C5F8B9">
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<mods:date id="061BA605FB9783DAB3D728A09950EE79">2021</mods:date>
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<mods:number id="BE66414F2351C388188482B31F564D39">2021-04-13</mods:number>
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<mods:number id="55D78D144B6B6A0AA6A7C81EC039B9A4">4954</mods:number>
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<mods:classification id="79297E425968B10E456DC0F0D5CA6193">journal article</mods:classification>
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<mods:identifier id="3DC79D61FC0BAEE8D569B55320FD690D" type="DOI">10.11646/zootaxa.4954.1.1</mods:identifier>
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<treatment id="DE47C351FFD2C171FF67D966FBC57B81" ID-DOI="http://doi.org/10.5281/zenodo.4701194" ID-GBIF-Taxon="181198285" ID-Zenodo-Dep="4701194" LSID="urn:lsid:zoobank.org:act:7420FE24-2D06-48EA-AB5E-EAE1C6BBA0CB" httpUri="http://treatment.plazi.org/id/DE47C351FFD2C171FF67D966FBC57B81" lastPageId="75" lastPageNumber="75" pageId="72" pageNumber="73">
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<subSubSection id="1EF421CCFFD2C172FF67D966FE2E7C02" box="[151,472,1851,1878]" pageId="72" pageNumber="73" type="nomenclature">
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<paragraph id="56517247FFD2C172FF67D966FE2E7C02" blockId="72.[151,472,1851,1914]" box="[151,472,1851,1878]" pageId="72" pageNumber="73">
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<heading id="0D19C52BFFD2C172FF67D966FE2E7C02" bold="true" box="[151,472,1851,1878]" fontSize="11" level="2" pageId="72" pageNumber="73" reason="4">
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<emphasis id="649AAE55FFD2C172FF67D966FE2E7C02" bold="true" box="[151,472,1851,1878]" pageId="72" pageNumber="73">
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<taxonomicName id="91EE09C4FFD2C172FF67D966FE787C02" ID-CoL="8KMZC" authority="Zahniser, 2021" authorityName="Zahniser" authorityYear="2021" box="[151,398,1851,1878]" class="Insecta" family="Cicadellidae" genus="Hecullus" kingdom="Animalia" order="Hemiptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="mexicanus" status="sp. nov.">
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<emphasis id="649AAE55FFD2C172FF67D966FE787C02" bold="true" box="[151,398,1851,1878]" italics="true" pageId="72" pageNumber="73">Hecullus mexicanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7FA9132EFFD2C172FE65D961FE2E7C02" box="[405,472,1852,1878]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1EF421CCFFD2C172FF67D93DFED87C2E" box="[151,302,1888,1914]" pageId="72" pageNumber="73" type="description">
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<paragraph id="56517247FFD2C172FF67D93DFED87C2E" blockId="72.[151,472,1851,1914]" box="[151,302,1888,1914]" pageId="72" pageNumber="73">
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(
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<figureCitation id="CED56EC2FFD2C172FF6FD93DFF0B7C2E" box="[159,253,1888,1914]" captionStart="FIGUIRE 10" captionStartId="27.[151,259,770,794]" captionTargetBox="[151,1436,181,747]" captionTargetId="figure-16@27.[151,1436,181,747]" captionTargetPageId="27" captionText="FIGUIRE 10. Distribution of Acrolithus, Hecalocorica, Hecullus mexicanus n. sp., and three species of Tenucephalus from Mexico and Panama." figureDoi="http://doi.org/10.5281/zenodo.4690803" httpUri="https://zenodo.org/record/4690803/files/figure.png" pageId="72" pageNumber="73">Figs. 10</figureCitation>
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,
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<figureCitation id="CED56EC2FFD2C172FEF9D93DFED07C2E" box="[265,294,1888,1914]" captionStart="FIGUIRE 48" captionStartId="77.[151,259,1471,1495]" captionTargetBox="[159,1435,310,1443]" captionTargetId="figure-67@77.[151,1435,309,1448]" captionTargetPageId="77" captionText="FIGUIRE 48. Hecullus mexicanus n. sp., images of female holotype. (A–C, J) standard views. (Q) habitus, ventral view. (R) sternite VII, pygofer, and ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4690925" httpUri="https://zenodo.org/record/4690925/files/figure.png" pageId="72" pageNumber="73">48</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="1EF421CCFFD2C170FF67D9F5FDAA7DE9" lastPageId="74" lastPageNumber="75" pageId="72" pageNumber="73" type="diagnosis">
