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<document ID-CLB-Dataset="9646" ID-DOI="http://dx.doi.org/10.3897/zookeys.984.53587" ID-GBIF-Dataset="9d291585-91c0-45e5-94d9-ee28795291ba" ID-PMC="PMC7658186" ID-Pensoft-Pub="1313-2970-984-83" ID-Pensoft-UUID="1448149CA44B51ED9EE57DC3A924C430" ID-PubMed="33223870" ID-ZooBank="D8B945AE620949578BB1D96DDBB5AA44" ModsDocID="1313-2970-984-83" checkinTime="1604561368050" checkinUser="pensoft" docAuthor="Irwin, Michael E., Winterton, Shaun L. &amp; Metz, Mark A." docDate="2020" docId="9357196FBE265A868402E475EBF46982" docLanguage="en" docName="ZooKeys 984: 83-132" docOrigin="ZooKeys 984" docSource="http://dx.doi.org/10.3897/zookeys.984.53587" docTitle="Jeanchazeauia nubilosus Irwin &amp; Winterton &amp; Metz 2020, sp. nov." docType="treatment" docUuid="F826CA19-F56D-4391-A6CC-EAA198ECE40C" docUuidSource="ZooBank" docVersion="6" id="1448149CA44B51ED9EE57DC3A924C430" lastPageNumber="83" masterDocId="1448149CA44B51ED9EE57DC3A924C430" masterDocTitle="New stiletto flies from New Caledonia (Therevidae, Agapophytinae)" masterLastPageNumber="132" masterPageNumber="83" pageNumber="83" updateTime="1732742644444" updateUser="ExternalLinkService">
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<mods:title id="DA4A1F71F97618717DD9C0970F721314">New stiletto flies from New Caledonia (Therevidae, Agapophytinae)</mods:title>
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<mods:name id="7F564B743210ED284F444ECB2327FDC0" type="personal">
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<mods:namePart id="2CE843DBC0FF49549B5336257605455F">Irwin, Michael E.</mods:namePart>
<mods:affiliation id="EED371EACA3E13B087161153CF450EC3">Emeritus, University of Illinois, Urbana-Champaign, Illinois, USA &amp; University of Arizona, Tucson, USA</mods:affiliation>
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<mods:namePart id="75A759F9041E975728F462397D3254AC">Winterton, Shaun L.</mods:namePart>
<mods:affiliation id="28E89E5728DA3DAC18EDD447C67F9C27">California State Collection of Arthropods, California Department of Food &amp; Agriculture, Sacramento, California, USA</mods:affiliation>
<mods:nameIdentifier id="3FC36AEB3A9A5E55D8C86FA74CCFBCBA" type="email">wintertonshaun@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F57BCEBD4C7C439E18A87EFCEDCAAD26">Metz, Mark A.</mods:namePart>
<mods:affiliation id="0826809F953C83B537BEA25E278E2152">USDA, ARS, Systematic Entomology Laboratory, Beltsville, Maryland, USA</mods:affiliation>
<mods:nameIdentifier id="018AC32EF1FE4AC9A7EEDBEFE97FB849" type="orcid">https://orcid.org/0000-0002-3535-535X</mods:nameIdentifier>
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<mods:date id="198B472B522A34BA09E882CDB3361DBF">2020</mods:date>
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<treatment id="9357196FBE265A868402E475EBF46982" ID-GBIF-Taxon="168756111" LSID="urn:lsid:zoobank.org:act:F826CA19-F56D-4391-A6CC-EAA198ECE40C" httpUri="http://treatment.plazi.org/id/9357196FBE265A868402E475EBF46982" lastPageNumber="83" pageId="0" pageNumber="83" scope_family="Therevidae" scope_subFamily="Agapophytinae">
<subSubSection id="267AD40B9BD8DA0A1A7EE9E896023938" pageId="0" pageNumber="83" type="nomenclature">
<paragraph id="7DBE73FD20EB5C18E7D4A35C49A95EB3" pageId="0" pageNumber="83">
<taxonomicName id="2904A7336B6057C1A78E9CA21C1F6DAC" LSID="9357196F-BE26-5A86-8402-E475EBF46982" authority="Irwin &amp; Winterton &amp; Metz, 2020" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" family="Agapophytinae" genus="Jeanchazeauia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Jeanchazeauia nubilosus" order="Therevidae" pageId="0" pageNumber="83" rank="species" species="nubilosus" status="sp. nov.">Jeanchazeauia nubilosus</taxonomicName>
<taxonomicNameLabel id="E238BEA2D927611001C8AA935E6B42FA" pageId="0" pageNumber="83">sp. nov.</taxonomicNameLabel>
<figureCitation id="B677D8ECB03A228E36277212F1ACE29A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Jeanchazeauia nubilosus sp. nov., female habitus (MEI 030203) (artistic rendering by J. Marie Metz)." figureDoi="10.3897/zookeys.984.53587.figure3" httpUri="https://binary.pensoft.net/fig/471319" pageId="0" pageNumber="83">Figs 3</figureCitation>
<figureCitation id="B99888E9B80AFA3D83DE7FC274E32722" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scutum and scutellum of Calophytus gen. nov. and Jeanchazeauia gen. nov. A C. schlingeri sp. nov. B C. grandiosus sp. nov. C C. matilei sp. nov. D C. webbi sp. nov. E J. nubilosus sp. nov. (figures not to scale) (drawings by J. Marie Metz)." figureDoi="10.3897/zookeys.984.53587.figure6" httpUri="https://binary.pensoft.net/fig/471322" pageId="0" pageNumber="83">, 6E</figureCitation>
<figureCitation id="6C57E50E908C2913A1A6E96B5E3E93FB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Wings of Calophytus gen. nov. and Jeanchazeauia gen. nov. A C. chazeaui sp. nov. B C. schlingeri sp. nov. C C. grandiosus sp. nov. D C. matilei sp. nov. E C. webbi sp. nov. F C. monteithi sp. nov. G J. amoa sp. nov. H J. nubilosus sp. nov. (female) (figures not to scale)." figureDoi="10.3897/zookeys.984.53587.figure7" httpUri="https://binary.pensoft.net/fig/471313" pageId="0" pageNumber="83">, 7H</figureCitation>
<figureCitation id="320C81693511798B424DAFA50988B7EB" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female abdominal tergites VI-VIII A Calophytus schlingeri sp. nov. B Jeanchazeauia amoa sp. nov. C Jeanchazeauia nubilosus sp. nov. D Jeanchazeauia rufinatus sp. nov. Scale line: 0.2 mm." figureDoi="10.3897/zookeys.984.53587.figure23" httpUri="https://binary.pensoft.net/fig/471333" pageId="0" pageNumber="83">, 23C</figureCitation>
<figureCitation id="E7FDFACBE4699558C2F14DFB6BA481C7" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Jeanchazeauia nubilosus sp. nov. A adult male (MEI 030207), lateral view B same, oblique view C adult female (MEI 030203), lateral view D same, oblique view. Body length: male: 8.8 mm; female: 9.2 mm." figureDoi="10.3897/zookeys.984.53587.figure25" httpUri="https://binary.pensoft.net/fig/471335" pageId="0" pageNumber="83">, 25</figureCitation>
<figureCitation id="A44C3C236B0D3D6283A017735345ADEF" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Jeanchazeauia nubilosus sp. nov. A cleared male genitalia A aedeagus, lateral view B epandrium C gonocoxites, ventral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.984.53587.figure26" httpUri="https://binary.pensoft.net/fig/471336" pageId="0" pageNumber="83">, 26</figureCitation>
