306 lines
39 KiB
XML
306 lines
39 KiB
XML
<document ID-DOI="10.3897/zookeys.12.167" ID-GBIF-Dataset="3026c846-a607-4826-b2a9-8f7a531baec5" ID-GBIF-Taxon="163971731" ID-ISSN="1313–2970" ID-Zenodo-Dep="576460" ID-ZooBank="B32EF8D5-C3C6-4C8C-8845-284125448425" LSID="urn:lsid:zoobank.org:act:5137C5B6-4F58-4BCC-A746-21705E195066" approvalRequired="25" approvalRequired_for_textStreams="25" checkinTime="1587995767728" checkinUser="plazi" docAuthor="Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & VandenSpiegel, Didier" docDate="2009" docId="936787D7FFB5FFBF8ABB405BFE5DFDA8" docLanguage="en" docName="ZK_article_1994.pdf" docOrigin="ZooKeys 12 (12)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Eutrichodesmus regularis Golovatch, Geoffroy, Mauries & VandenSpiegel 2009, sp. n." docType="treatment" docUuid="5137C5B6-4F58-4BCC-A746-21705E195066" docUuidSource="ZooBank" docVersion="3" lastPageId="11" lastPageNumber="12" masterDocId="6F5EFFAFFFB3FFB48A374557FFC6FF89" masterDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species" masterLastPageNumber="46" masterPageNumber="1" pageId="6" pageNumber="7" updateTime="1643565021146" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei</mods:namePart>
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<mods:affiliation>Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Ecologie & Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, Brunoy, France & D 362 Dc 0 - & CC</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mauriès, Jean-Paul</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Systématique et Evolution, Section Arthropodes, Paris, France & - & - 42 F & D 01 - 28 Ad- 43 F</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>VandenSpiegel, Didier</mods:namePart>
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<mods:affiliation>Musée Royal de l’Afrique centrale, Tervuren, Belgium & A & EE & F- 04802 E 68 Cb</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-06-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>12</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>12</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>46</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.12.167</mods:identifier>
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<mods:identifier type="GBIF-Dataset">3026c846-a607-4826-b2a9-8f7a531baec5</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576460</mods:identifier>
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||
<mods:identifier type="ZooBank">B32EF8D5-C3C6-4C8C-8845-284125448425</mods:identifier>
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||
</mods:mods>
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<treatment ID-GBIF-Taxon="163971731" LSID="urn:lsid:zoobank.org:act:5137C5B6-4F58-4BCC-A746-21705E195066" httpUri="http://treatment.plazi.org/id/936787D7FFB5FFBF8ABB405BFE5DFDA8" lastPageId="11" lastPageNumber="12" pageId="6" pageNumber="7">
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<subSubSection box="[140,1090,1292,1319]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph blockId="6.[140,1090,1292,1390]" box="[140,1090,1292,1319]" pageId="6" pageNumber="7">
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<heading box="[140,1090,1292,1319]" level="2" pageId="6" pageNumber="7">
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||
<emphasis bold="true" box="[140,1090,1292,1319]" pageId="6" pageNumber="7">
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||
<taxonomicName authority="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityName="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityYear="2009" box="[140,1013,1292,1319]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="regularis" status="sp. n.">
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<emphasis bold="true" box="[140,438,1292,1319]" italics="true" pageId="6" pageNumber="7">
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<emphasis box="[140,328,1292,1319]" inLineHeading="true" pageId="6" pageNumber="7" reason="1">Eutrichodesmus</emphasis>
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regularis
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</emphasis>
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Golovatch, Geoffroy, Mauriès & VandenSpiegel
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1025,1090,1293,1319]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="6" pageNumber="7" type="description">
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||
<paragraph blockId="6.[140,1090,1292,1390]" box="[140,959,1328,1355]" pageId="6" pageNumber="7">