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<paragraph id="56517247FFD2C172FF67D9F5FA6A7CB2" blockId="72.[151,1436,1960,2023]" pageId="72" pageNumber="73">
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<emphasis id="649AAE55FFD2C172FF67D9F5FEE47C96" bold="true" box="[151,274,1960,1986]" pageId="72" pageNumber="73">Diagnosis.</emphasis>
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<taxonomicName id="91EE09C4FFD2C172FEE8D9F4FE597C96" authorityName="Zahniser" authorityYear="2021" box="[280,431,1961,1986]" class="Insecta" family="Cicadellidae" genus="Hecullus" kingdom="Animalia" order="Hemiptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="mexicanus">
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<emphasis id="649AAE55FFD2C172FEE8D9F4FE597C96" box="[280,431,1961,1986]" italics="true" pageId="72" pageNumber="73">H. mexicanus</emphasis>
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</taxonomicName>
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can be distinguished from other
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<taxonomicName id="91EE09C4FFD2C172FCECD9F5FC767C96" authorityName="Oman" authorityYear="1949" box="[796,896,1960,1986]" class="Insecta" family="Cicadellidae" genus="Hecullus" kingdom="Animalia" order="Hemiptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis id="649AAE55FFD2C172FCECD9F5FC767C96" box="[796,896,1960,1986]" italics="true" pageId="72" pageNumber="73">Hecullus</emphasis>
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</taxonomicName>
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by its size (female,
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<quantity id="9116DFA2FFD2C172FBADD9F5FB407C97" box="[1117,1206,1960,1987]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="72" pageNumber="73" unit="mm" value="8.3">8.3 mm</quantity>
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), female head length 1.7–1.8x interocular width, female face with broad brown to dark brown band along anterolateral portion of fron-
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</paragraph>
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<paragraph id="56517247FFD3C173FF67DECAFBB07B81" blockId="73.[151,1436,151,213]" pageId="73" pageNumber="74">toclypeus continuing beneath eye and onto ventral side of thorax, face with brown mark from basolateral corner of clypellus to medial margin of lorum and widening along lorum margin toward apex.</paragraph>
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<caption id="029122CFFFD3C173FF67D9EAFCDC7CA0" ID-DOI="http://doi.org/10.5281/zenodo.4690911" ID-Zenodo-Dep="4690911" httpUri="https://zenodo.org/record/4690911/files/figure.png" pageId="73" pageNumber="74" startId="73.[151,259,1975,1999]" targetBox="[161,1431,238,1942]" targetPageId="73">
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<paragraph id="56517247FFD3C173FF67D9EAFCDC7CA0" blockId="73.[151,1437,1975,2036]" pageId="73" pageNumber="74">
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<emphasis id="649AAE55FFD3C173FF67D9EAFEDC7C9B" bold="true" box="[151,298,1975,1999]" pageId="73" pageNumber="74">FIGUIRE 44.</emphasis>
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<taxonomicName id="91EE09C4FFD3C173FEC0D9E5FE0D7C84" box="[304,507,1976,2000]" class="Insecta" family="Cicadellidae" genus="Hecullus" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bracteatus">
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<emphasis id="649AAE55FFD3C173FEC0D9E5FE0D7C84" box="[304,507,1976,2000]" italics="true" pageId="73" pageNumber="74">Hecullus bracteatus</emphasis>
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</taxonomicName>
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. (A–E, H–I) standard views; A1, B1 & C1, female; A2, B2 & C2, male; A3, female nymph. (Q) valve, subgenital plates, connective, and styles, ventral view.
