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<mods:title id="16FC257452B7BC5BC98C9CF983126EEC">Towards reliable identification of male Dicerura: descriptions of three new and seven poorly known species in the Palearctic region (Diptera: Cecidomyiidae, Porricondylinae)</mods:title>
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<treatment id="9E1D954CFFE3966DFF51F95DDE5AFD7E" ID-DOI="http://doi.org/10.5281/zenodo.6490455" ID-GBIF-Taxon="148671191" ID-Zenodo-Dep="6490455" LSID="urn:lsid:plazi:treatment:9E1D954CFFE3966DFF51F95DDE5AFD7E" httpUri="http://treatment.plazi.org/id/9E1D954CFFE3966DFF51F95DDE5AFD7E" lastPageId="7" lastPageNumber="92" pageId="6" pageNumber="91">
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<paragraph id="160B245AFFE3966CFF51F95DD828F91D" blockId="6.[151,548,1751,1811]" box="[151,548,1751,1777]" pageId="6" pageNumber="91">
<heading id="4D439336FFE3966CFF51F95DD828F91D" bold="true" box="[151,548,1751,1777]" fontSize="11" level="1" pageId="6" pageNumber="91" reason="1">
<taxonomicName id="D1B45FD9FFE3966CFF51F95DD828F91D" ID-CoL="35JVW" authority="Mamaev, 1966" authorityName="Mamaev" authorityYear="1966" box="[151,548,1751,1777]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="barbata">
<emphasis id="24C0F848FFE3966CFF51F95DD828F91D" bold="true" box="[151,548,1751,1777]" pageId="6" pageNumber="91">
<emphasis id="24C0F848FFE3966CFF51F95DDB64F91D" bold="true" box="[151,360,1751,1777]" italics="true" pageId="6" pageNumber="91">Dicerura barbata</emphasis>
<bibRefCitation id="722559ABFFE3966CFEA8F95DD828F91D" box="[366,548,1751,1777]" pageId="6" pageNumber="91" refId="ref11145" refString="Mamaev, B. M. (1966) New and little known Palaearctic gall midges of the tribe Porricondylini (Diptera, Cecidomyiidae). Acta Entomologica BohemoSlovaca, 63, 213 - 239 [in Russian]" type="journal article">Mamaev, 1966</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="160B245AFFE3966CFF51F970DB0AF8FF" blockId="6.[151,548,1751,1811]" box="[151,262,1786,1811]" pageId="6" pageNumber="91">
<figureCitation id="8E8F38DFFFE3966CFF51F970DB0AF8FF" box="[151,262,1786,1811]" captionStart="FIGURES 912" captionStartId="7.[151,264,1983,2005]" captionTargetBox="[175,1392,758,1907]" captionTargetId="figure@7.[151,1436,713,1961]" captionTargetPageId="7" captionText="FIGURES 912. Dicerura barbata, male, from Sweden. 9: Ninth tergite, dorsal. 10: Gonostylus, ventral. 11: Genitalia, ventral. 12: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455513/files/figure.png" pageId="6" pageNumber="91">Figs 912</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5EAE77D1FFE3966CFF51F8C9DEB0F807" pageId="6" pageNumber="91" type="reference_group">
<paragraph id="160B245AFFE3966CFF51F8C9DEB0F807" blockId="6.[151,1437,1858,2027]" pageId="6" pageNumber="91">
<materialsCitation id="A6DC2E07FFE3966CFF51F8C9DF6DF893" ID-GBIF-Occurrence="1915497327" country="Ukraine" location="Ukraine" pageId="6" pageNumber="91" specimenCount="1" typeStatus="holotype">
<taxonomicName id="D1B45FD9FFE3966CFF51F8C9DB68F8B6" ID-CoL="35JVW" authorityName="Mamaev" authorityYear="1966" box="[151,356,1858,1882]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="barbata">
<emphasis id="24C0F848FFE3966CFF51F8C9DB68F8B6" box="[151,356,1858,1882]" italics="true" pageId="6" pageNumber="91">Dicerura barbata</emphasis>
</taxonomicName>
was previously known only from the original material, referred to by
<bibRefCitation id="722559ABFFE3966CFB78F8C9DF7BF8B7" box="[1214,1399,1858,1883]" pageId="6" pageNumber="91" refId="ref11145" refString="Mamaev, B. M. (1966) New and little known Palaearctic gall midges of the tribe Porricondylini (Diptera, Cecidomyiidae). Acta Entomologica BohemoSlovaca, 63, 213 - 239 [in Russian]" type="journal article">Mamaev (1966)</bibRefCitation>
as consisting of four males (including the
<typeStatus id="C90F9AF8FFE3966CFDABF8ECD8DDF893" box="[621,721,1894,1919]" pageId="6" pageNumber="91" type="holotype">holotype</typeStatus>
) and a female, from two separate localities in
<collectingCountry id="6EA364CAFFE3966CFAC5F8ECDF6DF893" box="[1283,1377,1894,1919]" name="Ukraine" pageId="6" pageNumber="91">Ukraine</collectingCountry>
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. The morphological description in the same publication takes no account of the female; rather it is stated there that females of this species were unknown. We identified a male from northern Sweden as conspecific with
<taxonomicName id="D1B45FD9FFE3966CFAE7F825DF90F82A" ID-CoL="35JVW" authorityName="Mamaev" authorityYear="1966" box="[1313,1436,1966,1990]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="barbata">
<emphasis id="24C0F848FFE3966CFAE7F825DF90F82A" box="[1313,1436,1966,1990]" italics="true" pageId="6" pageNumber="91">D. barbata</emphasis>
</taxonomicName>
based on the genitalic drawing provided with the original description (
<bibRefCitation id="722559ABFFE3966CFC6CF859DEA3F807" box="[938,1199,2002,2027]" pageId="6" pageNumber="91" refId="ref11145" refString="Mamaev, B. M. (1966) New and little known Palaearctic gall midges of the tribe Porricondylini (Diptera, Cecidomyiidae). Acta Entomologica BohemoSlovaca, 63, 213 - 239 [in Russian]" type="journal article">Mamaev 1966: fig. 6.3</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5EAE77D1FFE2966DFF01FF1DDF15FE37" pageId="7" pageNumber="92" type="diagnosis">
<paragraph id="160B245AFFE2966DFF01FF1DDF90FF5C" blockId="7.[151,1436,151,659]" box="[199,1436,151,176]" pageId="7" pageNumber="92">
<emphasis id="24C0F848FFE2966DFF01FF1DDB4DFF5C" bold="true" box="[199,321,151,176]" pageId="7" pageNumber="92">Diagnosis.</emphasis>
The gonocoxites of
<taxonomicName id="D1B45FD9FFE2966DFDEBFF13D8A6FF5C" authorityName="Mamaev" authorityYear="1966" box="[557,682,152,176]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="barbata">
<emphasis id="24C0F848FFE2966DFDEBFF13D8A6FF5C" box="[557,682,152,176]" italics="true" pageId="7" pageNumber="92">D. barbata</emphasis>
</taxonomicName>
are quite unlike that of all other
<taxonomicName id="D1B45FD9FFE2966DFBE0FF12DE81FF43" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2013" box="[1062,1165,152,175]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="24C0F848FFE2966DFBE0FF12DE81FF43" box="[1062,1165,152,175]" italics="true" pageId="7" pageNumber="92">Dicerura</emphasis>
</taxonomicName>
in having an extremely
</paragraph>
<paragraph id="160B245AFFE2966DFF51FF36DF90FECE" blockId="7.[151,1436,151,659]" pageId="7" pageNumber="92">
large emargination ventrally, which leaves just a short, asetose intercoxal bridge; the emargination is partly filled with an ovate, microtrichose lobe (
<figureCitation id="8E8F38DFFFE2966DFD80FF68D8ACFF17" box="[582,672,226,251]" captionStart="FIGURES 912" captionStartId="7.[151,264,1983,2005]" captionTargetBox="[175,1392,758,1907]" captionTargetId="figure@7.[151,1436,713,1961]" captionTargetPageId="7" captionText="FIGURES 912. Dicerura barbata, male, from Sweden. 9: Ninth tergite, dorsal. 10: Gonostylus, ventral. 11: Genitalia, ventral. 12: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455513/files/figure.png" pageId="7" pageNumber="92">Fig. 11</figureCitation>
, ↓1). The large, subtriangular gonocoxal processes are densely microtrichose medially; the dorsal apodemes are conspicuously long and thin (
<figureCitation id="8E8F38DFFFE2966DFBC5FE83DE58FECD" box="[1027,1108,265,289]" captionStart="FIGURES 912" captionStartId="7.[151,264,1983,2005]" captionTargetBox="[175,1392,758,1907]" captionTargetId="figure@7.[151,1436,713,1961]" captionTargetPageId="7" captionText="FIGURES 912. Dicerura barbata, male, from Sweden. 9: Ninth tergite, dorsal. 10: Gonostylus, ventral. 11: Genitalia, ventral. 12: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455513/files/figure.png" pageId="7" pageNumber="92">Fig. 11</figureCitation>
, ↓2). The elongate gonostylus
</paragraph>
<paragraph id="160B245AFFE2966DFF51FEA7DF90FE81" blockId="7.[151,1436,151,659]" pageId="7" pageNumber="92">
has a small side lobe subapicomedially, which is covered apically and dorsally with dense, large microtrichia, a few setulae, and 12 short bristles (
<figureCitation id="8E8F38DFFFE2966DFE30FEDED845FE81" box="[502,585,340,365]" captionStart="FIGURES 912" captionStartId="7.[151,264,1983,2005]" captionTargetBox="[175,1392,758,1907]" captionTargetId="figure@7.[151,1436,713,1961]" captionTargetPageId="7" captionText="FIGURES 912. Dicerura barbata, male, from Sweden. 9: Ninth tergite, dorsal. 10: Gonostylus, ventral. 11: Genitalia, ventral. 12: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455513/files/figure.png" pageId="7" pageNumber="92">Fig. 10</figureCitation>
, ↓3). The apical fork of the ejaculatory apodeme is perfectly V-shaped (
<figureCitation id="8E8F38DFFFE2966DFAA8FEDEDF90FE81" box="[1390,1436,340,365]" captionStart="FIGURES 912" captionStartId="7.[151,264,1983,2005]" captionTargetBox="[175,1392,758,1907]" captionTargetId="figure@7.[151,1436,713,1961]" captionTargetPageId="7" captionText="FIGURES 912. Dicerura barbata, male, from Sweden. 9: Ninth tergite, dorsal. 10: Gonostylus, ventral. 11: Genitalia, ventral. 12: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455513/files/figure.png" pageId="7" pageNumber="92">Fig.</figureCitation>
</paragraph>
<paragraph id="160B245AFFE2966DFF51FEF3DF90FE54" blockId="7.[151,1436,151,659]" pageId="7" pageNumber="92">
12). The elongate, parallel-sided tegmen, which is membranous for the most part, has a broadly rounded apex and a single pair of small, sclerotized processes subapicolaterally (
<figureCitation id="8E8F38DFFFE2966DFC8DFE15D993FE5B" box="[843,927,415,440]" captionStart="FIGURES 912" captionStartId="7.[151,264,1983,2005]" captionTargetBox="[175,1392,758,1907]" captionTargetId="figure@7.[151,1436,713,1961]" captionTargetPageId="7" captionText="FIGURES 912. Dicerura barbata, male, from Sweden. 9: Ninth tergite, dorsal. 10: Gonostylus, ventral. 11: Genitalia, ventral. 12: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455513/files/figure.png" pageId="7" pageNumber="92">Fig. 12</figureCitation>
, ↓4). The apex of the ninth tergite is bilobed
</paragraph>
<paragraph id="160B245AFFE2966DFF51FE49DF15FE37" blockId="7.[151,1436,151,659]" box="[151,1305,451,476]" pageId="7" pageNumber="92">
and densely covered with short, thick microtrichia, especially along the edge and on the inside (
<figureCitation id="8E8F38DFFFE2966DFB00FE49DF07FE37" box="[1222,1291,451,476]" captionStart="FIGURES 912" captionStartId="7.[151,264,1983,2005]" captionTargetBox="[175,1392,758,1907]" captionTargetId="figure@7.[151,1436,713,1961]" captionTargetPageId="7" captionText="FIGURES 912. Dicerura barbata, male, from Sweden. 9: Ninth tergite, dorsal. 10: Gonostylus, ventral. 11: Genitalia, ventral. 12: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455513/files/figure.png" pageId="7" pageNumber="92">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5EAE77D1FFE2966DFF01FE6DDBB6FDA6" pageId="7" pageNumber="92" type="discussion">
<paragraph id="160B245AFFE2966DFF01FE6DDF90FDEC" blockId="7.[151,1436,151,659]" box="[199,1436,487,512]" pageId="7" pageNumber="92">
<emphasis id="24C0F848FFE2966DFF01FE6DDB37FDEC" bold="true" box="[199,315,487,512]" pageId="7" pageNumber="92">Remarks.</emphasis>
The only specimen of
<taxonomicName id="D1B45FD9FFE2966DFD93FE62D8D9FE13" authorityName="Mamaev" authorityYear="1966" box="[597,725,487,511]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="barbata">
<emphasis id="24C0F848FFE2966DFD93FE62D8D9FE13" box="[597,725,487,511]" italics="true" pageId="7" pageNumber="92">D. barbata</emphasis>
</taxonomicName>
examined here is remarkable for the presence of a distinct,
</paragraph>
<paragraph id="160B245AFFE2966DFF51FD81DF90FDC8" blockId="7.[151,1436,151,659]" box="[151,1436,523,552]" pageId="7" pageNumber="92">
complete M1+2, a vein usually regarded as vestigial in
<taxonomicName id="D1B45FD9FFE2966DFCC6FD86D96BFDCF" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2013" box="[768,871,524,547]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="24C0F848FFE2966DFCC6FD86D96BFDCF" box="[768,871,524,547]" italics="true" pageId="7" pageNumber="92">Dicerura</emphasis>
</taxonomicName>
.
<bibRefCitation id="722559ABFFE2966DFCB0FD86DE20FDC8" box="[886,1068,523,548]" pageId="7" pageNumber="92" refId="ref11145" refString="Mamaev, B. M. (1966) New and little known Palaearctic gall midges of the tribe Porricondylini (Diptera, Cecidomyiidae). Acta Entomologica BohemoSlovaca, 63, 213 - 239 [in Russian]" type="journal article">Mamaev (1966)</bibRefCitation>
did not mention this vein in his
</paragraph>
<paragraph id="160B245AFFE2966DFF51FDB8DBB6FDA6" blockId="7.[151,1436,151,659]" box="[151,442,562,587]" pageId="7" pageNumber="92">
description of
<taxonomicName id="D1B45FD9FFE2966DFEFEFDB9DBB8FDA6" authorityName="Mamaev" authorityYear="1966" box="[312,436,562,586]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="barbata">
<emphasis id="24C0F848FFE2966DFEFEFDB9DBB8FDA6" box="[312,436,562,586]" italics="true" pageId="7" pageNumber="92">D. barbata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="5EAE77D1FFE2966DFF01FDDFDE5AFD7E" pageId="7" pageNumber="92" type="materials_examined">
<paragraph id="160B245AFFE2966DFF01FDDFDF90FD82" blockId="7.[151,1436,151,659]" box="[199,1436,597,622]" pageId="7" pageNumber="92">
<materialsCitation id="A6DC2E07FFE2966DFF01FDDFDF90FD82" ID-GBIF-Occurrence="1915497330" box="[199,1436,597,622]" collectorName="Kaltisbacken NR" country="Sweden" location="Sweden" pageId="7" pageNumber="92" specimenCount="1">
<emphasis id="24C0F848FFE2966DFF01FDDFDBBDFD82" bold="true" box="[199,433,597,622]" pageId="7" pageNumber="92">Material examined.</emphasis>
<collectingCountry id="6EA364CAFFE2966DFE7EFDDFD819FD82" box="[440,533,597,622]" name="Sweden" pageId="7" pageNumber="92">Sweden</collectingCountry>
: male, Lule Lappmark, Jokkmokk,
<collectorName id="BB41418CFFE2966DFC63FDDFDE64FD82" box="[933,1128,597,622]" pageId="7" pageNumber="92">Kaltisbäcken NR</collectorName>
, herb-rich old-growth taiga
</materialsCitation>
</paragraph>
<paragraph id="160B245AFFE2966DFF51FDF1DE5AFD7E" blockId="7.[151,1436,151,659]" box="[151,1110,634,659]" pageId="7" pageNumber="92">
near stream,
<date id="620A029AFFE2966DFEE1FDF0DBB6FD7F" box="[295,442,634,659]" pageId="7" pageNumber="92" value="2016-07-10">10 July 2016</date>
, aspirator, M. Jaschhof (specimen no. CEC
<quantity id="D14C89BFFFE2966DFC66FDF0D9F8FD7F" box="[928,1012,634,659]" metricMagnitude="1" metricUnit="m" metricValue="3.5179" pageId="7" pageNumber="92" unit="in" value="1385.0">1385 in</quantity>
NHRS).
</paragraph>
</subSubSection>
</treatment>
</document>