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<document ID-CLB-Dataset="53615" ID-DOI="http://dx.doi.org/10.3897/dez.69.80492" ID-GBIF-Dataset="f61bc90c-fed3-41d4-b954-2b8c0f552848" ID-Pensoft-Pub="1860-1324-1-45" ID-Pensoft-UUID="0A5DABFBAA6551E194C35CEA6054B588" ID-ZooBank="C402CAE814FD402AAD0554646B1493E8" ModsDocID="1860-1324-69-1-45" checkinTime="1649136064731" checkinUser="pensoft" docAuthor="Morishita, Shunsuke &amp; Watanabe, Kyohei" docDate="2022" docId="8DDA9D88629C59378D2AAEA99213B44E" docLanguage="en" docName="DeutEntomolZeit 69(1): 45-64" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-04-04" docSource="http://dx.doi.org/10.3897/dez.69.80492" docTitle="Woldstedtius alpicola Morishita &amp; Watanabe 2022, sp. nov." docType="treatment" docUuid="E030CAC0-73B1-4BDC-BD63-F5E6BD3BAC4B" docUuidSource="ZooBank" docVersion="5" id="0A5DABFBAA6551E194C35CEA6054B588" lastPageNumber="45" masterDocId="0A5DABFBAA6551E194C35CEA6054B588" masterDocTitle="Revision of the genus Woldstedtius Carlson, 1979 (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan" masterLastPageNumber="64" masterPageNumber="45" pageNumber="45" updateTime="1732742606261" updateUser="ExternalLinkService">
<mods:mods id="162DF140D63ED36DB6A993ED9E1A4EDA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="04C0E7C4B4458BB5EDC9F55CF753302A">Revision of the genus Woldstedtius Carlson, 1979 (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan</mods:title>
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<mods:namePart id="81616E5A3A19D13805B576944CFDDE52">Morishita, Shunsuke</mods:namePart>
<mods:nameIdentifier id="D22D0F05D9C084EC9000F0353BAA5925" type="ORCID">https://orcid.org/0000-0001-5842-4066</mods:nameIdentifier>
<mods:affiliation id="3E31B0FE1CC032BEF8D096A371F10D29">Hatchodori 4 - 9, Toyohashi, Aichi 440 - 0806, Japan</mods:affiliation>
<mods:nameIdentifier id="2C398A1A5400F3FC8C976CA3ECB1EE6D" type="email">ezogengoroumodoki4.23@gmail.com</mods:nameIdentifier>
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<mods:namePart id="966ADB2A41B101026B4367A020120746">Watanabe, Kyohei</mods:namePart>
<mods:affiliation id="E4BE5A19509FAC0DB0EE9D0848FF091A">Kanagawa Prefectural Museum of Natural History, Iryuda 499, Odawara, Kanagawa 250 - 0031, Japan</mods:affiliation>
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<mods:title id="52C58633ADA275C19A18EF21A3472FB8">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="7349C8D0A367C61EDF5C3992F6CAEF6A">2022</mods:date>
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<mods:number id="E011ECC52256F0438AF7120ED627EB2B">69</mods:number>
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<mods:identifier id="0562BE981E643573916B28A3C3F94DEF" type="DOI">http://dx.doi.org/10.3897/dez.69.80492</mods:identifier>
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<treatment id="8DDA9D88629C59378D2AAEA99213B44E" ID-GBIF-Taxon="194444360" LSID="urn:lsid:zoobank.org:act:E030CAC0-73B1-4BDC-BD63-F5E6BD3BAC4B" httpUri="http://treatment.plazi.org/id/8DDA9D88629C59378D2AAEA99213B44E" lastPageNumber="45" pageId="0" pageNumber="45" scope_family="Ichneumonidae" scope_order="Hymenoptera" scope_subFamily="Diplazontinae">
<subSubSection id="2CF0D15E923A00FC90CAB6D565361623" pageId="0" pageNumber="45" type="nomenclature">
<paragraph id="C333924FDB6D5CEE82EB183D462E0981" pageId="0" pageNumber="45">
<taxonomicName id="044F0B4AC71F99367B36C1E0A89F3786" LSID="http://zoobank.org/E030CAC0-73B1-4BDC-BD63-F5E6BD3BAC4B" authority="Morishita &amp; Watanabe, 2022" authorityName="Morishita &amp; Watanabe" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Woldstedtius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Woldstedtius alpicola" order="Hymenoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="alpicola" status="sp. nov.">Woldstedtius alpicola</taxonomicName>
<taxonomicNameLabel id="E5335FD82D94CDFAEE85E88DCA581740" pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="4A098A5F85DCAB9E79A0A4CCA24B353F" pageId="0" pageNumber="45" type="description">
<paragraph id="5B26AEBABB01D3C8BBB24FE54474DF09" pageId="0" pageNumber="45">
<figureCitation id="71381DF8EF11384CE6BB1C6D5B0A0CFB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">Figs 1A-F</figureCitation>
<figureCitation id="E6304729044924DA965BCC73D53C6223" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Japanese Woldstedtius A, G. W. f. flavolineatus (Gravenhorst, 1829); B, I. W. kuroashii (Uchida, 1957); C, D, K. W. takagii (Uchida, 1957); E. W. alpicola sp. nov.; F. W. biguttatus (Gravenhorst, 1829); H. W. karafutensis (Uchida, 1957); J. W. punctatus sp. nov. (E, J. holotype; B, I. paratype) - A, B. head, lateral view; C. head, frontal view; D. propodeum, lateral view; E-K. T I, dorsal view." figureDoi="10.3897/dez.69.80492.figure9" httpUri="https://binary.pensoft.net/fig/667929" pageId="0" pageNumber="45">, 9E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="AAF5E47B2DBFDD00E056300953BF8BFA" pageId="0" pageNumber="45" type="type series">
<paragraph id="568A996CA19980689872A6D983840827" pageId="0" pageNumber="45">Type series.</paragraph>
<paragraph id="4437259E288830F96DB7ED5261CD2A57" pageId="0" pageNumber="45">
<emphasis id="0FB825B403FE8635E618F1686CDAB2A7" bold="true" italics="true" pageId="0" pageNumber="45">Holotype</emphasis>
: F, Japan, Honshu, Mie Pref., Inabe City, Mt. Fujiwaradake, 1 Jun 2021, S. Morishita leg. (KPMNH).
<emphasis id="2B487FF19EE1AE0DCECE84CCB336AD0F" bold="true" italics="true" pageId="0" pageNumber="45">Paratypes</emphasis>
: Japan: [Honshu] 1 F, Yamanashi Pref., Koshu City, Mt. Daibosatsu, Kaminikkawa-toge, 16 Jun 2007, K. Watanabe leg. (TMNH); 1 M, Kanagawa Pref., Yamakita Town, Mt. Oomuroyama-Mt. Kanyuudousan, 15 Jun 2008, H. Kawai leg. (KPMNH); 1 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, 13-25 Jul 2015, S. Shimizu leg. (MT) (KPMNH).
</paragraph>
<caption id="3091CC37909D2BA8FDB84A27D940EDB2" doi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45" start="Figure 1" startId="F1">
<paragraph id="DA0C196E30A365C2F5B2C0CE4AD40792" pageId="0" pageNumber="45">
<emphasis id="650EDBCE832075FA1F9AF17FA42080CB" bold="true" pageId="0" pageNumber="45">Figure 1.</emphasis>
<taxonomicName id="8A15BA0A83DC76A40205869CF49B5543" authorityName="Morishita &amp; Watanabe" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Woldstedtius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Woldstedtius alpicola" order="Hymenoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="7F77984501001AD3248B2DD68BD60C0A" italics="true" pageId="0" pageNumber="45">Woldstedtius alpicola</emphasis>
</taxonomicName>
sp. nov. (
<emphasis id="B37379CA4F3C7A0CFCCCDA867F458EAB" bold="true" pageId="0" pageNumber="45">A-D.</emphasis>
female, holotype;
<emphasis id="350B6CC6D77CA590811963BAA0AFE5F2" bold="true" pageId="0" pageNumber="45">E, F.</emphasis>
male, paratype) -
<emphasis id="70D023CADA827B72AF853BF149DC93AA" bold="true" pageId="0" pageNumber="45">A, E.</emphasis>
Habitus;
<emphasis id="84E4571DC140970D921B63FC262742E3" bold="true" pageId="0" pageNumber="45">B, F.</emphasis>
Head, frontal view;
<emphasis id="EE7FD66EFBEF35CCE0CDB206EDE7C926" bold="true" pageId="0" pageNumber="45">C.</emphasis>
Mesonotum, dorsal view;
<emphasis id="8CA9751CED4FE5D137FE77D8B109B4C9" bold="true" pageId="0" pageNumber="45">D.</emphasis>
Propodeum, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4EAC9F1062DD51D01DDF9F90E434DA37" pageId="0" pageNumber="45" type="description">
<paragraph id="D0B97FF2A1EA924886A595D86F154A9F" pageId="0" pageNumber="45">Description.</paragraph>
<paragraph id="3E75331848FA72B0B10B68F565CB48AA" pageId="0" pageNumber="45">Female (n = 3). Body length 8.5-9.2 (HT: 8.5) mm, polished, coriaceous and covered with silver setae.</paragraph>
<paragraph id="5069467723E0A74745A45C3BAA89020F" pageId="0" pageNumber="45">
<emphasis id="1E96F771986B06999CE5CFBB6FEE0F34" bold="true" italics="true" pageId="0" pageNumber="45">Head</emphasis>
0.51-0.54 (HT: 0.53)
<normalizedToken id="8AE3F8F34B961D4CD16C81AEB17A04BB" originalValue="×">x</normalizedToken>
as long as wide. Clypeus 2.0-2.35 (HT: 2.0)
<normalizedToken id="139771E7A10A35BA478F5D568C402BAA" originalValue="×">x</normalizedToken>
as broad as high, slightly convex basally in lateral view. Face 2.1-2.4 (HT: 2.4)
<normalizedToken id="976D5C199E51BA6ABC250E141D4A7912" originalValue="×">x</normalizedToken>
as broad as high, densely punctate, convex medially in lateral view and separated from clypeus by shallow clypeal sulcus. Inner orbits almost parallel (Fig.
<figureCitation id="E11E198B1F226E5409CF73CDF47C68FE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1B</figureCitation>
). Length of malar space 0.9-1.15 (HT: 0.9)
<normalizedToken id="85C76DDB58DFF1F0B2EC4814A4896A7A" originalValue="×">x</normalizedToken>
as long as basal mandibular width. POL 1.9-2.1 (HT: 1.9)
<normalizedToken id="16BFCECD44C7EEEA447DB5135B072633" originalValue="×">x</normalizedToken>
as long as OD. OOL 0.9-1.18 (HT: 0.9)
<normalizedToken id="962EAAE4A909AA640BE11C401E8039B9" originalValue="×">x</normalizedToken>
as long as OD. POL 1.86-1.95 (HT: 1.86)
<normalizedToken id="A5C32C19218FF378080CF5752FFB03DC" originalValue="×">x</normalizedToken>
as long as OOL. Antenna with 24-25 (HT: 24) flagellomeres. FL I 1.29-1.35 (HT: 1.35)
<normalizedToken id="4826DE34124C11DB28DCF064578A8296" originalValue="×">x</normalizedToken>
as long as FL II. MP IV1.42-1.53 (HT: 1.53)
<normalizedToken id="0C6ABC6B16572F6CD86DB8821AA196B8" originalValue="×">x</normalizedToken>
as long as MP V.
</paragraph>
<paragraph id="E3C296E158477569A2B46F92295EA043" pageId="0" pageNumber="45">
<emphasis id="094022D984668D0462D173833BD4DF9B" bold="true" pageId="0" pageNumber="45">
<emphasis id="D1D269CAFBA3CCCFC22DA949317EED01" italics="true" pageId="0" pageNumber="45">Mesosoma</emphasis>
.
</emphasis>
Lateral aspect of pronotum strigose anteriorly. Mesoscutum finely and densely punctate (separated by ca.1.0
<normalizedToken id="9E31ACBF450A5AEB52E9C6C435A831BD" originalValue="×">x</normalizedToken>
their diameter) (Fig.
<figureCitation id="7A32AFDAF59511169ECA6377BD290065" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1C</figureCitation>
). Scutellum finely and sparsely punctate (separated by ca.1.5-2.5
<normalizedToken id="5454F9529E613C7A0825628934B74C7A" originalValue="×">x</normalizedToken>
their diameter) (Fig.
<figureCitation id="8959B2B674F03F4D966E72C6526C7A41" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1C</figureCitation>
). Anterior part and lower half of mesopleuron coarsely and densely punctate. Speculum smooth. Sternaulus weakly impressed. Propodeum finely rugulose (Fig.
<figureCitation id="0226339DBF14074872F01710F2475A34" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1D</figureCitation>
), rounded in lateral view and pleural carina entirely present. Fore wing length 7.4-8.5 (HT: 7.4) mm. Nervellus intercepted below middle. Hind femur 4.7-5.3 (HT: 4.7)
<normalizedToken id="15026C200F3C629393A60B2DDAA1F782" originalValue="×">x</normalizedToken>
as long as maximum depth in lateral view. Hind tibia 7.5-8.0 (HT: 8.0)
<normalizedToken id="0B810DCE4534647C10EA643400C05059" originalValue="×">x</normalizedToken>
as long as maximum depth in lateral view. Ratio of length of hind first to fifth tarsomeres 1.0: 0.6: 0.4: 0.2: 0.2-0.3 (HT: 0.2).
</paragraph>
<paragraph id="4A7436B34E7FE481182136264660EB28" pageId="0" pageNumber="45">
<emphasis id="CE50F0D5467788DA6BAAF2DDEBD3A483" bold="true" pageId="0" pageNumber="45">
<emphasis id="C34491582D68C8A0A07EF6CF41BFD264" italics="true" pageId="0" pageNumber="45">Metasoma</emphasis>
.
</emphasis>
T I rectangular in dorsal view (Fig.
<figureCitation id="350791681AD895E228B9C1B1AAF230F4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Japanese Woldstedtius A, G. W. f. flavolineatus (Gravenhorst, 1829); B, I. W. kuroashii (Uchida, 1957); C, D, K. W. takagii (Uchida, 1957); E. W. alpicola sp. nov.; F. W. biguttatus (Gravenhorst, 1829); H. W. karafutensis (Uchida, 1957); J. W. punctatus sp. nov. (E, J. holotype; B, I. paratype) - A, B. head, lateral view; C. head, frontal view; D. propodeum, lateral view; E-K. T I, dorsal view." figureDoi="10.3897/dez.69.80492.figure9" httpUri="https://binary.pensoft.net/fig/667929" pageId="0" pageNumber="45">9E</figureCitation>
), 1.4-1.5 (HT: 1.4)
<normalizedToken id="39232C88F920F57DB3116608BD23A8F1" originalValue="×">x</normalizedToken>
as long as maximum width, irregular rugulose laterally, sometimes striate between latero-median carinae. Latero-median carina present basal ca. 0.5 of T I (Fig.
<figureCitation id="B1EEF6F994DA973D51FD194FF5664941" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Japanese Woldstedtius A, G. W. f. flavolineatus (Gravenhorst, 1829); B, I. W. kuroashii (Uchida, 1957); C, D, K. W. takagii (Uchida, 1957); E. W. alpicola sp. nov.; F. W. biguttatus (Gravenhorst, 1829); H. W. karafutensis (Uchida, 1957); J. W. punctatus sp. nov. (E, J. holotype; B, I. paratype) - A, B. head, lateral view; C. head, frontal view; D. propodeum, lateral view; E-K. T I, dorsal view." figureDoi="10.3897/dez.69.80492.figure9" httpUri="https://binary.pensoft.net/fig/667929" pageId="0" pageNumber="45">9E</figureCitation>
). T II 0.6-0.73 (HT: 0.73)
<normalizedToken id="E01A3C196E866A5E903805A3E809B8F1" originalValue="×">x</normalizedToken>
as long as maximum width, striate anteriorly and strigose laterally.
</paragraph>
<paragraph id="82F50582106A6CC639B9457C85750839" pageId="0" pageNumber="45">
<emphasis id="583F2AE06726C3B257DD1B80F5EBBAA7" bold="true" pageId="0" pageNumber="45">
<emphasis id="B86093329C37EFD557EEFCF2196B5361" italics="true" pageId="0" pageNumber="45">Coloration</emphasis>
(Fig.
<figureCitation id="4DD6B284F979E0305980D5E8ACC6A2F6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1A-D</figureCitation>
).
</emphasis>
Body (excluding wings and legs) black. Face with a large yellow spot medially. Palpi, subtegular ridge and mesepisternum yellow. Mandible yellow, except for apex and base. Lateral aspect of pronotum with a yellow spot posteriorly. Mesoscutum with yellow shoulder marks. Tegula tinged with yellow anteriorly. Scutellum with a yellow spot apically. Wings hyaline. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Legs blackish-brown to black. Fore trochanter, trochantellus, apex of hind femur and base of hind tibia tinged with white. Fore femur, tibia and tarsomeres orange to brown. Apex of hind trochantellus and base of hind femur tinged with reddish-brown.
</paragraph>
<paragraph id="637D3F88E51AEA0BFA74D6F3952FF28B" pageId="0" pageNumber="45">
<emphasis id="7DE98ECFCF51146FBEA11F63387E45CA" bold="true" pageId="0" pageNumber="45">Male (n = 1).</emphasis>
Similar to female. Body length (excluding antennae) 8.1 mm. Face 2.5
<normalizedToken id="242454AEA59520DE822ABDBA8CDD42C3" originalValue="×">x</normalizedToken>
as broad as high. Length of malar space 0.8
<normalizedToken id="025491FE2454AF3AD73C643D6E0F3086" originalValue="×">x</normalizedToken>
as long as basal mandibular width. POL 2.2
<normalizedToken id="EE6558C04B34BFFA4F697F368682CF32" originalValue="×">x</normalizedToken>
as long as OD. OOL 1.2
<normalizedToken id="F95D94A8E02F04888B15DC5A5E6878B2" originalValue="×">x</normalizedToken>
as long as OD. POL 2.1
<normalizedToken id="62705559DB15B752A20386E81CCBAFB5" originalValue="×">x</normalizedToken>
as long as OOL. Inner orbits weakly divergent downward (Fig.
<figureCitation id="04E6FA323EE643EF81B0CCAF58E461F7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1F</figureCitation>
). Fore wing length 6.3 mm.
</paragraph>
<paragraph id="5D9F045C529CE508A153EC0FE0BE9AA2" pageId="0" pageNumber="45">
<emphasis id="AB1634A604C679A277279B9A366D4E83" bold="true" pageId="0" pageNumber="45">
<emphasis id="A7F93BC11D92BC6A8AE0407F0364F16E" italics="true" pageId="0" pageNumber="45">Coloration</emphasis>
(Fig.
<figureCitation id="B200CDCB07CFE3079E9B181FD4D07A84" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1E, F</figureCitation>
).
</emphasis>
Body (excluding wings and legs) black. Clypeus, palpi, face, ventral surface of antenna, malar space, propleuron, epicnemium, mesosternum, tegula, subtegular ridge and mesepisternum yellow. Gena tinged with yellow ventrally. Mandible yellow, except for apex. Lateral aspect of pronotum tinged with yellow ventrally and posteriorly. Mesopleuron with a large yellow marking, it enlarged anteriorly. Scutellum with a yellow spot apically. T IV to T VII with a transverse yellow band posteriorly. Wings hyaline. Veins and pterostigma brown to blackish-brown except for yellow wing base. Legs yellow to yellowish-brown. Hind coxa and trochanter each with a blackish-brown stripe dorsally. Hind trochantellus and tibia tinged with blackish-brown.
</paragraph>
</subSubSection>
<subSubSection id="2C55C2DCAB43E67A586D922D9F830FF8" pageId="0" pageNumber="45" type="distribution">
<paragraph id="D3BD7B2406C89F13D586D211FC15D40E" pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph id="7C7F381F83D6EFD6E4856E33E357C93D" pageId="0" pageNumber="45">Japan (Honshu).</paragraph>
</subSubSection>
<subSubSection id="F7421883E4213342FDE103EECE91DD90" pageId="0" pageNumber="45" type="bionomics">
<paragraph id="8758A8E7397C99BC52677F4A622C4FA1" pageId="0" pageNumber="45">Bionomics.</paragraph>
<paragraph id="6A3887428F24302F714605D334AFC9E4" pageId="0" pageNumber="45">Host unknown. Adults were collected in broad-leaved forest at altitudes of ca. 1,000-2,000 meters.</paragraph>
</subSubSection>
<subSubSection id="6D176BD0CB1CD390182F4C06D742C30A" pageId="0" pageNumber="45" type="etymology">
<paragraph id="72E205F4D5566C52B962BD490020ECB9" pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph id="1A13801641AC6B0E24D6465F9A97BBE1" pageId="0" pageNumber="45">The species name refers that this species inhabits alpine region.</paragraph>
</subSubSection>
<subSubSection id="671229AD910CA5DC8ABAC2C58B8E6065" pageId="0" pageNumber="45" type="remarks">
<paragraph id="752ABAEE9D41641BBF0205762CB2D3F1" pageId="0" pageNumber="45">Remarks.</paragraph>
<paragraph id="B53BE1BCEA3B9CBC3EC835F5DBB6021A" pageId="0" pageNumber="45">
This species resembles
<taxonomicName id="8C902EF39E6FF5CD4A6979CE86FD257F" lsidName="W. kuroashii" pageId="0" pageNumber="45" rank="species" species="kuroashii">
<emphasis id="2EB74BED66D7293AF9A9C2A57E576E35" italics="true" pageId="0" pageNumber="45">W. kuroashii</emphasis>
</taxonomicName>
but can be distinguished from the latter by the following combination of character states: pleural carina of propodeum entirely present in both sexes (absent posteriorly in both sexes of
<taxonomicName id="49AE9FEEC480754D99AC16626367118D" lsidName="W. kuroashii" pageId="0" pageNumber="45" rank="species" species="kuroashii">
<emphasis id="53F5F60CE7EC9A12967F14EB8E9C51FB" italics="true" pageId="0" pageNumber="45">W. kuroashii</emphasis>
</taxonomicName>
); propodeum finely rugulose in both sexes (coriaceous in both sexes of
<taxonomicName id="A21B233C754244FB5044C51BCB7EAD3A" lsidName="W. kuroashii" pageId="0" pageNumber="45" rank="species" species="kuroashii">
<emphasis id="67BAFB5E4A0E77F91904B1E45D1D380F" italics="true" pageId="0" pageNumber="45">W. kuroashii</emphasis>
</taxonomicName>
); T I 1.4-1.5
<normalizedToken id="D863CD0084C2BC7F1263579B7A186A6F" originalValue="×">x</normalizedToken>
as long as maximum width in both sexes (1.1-1.25 in females, 1.14-1.26 in males of
<taxonomicName id="9B2FFFB2FE0EC58D36F86B45ED6DD063" lsidName="W. kuroashii" pageId="0" pageNumber="45" rank="species" species="kuroashii">
<emphasis id="86E69A52EFE15860E848017D0E06636F" italics="true" pageId="0" pageNumber="45">W. kuroashii</emphasis>
</taxonomicName>
). This species also resembles a Korean species,
<taxonomicName id="ADE83FC60B2595965F2C4F736CFA79FB" lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="620802051DCDFB485B1B0F2B16DA0EDC" italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
Balueva &amp; Lee, 2016 (male is unknown), but can be distinguished by the following combination of character states in females: a large yellow median spot of face present (absent in
<taxonomicName id="BDE0760AB82F98414F7C7312B2B540E9" lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="7716027AEB5083C99368CB40BA666B7C" italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
); antenna with 24-25 flagellomeres (22-23 in
<taxonomicName id="154FA29BADD0851A85571CF8181EDDED" lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="B02BBDC9A143198C0E8C5D3DC1A59880" italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
); scutellum finely and sparsely punctate on coriaceous background (entirely coriaceous in
<taxonomicName id="E5FAB98ECC2B80CBCB9714F770D34E81" lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="E73CDA52052673556AB20722F1A913BE" italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
); propodeum finely rugulose (coriaceous in
<taxonomicName id="CF384A6AE26D3EE1E61D107170B810BB" lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="CED4B5EE1169CA7EE12C19C0BE8D235C" italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>