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</paragraph>
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||
</caption>
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<caption id="029122CFFFD0C170FF67DAA9FD147E58" ID-DOI="http://doi.org/10.5281/zenodo.4690915" ID-Zenodo-Dep="4690915" box="[151,738,1268,1293]" httpUri="https://zenodo.org/record/4690915/files/figure.png" pageId="74" pageNumber="75" startId="74.[151,259,1268,1292]" targetBox="[170,1411,183,1242]" targetPageId="74">
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<paragraph id="56517247FFD0C170FF67DAA9FD147E58" blockId="74.[151,738,1268,1293]" box="[151,738,1268,1293]" pageId="74" pageNumber="75">
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<emphasis id="649AAE55FFD0C170FF67DAA9FEDC7E58" bold="true" box="[151,298,1268,1292]" pageId="74" pageNumber="75">FIGUIRE 45.</emphasis>
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<taxonomicName id="91EE09C4FFD0C170FEC0DAA9FE0D7E58" box="[304,507,1268,1292]" class="Insecta" family="Cicadellidae" genus="Hecullus" kingdom="Animalia" order="Hemiptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="bracteatus">
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<emphasis id="649AAE55FFD0C170FEC0DAA9FE0D7E58" box="[304,507,1268,1292]" italics="true" pageId="74" pageNumber="75">Hecullus bracteatus</emphasis>
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</taxonomicName>
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. (J–P) standard views.
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||
</paragraph>
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</caption>
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<paragraph id="56517247FFD0C170FF37DB66FD677E2D" blockId="74.[151,1437,1339,2013]" pageId="74" pageNumber="75">
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<emphasis id="649AAE55FFD0C170FF37DB66FEFD7E01" bold="true" box="[199,267,1339,1365]" pageId="74" pageNumber="75">Body.</emphasis>
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Female
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<quantity id="9116DFA2FFD0C170FE9ADB66FE347E01" box="[362,450,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="74" pageNumber="75" unit="mm" value="8.3">8.3 mm</quantity>
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; head length 1.7–1.8x interocular width. Forewing apex truncate; with many extra crossveins, but venation not completely reticulate.
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</paragraph>
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<paragraph id="56517247FFD0C170FF37DBDFFB097D79" blockId="74.[151,1437,1339,2013]" pageId="74" pageNumber="75">
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<emphasis id="649AAE55FFD0C170FF37DBDFFEE67EC8" bold="true" box="[199,272,1410,1436]" pageId="74" pageNumber="75">Color.</emphasis>
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Female face with broad brown to dark brown band along anterolateral portion of frontoclypeus continuing beneath eye and onto ventral side of thorax, brown bands weak at apex of frontoclypeus; frontoclypeus ivory medially; face with brown mark from basolateral corner of clypellus to medial margin of lorum and widening along lorum margin toward apex. Legs ivory with distinct brown markings basally on anterior side of profemur and subapically and apically on anterior side of meso- and metafemur; with other scattered brown markings.
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</paragraph>
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<paragraph id="56517247FFD0C170FF37D86AFE7E7D05" blockId="74.[151,1437,1339,2013]" box="[199,392,1591,1617]" pageId="74" pageNumber="75">
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<emphasis id="649AAE55FFD0C170FF37D86AFEFD7D05" bold="true" box="[199,267,1591,1617]" pageId="74" pageNumber="75">Male.</emphasis>
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Unknown.
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</paragraph>
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<paragraph id="56517247FFD0C170FF37D806FDAA7DE9" blockId="74.[151,1437,1339,2013]" pageId="74" pageNumber="75">
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<emphasis id="649AAE55FFD0C170FF37D806FED37D21" bold="true" box="[199,293,1627,1653]" pageId="74" pageNumber="75">Female.</emphasis>
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Sternite VII relatively deeply excavated medially, forming pair of bluntly pointed lateral lobes; on each side, deepest excavation reaching to near midlength of sternite; medially with distinct medial rounded lobe; margin laterad of median lobe slightly undulate.
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</paragraph>
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</subSubSection>
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<subSubSection id="1EF421CCFFD0C170FF37D89BFCF07C51" pageId="74" pageNumber="75" type="materials_examined">
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<paragraph id="56517247FFD0C170FF37D89BFCF07C51" blockId="74.[151,1437,1339,2013]" pageId="74" pageNumber="75">
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<emphasis id="649AAE55FFD0C170FF37D89BFE437DB4" bold="true" box="[199,437,1734,1760]" pageId="74" pageNumber="75">Material examined.</emphasis>
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<materialsCitation id="E686781AFFD0C170FE31D89AFCF07C51" ID-GBIF-Occurrence="3080386345" collectingDate="2005-11-10" collectionCode="INHS" collectorName="R. Rakitov" country="Mexico" elevation="2350" latitude="18.63385" location="Azumbilla" longLatPrecision="1" longitude="-97.34344" pageId="74" pageNumber="75" specimenCount="1" specimenCount-female="1" stateProvince="Puebla" typeStatus="holotype">
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<typeStatus id="8955CCE5FFD0C170FE31D89AFDAF7DB5" box="[449,601,1735,1761]" pageId="74" pageNumber="75" type="holotype">HOLOTYPE</typeStatus>
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:
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<specimenCount id="40E8B9CEFFD0C170FD98D89AFD7D7DB5" box="[616,651,1735,1761]" pageId="74" pageNumber="75" type="female">1♀</specimenCount>
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,
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<collectingCountry id="2EF932D7FFD0C170FD6ED89AFCF97DB5" box="[670,783,1735,1761]" name="Mexico" pageId="74" pageNumber="75">MEXICO</collectingCountry>
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,
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<collectingRegion id="942ABCA5FFD0C170FCEFD89AFC987DB5" box="[799,878,1735,1761]" country="Mexico" name="Puebla" pageId="74" pageNumber="75">Puebla</collectingRegion>
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,
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<quantity id="9116DFA2FFD0C170FC8ED89AFC277DB5" box="[894,977,1735,1761]" metricMagnitude="3" metricUnit="m" metricValue="6.3" pageId="74" pageNumber="75" unit="km" value="6.3">6.3 km</quantity>
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E of
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<location id="5331249CFFD0C170FBEAD89AFB657DB5" LSID="urn:lsid:plazi:treatment:DE47C351FFD2C171FF67D966FBC57B81:5331249CFFD0C170FBEAD89AFB657DB5" box="[1050,1171,1735,1761]" country="Mexico" latitude="18.63385" longLatPrecision="1" longitude="-97.34344" name="Azumbilla" pageId="74" pageNumber="75" stateProvince="Puebla">Azumbilla</location>
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,
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<quantity id="9116DFA2FFD0C170FB53D89BFAF47DB5" box="[1187,1282,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="74" pageNumber="75" unit="m" value="2350.0">
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<elevation id="DDC39574FFD0C170FB53D89BFAF47DB5" box="[1187,1282,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="74" pageNumber="75" unit="m" value="2350.0">2350 m</elevation>
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</quantity>
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.,
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<geoCoordinate id="33DA1480FFD0C170FAE6D89BFA6B7DB5" box="[1302,1437,1734,1761]" degrees="18.63385" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="1" value="18.63385">18.63385ºN</geoCoordinate>
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<geoCoordinate id="33DA1480FFD0C170FF67D8B6FED57C51" box="[151,291,1771,1797]" degrees="97.34344" direction="west" orientation="longitude" pageId="74" pageNumber="75" precision="1" value="-97.34344">97.34344ºW</geoCoordinate>
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,
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<date id="22505487FFD0C170FEDFD8B7FE437C51" box="[303,437,1770,1797]" pageId="74" pageNumber="75" value="2005-11-10">
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<collectingDate id="3214AD6FFFD0C170FEDFD8B7FE437C51" box="[303,437,1770,1797]" pageId="74" pageNumber="75" value="2005-11-10">10-XI-2005</collectingDate>
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</date>
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,
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<collectorName id="FB1B1791FFD0C170FE31D8B6FDCA7C51" box="[449,572,1771,1797]" pageId="74" pageNumber="75">R. Rakitov</collectorName>
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, #05-238 [
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<collectionCode id="30FFEA82FFD0C170FD42D8B7FD0D7C51" box="[690,763,1770,1797]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="74" pageNumber="75" type="Other type of research institution/biorepository">INHS</collectionCode>
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].