<figureCitation id="6D16784D0F3DA34EF20A925852D3055C" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Jeanchazeauia spp. genitalia. Jeanchazeauia nubilosus sp. nov. A epandrium B gonocoxites, ventral view C same, lateral view D aedeagus, lateral view E same, dorsal view F female genitalia, lateral view, internal structures shown. Jeanchazeauia rufinatus sp. nov. G female tergites VII and VIII, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.984.53587.figure27" httpUri="https://binary.pensoft.net/fig/471337" pageId="0" pageNumber="83">, 27A-F</figureCitation>
<figureCitation id="788D2DB3B9241104D230D53C2DCFA86A" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Distribution of Jeanchazeauia spp. collecting records throughout New Caledonia. Provinces are labelled and ecoregions delineated by colour." figureDoi="10.3897/zookeys.984.53587.figure30" httpUri="https://binary.pensoft.net/fig/471340" pageId="0" pageNumber="83">, 30</figureCitation>
<figureCitation id="0F2D0C65537A5FEA1D4A5E8AEA3008AC" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Natural habitats of New Caledonian agapophytine stiletto flies A Riviere Bleue Reserve, sclerophyll forest and maquis scrub, habitat of C. chazeaui sp. nov., C. grandiosus sp. nov., C. schlingeri sp. nov., and J. nubilosus sp. nov. B Pindai Peninsula, tropical dry forest, habitat of C. webbi sp. nov. C Mount Dzumac, rainforest, habitat of J. nubilosus sp. nov. D Mount Dore, rainforest and dry sclerophyll forest E Mount Dore, Malaise trap in flyway (photographs by Michael E. Irwin)." figureDoi="10.3897/zookeys.984.53587.figure31" httpUri="https://binary.pensoft.net/fig/471341" pageId="0" pageNumber="83">, 31A, C</figureCitation>
<figureCitation id="794E42A4CB2C7EDEE353FF28747D8671" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Natural habitats of New Caledonian agapophytine stiletto flies A Mount Aoupinie, rainforest, habitat of C. monteithi sp. nov. B Mount Ningua, rainforest, habitat of J. nubilosus sp. nov. C Mount Mandjelia, habitat of C. grandiosus sp. nov. (photographs by Michael E. Irwin)." figureDoi="10.3897/zookeys.984.53587.figure32" httpUri="https://binary.pensoft.net/fig/471342" pageId="0" pageNumber="83">, 32B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="79B586F687066B14F7EB37FC1BA2DFBF" pageId="0" pageNumber="83" type="diagnosis">
<paragraph id="B6E93239AA698D64D785D95B07CABFFE" pageId="0" pageNumber="83">Diagnosis.</paragraph>
<paragraph id="1D63DF5BF88562B61E12CAA335BBC886" pageId="0" pageNumber="83">Black macrosetae on legs; frons dark, sparsely pubescent; female abdomen dark with white bands; female wing with strong banding, irregular basally, wing apex dark; male wing with uniform infuscation.</paragraph>
</subSubSection>
<subSubSection id="B4ED51A4B7AAB3891C979F9C6E745BD1" pageId="0" pageNumber="83" type="description">
<paragraph id="9BC1D5B8869363D79765085829124E0A" pageId="0" pageNumber="83">Description.</paragraph>
<paragraph id="DA35ECBCAFD611C27AE337A843425F28" pageId="0" pageNumber="83">
Length 8.3-9.6 mm.
<emphasis id="8A10D44829D44ECF611C136802B70BE9" italics="true" pageId="0" pageNumber="83">Head</emphasis>
. Black to brown; mostly silver pubescent, upper frons sparsely pubescent, admixed with black setae, longer in male, a few setae dorsolateral to antennal bases, eyes separated by width of ocellar tubercle in both sexes; parafacial black with silver pubescence. Occiput black, matte pubescent, more bronze coloured along postocular ridge, silver laterally and ventrally. Postocular macrosetae mostly black, in a single row in both sexes, multiple irregular rows laterally, replaced by white finer setae laterally and on gena. Scape
<normalizedToken id="660F12A55ED6FE9D7F23B349AC3276FA" originalValue="½×">1/2x</normalizedToken>
head length; dark brown, sparsely silver pubescent; long, black, fine filiform setose except medial surface bare of setae. Basal flagellomere
<normalizedToken id="F1DEA8E7A41A0CC6A541B5E5A60732DF" originalValue="½×">1/2x</normalizedToken>
length of scape, elongate and gradually tapering to a blunt point apically; sparsely silver pubescent with short, fine, black setae around base. Second flagellomere slightly conical, apex narrower than base; &lt;1/10
<normalizedToken id="D9B6106DA1138A3A7DD0F6642960B0D4" originalValue="×">x</normalizedToken>
length of basal flagellomere. Third flagellomere subequal in length to second, conical. Style small, spiculate. Palpus one segmented, cylindrical, dark brown basally, light brown apically; silver pubescent admixed with white, setae basally, brown apically. Mouthparts brown with brown setae.
<emphasis id="5CCB69171BCB3CA416826136E071D442" italics="true" pageId="0" pageNumber="83">Thorax</emphasis>
. Dark brown-black with black and silver pubescence; scutum brown pubescent with anterior postpronotal lobe and notopleuron sparsely silver pubescent; scutellum with black matte pubescence anteriorly and medially, bronze pubescent along posterior margin; pleuron silver pubescent with two brown glabrous vertical bands passing from the notopleuron at the prothoracic spiracle posteroventrally through the anterior katepisternum, and from the wing base posteroventrally through the meron; macrosetae black (np: 3, sa: 1, pa: 1, dc: 1, sc: 1); scutum with short, black setae; postpronotal lobe, cervical sclerite, proepisternum, and lateral prosternum with white setae; katatergite with mostly white setae, both sides with one dark brown seta.
<emphasis id="E464F59372F2BB89E89D295234D18F29" italics="true" pageId="0" pageNumber="83">Legs</emphasis>
. Dark brown; coxae with dense silver pubescence, otherwise sparsely pubescent and admixed with short, black setae; macrosetae mostly black with a few light brown macrosetae on hind tibia. Coxae with white setae; forecoxa with two, midcoxa with three, and hind coxa with five black, anteroventral, marginal macrosetae; hind coxa lacking lateral seta. Posterior surface of femora with long, white setae admixed with short, black setae. Hind femur with one or two subapical anteroventral macrosetae; series of minute posteroventral macrosetae barely evident along middle of femur.
<emphasis id="2284B8ABCCB426166D8C38B9F9BEC4C1" italics="true" pageId="0" pageNumber="83">Wing</emphasis>
. Male wing membrane with entirely grey infuscation, darker subapically near anterior margin and apices of cells br and bm; membrane completely covered with microtrichia. Female wing strongly banded, with extensive hyaline areas; pterostigma brown; venation brown, cell m3 widely open at wing margin. Haltere entirely dark brown, knob slightly lighter ventrally.
<emphasis id="CA0E54A6BD7FCAE29F9B02C37E5D515E" italics="true" pageId="0" pageNumber="83">Abdomen</emphasis>
. Dark brown-black, short, black, setae on all segments, tergites I and II laterally and sternites I-V with long, white setae; male tergites II-IV and intersegmental membrane on sternites II and III covered with silver velutum pubescence.
<emphasis id="4F6EE0D245B5F2109C5321CE161BC4D8" italics="true" pageId="0" pageNumber="83">Genitalia</emphasis>
. Male: tergite VIII emarginate medially; black setose laterally on posterior 1/2. Sternite VIII quadrate, slightly wider posteriorly, posterior margin black setose. Epandrium quadrate, emarginate anteriorly and posteriorly, lateral corners only slightly extended posteriorly; dark brown setae longer posterolaterally. Cerci bluntly pointed posteriorly, dark brown setae dorsally and apically. Subepandrial sclerite wide, 1/2 width of epandrium; partially sclerotised, lateral margins more strongly so. Gonocoxites rounded with subtriangular outer gonocoxal process; long, dark setae, densely spaced laterally and small patch ventromedially; inner gonocoxal process smoothly curving medially, acuminate, lacking setae apically; ventral lobe minutely setose ventrally, gonostylus curved dorsomedially with a broad, dorsal process at 2/3 of length with medial and lateral carinae basodorsally, dark brown setose ventrally and on medial face. Aedeagus with dorsal apodeme of parameral sheath
<normalizedToken id="86E6276971D4360AC517BA8170F7452D" originalValue="T">'T'</normalizedToken>
-shaped, narrow posteriorly and widened anteriorly with anterolateral corners curving ventrally and laterally; distiphallus wide basally with lateral carinae, narrow apically, distiphallus spinose laterally; ventral apodeme dorsoventrally flattened, bifurcations subparallel to ejaculatory apodeme; ejaculatory apodeme robust, roughly cylindrical; posterior end of ejaculatory apodeme broadened laterally and ventrally expanded to a point creating a basket-shaped posterior face, lateral ejaculatory apodeme short and subtriangular with lateral process; basiphallus membranous anteriorly. Female: tergites VII and VIII highly modified, much wider than long, with a long and narrow anteromedial process; posterior margin slightly emarginate; with elongate dark setae. Sternite VIII slightly longer than wide, convex ventrally, posterior lobes tapering sharply posteriorly, separate, with a median aedeagal guide; extensive elongate, orange setae, bare at extreme lateral margin. Median lobe of tergite IX reduced, very short and not sclerotised. Acanthophorite with acanthophorite spines dark brown, A2 series spines indistinguishable from rest of acanthophorite setae. Sternite X quadrate. Cercus slightly laterally flattened, extended posteriorly, longer than wide, membranous, slightly sclerotised; furca longer than wide, semicircular anteriorly, tapered to a point posteriorly; origin of spermathecal ducts occurring immediately adjacent to furcal membrane; three spermathecae, much longer than wide, saclike, wider apically than basally; spermathecal duct thickened basally, extremely narrowed basal to spermathecae; bifurcation of spermathecal sac duct with spermathecal ducts very close to furcal membrane; spermathecal sac short, 2.5
<normalizedToken id="4A22845A2092F75C48D4CD7D125C63F6" originalValue="×">x</normalizedToken>
furcal lengths; sac ovoid, longer than wide;
<normalizedToken id="6F49A8433AB540970DE47C76F3B28FC1" originalValue="¾×">3/4x</normalizedToken>
length of spermathecal sac duct.
</paragraph>
<caption id="7AF59ACE3400C3F870A76E5E1A06EC50" doi="10.3897/zookeys.984.53587.figure3" httpUri="https://binary.pensoft.net/fig/471319" pageId="0" pageNumber="83" start="Figure 3" startId="F3">
<paragraph id="5C3C0C3818B2421112B65DBD83140B28" pageId="0" pageNumber="83">
<emphasis id="D7F089BF27159D302B9EA6F46470DAFC" bold="true" pageId="0" pageNumber="83">Figure 3.</emphasis>
<taxonomicName id="4427197F760805CB87C2F763ECDCFD4F" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Jeanchazeauia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jeanchazeauia nubilosus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="nubilosus">
<emphasis id="C4D352BB5651D1891522577A537E5A63" italics="true" pageId="0" pageNumber="83">Jeanchazeauia nubilosus</emphasis>
</taxonomicName>
sp. nov., female habitus (MEI030203) (artistic rendering by J. Marie Metz).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8000E1EE4A30D2E1FC9760584A08B74A" pageId="0" pageNumber="83" type="etymology">
<paragraph id="271939C53DEFB4FC21B496133339333B" pageId="0" pageNumber="83">Etymology.</paragraph>
<paragraph id="0767548F7F98CA372E60116FC97347F5" pageId="0" pageNumber="83">
Derived from the Latin
<emphasis id="8D3B6AAFDE461A503C38DCA6692B4574" italics="true" pageId="0" pageNumber="83">nubilus</emphasis>
, cloudy, and -
<emphasis id="31935EEA1B6A61399C977F57368D8666" italics="true" pageId="0" pageNumber="83">osus</emphasis>
, full of; referring to the dark wing infuscation of the male.
</paragraph>
<caption id="AF8D237504CE56658974652CC201D9B2" doi="10.3897/zookeys.984.53587.figure4" httpUri="https://binary.pensoft.net/fig/471320" pageId="0" pageNumber="83" start="Figure 4" startId="F4">
<paragraph id="3CED6560944A4451A05233A51B8660F9" pageId="0" pageNumber="83">
<emphasis id="7F50EC4BCEF2AF5786196E1C09DD2EAB" bold="true" pageId="0" pageNumber="83">Figure 4.</emphasis>
Phylogenetic placement of
<taxonomicName id="9E392D0383085A54082AFD09D4B18A1D" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="8B5AC8BB7C82F93C87F70602284B02D8" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
</taxonomicName>
gen. nov. and
<taxonomicName id="84DFAB23AC456C57562F2B4AF0EAF4E8" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Jeanchazeauia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jeanchazeauia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="5C42E6FAE5ECE5FB85F406640394DEFE" italics="true" pageId="0" pageNumber="83">Jeanchazeauia</emphasis>
</taxonomicName>
gen. nov. in
<taxonomicName id="1CF3900DA47D374193621A3FF7BCF548" lsidName="" pageId="0" pageNumber="83" rank="subFamily" subFamily="Agapophytinae">Agapophytinae</taxonomicName>
based on supermatrix analysis of DNA sequence data. Figure modified after
<bibRefCitation id="50FCD9C4E2B248F5200F0F4C3D7E23DC" DOI="https://doi.org/10.1111/syen.12147" author="Winterton, SL" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="83" pagination="144 - 161" refId="B18" refString="Winterton, SL, Hardy, NB, Gaimari, SD, Hauser, M, Hill, HN, Holston, KC, Irwin, ME, Lambkin, CL, Metz, MA, Turco, F, Webb, D, Yang, L, Yeates, DK, Wiegmann, BM, 2016. The phylogeny of stiletto flies (Diptera: Therevidae). Systematic Entomology 41: 144 - 161, DOI: https://doi.org/10.1111/syen.12147" title="The phylogeny of stiletto flies (Diptera: Therevidae)." url="https://doi.org/10.1111/syen.12147" volume="41" year="2016">Winterton et al. (2016</bibRefCitation>
: fig. 3).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F70E22809D6515D45E3977AAE3B97437" pageId="0" pageNumber="83" type="comments">
<paragraph id="1F8E710D0E681213D144FE061D912D64" pageId="0" pageNumber="83">Comments.</paragraph>
<paragraph id="8F752DBA7C42DFCD0DC125C5677722AA" pageId="0" pageNumber="83">
<taxonomicName id="C3BEE15497480F1151E4598D3498A1EE" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Jeanchazeauia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jeanchazeauia nubilosus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="nubilosus">
<emphasis id="84720848D6D4E7B93D686C235889BCD1" italics="true" pageId="0" pageNumber="83">Jeanchazeauia nubilosus</emphasis>
</taxonomicName>
sp. nov. exhibits remarkable sexual dimorphism in wing pattern, but little to no other differences in the body or head. The male has largely uniform dark infuscate wings, while the wings of the female are strongly banded. It is not known if this condition extends to other species in the genus where the males are unknown, and such dramatic sexual dimorphism in wing patterning is rare in stiletto flies to this degree.
<taxonomicName id="A4D9C1F7BF5A70731E9415A115DD09C4" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Jeanchazeauia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jeanchazeauia nubilosus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="nubilosus">
<emphasis id="ED70BF92DDAB9DB7C3CA73AB14FE2E99" italics="true" pageId="0" pageNumber="83">Jeanchazeauia nubilosus</emphasis>
</taxonomicName>
sp. nov. has been collected in dry to humid tropical forest at higher elevations on several mountains in New Caledonia (e.g., Figs
<figureCitation id="9C312FDE220A93AE462A5A4D6243FA91" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Natural habitats of New Caledonian agapophytine stiletto flies A Riviere Bleue Reserve, sclerophyll forest and maquis scrub, habitat of C. chazeaui sp. nov., C. grandiosus sp. nov., C. schlingeri sp. nov., and J. nubilosus sp. nov. B Pindai Peninsula, tropical dry forest, habitat of C. webbi sp. nov. C Mount Dzumac, rainforest, habitat of J. nubilosus sp. nov. D Mount Dore, rainforest and dry sclerophyll forest E Mount Dore, Malaise trap in flyway (photographs by Michael E. Irwin)." figureDoi="10.3897/zookeys.984.53587.figure31" httpUri="https://binary.pensoft.net/fig/471341" pageId="0" pageNumber="83">31A, C</figureCitation>
,
<figureCitation id="B2DDF02BC5BB950B73DD96DAB0294480" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Natural habitats of New Caledonian agapophytine stiletto flies A Mount Aoupinie, rainforest, habitat of C. monteithi sp. nov. B Mount Ningua, rainforest, habitat of J. nubilosus sp. nov. C Mount Mandjelia, habitat of C. grandiosus sp. nov. (photographs by Michael E. Irwin)." figureDoi="10.3897/zookeys.984.53587.figure32" httpUri="https://binary.pensoft.net/fig/471342" pageId="0" pageNumber="83">32B</figureCitation>
). GenBank sequences for this species (see
<bibRefCitation id="049430A0B71A50C38AF9141BA7E7EB98" DOI="https://doi.org/10.1111/syen.12147" author="Winterton, SL" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="83" pagination="144 - 161" refId="B18" refString="Winterton, SL, Hardy, NB, Gaimari, SD, Hauser, M, Hill, HN, Holston, KC, Irwin, ME, Lambkin, CL, Metz, MA, Turco, F, Webb, D, Yang, L, Yeates, DK, Wiegmann, BM, 2016. The phylogeny of stiletto flies (Diptera: Therevidae). Systematic Entomology 41: 144 - 161, DOI: https://doi.org/10.1111/syen.12147" title="The phylogeny of stiletto flies (Diptera: Therevidae)." url="https://doi.org/10.1111/syen.12147" volume="41" year="2016">Winterton et al. 2016</bibRefCitation>
: table S1): KM885007 (28S rDNA), KM879119 (EF1a).
</paragraph>
<caption id="9C865FF4757AC241559E0ECC0931FA7C" doi="10.3897/zookeys.984.53587.figure5" httpUri="https://binary.pensoft.net/fig/471321" pageId="0" pageNumber="83" start="Figure 5" startId="F5">
<paragraph id="D8E5436DC10B32D254FB122CC741EBA7" pageId="0" pageNumber="83">
<emphasis id="42EE5B8E2C6437A6F4C5E27BF2386A10" bold="true" pageId="0" pageNumber="83">Figure 5.</emphasis>
Lateral view of antennae of
<taxonomicName id="E23CA2B22E1AF6D83A4041B451EDAD12" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="ADED3E58DD64774D4257A4003595D70C" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
</taxonomicName>
gen. nov.
<emphasis id="D1A85F00CB454146BDB38C25F5260DA1" bold="true" pageId="0" pageNumber="83">A</emphasis>
<taxonomicName id="3C5E4DB13BF8AC9DCD28B94A97B43F52" lsidName="C. chazeaui" pageId="0" pageNumber="83" rank="species" species="chazeaui">
<emphasis id="D8586CCDF396D087E1D81C8F1EC79E14" italics="true" pageId="0" pageNumber="83">C. chazeaui</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="7511BAC2DCEFA61E0675D8C7A6F4AFDF" bold="true" pageId="0" pageNumber="83">B</emphasis>
<taxonomicName id="50114F1F9CC3434A0B853DB0AB706880" lsidName="C. grandiosus" pageId="0" pageNumber="83" rank="species" species="grandiosus">
<emphasis id="5B92628A52FBFB8992855B1BEFED2C13" italics="true" pageId="0" pageNumber="83">C. grandiosus</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="B104380AFD24134C27CC32A88E52C261" bold="true" pageId="0" pageNumber="83">C</emphasis>
<taxonomicName id="C30D37468500138C9A6CC9D92F40DEEC" lsidName="C. schlingeri" pageId="0" pageNumber="83" rank="species" species="schlingeri">
<emphasis id="CBC24F8068F4DC98A4A904A05DE1C6B3" italics="true" pageId="0" pageNumber="83">C. schlingeri</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="5703A44485785217E24C9CF02154527E" bold="true" pageId="0" pageNumber="83">D</emphasis>
<taxonomicName id="9B654BEF49B6AA46730D73D119D0D292" lsidName="C. matilei" pageId="0" pageNumber="83" rank="species" species="matilei">
<emphasis id="127DC4A9B43EC23FA366847D2D33EABA" italics="true" pageId="0" pageNumber="83">C. matilei</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="604E0B0B039F32DB7559C20F9028C2AA" bold="true" pageId="0" pageNumber="83">E</emphasis>
<taxonomicName id="DF972CDC7F22BB18CC5A46245B8E5E57" lsidName="C. webbi" pageId="0" pageNumber="83" rank="species" species="webbi">
<emphasis id="1A2352160053225E3A14EC95AE3C839B" italics="true" pageId="0" pageNumber="83">C. webbi</emphasis>
</taxonomicName>
sp. nov. (figures not to scale) (drawings by J. Marie Metz).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7A4A958A6915CBBF8C42E8767BEF232E" pageId="0" pageNumber="83" type="materials_examined">
<paragraph id="4DBA1B900FBFB0C0F3E5793E4039E9E3" pageId="0" pageNumber="83">Specimens examined.</paragraph>
<paragraph id="845DDF9B1DB40F1D15570203E4B1AE9D" pageId="0" pageNumber="83">
<emphasis id="F8C336B5DBF0D274D987070B8C3D7604" bold="true" italics="true" pageId="0" pageNumber="83">Holotype</emphasis>
male, New Caledonia: Province Nord: Mt. Dzumac [Dumbea, approximated as
<geoCoordinate id="FE42CF90F5BAB6D31A99C8EF7CF95D1F" degrees="22.106" direction="south" orientation="latitude" precision="55" value="-22.106">-22.106</geoCoordinate>
,
<geoCoordinate id="337C84182B1CA109AC565079EE8B01B7" degrees="166.458" direction="east" orientation="longitude" precision="55" value="166.458">166.458</geoCoordinate>
], black light (UV), 27-28.II.1984, M. Pogue, M. Epstein (MEI030206, MNHN).
</paragraph>
<caption id="CD17914892D2231F89E4EB990FA672FB" doi="10.3897/zookeys.984.53587.figure6" httpUri="https://binary.pensoft.net/fig/471322" pageId="0" pageNumber="83" start="Figure 6" startId="F6">
<paragraph id="C84B81B1D542261502CB202F3EB8872E" pageId="0" pageNumber="83">
<emphasis id="D99EAFC58F30F7A42E585717C9F2E942" bold="true" pageId="0" pageNumber="83">Figure 6.</emphasis>
Scutum and scutellum of
<taxonomicName id="B63785AE7198EF326446092773A3E1BB" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="C93DA98CC5F5F0AAA3FCF2A9A5131E56" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
</taxonomicName>
gen. nov. and
<taxonomicName id="C13A7F0BA2673BBDF16483B938EB64AF" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Jeanchazeauia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jeanchazeauia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="3037B45305E9ED17E01146AE85EA6033" italics="true" pageId="0" pageNumber="83">Jeanchazeauia</emphasis>
</taxonomicName>
gen. nov.
<emphasis id="654DF3FB7B75C9696E4DB8F62EAFCB65" bold="true" pageId="0" pageNumber="83">A</emphasis>
<taxonomicName id="7152DBBA54748E44F04B593B53BB68AA" lsidName="C. schlingeri" pageId="0" pageNumber="83" rank="species" species="schlingeri">
<emphasis id="E03A4E138951C02297F7A0A3127EA03C" italics="true" pageId="0" pageNumber="83">C. schlingeri</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="F9B8FE79F5A149CBF8FFDEBF258DA2ED" bold="true" pageId="0" pageNumber="83">B</emphasis>
<taxonomicName id="2AD0384AA9F90C3E06E540932525BB05" lsidName="C. grandiosus" pageId="0" pageNumber="83" rank="species" species="grandiosus">
<emphasis id="731F71A4C8B5695DFCD3BFF099D92DC5" italics="true" pageId="0" pageNumber="83">C. grandiosus</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="7953E5A5A6C29732B72FE652E69405E3" bold="true" pageId="0" pageNumber="83">C</emphasis>
<taxonomicName id="7153E7D4233F88BBC1C6A53D956187AA" lsidName="C. matilei" pageId="0" pageNumber="83" rank="species" species="matilei">
<emphasis id="091009471D2D57BC1C9285AE29D082BC" italics="true" pageId="0" pageNumber="83">C. matilei</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="79777B1AB82E1F0929D103885CC4C007" bold="true" pageId="0" pageNumber="83">D</emphasis>
<taxonomicName id="934E59759E30D4F1F95560AF0F860748" lsidName="C. webbi" pageId="0" pageNumber="83" rank="species" species="webbi">
<emphasis id="6DAC2E989AF7CAB669084E756E2CA051" italics="true" pageId="0" pageNumber="83">C. webbi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="27617E6D8CDFFC2A27CC55CB40464539" bold="true" pageId="0" pageNumber="83">E</emphasis>
<taxonomicName id="376CE54EB218AAD637D17772FDFA3DAC" lsidName="J. nubilosus" pageId="0" pageNumber="83" rank="species" species="nubilosus">
<emphasis id="57EA786D0745E6BBF1F5ED09BCA638F2" italics="true" pageId="0" pageNumber="83">J. nubilosus</emphasis>
</taxonomicName>
sp. nov. (figures not to scale) (drawings by J. Marie Metz).
</paragraph>
</caption>
<caption id="104365AAE19F4C3C1AB42C9D00233793" doi="10.3897/zookeys.984.53587.figure13" httpUri="https://binary.pensoft.net/fig/471323" pageId="0" pageNumber="83" start="Figure 13" startId="F13">
<paragraph id="58D3F1F137A1E0B30E0F3D4F23D8B8B0" pageId="0" pageNumber="83">
<emphasis id="44B55959CE4F4BA1AEC06176EC2A7D80" bold="true" pageId="0" pageNumber="83">Figure 13.</emphasis>
<taxonomicName id="7F30FC25D344A4BE90C62ED86B38D3FA" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus matilei" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="matilei">
<emphasis id="09B7B590F2AA838BF00281990C8E8D7C" italics="true" pageId="0" pageNumber="83">Calophytus matilei</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="0F9DA82EA773C0416E20798DCC75C2B9" bold="true" pageId="0" pageNumber="83">A</emphasis>
adult male (MEI135018), oblique view
<emphasis id="5C3D2FCCDAFC167150C8B39532DCD428" bold="true" pageId="0" pageNumber="83">B</emphasis>
same, lateral view
<emphasis id="2E66A1634ABA2C021B612F5CB2195A56" bold="true" pageId="0" pageNumber="83">C</emphasis>
adult female (MEI071892), oblique view
<emphasis id="D2CCAA6E5954A327F64667A9771D205C" bold="true" pageId="0" pageNumber="83">D</emphasis>
same, lateral view. Body length: male: 8.0 mm; female: 9.4 mm.
</paragraph>
</caption>
<caption id="3F6119ED6DC47DCBDD82945CCA5992CF" doi="10.3897/zookeys.984.53587.figure14" httpUri="https://binary.pensoft.net/fig/471324" pageId="0" pageNumber="83" start="Figure 14" startId="F14">
<paragraph id="980C9A531D82CFF77DFCFFAB66A0A92D" pageId="0" pageNumber="83">
<emphasis id="5A00E4E959997C1CE26DCCB3183C4D86" bold="true" pageId="0" pageNumber="83">Figure 14.</emphasis>
<taxonomicName id="992F1133AC172BF89C42AAF725BA1646" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus matilei" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="matilei">
<emphasis id="BE67A2DB08D95BF0592C7B40748EE566" italics="true" pageId="0" pageNumber="83">Calophytus matilei</emphasis>
</taxonomicName>
sp. nov., cleared male genitalia
<emphasis id="199E5ACE600C9DAC4CA5C0CF4834A82A" bold="true" pageId="0" pageNumber="83">A</emphasis>
aedeagus, lateral view
<emphasis id="0AE303F7C6362EC5C2DF20A39734FBD4" bold="true" pageId="0" pageNumber="83">B</emphasis>
gonocoxites, ventral view
<emphasis id="27C4C009BEAC56E31CB6489434AEF020" bold="true" pageId="0" pageNumber="83">C</emphasis>
epandrium. Scale bar: 0.2 mm.
</paragraph>
</caption>
<caption id="D85E8493FA7E6921F24256BA6686AEAB" doi="10.3897/zookeys.984.53587.figure15" httpUri="https://binary.pensoft.net/fig/471325" pageId="0" pageNumber="83" start="Figure 15" startId="F15">
<paragraph id="997947CB21C51AFF2DB999290444CC78" pageId="0" pageNumber="83">
<emphasis id="4788C12DE4C95096C4EDA619B58DB221" bold="true" pageId="0" pageNumber="83">Figure 15.</emphasis>
<taxonomicName id="FC8B72DAB3ADC090D56C00DE39F3E696" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus monteithi" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="77421066FC214D3DE67FAAC1B3B53047" italics="true" pageId="0" pageNumber="83">Calophytus monteithi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="2478AE533516CAEBFD060603A4F48D95" bold="true" pageId="0" pageNumber="83">A</emphasis>
adult female (MEI138464), oblique view
<emphasis id="207573F5F76229B97EC73C585049B34F" bold="true" pageId="0" pageNumber="83">B</emphasis>
same, lateral view. Body length: 10.3 mm.
</paragraph>
</caption>
<paragraph id="D52DF34350B2D039D9E80067A73FD5FE" pageId="0" pageNumber="83">
<emphasis id="F39D55773C2FB466B45F9E5A8093C226" bold="true" italics="true" pageId="0" pageNumber="83">Paratypes</emphasis>
. New Caledonia: Province Nord: 1 male, 13 km by road from Ouego[a] to Mont
<normalizedToken id="C72ABD199886B55536958B965A10931C" originalValue="Mandjanié">Mandjanie</normalizedToken>
, Malaise trap along stream [
<geoCoordinate id="FAD829A752371FE2A832DEFC14B578BE" degrees="20.374" direction="south" orientation="latitude" precision="55" value="-20.374">-20.374</geoCoordinate>
,
<geoCoordinate id="205779ED2C044CEBE0FBC5DCA719E7B4" degrees="164.482" direction="east" orientation="longitude" precision="55" value="164.482">164.482</geoCoordinate>
], 175 m, 26.XI.1992, D.W. Webb, E.I. &amp; M. Schlinger, (MEI030207, CSCA). Province Sud: 1 male,
<normalizedToken id="A592D56BDB1EAB14FAF83D9AEC52E227" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="0A3A9563366561D800FD6E817C3F5B46" originalValue="Spéciale">Speciale</normalizedToken>
de Botanique, Mount Ningua, Malaise trap [
<geoCoordinate id="B0830F0A8128E763D6D0EB9B48BFE55F" degrees="21.735" direction="south" orientation="latitude" precision="55" value="-21.735">-21.735</geoCoordinate>
,
<geoCoordinate id="582CDC8F48DBF1B2A8A274B061BCDA68" degrees="166.142" direction="east" orientation="longitude" precision="55" value="166.142">166.142</geoCoordinate>
], 1100 m, 12-21.XII.2000, L.J. Boutin, M.E. Irwin (MEI131365, CSCA); 3 females,
<normalizedToken id="2B67488737637F43A63E5170E666DEE8" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="E14FAAEA59216BD159AD990B5F1F5FD9" originalValue="Spéciale">Speciale</normalizedToken>
de Botanique, Mount Ningua, Malaise trap [-21.735, 166.142], 1100 m, various dates: 9-21.XII.2000, D.W. Webb, E.I. Schlinger, M.E. Irwin (MEI131362-4, CSCA); 2 females, Ningua Reserve Camp, Malaise trap, rainforest [-21.75, 166.15], 1100 m, 12-13.XI.2001, 27.XI-29.I.2002, Burwell, Monteith (MEI138461-2, QM); 2 females, 17 km NNE
<normalizedToken id="818917B664D7AF3DF799B0D411781EAB" originalValue="Nouméa">Noumea</normalizedToken>
, Mt. Koghis, Malaise trap in tropical forest [
<geoCoordinate id="7D68DAC99988882869C23CF5402C1062" degrees="22.167" direction="south" orientation="latitude" precision="55" value="-22.167">-22.167</geoCoordinate>
,
<geoCoordinate id="23441AE2F832312A7AA92A7AD9D34A58" degrees="166.533" direction="east" orientation="longitude" precision="55" value="166.533">166.533</geoCoordinate>
], 500 m, 5-15.XI.1992, D.W. Webb (MEI030202-3, CSCA); 2 females, 17 km NNE
<normalizedToken id="92366233CC404E4536BA35600B15BC0F" originalValue="Nouméa">Noumea</normalizedToken>
, Mt. Koghis, Malaise trap across forest stream [
<geoCoordinate id="AFA911279E01DC862193679D3E4BE4D6" degrees="22.167" direction="south" orientation="latitude" precision="55" value="-22.167">-22.167</geoCoordinate>
,
<geoCoordinate id="3CC3783352BC83C11AB77AA7FF52D358" degrees="166.533" direction="east" orientation="longitude" precision="55" value="166.533">166.533</geoCoordinate>
], 500 m, 23-26.XII.1991, M.E. Irwin, D.W. Webb (MEI030195-6, MEI); 1 female,
<normalizedToken id="5C9259C367337941993B08570E3CAF3D" originalValue="Nouméa">Noumea</normalizedToken>
, Mt. Koghis, Malaise trap [
<geoCoordinate id="3AD9B035E9C28D3E4A9C819886AFE1BF" degrees="22.167" direction="south" orientation="latitude" precision="55" value="-22.167">-22.167</geoCoordinate>
,
<geoCoordinate id="DB8790A625ABED9322A278977360EBF6" degrees="166.533" direction="east" orientation="longitude" precision="55" value="166.533">166.533</geoCoordinate>
], 500 m, 4.XII.1963, R. Straatman (MEI030197, BPBM); 1 female, 17 km NNE
<normalizedToken id="CF16DC693FE3905200E834A4099576C2" originalValue="Nouméa">Noumea</normalizedToken>
, Mount Khogis, Malaise trap across path in rainforest [
<geoCoordinate id="ED79D555CB6344572311AB2325B526BE" degrees="22.176" direction="south" orientation="latitude" precision="55" value="-22.176">-22.176</geoCoordinate>
,
<geoCoordinate id="B1B5AF72D14BA63142DA794F8AE74D15" degrees="166.505" direction="east" orientation="longitude" precision="55" value="166.505">166.505</geoCoordinate>
], 425 m, 8.I.1996, M.E. Irwin, D.W. Webb, E.I. Schlinger, (MEI071888, CSCA); 3 females, 30 km NW Yate,
<normalizedToken id="4F7D66D5636E9F19FA06564CA3628B55" originalValue="Rivière">Riviere</normalizedToken>
Bleue, Malaise trap [
<geoCoordinate id="83D1AE7D00DC8316197DB99306F0D6C2" degrees="22.117" direction="south" orientation="latitude" precision="55" value="-22.117">-22.117</geoCoordinate>
,
<geoCoordinate id="46990849D3CB3CF903CCC6FBB19B95E2" degrees="166.658" direction="east" orientation="longitude" precision="55" value="166.658">166.658</geoCoordinate>
], 12-25.XI.1986, 11-27.X.1988, L.B. de Larbogne, J. Chazeau, A. &amp; S. Tillier, (MEI030198-200, MNHN); 1 female,
<normalizedToken id="1530441FFE9EF7380C6D60BA7C28B580" originalValue="Rivière">Riviere</normalizedToken>
Bleue Provincial Park, 19.6 km
<normalizedToken id="64F5DA8B68EE7E10E6F16D918B11E843" originalValue="Rivière">Riviere</normalizedToken>
Bleue Road, Malaise trap across forest path [
<geoCoordinate id="6704427CCFBF90602070BAC2FA1E3A96" degrees="22.117" direction="south" orientation="latitude" precision="55" value="-22.117">-22.117</geoCoordinate>
,
<geoCoordinate id="EF7E63362605E59FED7167992AAC452A" degrees="166.658" direction="east" orientation="longitude" precision="55" value="166.658">166.658</geoCoordinate>
], 183 m, 18-20.XI.1992, D.W. Webb (MEI030201, CSCA).
</paragraph>
<caption id="BF1E6580E381B6B0BD7D85DE2CF44779" doi="10.3897/zookeys.984.53587.figure16" httpUri="https://binary.pensoft.net/fig/471326" pageId="0" pageNumber="83" start="Figure 16" startId="F16">
<paragraph id="CE608B1282245D85046BD33871CC801B" pageId="0" pageNumber="83">
<emphasis id="49101A2FDC53E85F33A56BBCA42C75E7" bold="true" pageId="0" pageNumber="83">Figure 16.</emphasis>
<taxonomicName id="3898A5B7A8DA3C2D335A486E466389DD" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus schlingeri" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="4A6621ABF745550A254555FA57AE1319" italics="true" pageId="0" pageNumber="83">Calophytus schlingeri</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="078B1399A5B0702D2F4C76E15F926E9F" bold="true" pageId="0" pageNumber="83">A</emphasis>
adult male (MEI030158), anterolateral view
<emphasis id="1BB43E92704E3205D2C1E0C1994FF69C" bold="true" pageId="0" pageNumber="83">B</emphasis>
same, oblique view
<emphasis id="B037B655FEBF9EEE4BE5B4F8D5901377" bold="true" pageId="0" pageNumber="83">C</emphasis>
adult female (MEI030172), lateral view
<emphasis id="1DAC7E92CD965650020E59F22B449C31" bold="true" pageId="0" pageNumber="83">D</emphasis>
same, oblique view. Body length: male: 8.6 mm; female: 9.8 mm.
</paragraph>
</caption>
<caption id="D5DDC8495FF9A4D7E21242EB8C623AE8" doi="10.3897/zookeys.984.53587.figure17" httpUri="https://binary.pensoft.net/fig/471327" pageId="0" pageNumber="83" start="Figure 17" startId="F17">
<paragraph id="60A25D51E947C18CF93F3E710FCA60F4" pageId="0" pageNumber="83">
<emphasis id="8DF33AA42CADF8E045540EB502B5416B" bold="true" pageId="0" pageNumber="83">Figure 17.</emphasis>
<taxonomicName id="3BD231BD23F7D511D2E5DC5302A88FE3" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus schlingeri" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="8609C1A1BADEDAC8E44D5BE32FC10FCF" italics="true" pageId="0" pageNumber="83">Calophytus schlingeri</emphasis>
</taxonomicName>
sp. nov., cleared male genitalia
<emphasis id="048C0345DDBA36BECE480FB929186FFB" bold="true" pageId="0" pageNumber="83">A</emphasis>
epandrium
<emphasis id="5972B0F480B4DC3E4C8FE42731DDC707" bold="true" pageId="0" pageNumber="83">B</emphasis>
gonocoxites and aedeagus, ventral view
<emphasis id="87CCDFF07E811D80E5B2966490CDE97D" bold="true" pageId="0" pageNumber="83">C</emphasis>
aedeagus, dorsal view
<emphasis id="D5161E53CBE87C68781DDF47074C5447" bold="true" pageId="0" pageNumber="83">D</emphasis>
gonocoxites, ventral view. Scale bar: 0.2 mm.
</paragraph>
</caption>
<caption id="B756DE2AE41DAF2A3D8B780E0F3ACE69" doi="10.3897/zookeys.984.53587.figure18" httpUri="https://binary.pensoft.net/fig/471328" pageId="0" pageNumber="83" start="Figure 18" startId="F18">
<paragraph id="C01C0A764A6166B5B0D7F330F473584F" pageId="0" pageNumber="83">
<emphasis id="4C671BBB1BA45C8ED679A74157570F8F" bold="true" pageId="0" pageNumber="83">Figure 18.</emphasis>
<taxonomicName id="ABC8EEBEC7949EC8A747AFF81C934E8A" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus schlingeri" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="E53EC6D5FEDBFDA957F03E4108D1DA27" italics="true" pageId="0" pageNumber="83">Calophytus schlingeri</emphasis>
</taxonomicName>
sp. nov., genitalia
<emphasis id="90553B7E822D330D15F1F63D814163BB" bold="true" pageId="0" pageNumber="83">A</emphasis>
epandrium
<emphasis id="BBD5DCE2470398612B9640F93656F1F0" bold="true" pageId="0" pageNumber="83">B</emphasis>
gonocoxites, ventral view
<emphasis id="A9B5AD58E487024FD3476F5AC7D936C3" bold="true" pageId="0" pageNumber="83">C</emphasis>
same, lateral view
<emphasis id="87485033839824FE15F4C021C1800C7D" bold="true" pageId="0" pageNumber="83">D</emphasis>
aedeagus, lateral view
<emphasis id="9309F273A47A392FA2A8340A391CA7CC" bold="true" pageId="0" pageNumber="83">E</emphasis>
same, dorsal view
<emphasis id="0BD2788C36AC44F44B9C849FA635DF9E" bold="true" pageId="0" pageNumber="83">F</emphasis>
female genitalia, lateral view, internal structures shown. Scale line: 0.2 mm. Abbreviations:
<emphasis id="D93DC070FEC8C25C9192145F3C2E4752" italics="true" pageId="0" pageNumber="83">ag</emphasis>
, accessory gland;
<emphasis id="2872E5DED2FE5D379BE77AD0F7B629EA" italics="true" pageId="0" pageNumber="83">d</emphasis>
, distiphallus;
<emphasis id="91068A02E91EE6089806563C268CEE09" italics="true" pageId="0" pageNumber="83">da</emphasis>
, dorsal apodeme of parameral sheath;
<emphasis id="75AA5AF32B3E65BA5F41BC284ECCCBE7" italics="true" pageId="0" pageNumber="83">ea</emphasis>
, ejaculatory apodeme;
<emphasis id="3CCA7A9A4A7672C36E28F8CD01540782" italics="true" pageId="0" pageNumber="83">c</emphasis>
, cercus;
<emphasis id="AD69EFFC113C85B2ACE94D2250FB12BA" italics="true" pageId="0" pageNumber="83">A1</emphasis>
, primary acanthophorite spines;
<emphasis id="34C8011149571068445C02335EA11424" italics="true" pageId="0" pageNumber="83">A2</emphasis>
secondary acanthophorite spines;
<emphasis id="19B42E8B637C886CBFB177E6A85650AE" italics="true" pageId="0" pageNumber="83">ga</emphasis>
, gonocoxal apodeme;
<emphasis id="AA828E3EF0FE6A1E06F2CC5F901ADF7D" italics="true" pageId="0" pageNumber="83">gs</emphasis>
, gonostylus;
<emphasis id="43146C0CEA3277BB52CA58A1DCFCE08A" italics="true" pageId="0" pageNumber="83">igp</emphasis>
, inner gonocoxal process (articulated);
<emphasis id="A0706C87AE681C7016488628CF9727E7" italics="true" pageId="0" pageNumber="83">lea</emphasis>
, lateral ejaculatory apodeme;
<emphasis id="57C8E7A9A49DE8F8FF8A7091C132F0C0" italics="true" pageId="0" pageNumber="83">ogp</emphasis>
, outer gonocoxal process;
<emphasis id="6EF4D55C664294769C805D8E2AF9B478" italics="true" pageId="0" pageNumber="83">s8</emphasis>
, sternite VIII;
<emphasis id="04C070A667C7C993E54354C26129B36D" italics="true" pageId="0" pageNumber="83">s10</emphasis>
, sternite X;
<emphasis id="AF93A9B524BB4D0FE9AA083744171654" italics="true" pageId="0" pageNumber="83">s</emphasis>
, spermatheca;
<emphasis id="B94F9A30EFAFB8402410CD8AA763D02E" italics="true" pageId="0" pageNumber="83">ss</emphasis>
, spermathecal sac;
<emphasis id="A76F7272A83D173C7F7F9F1A35D99246" italics="true" pageId="0" pageNumber="83">ssd</emphasis>
, spermathecal sac duct;
<emphasis id="8EB5C1A1C3CC37ACF3B0FBAA26BB34C7" italics="true" pageId="0" pageNumber="83">sd</emphasis>
, spermathecal duct;
<emphasis id="8A0C8DC148035A7AB59602652F77EFBA" italics="true" pageId="0" pageNumber="83">vl</emphasis>
, ventral lobe.
</paragraph>
</caption>
<caption id="2D50AE68F7A07A451F263A9296D329FA" doi="10.3897/zookeys.984.53587.figure19" httpUri="https://binary.pensoft.net/fig/471329" pageId="0" pageNumber="83" start="Figure 19" startId="F19">
<paragraph id="4C566C388FFC47D35F60582CBB16A1A3" pageId="0" pageNumber="83">
<emphasis id="41C9C143D2D832DFEFBBA3F8F1B28F25" bold="true" pageId="0" pageNumber="83">Figure 19.</emphasis>
<taxonomicName id="EF9E25BE02659AA5CB9E1C116A1F48C7" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus webbi" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="webbi">
<emphasis id="07D01210685CCDA21B448BB0C30240D2" italics="true" pageId="0" pageNumber="83">Calophytus webbi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="65BFFDB0E871A0B65604EE14CC3100D8" bold="true" pageId="0" pageNumber="83">A</emphasis>
adult male (MEI030179), lateral view
<emphasis id="0FB1DD4835CBDACCA72DF45DD480B18F" bold="true" pageId="0" pageNumber="83">B</emphasis>
same, oblique view. Body length: 6.3 mm.
</paragraph>
</caption>
<caption id="E9CC9FAA12AA726A2A133770F6C40118" doi="10.3897/zookeys.984.53587.figure20" httpUri="https://binary.pensoft.net/fig/471330" pageId="0" pageNumber="83" start="Figure 20" startId="F20">
<paragraph id="ADFB4522B1DF853BA7B501DC01ED501D" pageId="0" pageNumber="83">
<emphasis id="CB778BC05499A4970C2F0999DBF694A6" bold="true" pageId="0" pageNumber="83">Figure 20.</emphasis>
<taxonomicName id="3BB315000772DF8A57CBEFBF47FE9445" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus webbi" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="webbi">
<emphasis id="54F24190A46D343691DB8608D9E55C9E" italics="true" pageId="0" pageNumber="83">Calophytus webbi</emphasis>
</taxonomicName>
sp. nov. adult female head, dorsolateral view.
</paragraph>
</caption>
<caption id="0A5354C8002673D2D6CBBBC25C8FCDF7" doi="10.3897/zookeys.984.53587.figure21" httpUri="https://binary.pensoft.net/fig/471331" pageId="0" pageNumber="83" start="Figure 21" startId="F21">
<paragraph id="C2B00032C64C94674B182729102C209C" pageId="0" pageNumber="83">
<emphasis id="F5162DB4685F5EDAA55CE4CEE9540A69" bold="true" pageId="0" pageNumber="83">Figure 21.</emphasis>
<taxonomicName id="C737C9515C8BCF327124334122E4A091" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus webbi" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="webbi">
<emphasis id="BC51918C8CB977A5A806C78C0ED594A9" italics="true" pageId="0" pageNumber="83">Calophytus webbi</emphasis>
</taxonomicName>
sp. nov., cleared male genitalia
<emphasis id="4EC073A02FFE71619814C993846A308C" bold="true" pageId="0" pageNumber="83">A</emphasis>
epandrium and tergite VIII
<emphasis id="BCE2704415D3A870C1454C821BCEF73D" bold="true" pageId="0" pageNumber="83">B</emphasis>
gonocoxites and aedeagus, dorsal view with epandrium removed. Scale bar: 0.2 mm.
</paragraph>
</caption>
<caption id="B140C67DE23AEBEEEC3343EE975D7D61" doi="10.3897/zookeys.984.53587.figure22" httpUri="https://binary.pensoft.net/fig/471332" pageId="0" pageNumber="83" start="Figure 22" startId="F22">
<paragraph id="193881B709B8B12D7A4C0824D27E0CE1" pageId="0" pageNumber="83">
<emphasis id="BA3604A9AB311EFDE137A501F30F98C6" bold="true" pageId="0" pageNumber="83">Figure 22.</emphasis>
Distribution of
<taxonomicName id="EFBAC9D5826DFCC8DA2E4CAC0D9E8FC2" authorityName="Irwin &amp; Winterton &amp; Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="5C2FA9400BB442BE58F30A1914631776" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
</taxonomicName>
spp. collecting records throughout New Caledonia. Provinces are labelled and ecoregions delineated by colour.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>