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<uri box="[140,959,1328,1355]" pageId="6" pageNumber="7">
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urn:lsid:zoobank.org:act:
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<uuid box="[419,959,1328,1355]" pageId="6" pageNumber="7">5137C5B6-4F58-4BCC-A746-21705E195066</uuid>
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</uri>
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</paragraph>
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<paragraph blockId="6.[140,1090,1292,1390]" box="[140,231,1363,1390]" pageId="6" pageNumber="7">
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<figureCitation box="[140,231,1363,1390]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStart-3="Figure 7" captionStartId-0="7.[140,212,1622,1644]" captionStartId-1="8.[140,212,1498,1520]" captionStartId-2="9.[140,212,1558,1580]" captionStartId-3="10.[140,212,1622,1644]" captionTargetBox-0="[140,1108,623,1602]" captionTargetBox-1="[140,1108,165,1478]" captionTargetBox-2="[140,1108,478,1538]" captionTargetBox-3="[140,1108,308,1602]" captionTargetId-0="figure@7.[140,1108,622,1602]" captionTargetId-1="figure@8.[140,1108,165,1478]" captionTargetId-2="figure@9.[140,1108,478,1538]" captionTargetId-3="figure@10.[140,1108,308,1603]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Figure 4. Eutrichodesmus regularis sp. n., ♁ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." captionText-1="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." captionText-2="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." captionText-3="Figure 7. Eutrichodesmus regularis sp. n., ♁ paratypes; A-F, gonopods, frontoventral, submesal, mesal, mesal, mesal and lateral views, respectively. – Scale bars: A-C, 0.1 mm; D, 0.05 mm; E, F, 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3769977" figureDoi-1="http://doi.org/10.5281/zenodo.3769979" figureDoi-2="http://doi.org/10.5281/zenodo.3769981" figureDoi-3="http://doi.org/10.5281/zenodo.3769983" httpUri-0="https://zenodo.org/record/3769977/files/figure.png" httpUri-1="https://zenodo.org/record/3769979/files/figure.png" httpUri-2="https://zenodo.org/record/3769981/files/figure.png" httpUri-3="https://zenodo.org/record/3769983/files/figure.png" pageId="6" pageNumber="7">Figs 4-7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="7" type="materials_examined">
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<paragraph blockId="6.[140,1108,1433,1636]" pageId="6" pageNumber="7">
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<emphasis bold="true" box="[140,311,1433,1460]" pageId="6" pageNumber="7">
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<emphasis box="[140,197,1434,1460]" inLineHeading="true" pageId="6" pageNumber="7" reason="1">Type</emphasis>
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material:
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2610648304" collectingDate="2003-08-11" collectorName="L. Deharveng & Bedos" country="Vietnam" location="Hang Ta Phin Cave" municipality="Sa Pa" pageId="6" pageNumber="7" specimenCount="5" specimenCount-female="3" stateProvince="Lao Cai" typeStatus="holotype">
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<collectingCountry box="[321,420,1434,1460]" name="Vietnam" pageId="6" pageNumber="7">Vietnam</collectingCountry>
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,
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<collectingRegion box="[435,595,1434,1460]" country="Vietnam" name="Lao Cai" pageId="6" pageNumber="7">Lao Cai Prov.</collectingRegion>
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,
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<collectingMunicipality box="[610,675,1434,1460]" pageId="6" pageNumber="7">Sa Pa</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:936787D7FFB5FFBF8ABB405BFE5DFDA8:1E11601AFFB5FFB2888540CDFC5FFA3D" box="[690,921,1434,1460]" country="Vietnam" municipality="Sa Pa" name="Hang Ta Phin Cave" pageId="6" pageNumber="7" stateProvince="Lao Cai">Hang Ta Phin Cave</location>
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, hand collected,
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<date box="[140,291,1469,1495]" pageId="6" pageNumber="7" value="2003-08-11">
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<collectingDate box="[140,291,1469,1495]" pageId="6" pageNumber="7" value="2003-08-11">11.VIII.2003</collectingDate>
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</date>
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, leg.
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<collectorName box="[353,507,1469,1496]" pageId="6" pageNumber="7">L. Deharveng</collectorName>
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and
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<collectorName box="[569,635,1469,1495]" pageId="6" pageNumber="7">Bedos</collectorName>
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(Vn0308-009),
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<typeStatus box="[825,922,1469,1495]" pageId="6" pageNumber="7">holotype</typeStatus>
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♁ (MNHN JC 318),
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<typeStatus box="[205,312,1505,1531]" pageId="6" pageNumber="7">paratypes</typeStatus>
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: 1 ♁,
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<specimenCount box="[375,409,1504,1531]" pageId="6" pageNumber="7" type="female">2 ♀</specimenCount>
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,
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<specimenCount box="[429,523,1504,1530]" pageId="6" pageNumber="7" type="generic">1 subad.</specimenCount>
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,
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<specimenCount box="[533,592,1504,1530]" pageId="6" pageNumber="7" type="generic">1 juv.</specimenCount>
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(MNHN JC 318),
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<specimenCount box="[807,842,1504,1531]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
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(ZMUM), 1 ♁ (SEM)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="7" type="etymology">
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<paragraph blockId="6.[140,1108,1433,1636]" pageId="6" pageNumber="7">
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<emphasis bold="true" box="[185,263,1539,1565]" pageId="6" pageNumber="7">
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<emphasis box="[185,258,1539,1565]" inLineHeading="true" pageId="6" pageNumber="7" reason="1">Name</emphasis>
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:
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</emphasis>
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To emphasize the highly regular, isostictic (i.e. regular rows not only transversely, but also longitudinally) and almost undifferentiated pattern of metatergal tuberculation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="diagnosis">
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<paragraph blockId="7.[140,1107,166,581]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[185,312,166,192]" pageId="7" pageNumber="8">
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<emphasis box="[185,307,166,192]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">Diagnosis</emphasis>
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:
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</emphasis>
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Differs from congeners by the perfect volvation, coupled with faint metatergal lobulations, the especially regular, nearly undifferentiated and isostictic pattern of metatergal tuberculation, the peculiar, phylloid tergal setae and a few minor details of gonopod structure (in particular, the shape of the telopodite and distofemoral process).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8" type="description">
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<paragraph blockId="7.[140,1107,166,581]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[185,335,342,368]" pageId="7" pageNumber="8">
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<emphasis box="[185,330,342,368]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">Description</emphasis>
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:
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</emphasis>
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Length of adults of both sexes ca 9.0-10.0 mm, width
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<quantity box="[963,1103,343,369]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="7" pageNumber="8" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7-1.8 mm</quantity>
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, body broadest at segment 3 or 4.
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<typeStatus box="[515,622,378,404]" pageId="7" pageNumber="8">Holotype</typeStatus>
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ca
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<quantity box="[660,743,378,404]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="10.0">10 mm</quantity>
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long and
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<quantity box="[856,945,378,404]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="8" unit="mm" value="1.7">1.7 mm</quantity>
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wide. Coloration uniformly pallid, shown pinkish because of a photographic artifact (
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<figureCitation box="[961,1029,413,439]" captionStart="Figure 4" captionStartId="7.[140,212,1622,1644]" captionTargetBox="[140,1108,623,1602]" captionTargetId="figure@7.[140,1108,622,1602]" captionTargetPageId="7" captionText="Figure 4. Eutrichodesmus regularis sp. n., ♁ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi="http://doi.org/10.5281/zenodo.3769977" httpUri="https://zenodo.org/record/3769977/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
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).
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</paragraph>
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<paragraph blockId="7.[140,1107,166,581]" lastBlockId="10.[140,1108,167,264]" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
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Adults with 20 segments, pattern of conglobation typical of “doratodesmids” (
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<figureCitation box="[1034,1103,448,474]" captionStart="Figure 4" captionStartId="7.[140,212,1622,1644]" captionTargetBox="[140,1108,623,1602]" captionTargetId="figure@7.[140,1108,622,1602]" captionTargetPageId="7" captionText="Figure 4. Eutrichodesmus regularis sp. n., ♁ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi="http://doi.org/10.5281/zenodo.3769977" httpUri="https://zenodo.org/record/3769977/files/figure.png" pageId="7" pageNumber="8">Figs 4</figureCitation>
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,
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<figureCitation box="[140,176,483,510]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="7" pageNumber="8">5A</figureCitation>
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). Head, antennae (
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||
<figureCitation box="[388,473,483,510]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="7" pageNumber="8">Fig. 5E</figureCitation>
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), tegument (
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||
<figureCitation box="[616,683,483,510]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="7" pageNumber="8">Figs 6</figureCitation>
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A-C), sterna (
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<figureCitation box="[837,923,483,510]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="7" pageNumber="8">Fig. 6E</figureCitation>
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), gonopod aperture (
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<figureCitation box="[198,278,519,546]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="7" pageNumber="8">Fig. 6F</figureCitation>
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) and many other characters (
|
||
<figureCitation box="[595,675,518,545]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="7" pageNumber="8">Figs 5F</figureCitation>
|
||
,
|
||
<figureCitation box="[685,723,519,546]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="7" pageNumber="8">6D</figureCitation>
|
||
) much as in
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[863,991,519,545]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="distinctus" status="sp. n.">
|
||
<emphasis box="[863,991,519,545]" italics="true" pageId="7" pageNumber="8">E. distinctus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[997,1057,519,545]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
|
||
, but collum more regularly tuberculate and slightly flattened mid-dorsally (
|
||
<figureCitation box="[908,991,554,581]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="7" pageNumber="8">Fig. 5B</figureCitation>
|
||
). Metater- ga behind collum with three transverse, very regular and isostictic rows of subequallysized, rounded, flat, nearly undifferentiated tubercles (
|
||
<figureCitation box="[749,818,202,229]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="9" pageNumber="10">Figs 5</figureCitation>
|
||
A-D), each crowned with a short, 2-segmented, phylloid seta (
|
||
<figureCitation box="[555,646,237,264]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="9" pageNumber="10">Fig. 6C</figureCitation>
|
||
). Paraterga directed ventrolaterad, rather broad, slightly surpassing level of venter, slightly interrupting general outline of convex dorsum (
|
||
<figureCitation box="[239,360,307,334]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="9" pageNumber="10">Figs 5A, C</figureCitation>
|
||
,
|
||
<figureCitation box="[372,411,307,334]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="9" pageNumber="10">6D</figureCitation>
|
||
); paraterga 2 strongly enlarged, each margin nearly entire, with only very faint traces of a series of small lobulations anterolaterally (
|
||
<figureCitation box="[902,972,343,369]" captionStart="Figure 4" captionStartId="7.[140,212,1622,1644]" captionTargetBox="[140,1108,623,1602]" captionTargetId="figure@7.[140,1108,622,1602]" captionTargetPageId="7" captionText="Figure 4. Eutrichodesmus regularis sp. n., ♁ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi="http://doi.org/10.5281/zenodo.3769977" httpUri="https://zenodo.org/record/3769977/files/figure.png" pageId="9" pageNumber="10">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation box="[987,1093,342,369]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="9" pageNumber="10">5A, B, D</figureCitation>
|
||
). Following paraterga broadly rounded and slightly 3- or 4-lobate laterally and at least distinctly bilobate caudolaterally (
|
||
<figureCitation box="[478,636,413,440]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="9" pageNumber="10">Figs 5A, C, D</figureCitation>
|
||
,
|
||
<figureCitation box="[647,680,413,440]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="9" pageNumber="10">6F</figureCitation>
|
||
). Limbus very faintly crenulate, almost hidden by nearby abundant microvilli (
|
||
<figureCitation box="[577,664,167,194]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
|
||
). Pore formula normal, ozopores poorly visible, located dorsally near tip of ventrocaudal lobulation (
|
||
<figureCitation box="[798,924,202,229]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="10" pageNumber="11">Figs 5C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769977" ID-Zenodo-Dep="3769977" httpUri="https://zenodo.org/record/3769977/files/figure.png" pageId="7" pageNumber="8" startId="7.[140,212,1622,1644]" targetBox="[140,1108,623,1602]" targetPageId="7">
|
||
<paragraph blockId="7.[140,1107,1622,1676]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[140,240,1622,1644]" pageId="7" pageNumber="8">Figure 4.</emphasis>
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[247,466,1622,1643]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="regularis" status="sp. n.">
|
||
<emphasis box="[247,466,1622,1643]" italics="true" pageId="7" pageNumber="8">Eutrichodesmus regularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[473,526,1623,1644]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
|
||
, ♁ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769979" ID-Zenodo-Dep="3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="8" pageNumber="9" startId="8.[140,212,1498,1520]" targetBox="[140,1108,165,1478]" targetPageId="8">
|
||
<paragraph blockId="8.[140,1108,1498,1584]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[140,237,1498,1520]" pageId="8" pageNumber="9">Figure 5.</emphasis>
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[242,459,1498,1519]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="regularis" status="sp. n.">
|
||
<emphasis box="[242,459,1498,1519]" italics="true" pageId="8" pageNumber="9">Eutrichodesmus regularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[464,515,1498,1519]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
|
||
, ♁ paratype;
|
||
<emphasis bold="true" box="[638,657,1498,1520]" pageId="8" pageNumber="9">A</emphasis>
|
||
, habitus, lateral view;
|
||
<emphasis bold="true" box="[860,902,1498,1520]" pageId="8" pageNumber="9">B, E</emphasis>
|
||
, anterior part of body, frontodorsal and ventral views, respectively;
|
||
<emphasis bold="true" box="[557,576,1530,1552]" pageId="8" pageNumber="9">C</emphasis>
|
||
, midbody segments, dorsal view;
|
||
<emphasis bold="true" box="[896,939,1530,1552]" pageId="8" pageNumber="9">D, F</emphasis>
|
||
, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769981" ID-Zenodo-Dep="3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="9" pageNumber="10" startId="9.[140,212,1558,1580]" targetBox="[140,1108,478,1538]" targetPageId="9">
|
||
<paragraph blockId="9.[140,1113,1558,1676]" pageId="9" pageNumber="10">
|
||
<emphasis bold="true" box="[140,238,1558,1580]" pageId="9" pageNumber="10">Figure 6.</emphasis>
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[243,461,1558,1579]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="regularis" status="sp. n.">
|
||
<emphasis box="[243,461,1558,1579]" italics="true" pageId="9" pageNumber="10">Eutrichodesmus regularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[467,518,1559,1580]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
, ♁ paratype;
|
||
<emphasis bold="true" box="[643,662,1558,1580]" pageId="9" pageNumber="10">A</emphasis>
|
||
, texture of tegument, dorsal view;
|
||
<emphasis bold="true" box="[982,999,1558,1580]" pageId="9" pageNumber="10">B</emphasis>
|
||
, metatergal tubercle, dorsal view;
|
||
<emphasis bold="true" box="[338,357,1590,1612]" pageId="9" pageNumber="10">C</emphasis>
|
||
, metatergal seta, dorsal view;
|
||
<emphasis bold="true" box="[628,647,1590,1612]" pageId="9" pageNumber="10">D</emphasis>
|
||
, cross-section of a midbody segment, caudal view;
|
||
<emphasis bold="true" box="[140,155,1622,1644]" pageId="9" pageNumber="10">E</emphasis>
|
||
, midbody legs, caudal view;
|
||
<emphasis bold="true" box="[428,442,1622,1644]" pageId="9" pageNumber="10">F</emphasis>
|
||
, segments 7 and 8, ventral view;
|
||
<emphasis bold="true" box="[759,779,1622,1644]" pageId="9" pageNumber="10">G</emphasis>
|
||
, left gonopod
|
||
<emphasis box="[918,976,1623,1644]" italics="true" pageId="9" pageNumber="10">in situ</emphasis>
|
||
, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="10.[140,1108,167,264]" box="[185,975,237,264]" pageId="10" pageNumber="11">
|
||
Legs relatively long, slightly surpassing edge of paraterga (
|
||
<figureCitation box="[835,962,237,264]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="10" pageNumber="11">Figs 6D, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769983" ID-Zenodo-Dep="3769983" httpUri="https://zenodo.org/record/3769983/files/figure.png" pageId="10" pageNumber="11" startId="10.[140,212,1622,1644]" targetBox="[140,1108,308,1602]" targetPageId="10">
|
||
<paragraph blockId="10.[140,1108,1622,1676]" pageId="10" pageNumber="11">
|
||
<emphasis bold="true" box="[140,239,1622,1644]" pageId="10" pageNumber="11">Figure 7.</emphasis>
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[246,465,1622,1643]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="regularis" status="sp. n.">
|
||
<emphasis box="[246,465,1622,1643]" italics="true" pageId="10" pageNumber="11">Eutrichodesmus regularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[472,525,1623,1644]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
, ♁ paratypes;
|
||
<emphasis bold="true" box="[661,702,1622,1644]" pageId="10" pageNumber="11">A-F</emphasis>
|
||
, gonopods, frontoventral, submesal, mesal, mesal, mesal and lateral views, respectively. – Scale bars: A-C, 0.1 mm; D, 0.05 mm; E, F, 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="11.[140,1108,167,545]" pageId="11" pageNumber="12">
|
||
Gonopods (
|
||
<figureCitation box="[317,436,167,193]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="11" pageNumber="12">Figs 6F, G</figureCitation>
|
||
,
|
||
<figureCitation box="[449,461,167,193]" captionStart="Figure 7" captionStartId="10.[140,212,1622,1644]" captionTargetBox="[140,1108,308,1602]" captionTargetId="figure@10.[140,1108,308,1603]" captionTargetPageId="10" captionText="Figure 7. Eutrichodesmus regularis sp. n., ♁ paratypes; A-F, gonopods, frontoventral, submesal, mesal, mesal, mesal and lateral views, respectively. – Scale bars: A-C, 0.1 mm; D, 0.05 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769983" httpUri="https://zenodo.org/record/3769983/files/figure.png" pageId="11" pageNumber="12">7</figureCitation>
|
||
A-E) relatively complex. Coxae subquadrate, large, microtuberculate, nearly bare, with only a few, mostly short setae on and near base of a small ventro-apical lobe. Telopodite considerably longer than coxite, slightly stouter and setose in its basal half, with an inconspicuous, digitiform, simple, lateral, distally papillate, distofemoral process (dp) at about midway, more distally with another simple outgrowth with its base marking recurvature point of a relatively short seminal groove; acropodite (= solenomere) relatively stout, lamelliform, slightly curved along main axis, with two small denticles apically and a conspicuous pilose-spinulate pulvillus near base (
|
||
<figureCitation box="[937,1054,413,440]" captionStart="Figure 7" captionStartId="10.[140,212,1622,1644]" captionTargetBox="[140,1108,308,1602]" captionTargetId="figure@10.[140,1108,308,1603]" captionTargetPageId="10" captionText="Figure 7. Eutrichodesmus regularis sp. n., ♁ paratypes; A-F, gonopods, frontoventral, submesal, mesal, mesal, mesal and lateral views, respectively. – Scale bars: A-C, 0.1 mm; D, 0.05 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769983" httpUri="https://zenodo.org/record/3769983/files/figure.png" pageId="11" pageNumber="12">Figs 7E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="12" type="discussion">
|
||
<paragraph blockId="11.[140,1108,167,545]" pageId="11" pageNumber="12">
|
||
<emphasis bold="true" box="[185,298,448,474]" pageId="11" pageNumber="12">
|
||
<emphasis box="[185,293,448,474]" inLineHeading="true" pageId="11" pageNumber="12" reason="1">Remarks</emphasis>
|
||
:
|
||
</emphasis>
|
||
This pallid species shows highly peculiar, phylloid but very short, tergal setae and undifferentiated metatergal tubercles. It is also a typical “doratodesmid”, possibly another troglobite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |