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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.238.118032" ID-Pensoft-Pub="1314-2003-238-199" ID-Pensoft-UUID="2C202838490D5F6CA2296FE1958BBBE2" ModsDocID="1314-2003-238-199" checkinTime="1708767061112" checkinUser="pensoft" docAuthor="Sennikov, Alexander N. &amp; Tikhomirov, Valery N." docDate="2024" docId="A76A0B623B2E5E5DAEB581E7A8CD05B2" docLanguage="en" docName="PhytoKeys 238: 199-230" docOrigin="PhytoKeys 238" docPubDate="2024-02-23" docSource="http://dx.doi.org/10.3897/phytokeys.238.118032" docTitle="Cytisus hirsutus subsp. var. var. ciliatus Hazsl. in Verh. K. K. Zool. - Bot. Ges. Wien 1: 201 1852" docType="treatment" docVersion="1" id="2C202838490D5F6CA2296FE1958BBBE2" lastPageNumber="199" masterDocId="2C202838490D5F6CA2296FE1958BBBE2" masterDocTitle="Atlas Florae Europaeae notes, 35. Further critical notes on Cytisus sect. Tubocytisus (Fabaceae) in Europe" masterLastPageNumber="230" masterPageNumber="199" pageNumber="199" updateTime="1708767061112" updateUser="pensoft">
<mods:mods id="8BE7447C89C179BF5B67908D4E7C6EAF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A9E1A9A1079BBF1681414F0B04136A79">Atlas Florae Europaeae notes, 35. Further critical notes on Cytisus sect. Tubocytisus (Fabaceae) in Europe</mods:title>
</mods:titleInfo>
<mods:name id="5F8252A855DA4E0188E5C9820DE7CB98" type="personal">
<mods:role id="0DE2287C4C2EAADDBBAE6EF32802BEA7">
<mods:roleTerm id="2E4E5B36CC83090A65C38A16470AA457">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2E4DC2014C120ECD725968540C3F2FA3">Sennikov, Alexander N.</mods:namePart>
<mods:nameIdentifier id="09825E9FBF68A9394DB93A44BB00E603" type="ORCID">https://orcid.org/0000-0001-6664-7657</mods:nameIdentifier>
<mods:affiliation id="1417FCC02C300DF52466831C18D879AF">Botanical Museum, Finnish Museum of Natural History, University of Helsinki, Helsinki 00014, Finland</mods:affiliation>
<mods:nameIdentifier id="DF85C0B65546B90C035D4A3F23B244B9" type="email">alexander.sennikov@helsinki.fi</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="EAF4B1676B1ED42CF65C63A09F696FB7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="456F50C2B8223A7EA1094DDD71C3A5A7">Tikhomirov, Valery N.</mods:namePart>
<mods:nameIdentifier id="9787906932A924B45F8B0C94ED66142E" type="ORCID">https://orcid.org/0000-0003-1822-0557</mods:nameIdentifier>
<mods:affiliation id="6D920B449AB2C525B1C25B2B7046F6F5">Belarusian State University, Minsk, Belarus</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C7F54F74D713F6A2A5C5DDF0AC69C377">text</mods:typeOfResource>
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<mods:title id="A4B6F7610E1AC5D50D1EC71660D57AAD">PhytoKeys</mods:title>
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<mods:part id="5162126E0D86BA97BC1BCF893C8E4737">
<mods:date id="CDB0332658B4C8B14B7302EDC37A9E50">2024</mods:date>
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<mods:number id="CF867391CC4F6933FA8BFCF21780D479">2024-02-23</mods:number>
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<mods:detail id="2F1356F96EDB929773062435B33D069B" type="volume">
<mods:number id="D40E09A3E12BEC1B466BC5737BB141A5">238</mods:number>
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<mods:start id="44A0DD73B8E9026BA3AEDCC772015471">199</mods:start>
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<mods:url id="145157D3CCEDD62ED4C558CA3D5D558C">http://dx.doi.org/10.3897/phytokeys.238.118032</mods:url>
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<mods:classification id="E02DDAD66F92F0C5FA87E7D3A4E00E71">journal article</mods:classification>
<mods:identifier id="3201E7738EC7FD68EA4EE5AB4A6EFB67" type="DOI">http://dx.doi.org/10.3897/phytokeys.238.118032</mods:identifier>
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</mods:mods>
<treatment id="A76A0B623B2E5E5DAEB581E7A8CD05B2" LSID="urn:lsid:plazi:treatment:A76A0B623B2E5E5DAEB581E7A8CD05B2" httpUri="http://treatment.plazi.org/id/A76A0B623B2E5E5DAEB581E7A8CD05B2" lastPageNumber="199" pageId="0" pageNumber="199">
<subSubSection id="7E58FB1B3FA1544EBD09019FC1E7F455" pageId="0" pageNumber="199" type="nomenclature">
<paragraph id="BE2D3E23C8D1E2133F0B4D4C51E1971F" pageId="0" pageNumber="199">
1a.
<taxonomicName id="F090F7104EEEA4434699E18E42CC832D" ID-CoL="9DXBB" LSID="A76A0B62-3B2E-5E5D-AEB5-81E7A8CD05B2" authority="(Wahlenb.) Hazsl. in Verh. K. K. Zool. - Bot. Ges. Wien 1: 201 (1852)" authorityName="Hazsl. in Verh. K. K. Zool. - Bot. Ges. Wien 1: 201" authorityYear="1852" baseAuthorityName="Wahlenb." baseAuthorityYear="1852" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus hirsutus subsp. var. var. ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="variety" species="hirsutus" subSpecies="var." variety="ciliatus">Cytisus hirsutus var. ciliatus (Wahlenb.) Hazsl. in Verh. K.K. Zool.-Bot. Ges. Wien 1: 201 (1852)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="B0865B2172DF26D369A8990AEBF32B7A" pageId="0" pageNumber="199" type="reference_group">
<paragraph id="A844323011DCAE3B2E4459E0D2A77241" pageId="0" pageNumber="199">
<taxonomicName id="C1A44A8E44E3DA5D5C3629B2327760F3" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="ciliatus">- Cytisus ciliatus</taxonomicName>
Wahlenb., Fl. Carp.: 219 (1814) -
<taxonomicName id="2B4C9853C947AC349558DB16D6A75549" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus prostratus subsp. var. var. ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="variety" species="prostratus" subSpecies="var." variety="ciliatus">Cytisus prostratus var. ciliatus</taxonomicName>
(Wahlenb.) W.D.J.Koch, Syn. Deut. Schweiz. Fl. 1: 155 (1837) -
<taxonomicName id="1EB73AB9EFFC1FFFDB568D8E7AE191D4" authorityName="Hazsl. in Verh. K. K. Zool. - Bot. Ges. Wien 1: 201" authorityYear="1852" baseAuthorityName="Wahlenb." class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus hirsutus subsp. ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="subSpecies" species="hirsutus" subSpecies="ciliatus">Cytisus hirsutus subsp. ciliatus</taxonomicName>
(Wahlenb.) Simonk. in Math. Term.
<normalizedToken id="8735A040DA61BAD618AC24D470C1F76C" originalValue="Közlem">Koezlem</normalizedToken>
. 22: 376 (1888) -
<taxonomicName id="7BF74425B5D0CF642321759EEBE50970" class="Magnoliopsida" family="Fabaceae" genus="Chamaecytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chamaecytisus triflorus subsp. ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="subSpecies" species="triflorus" subSpecies="ciliatus">Chamaecytisus triflorus subsp. ciliatus</taxonomicName>
(Wahlenb.) Holub in
<normalizedToken id="21523E0A0E11AF114C6E066020DC8A7C" originalValue="Bertová">Bertova</normalizedToken>
, Fl. Slovenska IV(4): 38 (1988).
</paragraph>
<paragraph id="C07F2F5F50B809F6C850E8A3FC7A32F4" pageId="0" pageNumber="199">
<taxonomicName id="97438B2C0B1EC5148535172F0D53098F" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus glaber" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="glaber">= Cytisus glaber</taxonomicName>
L.f., Suppl. Pl.: 328. 1782, non Lam. 1779, nom. illeg. (Art. 53.1) -
<taxonomicName id="A2C611ACAB0A6A852D826F69791EAF8E" class="Magnoliopsida" family="Fabaceae" genus="Chamaecytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chamaecytisus glaber" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="glaber">Chamaecytisus glaber</taxonomicName>
Rothm. in Feddes Repert. Spec. Nov. Regni Veg. 53: 143 (1944). Type. Not designated.
</paragraph>
<paragraph id="939FD54C20E12BA46A217038B615DEAC" pageId="0" pageNumber="199">
<taxonomicName id="69069DF47F46158A206926AAB5D02288" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus serotinus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="serotinus">= Cytisus serotinus</taxonomicName>
Kit. ex DC., Prodr. 2: 156 (1825) -
<taxonomicName id="3CBF11A0F700E0BCC8164408C89D9B4A" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus hirsutus subsp. var. var. serotinus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="variety" species="hirsutus" subSpecies="var." variety="serotinus">Cytisus hirsutus var. serotinus</taxonomicName>
(Kit. ex DC.)
<normalizedToken id="1E745EE61A028FD6FD8D786F3421B661" originalValue="Soó">Soo</normalizedToken>
in
<normalizedToken id="336BD580A47D042A02EC8E69792F04D9" originalValue="Veröff">Veroeff</normalizedToken>
. Geobot. Inst.
<normalizedToken id="F113945D010AF9896F0E48A541CC7C7C" originalValue="Rübel">Ruebel</normalizedToken>
<normalizedToken id="BF5F9A3906634A4BD22F5070643A1979" originalValue="Zürich">Zuerich</normalizedToken>
6: 254 (1930). Type. Western Ukraine (Mukachevo) or Romania (Satu Mare). Locality unknown, 1815,
<emphasis id="90D485804314ED83539485D06DF08670" italics="true" pageId="0" pageNumber="199">P. Kitaibel</emphasis>
(holotype G-DC barcode G00477721; isotypes BM barcode BM000750883, M barcode M0210789).
</paragraph>
</subSubSection>
<subSubSection id="B0301893C53A67EF76EC2605606C74E2" pageId="0" pageNumber="199" type="materials_examined">
<paragraph id="9C1EEAF63F46A9010F8AD4809BCB4D41" pageId="0" pageNumber="199">
<typeStatus id="965E85B1E156B666F9036F62D55626C9">Type</typeStatus>
.
</paragraph>
<paragraph id="AEF2B1C6882DC35EBDCB51B254E1A08E" pageId="0" pageNumber="199">
<materialsCitation id="3170CDCE30A43746CC424FC97F19CDE4" collectingDate="1813-07-30" collectorName="G. Wahlenberg, Fig." country="Slovakia" location="Zilinsky kraj" specimenCount="1" stateProvince="Zilinsky kraj" typeStatus="lectotype">
<collectingCountry id="7E7D520C0C5ADBB3F51CF8593605503A" name="Slovakia">Slovakia</collectingCountry>
.
<collectingRegion id="A30019168AEE2F784DD1425DD38B34B4" country="Slovakia" name="Zilinsky">
<normalizedToken id="C9637D38114D9B003BE5BF087FCCF734" originalValue="Žilinský">Zilinsky</normalizedToken>
kraj
</collectingRegion>
: &quot;Hradska hola&quot; [
<normalizedToken id="A5C9299461A6D19901AD14216526D571" originalValue="Hradská">Hradska</normalizedToken>
Hora],
<collectingDate id="F122DAE33ECC418A915101580F8BE7BD" value="1813-07-30">30.07.1813</collectingDate>
,
<emphasis id="420A98AAB33853EACB7B19F7DA551CED" italics="true" pageId="0" pageNumber="199">
<collectorName id="DFB28E4F970E0256D9C5BECE8EFDC8FB">G. Wahlenberg</collectorName>
</emphasis>
(
<typeStatus id="1A15185DA3656358CE93348001C46421" type="lectotype">lectotype</typeStatus>
UPS V-1016663, designated here).
<collectorName id="B2A96071B7C5666790E647E29145D90B">Fig.</collectorName>
<figureCitation id="FD6EB15B95FA79D3B0D1162FB246A2D5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lectotype of Cytisus ciliatus Wahlenb." figureDoi="10.3897/phytokeys.238.118032.figure4" httpUri="https://binary.pensoft.net/fig/991100" pageId="0" pageNumber="199">4</figureCitation>
</materialsCitation>
.
</paragraph>
<caption id="6299BE49D1CE9E1E473F20E66DF60988" doi="10.3897/phytokeys.238.118032.figure4" httpUri="https://binary.pensoft.net/fig/991100" pageId="0" pageNumber="199" start="Figure 4" startId="F4">
<paragraph id="744FF9D0AB91DEB1585CBA2B88361458" pageId="0" pageNumber="199">
<emphasis id="7410BBE0AAF7AD4A673609AEE45D3A1D" bold="true" pageId="0" pageNumber="199">Figure 4.</emphasis>
<typeStatus id="3E6E867B768C7045BD58FD4D183C3962" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="15ECEAE106772D9A185F406BD8339AF9" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="ciliatus">
<emphasis id="59A5AF4845221C7F905114AB1AFF79F3" italics="true" pageId="0" pageNumber="199">Cytisus ciliatus</emphasis>
</taxonomicName>
Wahlenb.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1D9D5A78F43B4A4AED32F0E84E45960C" pageId="0" pageNumber="199" type="distribution">
<paragraph id="0B260945267F81F3BF822E3FA4418598" pageId="0" pageNumber="199">Distribution.</paragraph>
<paragraph id="6330049C0CBBC9EB7B0E4ADFEF23F00A" pageId="0" pageNumber="199">
Europe: certainly present in Slovakia, Ukraine, Hungary, Romania and the Balkans; reported as &quot;
<taxonomicName id="0862A4C5D70873260FEA6F5DBC91DAEC" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. falcatus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="falcatus">
<emphasis id="AFBE09FE6949FEDD792DB618CFC42C4B" italics="true" pageId="0" pageNumber="199">C. falcatus</emphasis>
</taxonomicName>
&quot; from North Macedonia (
<bibRefCitation id="F06316B827370DF262990DF89D1B7D13" author="Micevski, K" journalOrPublisher="Hungarian Natural History Museum, Budapest" pageId="0" pageNumber="199" refId="B52" refString="Micevski, K, 2001. Flora of the Republic of Macedonia (Vol. 1(5)). Macedonian Academy of Sciences and Arts, Skopje, 1121-1430. [In Macedonian]" title="Flora of the Republic of Macedonia (Vol. 1 (5 )). Macedonian Academy of Sciences and Arts, Skopje, 1121 - 1430. [In Macedonian]" year="2001">Micevski 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="96389ACD50D13DC045B881300286ED15" pageId="0" pageNumber="199" type="distribution">
<paragraph id="8DEF2B93A8253CD649CCFFABA099E110" pageId="0" pageNumber="199">Notes on taxonomy and distribution.</paragraph>
<paragraph id="AF11F91B7E13E19386D708A03114394E" pageId="0" pageNumber="199">
This taxon was described from the vicinities of
<normalizedToken id="489356BEEF502055557068E401B57A95" originalValue="Liptovský">Liptovsky</normalizedToken>
<normalizedToken id="50ADBB4D2B2E09EBC2FCEF40C9364582" originalValue="Hrádok">Hradok</normalizedToken>
in present-day Slovakia (
<bibRefCitation id="463B27B8027DF275FEDB1D9473481818" author="Wahlenberg, G" journalOrPublisher="Vandenhoeck &amp; Ruprecht, Goettingen" pageId="0" pageNumber="199" refId="B79" refString="Wahlenberg, G, 1814. Flora Carpatorum Principalium. Vandenhoeck &amp; Ruprecht, Goettingen" title="Flora Carpatorum Principalium." year="1814">Wahlenberg 1814</bibRefCitation>
) and occurs in the mountains surrounding the Pannonian Plain and in the Balkans (
<bibRefCitation id="B8862200DAA0DD98E5B098E6B1548F7A" author="Holub, J" editor="Bertova, L" journalOrPublisher="IV / 4). Veda, Bratislava" pageId="0" pageNumber="199" pagination="32 - 60" refId="B38" refString="Holub, J, Bertova, L, 1988. Chamaecytisus Link. In: Bertova, L, Ed., Flora Slovenska (Vol. IV/4). Veda, Bratislava: 32 - 60" title="Chamaecytisus Link." volumeTitle="Flora Slovenska (Vol." year="1988">
Holub and
<normalizedToken id="CC106F3062FECB5CCB7A6C3CA36CF91B" originalValue="Bertová">Bertova</normalizedToken>
1988
</bibRefCitation>
;
<bibRefCitation id="AF303CB8826D20DA31B61A09B5D62A36" author="Pifko, D" editor="Kiraly, G" journalOrPublisher="Aggteleki Nemzeti Park Igazgatosag, Josvafo" pageId="0" pageNumber="199" pagination="239 - 241" refId="B56" refString="Pifko, D, 2009. Chamaecytisus Link. In: Kiraly, G, Ed., Uj magyar fueveszkoenyv: Hatarozokulcsok. Aggteleki Nemzeti Park Igazgatosag, Josvafo: 239 - 241" title="Chamaecytisus Link." volumeTitle="Uj magyar fueveszkoenyv: Hatarozokulcsok." year="2009">
<normalizedToken id="A26C6213AD3F4C3A81F84F75A5E516C3" originalValue="Pifkó">Pifko</normalizedToken>
2009
</bibRefCitation>
and our data).
<taxonomicName id="E14CC00271A9EF170C662066242068D3" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="ciliatus">
<emphasis id="24CA92F80262BAF1EFB1484E91BE93CB" italics="true" pageId="0" pageNumber="199">Cytisus ciliatus</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="B313264368ADA592E54F22310BD6C93E" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. hirsutus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="hirsutus">
<emphasis id="745A606113E87A5F5D753B3FB9707D16" italics="true" pageId="0" pageNumber="199">C. hirsutus</emphasis>
</taxonomicName>
, but differs from the latter by the upper side of its leaf laminae and by pod surfaces being glabrous or nearly so (vs. regularly hairy). So far, we have no evidence that the distribution of hairy and glabrous plants of
<taxonomicName id="E539DA9CFBD84A826F19C774A224D94A" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. hirsutus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="hirsutus">
<emphasis id="F05FEC89DB88B4EE7691C7110476C41A" italics="true" pageId="0" pageNumber="199">C. hirsutus</emphasis>
</taxonomicName>
is separate; this distinction denotes the same casual loss of pubescence as observed in some other species of
<taxonomicName id="913EB96CABDBCE405B8C92CA44661B58" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="genus">
<emphasis id="584D4AF504ACA73C4EEB9772D5E2819B" italics="true" pageId="0" pageNumber="199">Cytisus</emphasis>
</taxonomicName>
(
<taxonomicName id="D4713EFD70ED2C80F3B6FCA2ABA4DA2D" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. ruthenicus subsp. var. var. zingeri" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="variety" species="ruthenicus" subSpecies="var." variety="zingeri">C. ruthenicus var. zingeri</taxonomicName>
Nenjukov:
<bibRefCitation id="D591B690F76C9BF80AA61AD50857D2C9" author="Sennikov, AN" journalOrPublisher="Memoranda Societatis pro Fauna et Flora Fennica" pageId="0" pageNumber="199" pagination="48 - 63" refId="B64" refString="Sennikov, AN, Tikhomirov, VN, Birykova, OV, Mininzon, IL, 2021. Personalities of Russian amateur botany, 3. Ivan Markelovitsch Schvetzov (1852-1924) and Theodor Stepanovitsch Nenukow (1883-1934), prominent experts in the flora of Nizhni Novgorod Region, and the taxonomic identity of plant species described by Nenukow. Memoranda Societatis pro Fauna et Flora Fennica 97: 48 - 63" title="Personalities of Russian amateur botany, 3. Ivan Markelovitsch Schvetzov (1852 - 1924) and Theodor Stepanovitsch Nenukow (1883 - 1934), prominent experts in the flora of Nizhni Novgorod Region, and the taxonomic identity of plant species described by Nenukow." volume="97" year="2021">Sennikov et al. (2021)</bibRefCitation>
;
<taxonomicName id="815EC0D6EA2D8653DCC20844EA38391C" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. polytrichus subsp. var. var. subglabratus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="variety" species="polytrichus" subSpecies="var." variety="subglabratus">C. polytrichus var. subglabratus</taxonomicName>
Val.N.Tikhom. &amp; Sennikov, see below) and corresponds to the rank of variety.
</paragraph>
<paragraph id="493CBA47F7FB292B8A1A0A1E0D33682D" pageId="0" pageNumber="199">
Some authors (
<bibRefCitation id="873DA50C9257BC918228B9264CAD326E" author="Bernard, C" journalOrPublisher="Phytologia Balcanica" pageId="0" pageNumber="199" refId="B8" refString="Bernard, C, 1977. Chamaecytisus glaber (L.f.) Rothm. In: Auquier P (Ed.) Societe pour l'Echange des plantes vasculaires de l'Europe occidentale et du bassin mediterraneen (Vol. 17): No. 8385. [Exsiccatae]" title="Chamaecytisus glaber (L. f.) Rothm. In: Auquier P (Ed.) Societe pour l'Echange des plantes vasculaires de l'Europe occidentale et du bassin mediterraneen (Vol. 17): No. 8385. [Exsiccatae]" year="1977">Bernard 1977</bibRefCitation>
) interpreted the name
<taxonomicName id="5004113E803156A77A9302D2E1181B9D" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus glaber" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="glaber">
<emphasis id="DA706F576C169080E1BAC126B33D9982" italics="true" pageId="0" pageNumber="199">Cytisus glaber</emphasis>
</taxonomicName>
as corresponding to
<taxonomicName id="DF92F38612E1B60143911330F052B069" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. hirsutus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="hirsutus">
<emphasis id="F632DF09C202A96ADDCDD8DCC55553CE" italics="true" pageId="0" pageNumber="199">C. hirsutus</emphasis>
</taxonomicName>
, which cannot be true because of its leaves glabrous above. Judging from the glabrous leaves of the plant and its occurrence in
<normalizedToken id="178492A6D4D7D7380701BD986B973C29" originalValue="“Austria”">&quot;Austria&quot;</normalizedToken>
,
<taxonomicName id="5BEB7DB5791A82656751472F579C498C" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. glaber" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="glaber">
<emphasis id="A71A312163B5E4264923798805C6B2DF" italics="true" pageId="0" pageNumber="199">C. glaber</emphasis>
</taxonomicName>
is an earlier (albeit illegitimate and therefore unusable) synonym of
<taxonomicName id="D903A3EFEE75AA2CD869CFE78B8A5BDE" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="ciliatus">
<emphasis id="FAEFFA83E39AAB0D59D5D9D177B822D2" italics="true" pageId="0" pageNumber="199">C. ciliatus</emphasis>
</taxonomicName>
Wahlenb. (
<taxonomicName id="3C3C20E63F01923BC03F7BE33B65D677" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. hirsutus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="hirsutus">
<emphasis id="EBE108B448B280D99C055BBE2825076C" italics="true" pageId="0" pageNumber="199">C. hirsutus</emphasis>
</taxonomicName>
s.l.).
<bibRefCitation id="84F65266755C0D38AB6CBBDEFEC0B85C" author="Tzvelev, NN" editor="Fedorov, AA" journalOrPublisher="6). Science Publishers, Leningrad" pageId="0" pageNumber="199" pagination="216 - 225" refId="B73" refString="Tzvelev, NN, 1987. Chamaecytisus Link. In: Fedorov, AA, Ed., Flora of Eastern Europe (Vol. 6). Science Publishers, Leningrad: 216 - 225" title="Chamaecytisus Link." volumeTitle="Flora of Eastern Europe (Vol." year="1987">Tzvelev (1987)</bibRefCitation>
formally accepted
<taxonomicName id="83B416FBC8E28D59532B8094040E9785" class="Magnoliopsida" family="Fabaceae" genus="Chamaecytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chamaecytisus glaber" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="glaber">
<emphasis id="BD102DA2843F5BDE18DFE1899C4DAC2D" italics="true" pageId="0" pageNumber="199">Chamaecytisus glaber</emphasis>
</taxonomicName>
(with
<taxonomicName id="A467CD1557CF555A1BB08E4897605D9C" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. elongatus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="elongatus">
<emphasis id="A6CBE6508B271FA297CE0682ED5CA209" italics="true" pageId="0" pageNumber="199">C. elongatus</emphasis>
</taxonomicName>
mis-added to its synonymy) and applied it to west Ukrainian and cultivated plants of Central European origin with erect stems, leaves glabrous above, lateral inflorescences and patent pubescence, which agrees with our interpretation.
</paragraph>
<paragraph id="DB39DC0563299D2F69D70427B4C5E2C7" pageId="0" pageNumber="199">
<taxonomicName id="4FAE7AA9AC1672F03711B16017D370AF" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus serotinus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="serotinus">
<emphasis id="77F255EB74FCB314316A81D8CFD89EDD" italics="true" pageId="0" pageNumber="199">Cytisus serotinus</emphasis>
</taxonomicName>
is a plant with the leaves glabrous above, which belongs to the
<taxonomicName id="3138EF2E0F66C967E00C0BF97C8C0665" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. hirsutus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="hirsutus">
<emphasis id="2AC2893CF8B055086A784DCBAB88C010" italics="true" pageId="0" pageNumber="199">C. hirsutus</emphasis>
</taxonomicName>
group. It was originally recognised due to its presumed late flowering season, but merely coincides with
<taxonomicName id="CD1C81435D2C8D20FFF8E01EE638339F" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="ciliatus">
<emphasis id="0E7C3B40BC57BB4405FFDA4C1C93DAC7" italics="true" pageId="0" pageNumber="199">C. ciliatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A3F4E4B522C5877E2340E3D6938BB517" pageId="0" pageNumber="199" type="notes on nomenclature">
<paragraph id="8E9417FE7E9EF7A508A6567D9F450B8E" pageId="0" pageNumber="199">Notes on nomenclature.</paragraph>
<paragraph id="08626739C12BFFB9B4AF485938F2E89C" pageId="0" pageNumber="199">
<bibRefCitation id="BA6B0F3FF5EC828D69E41872BA01F6DB" author="Wahlenberg, G" journalOrPublisher="Vandenhoeck &amp; Ruprecht, Goettingen" pageId="0" pageNumber="199" refId="B79" refString="Wahlenberg, G, 1814. Flora Carpatorum Principalium. Vandenhoeck &amp; Ruprecht, Goettingen" title="Flora Carpatorum Principalium." year="1814">Wahlenberg (1814)</bibRefCitation>
distinguished
<taxonomicName id="3FD8544CE65170CD12C2323165B2AC9C" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="ciliatus">
<emphasis id="861817E3E976E22A29AD76DC62D61E57" italics="true" pageId="0" pageNumber="199">Cytisus ciliatus</emphasis>
</taxonomicName>
from
<taxonomicName id="2257CA3D70C1F21B072414AB91B53F5D" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. hirsutus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="hirsutus">
<emphasis id="B5E542AFC2D3749BA1F288767B0E30D3" italics="true" pageId="0" pageNumber="199">C. hirsutus</emphasis>
</taxonomicName>
, which was the original name for his material, by the pubescence of its leaves and pods. In the collections of UPS, where the Herbarium of Wahlenberg is housed, two specimens of the original material were found, both corresponding to the original description and the provenance cited in the protologue. One specimen bears precise collection data, but the draft name of the taxon (
<taxonomicName id="07E6A74EADCD1F14D921FE99355CFA92" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. hirsutus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="hirsutus">
<emphasis id="C75D3D9B396E781600122C060068F88B" italics="true" pageId="0" pageNumber="199">C. hirsutus</emphasis>
</taxonomicName>
[...] glabris) written by Wahlenberg, whereas the second specimen bears the final plant name (
<taxonomicName id="ECEE45B2133D929C51E08DD6B52DAC1B" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. ciliatus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="ciliatus">
<emphasis id="6F7227DA5C4C59F1DE70E5E9390DFEEA" italics="true" pageId="0" pageNumber="199">C. ciliatus</emphasis>
</taxonomicName>
), but generalised collection data (&quot;e montibus Carpaticis&quot;) written by C.P. Thunberg. As both specimens correspond to the taxon as circumscribed by Wahlenberg and are undoubtedly linked with the protologue, we prefer the specimen with exact provenance from the
<normalizedToken id="F2D8E1C144B0ED857A00FEE4A1781705" originalValue="author’s">author's</normalizedToken>
collection as a lectotype.
</paragraph>
<paragraph id="F2FB1050B42074AB29E69B1B54843B83" pageId="0" pageNumber="199">
Despite all searches, we were not able to trace any herbarium material linked with the protologue of
<taxonomicName id="6E07BB7C61C8C656CD2035E26C4CC338" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. glaber" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="glaber">
<emphasis id="93A62C9C9840E14DECA7785059CA46F9" italics="true" pageId="0" pageNumber="199">C. glaber</emphasis>
</taxonomicName>
(Linnaeus filius 1782), in which a species with the leaves glabrous above and slightly hairy below was described from
<normalizedToken id="D22956DCE3B885A86F7554FD6776E37E" originalValue="“Austria”">&quot;Austria&quot;</normalizedToken>
. The only original element, an illustration of &quot;
<taxonomicName id="A03EFD8939F4D0D44433C3F05A95F63F" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cytisus glaber" order="Fabales" pageId="0" pageNumber="199" phylum="Tracheophyta" rank="species" species="glaber">Cytisus glaber</taxonomicName>
, siliqua angusta&quot; in
<bibRefCitation id="2FF35746447B4A1CF73657811C16C3C5" DOI="https://doi.org/10.5962/bhl.title.146639" author="Bauhin, J" journalOrPublisher="Impensis Joannis Regis, Basel" pageId="0" pageNumber="199" refId="B6" refString="Bauhin, J, Cherler, JH, 1650. Historia plantarum universalis (Vol. 1(2)). [Typographia Caldoriana,] Ebrodunum [Yverdon], 241-466. https://doi.org/10.5962/bhl.title.146639" title="Historia plantarum universalis (Vol. 1 (2 )). [Typographia Caldoriana,] Ebrodunum [Yverdon], 241 - 466." url="https://doi.org/10.5962/bhl.title.146639" year="1650">Bauhin and Cherler (1650</bibRefCitation>
: 373) was rejected by
<bibRefCitation id="B615CE8CE8E0444891053B8BE14D3516" DOI="https://doi.org/10.1080/00837792.1991.10670496" author="Cristofolini, G" journalOrPublisher="Webbia" pageId="0" pageNumber="199" pagination="187 - 219" refId="B15" refString="Cristofolini, G, 1991. Taxonomic revision of Cytisus Desf. sect. Tubocytisus DC. (Fabaceae). Webbia 45 (2): 187 - 219, DOI: https://doi.org/10.1080/00837792.1991.10670496" title="Taxonomic revision of Cytisus Desf. sect. Tubocytisus DC. (Fabaceae)." url="https://doi.org/10.1080/00837792.1991.10670496" volume="45" year="1991">Cristofolini (1991)</bibRefCitation>
as conflicting with the original description (calyces depicted as campanulate, whereas the protologue stated the calyx being &quot;oblongus subventricosus&quot;), although this presumed conflict may be explained by the crude nature of this drawing. So far, this species name remains untypified and interpreted on the basis of the protologue (
<bibRefCitation id="611D661EFF4C6036E60C5A842462FA78" author="Tzvelev, NN" editor="Fedorov, AA" journalOrPublisher="6). Science Publishers, Leningrad" pageId="0" pageNumber="199" pagination="216 - 225" refId="B73" refString="Tzvelev, NN, 1987. Chamaecytisus Link. In: Fedorov, AA, Ed., Flora of Eastern Europe (Vol. 6). Science Publishers, Leningrad: 216 - 225" title="Chamaecytisus Link." volumeTitle="Flora of Eastern Europe (Vol." year="1987">Tzvelev 1987</bibRefCitation>
).
</paragraph>
<paragraph id="83853EBBD0F9258D5631DD28E3D5B20F" pageId="0" pageNumber="199">
A later synonym belonging to the same taxon is
<taxonomicName id="15EA71965F7F5D87B8B7A0C486B46BAD" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. serotinus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="serotinus">
<emphasis id="770EA97B8C310388A9B00ECD967821DD" italics="true" pageId="0" pageNumber="199">C. serotinus</emphasis>
</taxonomicName>
Kit. ex DC. (
<bibRefCitation id="B9DBCE6510BDF86E19A84BEE0DFE7F7E" author="Candolle, AP" journalOrPublisher="Treuttel &amp; Wuertz, Paris" pageId="0" pageNumber="199" refId="B12" refString="Candolle, AP, 1825. Prodromus Sytematis Naturalis Regni Vegetabilis (Vol. 2). Treuttel &amp; Wuertz, Paris" title="Prodromus Sytematis Naturalis Regni Vegetabilis (Vol. 2)." year="1825">Candolle 1825</bibRefCitation>
), described from historical
<normalizedToken id="3E012598EC9CD8DE80762F26191CD643" originalValue="“Hungary”">&quot;Hungary&quot;</normalizedToken>
without a further specification.
<bibRefCitation id="3D0D28C061B9538E6E4D1AFCC82E913C" author="Pifko, D" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="199" pagination="21 - 32" refId="B55" refString="Pifko, D, 2005. Taxonomic revision and typification of Cytisus (Leguminosae) in the Herbarium Carpato-Pannonicum in Budapest (BP). Annales Historico-Naturales Musei Nationalis Hungarici 97: 21 - 32" title="Taxonomic revision and typification of Cytisus (Leguminosae) in the Herbarium Carpato-Pannonicum in Budapest (BP)." volume="97" year="2005">
<normalizedToken id="5E29B8356764BB38CC74E6755F612313" originalValue="Pifkó">Pifko</normalizedToken>
(2005)
</bibRefCitation>
designated a lectotype at BP; since no specimens were cited by Candolle as syntypes, his only specimen used for the original description is the holotype, and the lectotype at BP has no standing. The only original specimen in
<normalizedToken id="8C189E5666F75D11F9E21745BD89B96A" originalValue="Candolle’s">Candolle's</normalizedToken>
herbarium at G is lacking a precise provenance, which can be derived from comparisons with the main collections of P. Kitaibel kept at BP (
<bibRefCitation id="C20404570CB9175BAF409416303BB3CD" author="Javorka, S" journalOrPublisher="Akademiai Kiado, Budapest" pageId="0" pageNumber="199" refId="B40" refString="Javorka, S, 1957. Kitaibel Pal. Akademiai Kiado, Budapest" title="Kitaibel Pal." year="1957">
<normalizedToken id="8D0FAA44B5D67D796DA9F1FDB2EAA24D" originalValue="Jávorka">Javorka</normalizedToken>
1957
</bibRefCitation>
) and from the diaries of Kitaibel (
<bibRefCitation id="0A0C52F898E0A7B705048B750395070A" author="Gombocz, E [Ed.]" journalOrPublisher="Verlag des Ungarischen naturwissenschaftlichen Museum, Budapest" pageId="0" pageNumber="199" refId="B32" refString="Gombocz, E [Ed.], 1945. Diaria itinerum Pauli Kitaibelii (Vols 1-2). Verlag des Ungarischen naturwissenschaftlichen Museum, Budapest" title="Diaria itinerum Pauli Kitaibelii (Vols 1 - 2)." year="1945">Gombocz 1945</bibRefCitation>
;
<bibRefCitation id="4FDB720D1527DCA1585058FE9AB3E4E0" author="Lokoes, L [Ed.]" journalOrPublisher="Hungarian Natural History Museum, Budapest" pageId="0" pageNumber="199" refId="B51" refString="Lokoes, L [Ed.], 2001. Diaria itinerum Pauli Kitaibelii (Vol. 3). Hungarian Natural History Museum, Budapest" title="Diaria itinerum Pauli Kitaibelii (Vol. 3)." year="2001">
<normalizedToken id="1404D43DAB1F21498B64A1294DD24F9D" originalValue="Lőkös">Lokoes</normalizedToken>
2001
</bibRefCitation>
).
</paragraph>
<paragraph id="05924795E56D7DD4E3876338F66752F7" pageId="0" pageNumber="199">
Three specimens identified as
<taxonomicName id="B1735E395B6A2A95CE5DF77FE95BC171" class="Equisetopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. serotinus" order="Fabales" pageId="0" pageNumber="199" phylum="Magnoliophyta" rank="species" species="serotinus">
<emphasis id="C8218DAD0AD894C7393DFD33522DD6C7" italics="true" pageId="0" pageNumber="199">C. serotinus</emphasis>
</taxonomicName>
are preserved in the herbarium of Kitaibel at BP (
<bibRefCitation id="C394D397BC0B67A1008A045C805ED523" author="Pifko, D" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="199" pagination="21 - 32" refId="B55" refString="Pifko, D, 2005. Taxonomic revision and typification of Cytisus (Leguminosae) in the Herbarium Carpato-Pannonicum in Budapest (BP). Annales Historico-Naturales Musei Nationalis Hungarici 97: 21 - 32" title="Taxonomic revision and typification of Cytisus (Leguminosae) in the Herbarium Carpato-Pannonicum in Budapest (BP)." volume="97" year="2005">
<normalizedToken id="590623BAF41A1CCC4E6C84C1FCAEBEF4" originalValue="Pifkó">Pifko</normalizedToken>
2005
</bibRefCitation>
), collected near Mukachevo in present-day Ukraine and at
<normalizedToken id="9652EC863530D2D6FF1B42F8420D6A81" originalValue="Gödöllő">Goedoello</normalizedToken>
in present-day Hungary. Kitaibel (
<bibRefCitation id="87AD19BDF06595898ED4CFDA2D497223" author="Lokoes, L [Ed.]" journalOrPublisher="Hungarian Natural History Museum, Budapest" pageId="0" pageNumber="199" refId="B51" refString="Lokoes, L [Ed.], 2001. Diaria itinerum Pauli Kitaibelii (Vol. 3). Hungarian Natural History Museum, Budapest" title="Diaria itinerum Pauli Kitaibelii (Vol. 3)." year="2001">
<normalizedToken id="103FA1E037BE3DC48A9D8B7F9716C7DC" originalValue="Lőkös">Lokoes</normalizedToken>
2001
</bibRefCitation>
) also mentioned that he collected this species near
<normalizedToken id="0174C145C891C7DBBBE45D5ADA685A1C" originalValue="Szatmár">Szatmar</normalizedToken>
(now Satu Mare in Romania, near the border with Hungary and Ukraine). The specimen at G-DC is dated as received in 1815 and seemingly was collected during that year on the way from Mukachevo to Satu Mare (
<bibRefCitation id="823ABC86AD207E27E8F5A403F784650D" author="Lokoes, L [Ed.]" journalOrPublisher="Hungarian Natural History Museum, Budapest" pageId="0" pageNumber="199" refId="B51" refString="Lokoes, L [Ed.], 2001. Diaria itinerum Pauli Kitaibelii (Vol. 3). Hungarian Natural History Museum, Budapest" title="Diaria itinerum Pauli Kitaibelii (Vol. 3)." year="2001">
<normalizedToken id="C1293D936109D2041482BABFC36E098F" originalValue="Lőkös">Lokoes</normalizedToken>
2001
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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