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1EF421CCFFD0C170FF37D953FA6E7C19" pageId="74" pageNumber="75" type="distribution">
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<paragraph id="56517247FFD0C170FF37D953FA6E7C19" blockId="74.[151,1437,1339,2013]" pageId="74" pageNumber="75">
|
||
<emphasis id="649AAE55FFD0C170FF37D953FE977C7C" bold="true" box="[199,353,1806,1832]" pageId="74" pageNumber="75">Distribution.</emphasis>
|
||
This species is known only from a single female specimen from the state of
|
||
<collectingRegion id="942ABCA5FFD0C170FB45D952FAF27C7D" box="[1205,1284,1807,1833]" country="Mexico" name="Puebla" pageId="74" pageNumber="75">Puebla</collectingRegion>
|
||
,
|
||
<collectingCountry id="2EF932D7FFD0C170FAFFD952FA9E7C7D" box="[1295,1384,1807,1833]" name="Mexico" pageId="74" pageNumber="75">Mexico</collectingCountry>
|
||
. It is by far the southern-most record for the genus, around
|
||
<quantity id="9116DFA2FFD0C170FD04D96EFCA67C19" box="[756,848,1843,1869]" metricMagnitude="6" metricUnit="m" metricValue="1.9" pageId="74" pageNumber="75" unit="km" value="1900.0">1900km</quantity>
|
||
SE of the nearest record in southwest New
|
||
<collectingRegion id="942ABCA5FFD0C170FACBD96EFA627C19" box="[1339,1428,1843,1869]" country="Mexico" name="Mexico" pageId="74" pageNumber="75">Mexico</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1EF421CCFFD0C170FF37D90AFB257C25" box="[199,1235,1879,1905]" pageId="74" pageNumber="75" type="etymology">
|
||
<paragraph id="56517247FFD0C170FF37D90AFB257C25" blockId="74.[151,1437,1339,2013]" box="[199,1235,1879,1905]" pageId="74" pageNumber="75">
|
||
<emphasis id="649AAE55FFD0C170FF37D90AFEB87C25" bold="true" box="[199,334,1879,1905]" pageId="74" pageNumber="75">Etymology.</emphasis>
|
||
The adjectival species name highlight the finding of the new species in
|
||
<collectingCountry id="2EF932D7FFD0C170FB86D90AFB397C25" box="[1142,1231,1879,1905]" name="Mexico" pageId="74" pageNumber="75">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1EF421CCFFD0C171FF37D926FBC57B81" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75" type="discussion">
|
||
<paragraph id="56517247FFD0C171FF37D926FBC57B81" blockId="74.[151,1437,1339,2013]" lastBlockId="75.[151,1436,151,213]" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75">
|
||
<emphasis id="649AAE55FFD0C170FF37D926FECD7CC1" bold="true" box="[199,315,1915,1941]" pageId="74" pageNumber="75">Remarks.</emphasis>
|
||
Collection locality notes (R. Rakitov, pers. comm.) recorded the habitat as a dry meadow among pine-yucca forest, with
|
||
<taxonomicName id="91EE09C4FFD0C170FE6ED9C2FE0C7CED" box="[414,506,1951,1977]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="74" pageNumber="75" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="649AAE55FFD0C170FE6ED9C2FE0C7CED" box="[414,506,1951,1977]" italics="true" pageId="74" pageNumber="75">Opuntia</emphasis>
|
||
</taxonomicName>
|
||
and many dry forbs. It would be preferable to describe this species with at least one male specimen, but
|
||
<taxonomicName id="91EE09C4FFD0C170FE8AD99EFDE57C88" authorityName="Zahniser" authorityYear="2021" box="[378,531,1987,2012]" class="Insecta" family="Cicadellidae" genus="Hecullus" kingdom="Animalia" order="Hemiptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="mexicanus">
|
||
<emphasis id="649AAE55FFD0C170FE8AD99EFDE57C88" box="[378,531,1987,2012]" italics="true" pageId="74" pageNumber="75">H. mexicanus</emphasis>
|
||
</taxonomicName>
|
||
is clearly different and easily separated from the other two species in the genus based on the female. Its discovery substantially expands the known geographic range of the genus. Future collecting efforts directed near the
|
||
<typeStatus id="8955CCE5FFD1C171FE58DEE6FE2E7B81" box="[424,472,187,213]" pageId="75" pageNumber="76">type</typeStatus>
|
||
locality may reveal more specimens including males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |