treatments-xml/data/7C/3A/87/7C3A878E451A5A32FF74F921FACC366C.xml

<document id="A7CF5C28517CE3A022764AE3E257F980" ID-CLB-Dataset="31378" ID-DOI="10.11646/zootaxa.4350.3.1" ID-GBIF-Dataset="25b07a0d-7a27-460e-8813-713dbfb1888c" ID-ISSN="1175-5326" ID-Zenodo-Dep="1059179" ID-ZooBank="A3F42509-91EA-48BC-8C9D-253F5C57F20D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511167096685" checkinUser="plazi" docAuthor="Kluge, Nikita J., Gattolliat, Jean-Luc &amp; Salles, Frederico F." docDate="2017" docId="7C3A878E451A5A32FF74F921FACC366C" docLanguage="en" docName="zootaxa.4350.3.1.pdf" docOrigin="Zootaxa 4350 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crassabwa ludmilae Kluge &amp; Gattolliat &amp; Salles 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="428" masterDocId="8003FFF645085A29FFE3FF8CFFE83041" masterDocTitle="Redescription of the Afrotropical genus Crassabwa Lugo-Ortiz &amp; McCafferty 1996 (Ephemeroptera: Baetidae: Protopatellata)" masterLastPageNumber="435" masterPageNumber="401" pageNumber="419" updateTime="1698501443309" updateUser="ExternalLinkService">
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<mods:title id="724D365089F6481DEDAF15AE7B849F22">Redescription of the Afrotropical genus Crassabwa Lugo-Ortiz &amp; McCafferty 1996 (Ephemeroptera: Baetidae: Protopatellata)</mods:title>
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<mods:namePart id="6539ED63ADB17FD8BBEDBF23A76FD2EF">Kluge, Nikita J.</mods:namePart>
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<mods:namePart id="78B2BA0BB60281D26F99AF6520518352">Gattolliat, Jean-Luc</mods:namePart>
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<treatment id="7C3A878E451A5A32FF74F921FACC366C" ID-DOI="http://doi.org/10.5281/zenodo.6045787" ID-GBIF-Taxon="137453637" ID-Zenodo-Dep="6045787" LSID="urn:lsid:plazi:treatment:7C3A878E451A5A32FF74F921FACC366C" httpUri="http://treatment.plazi.org/id/7C3A878E451A5A32FF74F921FACC366C" lastPageId="27" lastPageNumber="428" pageId="18" pageNumber="419">
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<paragraph id="F42C3698451A5A3BFF74F921FE3D3689" blockId="18.[151,469,1709,1769]" box="[151,469,1709,1736]" pageId="18" pageNumber="419">
<heading id="AF6481F4451A5A3BFF74F921FE3D3689" bold="true" box="[151,469,1709,1736]" fontSize="11" level="1" pageId="18" pageNumber="419" reason="1">
<emphasis id="C6E7EA8A451A5A3BFF74F921FE3D3689" bold="true" box="[151,469,1709,1736]" pageId="18" pageNumber="419">
<taxonomicName id="33934D1B451A5A3BFF74F921FE633686" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[151,395,1709,1735]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A451A5A3BFF74F921FE633686" bold="true" box="[151,395,1709,1735]" italics="true" pageId="18" pageNumber="419">Crassabwa ludmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDD457F1451A5A3BFE70F922FE3D3689" box="[403,469,1710,1736]" pageId="18" pageNumber="419" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F42C3698451A5A3BFF74F95CFEDC36A8" blockId="18.[151,469,1709,1769]" box="[151,308,1744,1769]" pageId="18" pageNumber="419">
(
<figureCitation id="6CA82A1D451A5A3BFF7CF95CFEC336A8" box="[159,299,1744,1769]" captionStart-0="FIGURES 51 – 55" captionStart-1="FIGURES 56 – 59" captionStart-2="FIGURES 60 – 65" captionStart-3="FIGURES 66 – 75" captionStart-4="FIGURES 76 – 83" captionStart-5="FIGURES 84 – 95" captionStart-6="FIGURES 96 – 103" captionStart-7="FIGURES 104 – 107" captionStart-8="FIGURES 108 – 113" captionStartId-0="18.[151,264,1576,1599]" captionStartId-1="19.[151,264,1957,1980]" captionStartId-2="20.[151,264,1865,1888]" captionStartId-3="21.[151,264,1855,1878]" captionStartId-4="22.[151,264,1646,1669]" captionStartId-5="23.[151,264,1927,1950]" captionStartId-6="24.[151,264,1889,1912]" captionStartId-7="25.[151,264,1858,1881]" captionStartId-8="26.[151,264,1682,1705]" captionTargetBox-0="[163,1430,650,1552]" captionTargetBox-1="[151,1436,301,1936]" captionTargetBox-2="[157,1426,193,1824]" captionTargetBox-3="[186,1400,193,1812]" captionTargetBox-4="[154,1436,193,1625]" captionTargetBox-5="[155,1436,193,1897]" captionTargetBox-6="[163,1407,193,1851]" captionTargetBox-7="[161,1413,204,1831]" captionTargetBox-8="[151,1436,193,1653]" captionTargetId-0="figure@18.[151,1436,628,1555]" captionTargetId-1="figure@19.[151,1436,301,1936]" captionTargetId-2="figure@20.[151,1436,193,1844]" captionTargetId-3="figure@21.[170,1417,193,1835]" captionTargetId-4="figure@22.[151,1436,193,1625]" captionTargetId-5="figure@23.[151,1436,193,1907]" captionTargetId-6="figure@24.[163,1407,193,1851]" captionTargetId-7="figure@25.[160,1420,193,1835]" captionTargetId-8="figure@26.[151,1436,193,1653]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionTargetPageId-3="21" captionTargetPageId-4="22" captionTargetPageId-5="23" captionTargetPageId-6="24" captionTargetPageId-7="25" captionTargetPageId-8="26" captionText-0="FIGURES 51 – 55. Wings. 51 – 53, Crassabwa flava, hind wings of three individuals; 54 – 55, Crassabwa ludmilae sp. n., fore and hind wings on one scale." captionText-1="FIGURES 56 – 59. Crassabwa ludmilae sp. n., larvae. 56 – 57, female ultimolarvae; 58 – 59, exuviae of half of pronotum and mesonotum and of abdomen of ultimolarva ready to molt to subimago (with subimaginal right cercus inside)." captionText-2="FIGURES 60 – 65. Crassabwa ludmilae sp. n., holotype, ultimolarval exuviae. 60, abdominal terga and caudalii; 61, head with labrum and right mandible; 62, exuviae of half of pronotum and mesonotum; 63, fragment of cercus; 64, hind leg (triangle shows point where patella-tibial suture crosses inner margin); 65, abdominal terga IX and X." captionText-3="FIGURES 66 – 75. Crassabwa ludmilae sp. n., larva. 66, claw; 67, fore leg (anterior view); 68, paraproct; 69 – 70, right and left mandibles (ventral view); 71, labrum (at left ventral view, at right dorsal view); 72, maxilla; 73, labial palp (ventral view); 74, labium (at left ventral view, at right dorsal view); 75, hypopharynx and superlinguae." captionText-4="FIGURES 76 – 83. Crassabwa ludmilae sp. n. 76 – 77, right and left mandibles; 78 – 79, biting edges of left and right mandibles of penultimolarva with developing ultimolarval mandibles inside; 80 – 82, biting edges of left and right mandibles of ultimolarva; 83, exuviae of labium with focus on ventral side." captionText-5="FIGURES 84 – 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 – 91, tergalii II – VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 – 93 and 95, holotype)." captionText-6="FIGURES 96 – 103. Crassabwa ludmilae sp. n. 96 – 97, female imagoes; 98, postsubalar sclerite of subimaginal exuviae (arrow shows concavity of dorsal margin of posterior-dorsal process); 99, half of exuviae of subimaginal mesonotum; 100 – 101, male imago; 102, fragment of fore leg of male imago with apex of tibia and 1 st and 2 nd tarsomeres; 103, apex of tibia and tarsus of hind leg." captionText-7="FIGURES 104 – 107. Crassabwa ludmilae sp. n., male genitals. 104, subimaginal genitals developing under larval cuticle; 105, exuviae of subimaginal genitals; 106, genitals of male imago, ventral view (unistyligers and gonostyli unnaturally brought together because of wrong fixation; internal sclerites and muscles shown by interrupted lines, right gonostylar muscle not shown; large arrow shows median projection of dorsal wall of unistyliger); 107, penis enlarged, ventral view (gonoducts shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, places of attachment of muscles running to common base of paraprocts; pb, paired internal projection of penial bridge; usg, unistyliger." captionText-8="FIGURES 108 – 113. Crassabwa ludmilae sp. n., eggs extracted from holotype." httpUri-0="https://zenodo.org/record/1059201/files/figure.png" httpUri-1="https://zenodo.org/record/1059203/files/figure.png" httpUri-2="https://zenodo.org/record/1059205/files/figure.png" httpUri-3="https://zenodo.org/record/1059207/files/figure.png" httpUri-4="https://zenodo.org/record/1059209/files/figure.png" httpUri-5="https://zenodo.org/record/1059211/files/figure.png" httpUri-6="https://zenodo.org/record/1059213/files/figure.png" httpUri-7="https://zenodo.org/record/1059215/files/figure.png" httpUri-8="https://zenodo.org/record/1059217/files/figure.png" pageId="18" pageNumber="419">Figs 54–113</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BC896513451A5A3BFF74F894FABD3770" box="[151,1365,1816,1841]" pageId="18" pageNumber="419" type="etymology">
<paragraph id="F42C3698451A5A3BFF74F894FABD3770" blockId="18.[151,1437,1816,2021]" box="[151,1365,1816,1841]" pageId="18" pageNumber="419">
<emphasis id="C6E7EA8A451A5A3BFF74F894FEF63770" bold="true" box="[151,286,1816,1841]" pageId="18" pageNumber="419">Etymology.</emphasis>
This species is named in honor of Ludmila Sheyko, who collected mature larvae of this species.
</paragraph>
</subSubSection>
<subSubSection id="BC896513451A5A3FFF24F8B0FAE037D6" lastPageId="22" lastPageNumber="423" pageId="18" pageNumber="419" type="materials_examined">
<paragraph id="F42C3698451A5A3BFF24F8B0FA7537A5" blockId="18.[151,1437,1816,2021]" pageId="18" pageNumber="419">
<materialsCitation id="44FB3CC5451A5A3BFF24F8B0FBFD3738" ID-GBIF-Occurrence="1701220171" collectingDate="2014-08-07" collectionCode="ZIN" collectorName="N. Kluge &amp; L. Sheyko" country="Zambia" county="Luangwa Bridge" latitude="-15.0" location="Material" longLatPrecision="1286" longitude="30.216667" municipality="Luangwa river" pageId="18" pageNumber="419" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="C6E7EA8A451A5A3BFF24F8B0FE5B3714" bold="true" box="[199,435,1852,1877]" pageId="18" pageNumber="419">
<location id="F14C6043451A5A3BFF24F8B0FEC73714" LSID="urn:lsid:plazi:treatment:7C3A878E451A5A32FF74F921FACC366C:F14C6043451A5A3BFF24F8B0FEC73714" box="[199,303,1852,1877]" country="Zambia" county="Luangwa Bridge" latitude="-15.0" longLatPrecision="1286" longitude="30.216667" municipality="Luangwa river" name="Material" pageId="18" pageNumber="419">Material</location>
examined.
</emphasis>
<typeStatus id="2B28883A451A5A3BFE5EF8B0FDC23714" box="[445,554,1852,1877]" pageId="18" pageNumber="419" type="holotype">Holotype</typeStatus>
: L-S-I 
<specimenCount id="E295FD11451A5A3BFD98F8B1FD613714" box="[635,649,1853,1877]" pageId="18" pageNumber="419" type="female">♀</specimenCount>
{specimen [V](9)A2014}, 
<collectingCountry id="8C847608451A5A3BFC2BF8B1FBD13714" box="[968,1081,1853,1877]" name="Zambia" pageId="18" pageNumber="419">ZAMBIA</collectingCountry>
, 
<collectingMunicipality id="1448ACE2451A5A3BFBA4F8B1FB073714" box="[1095,1263,1852,1877]" pageId="18" pageNumber="419">Luangwa river</collectingMunicipality>
near 
<collectingCounty id="1D4D4E14451A5A3BFAD7F8B1FF0E3738" pageId="18" pageNumber="419">Luangwa Bridge</collectingCounty>
, 
<geoCoordinate id="91A7505F451A5A3BFF15F8ECFEA63738" box="[246,334,1888,1913]" direction="south" orientation="latitude" pageId="18" pageNumber="419" precision="925" value="-15.0">15°00'S</geoCoordinate>
/ 
<geoCoordinate id="91A7505F451A5A3BFE8CF8EDFE233738" box="[367,459,1889,1913]" direction="east" orientation="longitude" pageId="18" pageNumber="419" precision="925" value="30.216667">30°13'E</geoCoordinate>
, 
<collectingDate id="9069E9B0451A5A3BFE3FF8EDFD8B3738" box="[476,611,1889,1913]" pageId="18" pageNumber="419" value="2014-08-07">
<date id="802D1058451A5A3BFE3FF8EDFD8B3738" box="[476,611,1889,1913]" pageId="18" pageNumber="419" value="2014-08-07">7.VIII.2014</date>
</collectingDate>
, coll. 
<collectorName id="5966534E451A5A3BFD52F8EDFCF43738" box="[689,796,1888,1913]" pageId="18" pageNumber="419">N. Kluge</collectorName>
&amp; 
<collectorName id="5966534E451A5A3BFCA8F8EDFC2C3738" box="[843,964,1888,1913]" pageId="18" pageNumber="419">L. Sheyko</collectorName>
(
<collectionCode id="9282AE5D451A5A3BFC3BF8EDFBE43738" box="[984,1036,1889,1913]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="18" pageNumber="419">ZIN</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="44FB3CC5451A5A3BFBC6F8EDFD1937DC" ID-GBIF-Occurrence="1701220153" collectingDate="2014-08-02" collectingDateMax="2014-08-08" collectingDateMin="2014-08-02" collectionCode="ZIN" collectorName="N. Kluge &amp; L. Sheyko" country="Zambia" county="Luangwa Bridge" latitude="-15.0" location="Material" longLatPrecision="1286" longitude="30.216667" municipality="Luangwa river" pageId="18" pageNumber="419" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="2B28883A451A5A3BFBC6F8EDFB7E3738" box="[1061,1174,1889,1913]" pageId="18" pageNumber="419" type="paratype">Paratypes</typeStatus>
: the same locality, 
<collectingDate id="9069E9B0451A5A3BFA63F8EDFEF537DC" pageId="18" pageNumber="419" value="2014-08-02" valueMax="2014-08-08" valueMin="2014-08-02">
<date id="802D1058451A5A3BFA63F8EDFEF537DC" pageId="18" pageNumber="419" value="2014-08-02" valueMax="2014-08-08" valueMin="2014-08-02">2– 8.VIII.2014</date>
</collectingDate>
, coll. 
<collectorName id="5966534E451A5A3BFE8DF809FE3437DC" box="[366,476,1924,1949]" pageId="18" pageNumber="419">N. Kluge</collectorName>
&amp; 
<collectorName id="5966534E451A5A3BFDEEF809FD6237DC" box="[525,650,1924,1949]" pageId="18" pageNumber="419">L. Sheyko</collectorName>
: 1 S-I 
<specimenCount id="E295FD11451A5A3BFD3EF809FD1937DC" box="[733,753,1925,1949]" pageId="18" pageNumber="419" type="male">♂</specimenCount>
</materialsCitation>
, 1 L-S-I♀, 
<date id="802D1058451A5A3BFC66F809FC4637DD" box="[901,942,1925,1949]" pageId="18" pageNumber="419">1 I</date>
♀, L/S♂, 3 larvae (ZIN). 
<materialsCitation id="44FB3CC5451A5A3BFB0AF809FA703780" ID-GBIF-Occurrence="1701220169" collectingDate="1985-03-02" collectionCode="MZL, CZNC" collectorName="M. Elouard" country="Niger" location="Loc. Sassambaya" municipality="Niandan river" pageId="18" pageNumber="419" specimenCount="9" specimenCount-larva="9" typeStatus="paratype">
GUINEA: 
<location id="F14C6043451A5A3BFA89F809FECB3780" LSID="urn:lsid:plazi:treatment:7C3A878E451A5A32FF74F921FACC366C:F14C6043451A5A3BFA89F809FECB3780" country="Niger" municipality="Niandan river" name="Loc. Sassambaya" pageId="18" pageNumber="419">Loc. Sassambaya</location>
, basin of 
<collectingCountry id="8C847608451A5A3BFE44F824FE013780" box="[423,489,1960,1985]" name="Niger" pageId="18" pageNumber="419">Niger</collectingCountry>
, 
<collectingMunicipality id="1448ACE2451A5A3BFE1FF824FD493780" box="[508,673,1960,1985]" pageId="18" pageNumber="419">Niandan river</collectingMunicipality>
, 
<collectingDate id="9069E9B0451A5A3BFD56F824FCDD3780" box="[693,821,1960,1985]" pageId="18" pageNumber="419" value="1985-03-02">
<date id="802D1058451A5A3BFD56F824FCDD3780" box="[693,821,1960,1985]" pageId="18" pageNumber="419" value="1985-03-02">02.III.1985</date>
</collectingDate>
, coll. J.- 
<collectorName id="5966534E451A5A3BFC40F825FBD93780" box="[931,1073,1960,1985]" pageId="18" pageNumber="419">M. Elouard</collectorName>
: 
<specimenCount id="E295FD11451A5A3BFBA7F825FB4E3780" box="[1092,1190,1960,1985]" pageId="18" pageNumber="419" type="larva">9 larvae</specimenCount>
(
<collectionCode id="9282AE5D451A5A3BFB5FF825FB113780" box="[1212,1273,1961,1985]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="18" pageNumber="419">MZL</collectionCode>
and 
<collectionCode id="9282AE5D451A5A3BFAA3F825FA783780" box="[1344,1424,1961,1985]" pageId="18" pageNumber="419">CZNC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="44FB3CC5451A5A3BFF74F840FA7537A5" ID-GBIF-Occurrence="1701220155" box="[151,1437,1996,2021]" collectingDate="1988-03-11" collectionCode="MZL" collectorName="M. Elouard &amp; Loc." country="Niger" location="Dion-Niger" municipality="Le Kouan river" pageId="18" pageNumber="419" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
Tambicola-Beyla, Loc. Beyla (
<location id="F14C6043451A5A3BFE16F840FD9437A4" LSID="urn:lsid:plazi:treatment:7C3A878E451A5A32FF74F921FACC366C:F14C6043451A5A3BFE16F840FD9437A4" box="[501,636,1996,2021]" country="Niger" municipality="Le Kouan river" name="Dion-Niger" pageId="18" pageNumber="419">Dion-Niger</location>
), 
<collectingMunicipality id="1448ACE2451A5A3BFD72F841FCAB37A4" box="[657,835,1996,2021]" pageId="18" pageNumber="419">Le Kouan river</collectingMunicipality>
, 
<collectingDate id="9069E9B0451A5A3BFCACF841FC2737A4" box="[847,975,1997,2021]" pageId="18" pageNumber="419" value="1988-03-11">
<date id="802D1058451A5A3BFCACF841FC2737A4" box="[847,975,1997,2021]" pageId="18" pageNumber="419" value="1988-03-11">11.III.1988</date>
</collectingDate>
, coll. J.- 
<collectorName id="5966534E451A5A3BFBD3F841FB5037A4" box="[1072,1208,1996,2021]" pageId="18" pageNumber="419">M. Elouard</collectorName>
: 
<specimenCount id="E295FD11451A5A3BFB26F841FB0437A5" box="[1221,1260,1997,2021]" pageId="18" pageNumber="419" type="female">1 ♀</specimenCount>
L (
<collectionCode id="9282AE5D451A5A3BFAF0F841FABC37A4" box="[1299,1364,1997,2021]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="18" pageNumber="419">MZL</collectionCode>
); 
<collectorName id="5966534E451A5A3BFA8AF841FA7537A5" box="[1385,1437,1997,2020]" pageId="18" pageNumber="419">Loc.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="F42C3698451B5A3AFF74FF1BFE3C30B9" blockId="19.[151,1436,151,248]" pageId="19" pageNumber="420">
<materialsCitation id="44FB3CC5451B5A3AFF74FF1BFB2830F1" ID-GBIF-Occurrence="1701220166" box="[151,1216,151,176]" collectingDate="1988-03-10" collectionCode="MZL" collectorName="M. Elouard" country="Senegal" location="Diomagoudou" municipality="Baoule river" pageId="19" pageNumber="420" specimenCount="1">
<location id="F14C6043451B5A3AFF74FF1BFED330F1" LSID="urn:lsid:plazi:treatment:7C3A878E451A5A32FF74F921FACC366C:F14C6043451B5A3AFF74FF1BFED330F1" box="[151,315,151,176]" country="Senegal" municipality="Baoule river" name="Diomagoudou" pageId="19" pageNumber="420">Diomagoudou</location>
; basin of 
<collectingCountry id="8C847608451B5A3AFE4BFF1BFDEB30F1" box="[424,515,151,176]" name="Senegal" pageId="19" pageNumber="420">Senegal</collectingCountry>
, 
<collectingMunicipality id="1448ACE2451B5A3AFDECFF1BFD7630F1" box="[527,670,151,176]" pageId="19" pageNumber="420">Baoulé river</collectingMunicipality>
, 
<collectingDate id="9069E9B0451B5A3AFD4AFF14FCC230F1" box="[681,810,152,176]" pageId="19" pageNumber="420" value="1988-03-10">
<date id="802D1058451B5A3AFD4AFF14FCC230F1" box="[681,810,152,176]" pageId="19" pageNumber="420" value="1988-03-10">10.III.1988</date>
</collectingDate>
, coll. J.- 
<collectorName id="5966534E451B5A3AFC6BFF14FBE630F1" box="[904,1038,151,176]" pageId="19" pageNumber="420">M. Elouard</collectorName>
: 1 larva (
<collectionCode id="9282AE5D451B5A3AFB94FF14FB5030F1" box="[1143,1208,152,176]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="19" pageNumber="420">MZL</collectionCode>
)
</materialsCitation>
. MALI: Koulikoro, Loc. Tienfala, 
<materialsCitation id="44FB3CC5451B5A3AFEAEFF30FB343095" ID-GBIF-Occurrence="1701220176" box="[333,1244,188,213]" collectingDate="1985-03-20" collectionCode="MZL, CZNC" collectorName="M. Elouard" country="Niger" location="Niger river" pageId="19" pageNumber="420" specimenCount="35" specimenCount-larva="35">
<location id="F14C6043451B5A3AFEAEFF30FE3A3094" LSID="urn:lsid:plazi:treatment:7C3A878E451A5A32FF74F921FACC366C:F14C6043451B5A3AFEAEFF30FE3A3094" box="[333,466,188,213]" country="Niger" name="Niger river" pageId="19" pageNumber="420">
<collectingCountry id="8C847608451B5A3AFEAEFF30FE653094" box="[333,397,188,213]" name="Niger" pageId="19" pageNumber="420">Niger</collectingCountry>
river
</location>
, 
<collectingDate id="9069E9B0451B5A3AFE04FF30FD8F3094" box="[487,615,188,213]" pageId="19" pageNumber="420" value="1985-03-20">
<date id="802D1058451B5A3AFE04FF30FD8F3094" box="[487,615,188,213]" pageId="19" pageNumber="420" value="1985-03-20">20.III.1985</date>
</collectingDate>
, coll. J.- 
<collectorName id="5966534E451B5A3AFD35FF31FC8C3094" box="[726,868,188,213]" pageId="19" pageNumber="420">M. Elouard</collectorName>
: 
<specimenCount id="E295FD11451B5A3AFC9BFF30FC013094" box="[888,1001,188,213]" pageId="19" pageNumber="420" type="larva">35 larvae</specimenCount>
(
<collectionCode id="9282AE5D451B5A3AFBE3FF30FBD53095" box="[1024,1085,188,212]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="19" pageNumber="420">MZL</collectionCode>
and 
<collectionCode id="9282AE5D451B5A3AFB66FF30FB3D3095" box="[1157,1237,188,212]" pageId="19" pageNumber="420">CZNC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="44FB3CC5451B5A3AFB13FF31FE3830B9" ID-GBIF-Occurrence="1701220177" collectingDate="1985-06-26" collectionCode="MZL" collectorName="M. Elouard" country="Niger" location="Niger river" pageId="19" pageNumber="420" specimenCount="8" specimenCount-larva="8">
same data but 
<collectingDate id="9069E9B0451B5A3AFF74FF53FEF430B9" box="[151,284,223,248]" pageId="19" pageNumber="420" value="1985-06-26">
<date id="802D1058451B5A3AFF74FF53FEF430B9" box="[151,284,223,248]" pageId="19" pageNumber="420" value="1985-06-26">26.VI.1985</date>
</collectingDate>
: 
<specimenCount id="E295FD11451B5A3AFEFCFF53FE9F30B9" box="[287,375,223,248]" pageId="19" pageNumber="420" type="larva">8 larvae</specimenCount>
(
<collectionCode id="9282AE5D451B5A3AFE64FF6CFE2030B9" box="[391,456,224,248]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="19" pageNumber="420">MZL</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="A0EC6610451B5A3AFF74F829FB02379B" httpUri="https://zenodo.org/record/1059203/files/figure.png" pageId="19" pageNumber="420" targetBox="[151,1436,301,1936]" targetPageId="19">
<paragraph id="F42C3698451B5A3AFF74F829FB02379B" blockId="19.[151,1435,1957,2010]" pageId="19" pageNumber="420">
<emphasis id="C6E7EA8A451B5A3AFF74F829FEBE37FA" bold="true" box="[151,342,1957,1980]" pageId="19" pageNumber="420">FIGURES 56–59.</emphasis>
<taxonomicName id="33934D1B451B5A3AFEBDF829FDD937FA" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[350,561,1957,1979]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A451B5A3AFEBDF829FDD937FA" box="[350,561,1957,1979]" italics="true" pageId="19" pageNumber="420">Crassabwa ludmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDD457F1451B5A3AFDD9F82AFD9D37FA" box="[570,629,1958,1979]" pageId="19" pageNumber="420" rank="species">
<emphasis id="C6E7EA8A451B5A3AFDD9F82AFD9D37FA" bold="true" box="[570,629,1958,1979]" pageId="19" pageNumber="420">sp. n.</emphasis>
</taxonomicNameLabel>
, larvae. 56–57, female ultimolarvae; 58–59, exuviae of half of pronotum and mesonotum and of abdomen of ultimolarva ready to molt to subimago (with subimaginal right cercus inside).
</paragraph>
</caption>
<caption id="A0EC6610451C5A3DFF74F8C5FBA937DC" httpUri="https://zenodo.org/record/1059205/files/figure.png" pageId="20" pageNumber="421" targetBox="[157,1426,193,1824]" targetPageId="20">
<paragraph id="F42C3698451C5A3DFF74F8C5FBA937DC" blockId="20.[151,1436,1865,1949]" pageId="20" pageNumber="421">
<emphasis id="C6E7EA8A451C5A3DFF74F8C5FEBC371E" bold="true" box="[151,340,1865,1888]" pageId="20" pageNumber="421">FIGURES 60–65.</emphasis>
<taxonomicName id="33934D1B451C5A3DFEB9F8C6FDC23721" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[346,554,1866,1888]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A451C5A3DFEB9F8C6FDC23721" box="[346,554,1866,1888]" italics="true" pageId="20" pageNumber="421">Crassabwa ludmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDD457F1451C5A3DFDD2F8C6FD82371E" box="[561,618,1866,1887]" pageId="20" pageNumber="421" rank="species">
<emphasis id="C6E7EA8A451C5A3DFDD2F8C6FD82371E" bold="true" box="[561,618,1866,1887]" pageId="20" pageNumber="421">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype, ultimolarval exuviae. 60, abdominal terga and caudalii; 61, head with labrum and right mandible; 62, exuviae of half of pronotum and mesonotum; 63, fragment of cercus; 64, hind leg (triangle shows point where patella-tibial suture crosses inner margin); 65, abdominal terga IX and X.
</paragraph>
</caption>
<caption id="A0EC6610451D5A3CFF74F8B3FC1D37D2" httpUri="https://zenodo.org/record/1059207/files/figure.png" pageId="21" pageNumber="422" targetBox="[186,1400,193,1812]" targetPageId="21">
<paragraph id="F42C3698451D5A3CFF74F8B3FC1D37D2" blockId="21.[151,1436,1855,1939]" pageId="21" pageNumber="422">
<emphasis id="C6E7EA8A451D5A3CFF74F8B3FEBC3714" bold="true" box="[151,340,1855,1878]" pageId="21" pageNumber="422">FIGURES 66–75.</emphasis>
<taxonomicName id="33934D1B451D5A3CFEB9F8CCFDC33717" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[346,555,1856,1878]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A451D5A3CFEB9F8CCFDC33717" box="[346,555,1856,1878]" italics="true" pageId="21" pageNumber="422">Crassabwa ludmilae</emphasis>
</taxonomicName>
<emphasis id="C6E7EA8A451D5A3CFDD2F8CCFD823714" bold="true" box="[561,618,1856,1877]" pageId="21" pageNumber="422">
<taxonomicNameLabel id="DDD457F1451D5A3CFDD2F8CCFD823714" box="[561,618,1856,1877]" pageId="21" pageNumber="422" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, larva. 66, claw; 67, fore leg (anterior view); 68, paraproct; 69–70, right and left mandibles (ventral view); 71, labrum (at left ventral view, at right dorsal view); 72, maxilla; 73, labial palp (ventral view); 74, labium (at left ventral view, at right dorsal view); 75, hypopharynx and superlinguae.
</paragraph>
</caption>
<caption id="A0EC6610451E5A3FFF74F9E2FCE03683" httpUri="https://zenodo.org/record/1059209/files/figure.png" pageId="22" pageNumber="423" targetBox="[154,1436,193,1625]" targetPageId="22">
<paragraph id="F42C3698451E5A3FFF74F9E2FCE03683" blockId="22.[151,1436,1646,1730]" pageId="22" pageNumber="423">
<emphasis id="C6E7EA8A451E5A3FFF74F9E2FEBC36C5" bold="true" box="[151,340,1646,1669]" pageId="22" pageNumber="423">FIGURES 76–83.</emphasis>
<taxonomicName id="33934D1B451E5A3FFEB9F9E3FDC336C4" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[346,555,1647,1669]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A451E5A3FFEB9F9E3FDC336C4" box="[346,555,1647,1669]" italics="true" pageId="22" pageNumber="423">Crassabwa ludmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDD457F1451E5A3FFDD1F9E3FD8336C5" box="[562,619,1647,1668]" pageId="22" pageNumber="423" rank="species">
<emphasis id="C6E7EA8A451E5A3FFDD1F9E3FD8336C5" bold="true" box="[562,619,1647,1668]" pageId="22" pageNumber="423">sp. n.</emphasis>
</taxonomicNameLabel>
76–77, right and left mandibles; 78–79, biting edges of left and right mandibles of penultimolarva with developing ultimolarval mandibles inside; 80–82, biting edges of left and right mandibles of ultimolarva; 83, exuviae of labium with focus on ventral side.
</paragraph>
</caption>
<paragraph id="F42C3698451E5A3FFF24F962FAE037D6" blockId="22.[151,1436,1774,2016]" pageId="22" pageNumber="423">
<emphasis id="C6E7EA8A451E5A3FFF24F962FE5E3746" bold="true" box="[199,438,1774,1799]" pageId="22" pageNumber="423">
Descriptions. 
<emphasis id="C6E7EA8A451E5A3FFE88F963FE473747" bold="true" box="[363,431,1775,1798]" italics="true" pageId="22" pageNumber="423">Larva</emphasis>
.
</emphasis>
CUTICULAR COLORATION. As characterized above [see 
<taxonomicName id="33934D1B451E5A3FFB8EF963FB033746" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[1133,1259,1775,1799]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A451E5A3FFB8EF963FB033746" box="[1133,1259,1775,1799]" italics="true" pageId="22" pageNumber="423">Crassabwa</emphasis>
</taxonomicName>
(1)]. Cuticle of fore protoptera in last larval instar with dark lines along all veins: convex longitudinal veins, concave longitudinal veins and cross veins (
<figureCitation id="6CA82A1D451E5A3FFE74F8BAFDF1370E" box="[407,537,1846,1871]" captionStart="FIGURES 56 – 59" captionStartId="19.[151,264,1957,1980]" captionTargetBox="[151,1436,301,1936]" captionTargetId="figure@19.[151,1436,301,1936]" captionTargetPageId="19" captionText="FIGURES 56 – 59. Crassabwa ludmilae sp. n., larvae. 56 – 57, female ultimolarvae; 58 – 59, exuviae of half of pronotum and mesonotum and of abdomen of ultimolarva ready to molt to subimago (with subimaginal right cercus inside)." httpUri="https://zenodo.org/record/1059203/files/figure.png" pageId="22" pageNumber="423">Figs 56–58</figureCitation>
, 
<figureCitation id="6CA82A1D451E5A3FFDC5F8BAFDAE370E" box="[550,582,1846,1871]" captionStart="FIGURES 60 – 65" captionStartId="20.[151,264,1865,1888]" captionTargetBox="[157,1426,193,1824]" captionTargetId="figure@20.[151,1436,193,1844]" captionTargetPageId="20" captionText="FIGURES 60 – 65. Crassabwa ludmilae sp. n., holotype, ultimolarval exuviae. 60, abdominal terga and caudalii; 61, head with labrum and right mandible; 62, exuviae of half of pronotum and mesonotum; 63, fragment of cercus; 64, hind leg (triangle shows point where patella-tibial suture crosses inner margin); 65, abdominal terga IX and X." httpUri="https://zenodo.org/record/1059205/files/figure.png" pageId="22" pageNumber="423">62</figureCitation>
); in previous larval instars, dark lines only along convex longitudinal veins (as in 
<figureCitation id="6CA82A1D451E5A3FFF38F8D7FE9E3732" box="[219,374,1883,1908]" captionStart="FIGURES 114 – 118" captionStartId="28.[151,264,1910,1933]" captionTargetBox="[151,1436,193,1889]" captionTargetId="figure@28.[151,1436,193,1889]" captionTargetPageId="28" captionText="FIGURES 114 – 118. Crassabwa ameliae sp. n. 114 – 116, last instar larvae; 117 – 118, larva of penultimate instar." httpUri="https://zenodo.org/record/1059219/files/figure.png" pageId="22" pageNumber="423">Figs 117–118</figureCitation>
). Coloration of thorax and abdomen varies from nearly uniform (
<figureCitation id="6CA82A1D451E5A3FFBAEF8D7FB4B3735" box="[1101,1187,1883,1908]" captionStart="FIGURES 56 – 59" captionStartId="19.[151,264,1957,1980]" captionTargetBox="[151,1436,301,1936]" captionTargetId="figure@19.[151,1436,301,1936]" captionTargetPageId="19" captionText="FIGURES 56 – 59. Crassabwa ludmilae sp. n., larvae. 56 – 57, female ultimolarvae; 58 – 59, exuviae of half of pronotum and mesonotum and of abdomen of ultimolarva ready to molt to subimago (with subimaginal right cercus inside)." httpUri="https://zenodo.org/record/1059203/files/figure.png" pageId="22" pageNumber="423">Figs 57</figureCitation>
, 
<figureCitation id="6CA82A1D451E5A3FFB4CF8D7FB243735" box="[1199,1228,1883,1908]" captionStart="FIGURES 60 – 65" captionStartId="20.[151,264,1865,1888]" captionTargetBox="[157,1426,193,1824]" captionTargetId="figure@20.[151,1436,193,1844]" captionTargetPageId="20" captionText="FIGURES 60 – 65. Crassabwa ludmilae sp. n., holotype, ultimolarval exuviae. 60, abdominal terga and caudalii; 61, head with labrum and right mandible; 62, exuviae of half of pronotum and mesonotum; 63, fragment of cercus; 64, hind leg (triangle shows point where patella-tibial suture crosses inner margin); 65, abdominal terga IX and X." httpUri="https://zenodo.org/record/1059205/files/figure.png" pageId="22" pageNumber="423">60</figureCitation>
) to light with very contrastingly darkened posterior part of mesonotum and abdominal terga III, VI and IX (
<figureCitation id="6CA82A1D451E5A3FFB9BF8F2FB1337D6" box="[1144,1275,1918,1943]" captionStart="FIGURES 56 – 59" captionStartId="19.[151,264,1957,1980]" captionTargetBox="[151,1436,301,1936]" captionTargetId="figure@19.[151,1436,301,1936]" captionTargetPageId="19" captionText="FIGURES 56 – 59. Crassabwa ludmilae sp. n., larvae. 56 – 57, female ultimolarvae; 58 – 59, exuviae of half of pronotum and mesonotum and of abdomen of ultimolarva ready to molt to subimago (with subimaginal right cercus inside)." httpUri="https://zenodo.org/record/1059203/files/figure.png" pageId="22" pageNumber="423">Figs 56, 59</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BC896513451E5A32FF24F82FFACC366C" lastPageId="27" lastPageNumber="428" pageId="22" pageNumber="423" type="description">
<paragraph id="F42C3698451E5A3FFF24F82FFAB7379E" blockId="22.[151,1436,1774,2016]" pageId="22" pageNumber="423">
HYPODERMAL COLORATION. As characterized above [see 
<taxonomicName id="33934D1B451E5A3FFC4CF82FFBC537FA" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[943,1069,1955,1979]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A451E5A3FFC4CF82FFBC537FA" box="[943,1069,1955,1979]" italics="true" pageId="22" pageNumber="423">Crassabwa</emphasis>
</taxonomicName>
(2)]. Before molt to subimago larva gets hypodermal coloration as in imago, including red coloration of proximal parts of cerci (see below).
</paragraph>
<caption id="A0EC6610451F5A3EFF74F80BFC38379A" httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="23" pageNumber="424" targetBox="[155,1436,193,1897]" targetPageId="23">
<paragraph id="F42C3698451F5A3EFF74F80BFC38379A" blockId="23.[151,1436,1927,2011]" pageId="23" pageNumber="424">
<emphasis id="C6E7EA8A451F5A3EFF74F80BFEBD37DC" bold="true" box="[151,341,1927,1950]" pageId="23" pageNumber="424">FIGURES 84–95.</emphasis>
<taxonomicName id="33934D1B451F5A3EFEBFF804FDC637DF" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[348,558,1928,1950]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A451F5A3EFEBFF804FDC637DF" box="[348,558,1928,1950]" italics="true" pageId="23" pageNumber="424">Crassabwa ludmilae</emphasis>
</taxonomicName>
<emphasis id="C6E7EA8A451F5A3EFDD6F804FD8737DC" bold="true" box="[565,623,1928,1949]" pageId="23" pageNumber="424">
<taxonomicNameLabel id="DDD457F1451F5A3EFDD6F804FD8737DC" box="[565,623,1928,1949]" pageId="23" pageNumber="424" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86–91, tergalii II–VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84–93 and 95, holotype).
</paragraph>
</caption>
<caption id="A0EC661045105A31FF74F8EDFF183795" httpUri="https://zenodo.org/record/1059213/files/figure.png" pageId="24" pageNumber="425" targetBox="[163,1407,193,1851]" targetPageId="24">
<paragraph id="F42C369845105A31FF74F8EDFF183795" blockId="24.[151,1436,1889,2004]" pageId="24" pageNumber="425">
<emphasis id="C6E7EA8A45105A31FF74F8EDFE883736" bold="true" box="[151,352,1889,1912]" pageId="24" pageNumber="425">FIGURES 96–103.</emphasis>
<taxonomicName id="33934D1B45105A31FE85F8EEFDD03739" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[358,568,1890,1912]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="425" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A45105A31FE85F8EEFDD03739" box="[358,568,1890,1912]" italics="true" pageId="24" pageNumber="425">Crassabwa ludmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDD457F145105A31FDDDF8EEFD9F3736" box="[574,631,1890,1911]" pageId="24" pageNumber="425" rank="species">
<emphasis id="C6E7EA8A45105A31FDDDF8EEFD9F3736" bold="true" box="[574,631,1890,1911]" pageId="24" pageNumber="425">sp. n.</emphasis>
</taxonomicNameLabel>
96–97, female imagoes; 98, postsubalar sclerite of subimaginal exuviae (arrow shows concavity of dorsal margin of posterior-dorsal process); 99, half of exuviae of subimaginal mesonotum; 100–101, male imago; 102, fragment of fore leg of male imago with apex of tibia and 1 
<taxonomicName id="33934D1B45105A31FC91F813FC5937F4" box="[882,945,1951,1973]" pageId="24" pageNumber="425" rank="subSpecies" subSpecies="and">st and</taxonomicName>
2nd tarsomeres; 103, apex of tibia and tarsus of hind leg.
</paragraph>
</caption>
<caption id="A0EC661045115A30FF74F8CEFB4E37B3" httpUri="https://zenodo.org/record/1059215/files/figure.png" pageId="25" pageNumber="426" targetBox="[161,1413,204,1831]" targetPageId="25">
<paragraph id="F42C369845115A30FF74F8CEFB4E37B3" blockId="25.[151,1436,1858,2034]" pageId="25" pageNumber="426">
<emphasis id="C6E7EA8A45115A30FF74F8CEFE843719" bold="true" box="[151,364,1858,1881]" pageId="25" pageNumber="426">FIGURES 104–107.</emphasis>
<taxonomicName id="33934D1B45115A30FE91F8CFFDAB3718" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[370,579,1859,1881]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="426" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A45115A30FE91F8CFFDAB3718" box="[370,579,1859,1881]" italics="true" pageId="25" pageNumber="426">Crassabwa ludmilae</emphasis>
</taxonomicName>
<emphasis id="C6E7EA8A45115A30FDAAF8CFFD693719" bold="true" box="[585,641,1859,1880]" pageId="25" pageNumber="426">
<taxonomicNameLabel id="DDD457F145115A30FDAAF8CFFD693719" box="[585,641,1859,1880]" pageId="25" pageNumber="426" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male genitals. 104, subimaginal genitals developing under larval cuticle; 105, exuviae of subimaginal genitals; 106, genitals of male imago, ventral view (unistyligers and gonostyli unnaturally brought together because of wrong fixation; internal sclerites and muscles shown by interrupted lines, right gonostylar muscle not shown; large arrow shows median projection of dorsal wall of unistyliger); 107, penis enlarged, ventral view (gonoducts shown by dotted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; mIX-X, places of attachment of muscles running to common base of paraprocts; pb, paired internal projection of penial bridge; usg, unistyliger.
</paragraph>
</caption>
<caption id="A0EC661045125A33FF74F91EFC5536E9" box="[151,957,1682,1705]" httpUri="https://zenodo.org/record/1059217/files/figure.png" pageId="26" pageNumber="427" targetBox="[151,1436,193,1653]" targetPageId="26">
<paragraph id="F42C369845125A33FF74F91EFC5536E9" blockId="26.[151,957,1682,1705]" box="[151,957,1682,1705]" pageId="26" pageNumber="427">
<emphasis id="C6E7EA8A45125A33FF74F91EFE8436E6" bold="true" box="[151,364,1682,1705]" pageId="26" pageNumber="427">FIGURES 108–113.</emphasis>
<taxonomicName id="33934D1B45125A33FE92F91EFDAA36E9" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[369,578,1682,1704]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="427" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A45125A33FE92F91EFDAA36E9" box="[369,578,1682,1704]" italics="true" pageId="26" pageNumber="427">Crassabwa ludmilae</emphasis>
</taxonomicName>
<emphasis id="C6E7EA8A45125A33FDAAF91FFD6A36E9" bold="true" box="[585,642,1683,1704]" pageId="26" pageNumber="427">
<taxonomicNameLabel id="DDD457F145125A33FDAAF91FFD6A36E9" box="[585,642,1683,1704]" pageId="26" pageNumber="427" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, eggs extracted from holotype.
</paragraph>
</caption>
<paragraph id="F42C369845125A33FF24F95AFC943787" blockId="26.[151,1436,1749,2026]" pageId="26" pageNumber="427">
STRUCTURE. Head and mouth parts as characterized above [see 
<taxonomicName id="33934D1B45125A33FC20F959FBA936AC" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[963,1089,1749,1773]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45125A33FC20F959FBA936AC" box="[963,1089,1749,1773]" italics="true" pageId="26" pageNumber="427">Crassabwa</emphasis>
</taxonomicName>
(3)–(10)]. Left mandible with dense setae between prostheca and mola; its mola proximad of distal molar projection forms 3 or 4 transverse ridges, which in dorsal or ventral view look as protuberances; fine molar processes do not form integral row, but are concentrated on tops of these protuberances (
<figureCitation id="6CA82A1D45125A33FD5AF8CDFCE73718" box="[697,783,1857,1882]" captionStart="FIGURES 66 – 75" captionStartId="21.[151,264,1855,1878]" captionTargetBox="[186,1400,193,1812]" captionTargetId="figure@21.[170,1417,193,1835]" captionTargetPageId="21" captionText="FIGURES 66 – 75. Crassabwa ludmilae sp. n., larva. 66, claw; 67, fore leg (anterior view); 68, paraproct; 69 – 70, right and left mandibles (ventral view); 71, labrum (at left ventral view, at right dorsal view); 72, maxilla; 73, labial palp (ventral view); 74, labium (at left ventral view, at right dorsal view); 75, hypopharynx and superlinguae." httpUri="https://zenodo.org/record/1059207/files/figure.png" pageId="26" pageNumber="427">Figs 70</figureCitation>
, 
<figureCitation id="6CA82A1D45125A33FCF8F8CEFC523718" box="[795,954,1857,1881]" captionStart="FIGURES 76 – 83" captionStartId="22.[151,264,1646,1669]" captionTargetBox="[154,1436,193,1625]" captionTargetId="figure@22.[151,1436,193,1625]" captionTargetPageId="22" captionText="FIGURES 76 – 83. Crassabwa ludmilae sp. n. 76 – 77, right and left mandibles; 78 – 79, biting edges of left and right mandibles of penultimolarva with developing ultimolarval mandibles inside; 80 – 82, biting edges of left and right mandibles of ultimolarva; 83, exuviae of labium with focus on ventral side." httpUri="https://zenodo.org/record/1059209/files/figure.png" pageId="26" pageNumber="427">77, 78, 80–81</figureCitation>
). Right mandible with dense setae between prostheca and mola; its mola bears separate distal protuberance and 4 or 5 transverse ridges, which in dorsal or ventral view look as protuberances (
<figureCitation id="6CA82A1D45125A33FDDEF805FD7F37E3" box="[573,663,1929,1954]" captionStart="FIGURES 66 – 75" captionStartId="21.[151,264,1855,1878]" captionTargetBox="[186,1400,193,1812]" captionTargetId="figure@21.[170,1417,193,1835]" captionTargetPageId="21" captionText="FIGURES 66 – 75. Crassabwa ludmilae sp. n., larva. 66, claw; 67, fore leg (anterior view); 68, paraproct; 69 – 70, right and left mandibles (ventral view); 71, labrum (at left ventral view, at right dorsal view); 72, maxilla; 73, labial palp (ventral view); 74, labium (at left ventral view, at right dorsal view); 75, hypopharynx and superlinguae." httpUri="https://zenodo.org/record/1059207/files/figure.png" pageId="26" pageNumber="427">Figs 69</figureCitation>
, 
<figureCitation id="6CA82A1D45125A33FD45F805FCC937E0" box="[678,801,1929,1954]" captionStart="FIGURES 76 – 83" captionStartId="22.[151,264,1646,1669]" captionTargetBox="[154,1436,193,1625]" captionTargetId="figure@22.[151,1436,193,1625]" captionTargetPageId="22" captionText="FIGURES 76 – 83. Crassabwa ludmilae sp. n. 76 – 77, right and left mandibles; 78 – 79, biting edges of left and right mandibles of penultimolarva with developing ultimolarval mandibles inside; 80 – 82, biting edges of left and right mandibles of ultimolarva; 83, exuviae of labium with focus on ventral side." httpUri="https://zenodo.org/record/1059209/files/figure.png" pageId="26" pageNumber="427">76, 79, 82</figureCitation>
). Inner-apical projection of 2nd segment of labial palp large, sometimes nearly as large as 3rd segment (
<figureCitation id="6CA82A1D45125A33FD5FF821FCAC3787" box="[700,836,1965,1990]" captionStart="FIGURES 66 – 75" captionStartId="21.[151,264,1855,1878]" captionTargetBox="[186,1400,193,1812]" captionTargetId="figure@21.[170,1417,193,1835]" captionTargetPageId="21" captionText="FIGURES 66 – 75. Crassabwa ludmilae sp. n., larva. 66, claw; 67, fore leg (anterior view); 68, paraproct; 69 – 70, right and left mandibles (ventral view); 71, labrum (at left ventral view, at right dorsal view); 72, maxilla; 73, labial palp (ventral view); 74, labium (at left ventral view, at right dorsal view); 75, hypopharynx and superlinguae." httpUri="https://zenodo.org/record/1059207/files/figure.png" pageId="26" pageNumber="427">Figs. 73–74</figureCitation>
, 
<figureCitation id="6CA82A1D45125A33FCB3F822FC873787" box="[848,879,1966,1990]" captionStart="FIGURES 76 – 83" captionStartId="22.[151,264,1646,1669]" captionTargetBox="[154,1436,193,1625]" captionTargetId="figure@22.[151,1436,193,1625]" captionTargetPageId="22" captionText="FIGURES 76 – 83. Crassabwa ludmilae sp. n. 76 – 77, right and left mandibles; 78 – 79, biting edges of left and right mandibles of penultimolarva with developing ultimolarval mandibles inside; 80 – 82, biting edges of left and right mandibles of ultimolarva; 83, exuviae of labium with focus on ventral side." httpUri="https://zenodo.org/record/1059209/files/figure.png" pageId="26" pageNumber="427">83</figureCitation>
).
</paragraph>
<paragraph id="F42C369845125A32FF24F85EFDAA3101" blockId="26.[151,1436,1749,2026]" lastBlockId="27.[151,1437,151,1581]" lastPageId="27" lastPageNumber="428" pageId="26" pageNumber="427">
Legs as characterized above [see 
<taxonomicName id="33934D1B45125A33FDDCF85DFD5537A8" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[575,701,2001,2025]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45125A33FDDCF85DFD5537A8" box="[575,701,2001,2025]" italics="true" pageId="26" pageNumber="427">Crassabwa</emphasis>
</taxonomicName>
(12)–(16), (19)] (
<figureCitation id="6CA82A1D45125A33FC65F85DFC3537AB" box="[902,989,2001,2026]" captionStart="FIGURES 60 – 65" captionStartId="20.[151,264,1865,1888]" captionTargetBox="[157,1426,193,1824]" captionTargetId="figure@20.[151,1436,193,1844]" captionTargetPageId="20" captionText="FIGURES 60 – 65. Crassabwa ludmilae sp. n., holotype, ultimolarval exuviae. 60, abdominal terga and caudalii; 61, head with labrum and right mandible; 62, exuviae of half of pronotum and mesonotum; 63, fragment of cercus; 64, hind leg (triangle shows point where patella-tibial suture crosses inner margin); 65, abdominal terga IX and X." httpUri="https://zenodo.org/record/1059205/files/figure.png" pageId="26" pageNumber="427">Figs 64</figureCitation>
, 
<figureCitation id="6CA82A1D45125A33FC0AF85DFBE037AB" box="[1001,1032,2001,2026]" captionStart="FIGURES 66 – 75" captionStartId="21.[151,264,1855,1878]" captionTargetBox="[186,1400,193,1812]" captionTargetId="figure@21.[170,1417,193,1835]" captionTargetPageId="21" captionText="FIGURES 66 – 75. Crassabwa ludmilae sp. n., larva. 66, claw; 67, fore leg (anterior view); 68, paraproct; 69 – 70, right and left mandibles (ventral view); 71, labrum (at left ventral view, at right dorsal view); 72, maxilla; 73, labial palp (ventral view); 74, labium (at left ventral view, at right dorsal view); 75, hypopharynx and superlinguae." httpUri="https://zenodo.org/record/1059207/files/figure.png" pageId="26" pageNumber="427">67</figureCitation>
). Fine colorless setae forming cross row near base of tibia and dispersed along posterior side of tibia and tarsus [see 
<taxonomicName id="33934D1B45135A32FBA2FF14FB5730F1" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[1089,1215,152,176]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FBA2FF14FB5730F1" box="[1089,1215,152,176]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(14)] simple (nonbifurcate); oblique row on anterior side of tibia consists of 3 such setae. Claw relatively short, with distal portion (from distal denticle to apex) shorter than remainder part (from base to distal denticle); denticles on claw equal in two rows [see 
<taxonomicName id="33934D1B45135A32FEA5FE88FE2C315D" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[326,452,260,284]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FEA5FE88FE2C315D" box="[326,452,260,284]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(19)]: two enlarged distal denticles in posterior row as large as two enlarged distal denticles in anterior row (
<figureCitation id="6CA82A1D45135A32FE54FEABFDE23101" box="[439,522,295,320]" captionStart="FIGURES 66 – 75" captionStartId="21.[151,264,1855,1878]" captionTargetBox="[186,1400,193,1812]" captionTargetId="figure@21.[170,1417,193,1835]" captionTargetPageId="21" captionText="FIGURES 66 – 75. Crassabwa ludmilae sp. n., larva. 66, claw; 67, fore leg (anterior view); 68, paraproct; 69 – 70, right and left mandibles (ventral view); 71, labrum (at left ventral view, at right dorsal view); 72, maxilla; 73, labial palp (ventral view); 74, labium (at left ventral view, at right dorsal view); 75, hypopharynx and superlinguae." httpUri="https://zenodo.org/record/1059207/files/figure.png" pageId="27" pageNumber="428">Fig. 66</figureCitation>
, 
<figureCitation id="6CA82A1D45135A32FDF6FEABFDDC3101" box="[533,564,295,320]" captionStart="FIGURES 84 – 95" captionStartId="23.[151,264,1927,1950]" captionTargetBox="[155,1436,193,1897]" captionTargetId="figure@23.[151,1436,193,1907]" captionTargetPageId="23" captionText="FIGURES 84 – 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 – 91, tergalii II – VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 – 93 and 95, holotype)." httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="27" pageNumber="428">95</figureCitation>
).
</paragraph>
<paragraph id="F42C369845135A32FF24FEC0FA9B327C" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="428">
Abdomen as characterized above [see 
<taxonomicName id="33934D1B45135A32FD61FEC0FCE83125" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[642,768,332,356]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FD61FEC0FCE83125" box="[642,768,332,356]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(22)–(24)]; all scales of terga short (
<figureCitation id="6CA82A1D45135A32FB4DFEC0FAED3125" box="[1198,1285,332,357]" captionStart="FIGURES 84 – 95" captionStartId="23.[151,264,1927,1950]" captionTargetBox="[155,1436,193,1897]" captionTargetId="figure@23.[151,1436,193,1907]" captionTargetPageId="23" captionText="FIGURES 84 – 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 – 91, tergalii II – VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 – 93 and 95, holotype)." httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="27" pageNumber="428">Fig. 94</figureCitation>
); sterna I–III without denticles, sternum IV with vestigial denticles, sterna V–IX with regular row of pointed denticles (
<figureCitation id="6CA82A1D45135A32FA8CFEE3FF2D31ED" captionStart="FIGURES 104 – 107" captionStartId="25.[151,264,1858,1881]" captionTargetBox="[161,1413,204,1831]" captionTargetId="figure@25.[160,1420,193,1835]" captionTargetPageId="25" captionText="FIGURES 104 – 107. Crassabwa ludmilae sp. n., male genitals. 104, subimaginal genitals developing under larval cuticle; 105, exuviae of subimaginal genitals; 106, genitals of male imago, ventral view (unistyligers and gonostyli unnaturally brought together because of wrong fixation; internal sclerites and muscles shown by interrupted lines, right gonostylar muscle not shown; large arrow shows median projection of dorsal wall of unistyliger); 107, penis enlarged, ventral view (gonoducts shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, places of attachment of muscles running to common base of paraprocts; pb, paired internal projection of penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/1059215/files/figure.png" pageId="27" pageNumber="428">Fig. 104</figureCitation>
). Tergalius of first pair wide, asymmetric, with anal margin more convex than costal margin (
<figureCitation id="6CA82A1D45135A32FB0FFE18FAA931EC" box="[1260,1345,404,429]" captionStart="FIGURES 84 – 95" captionStartId="23.[151,264,1927,1950]" captionTargetBox="[155,1436,193,1897]" captionTargetId="figure@23.[151,1436,193,1907]" captionTargetPageId="23" captionText="FIGURES 84 – 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 – 91, tergalii II – VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 – 93 and 95, holotype)." httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="27" pageNumber="428">Fig. 85</figureCitation>
); in last larval instar nearly half of tergalius is bent ventrally (
<figureCitation id="6CA82A1D45135A32FD15FE3BFCA03191" box="[758,840,439,464]" captionStart="FIGURES 84 – 95" captionStartId="23.[151,264,1927,1950]" captionTargetBox="[155,1436,193,1897]" captionTargetId="figure@23.[151,1436,193,1907]" captionTargetPageId="23" captionText="FIGURES 84 – 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 – 91, tergalii II – VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 – 93 and 95, holotype)." httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="27" pageNumber="428">Fig. 84</figureCitation>
) [see 
<taxonomicName id="33934D1B45135A32FC6DFE34FBE43191" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[910,1036,440,464]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FC6DFE34FBE43191" box="[910,1036,440,464]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(25)]. Tergalii II–V subequal (
<figureCitation id="6CA82A1D45135A32FA89FE3BFF0831B4" captionStart="FIGURES 84 – 95" captionStartId="23.[151,264,1927,1950]" captionTargetBox="[155,1436,193,1897]" captionTargetId="figure@23.[151,1436,193,1907]" captionTargetPageId="23" captionText="FIGURES 84 – 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 – 91, tergalii II – VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 – 93 and 95, holotype)." httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="27" pageNumber="428">Figs 85–89</figureCitation>
), tergalii VI–VII gradually smaller, with anterior margin more convex (
<figureCitation id="6CA82A1D45135A32FBEAFE50FB6631B5" box="[1033,1166,476,501]" captionStart="FIGURES 84 – 95" captionStartId="23.[151,264,1927,1950]" captionTargetBox="[155,1436,193,1897]" captionTargetId="figure@23.[151,1436,193,1907]" captionTargetPageId="23" captionText="FIGURES 84 – 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 – 91, tergalii II – VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 – 93 and 95, holotype)." httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="27" pageNumber="428">Figs 90–91</figureCitation>
). Brown band on dorsal cuticle of tergalius [see 
<taxonomicName id="33934D1B45135A32FE44FD8CFDCD3259" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[423,549,512,536]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FE44FD8CFDCD3259" box="[423,549,512,536]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(26)] either narrow, invisible on background of main trachea (
<figureCitation id="6CA82A1D45135A32FB0FFE73FA983259" box="[1260,1392,511,536]" captionStart="FIGURES 84 – 95" captionStartId="23.[151,264,1927,1950]" captionTargetBox="[155,1436,193,1897]" captionTargetId="figure@23.[151,1436,193,1907]" captionTargetPageId="23" captionText="FIGURES 84 – 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 – 91, tergalii II – VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 – 93 and 95, holotype)." httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="27" pageNumber="428">Figs 92–93</figureCitation>
), or absent. Tenth abdominal segment and caudalii as described above [see 
<taxonomicName id="33934D1B45135A32FC51FDA8FBD8327D" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[946,1072,548,572]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FC51FDA8FBD8327D" box="[946,1072,548,572]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(28), (29)] (
<figureCitation id="6CA82A1D45135A32FB59FDA8FA8E327C" box="[1210,1382,548,573]" captionStart="FIGURES 60 – 65" captionStartId="20.[151,264,1865,1888]" captionTargetBox="[157,1426,193,1824]" captionTargetId="figure@20.[151,1436,193,1844]" captionTargetPageId="20" captionText="FIGURES 60 – 65. Crassabwa ludmilae sp. n., holotype, ultimolarval exuviae. 60, abdominal terga and caudalii; 61, head with labrum and right mandible; 62, exuviae of half of pronotum and mesonotum; 63, fragment of cercus; 64, hind leg (triangle shows point where patella-tibial suture crosses inner margin); 65, abdominal terga IX and X." httpUri="https://zenodo.org/record/1059205/files/figure.png" pageId="27" pageNumber="428">Figs 60, 63, 65</figureCitation>
).
</paragraph>
<paragraph id="F42C369845135A32FF24FDCBFAAD3354" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="428">
<emphasis id="C6E7EA8A45135A32FF24FDCBFEAB3221" bold="true" box="[199,323,583,608]" pageId="27" pageNumber="428">
<emphasis id="C6E7EA8A45135A32FF24FDCBFED43221" bold="true" box="[199,316,583,608]" italics="true" pageId="27" pageNumber="428">Subimago</emphasis>
.
</emphasis>
CUTICULAR COLORATION. Head ocher with brown. Pronotum light brown. Mesonotum light brownish with colorless areas, with sutures in antero-lateral part contrastingly dark brown and with dark brown macula on posterior end of submedioscutum; parascutellum with contrasting dark brown macula on lateroposterior corner (
<figureCitation id="6CA82A1D45135A32FE87FD38FE56328D" box="[356,446,692,717]" captionStart="FIGURES 96 – 103" captionStartId="24.[151,264,1889,1912]" captionTargetBox="[163,1407,193,1851]" captionTargetId="figure@24.[163,1407,193,1851]" captionTargetPageId="24" captionText="FIGURES 96 – 103. Crassabwa ludmilae sp. n. 96 – 97, female imagoes; 98, postsubalar sclerite of subimaginal exuviae (arrow shows concavity of dorsal margin of posterior-dorsal process); 99, half of exuviae of subimaginal mesonotum; 100 – 101, male imago; 102, fragment of fore leg of male imago with apex of tibia and 1 st and 2 nd tarsomeres; 103, apex of tibia and tarsus of hind leg." httpUri="https://zenodo.org/record/1059213/files/figure.png" pageId="27" pageNumber="428">Fig. 99</figureCitation>
). Pleura and sterna of thorax light brownish with colorless membranes and certain sclerites dark brown (
<figureCitation id="6CA82A1D45135A32FE6CFD5BFE0E32B1" box="[399,486,727,752]" captionStart="FIGURES 96 – 103" captionStartId="24.[151,264,1889,1912]" captionTargetBox="[163,1407,193,1851]" captionTargetId="figure@24.[163,1407,193,1851]" captionTargetPageId="24" captionText="FIGURES 96 – 103. Crassabwa ludmilae sp. n. 96 – 97, female imagoes; 98, postsubalar sclerite of subimaginal exuviae (arrow shows concavity of dorsal margin of posterior-dorsal process); 99, half of exuviae of subimaginal mesonotum; 100 – 101, male imago; 102, fragment of fore leg of male imago with apex of tibia and 1 st and 2 nd tarsomeres; 103, apex of tibia and tarsus of hind leg." httpUri="https://zenodo.org/record/1059213/files/figure.png" pageId="27" pageNumber="428">Fig. 98</figureCitation>
). Legs light brown, fore tibiae and tarsi darker brown. Abdominal terga and sterna light brownish. Sternum IX of male subimago lighter than others; gonostyli dark brown. Cerci dark brown.
</paragraph>
<paragraph id="F42C369845135A32FF24FCACFAAC3379" blockId="27.[151,1437,151,1581]" box="[199,1348,799,824]" pageId="27" pageNumber="428">HYPODERMAL COLORATION. As in imago, but poorly visible through brown subimaginal cuticle.</paragraph>
<paragraph id="F42C369845135A32FF24FCC8FBA7343D" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="428">
<emphasis id="C6E7EA8A45135A32FF24FCC8FEB1331D" bold="true" box="[199,345,835,860]" italics="true" pageId="27" pageNumber="428">Imago, male</emphasis>
(
<figureCitation id="6CA82A1D45135A32FE8AFCC8FDE2331D" box="[361,522,836,861]" captionStart="FIGURES 96 – 103" captionStartId="24.[151,264,1889,1912]" captionTargetBox="[163,1407,193,1851]" captionTargetId="figure@24.[163,1407,193,1851]" captionTargetPageId="24" captionText="FIGURES 96 – 103. Crassabwa ludmilae sp. n. 96 – 97, female imagoes; 98, postsubalar sclerite of subimaginal exuviae (arrow shows concavity of dorsal margin of posterior-dorsal process); 99, half of exuviae of subimaginal mesonotum; 100 – 101, male imago; 102, fragment of fore leg of male imago with apex of tibia and 1 st and 2 nd tarsomeres; 103, apex of tibia and tarsus of hind leg." httpUri="https://zenodo.org/record/1059213/files/figure.png" pageId="27" pageNumber="428">Figs 100–101</figureCitation>
). Head, thorax and abdomen with hypodermal coloration forming ocher, orange, red or reddish-brownish areas [see 
<taxonomicName id="33934D1B45135A32FDC2FCE4FD7733C1" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[545,671,872,896]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FDC2FCE4FD7733C1" box="[545,671,872,896]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(2)]. Turbinate eyes wide; facetted surface orange, bordered by thin brown line; stem orange. Thorax orange, equally dark on dorsal and ventral sides. Wings with membrane and veins colorless, as characterized above [see 
<taxonomicName id="33934D1B45135A32FDBAFC3CFD3F3389" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[601,727,944,968]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FDBAFC3CFD3F3389" box="[601,727,944,968]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(20), (21)] (
<figureCitation id="6CA82A1D45135A32FC8CFC23FC103389" box="[879,1016,943,968]" captionStart="FIGURES 51 – 55" captionStartId="18.[151,264,1576,1599]" captionTargetBox="[163,1430,650,1552]" captionTargetId="figure@18.[151,1436,628,1555]" captionTargetPageId="18" captionText="FIGURES 51 – 55. Wings. 51 – 53, Crassabwa flava, hind wings of three individuals; 54 – 55, Crassabwa ludmilae sp. n., fore and hind wings on one scale." httpUri="https://zenodo.org/record/1059201/files/figure.png" pageId="27" pageNumber="428">Figs 54–55</figureCitation>
). Legs [see 
<taxonomicName id="33934D1B45135A32FB69FC3CFAE03389" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[1162,1288,944,968]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FB69FC3CFAE03389" box="[1162,1288,944,968]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(17)] ocher, without dark coloration on proximal portion of tibia. Abdominal terga orange with ocher. Abdominal sterna lighter, mainly ocher, without dark lateral sigmoid marks. Cerci near base intensively colored by deep red, in remainder part colorless [see 
<taxonomicName id="33934D1B45135A32FE9AFB90FE1F3475" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[377,503,1052,1076]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FE9AFB90FE1F3475" box="[377,503,1052,1076]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(2)]. Genitals (
<figureCitation id="6CA82A1D45135A32FD51FB90FCBC3474" box="[690,852,1052,1077]" captionStart="FIGURES 104 – 107" captionStartId="25.[151,264,1858,1881]" captionTargetBox="[161,1413,204,1831]" captionTargetId="figure@25.[160,1420,193,1835]" captionTargetPageId="25" captionText="FIGURES 104 – 107. Crassabwa ludmilae sp. n., male genitals. 104, subimaginal genitals developing under larval cuticle; 105, exuviae of subimaginal genitals; 106, genitals of male imago, ventral view (unistyligers and gonostyli unnaturally brought together because of wrong fixation; internal sclerites and muscles shown by interrupted lines, right gonostylar muscle not shown; large arrow shows median projection of dorsal wall of unistyliger); 107, penis enlarged, ventral view (gonoducts shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, places of attachment of muscles running to common base of paraprocts; pb, paired internal projection of penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/1059215/files/figure.png" pageId="27" pageNumber="428">Figs 106–107</figureCitation>
) poorly colored, ocher. Unistyligers with median projections [see 
<taxonomicName id="33934D1B45135A32FEB0FBCCFE393419" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[339,465,1088,1112]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FEB0FBCCFE393419" box="[339,465,1088,1112]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(31)]. Median side of first segment of gonostylus evenly convex, without concavity, entirely covered by small spines (
<figureCitation id="6CA82A1D45135A32FDECFBE8FD99343C" box="[527,625,1124,1149]" captionStart="FIGURES 104 – 107" captionStartId="25.[151,264,1858,1881]" captionTargetBox="[161,1413,204,1831]" captionTargetId="figure@25.[160,1420,193,1835]" captionTargetPageId="25" captionText="FIGURES 104 – 107. Crassabwa ludmilae sp. n., male genitals. 104, subimaginal genitals developing under larval cuticle; 105, exuviae of subimaginal genitals; 106, genitals of male imago, ventral view (unistyligers and gonostyli unnaturally brought together because of wrong fixation; internal sclerites and muscles shown by interrupted lines, right gonostylar muscle not shown; large arrow shows median projection of dorsal wall of unistyliger); 107, penis enlarged, ventral view (gonoducts shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, places of attachment of muscles running to common base of paraprocts; pb, paired internal projection of penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/1059215/files/figure.png" pageId="27" pageNumber="428">Fig. 106</figureCitation>
). Gonovectes wide and pointed (
<figureCitation id="6CA82A1D45135A32FC03FBE8FBAA343D" box="[992,1090,1124,1149]" captionStart="FIGURES 104 – 107" captionStartId="25.[151,264,1858,1881]" captionTargetBox="[161,1413,204,1831]" captionTargetId="figure@25.[160,1420,193,1835]" captionTargetPageId="25" captionText="FIGURES 104 – 107. Crassabwa ludmilae sp. n., male genitals. 104, subimaginal genitals developing under larval cuticle; 105, exuviae of subimaginal genitals; 106, genitals of male imago, ventral view (unistyligers and gonostyli unnaturally brought together because of wrong fixation; internal sclerites and muscles shown by interrupted lines, right gonostylar muscle not shown; large arrow shows median projection of dorsal wall of unistyliger); 107, penis enlarged, ventral view (gonoducts shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, places of attachment of muscles running to common base of paraprocts; pb, paired internal projection of penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/1059215/files/figure.png" pageId="27" pageNumber="428">Fig. 107</figureCitation>
).
</paragraph>
<paragraph id="F42C369845135A32FF24FB04FE953484" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="428">
<emphasis id="C6E7EA8A45135A32FF24FB04FE8634E1" bold="true" box="[199,366,1159,1184]" italics="true" pageId="27" pageNumber="428">Imago, female</emphasis>
(
<figureCitation id="6CA82A1D45135A32FE9DFB0BFDEA34E1" box="[382,514,1159,1184]" captionStart="FIGURES 96 – 103" captionStartId="24.[151,264,1889,1912]" captionTargetBox="[163,1407,193,1851]" captionTargetId="figure@24.[163,1407,193,1851]" captionTargetPageId="24" captionText="FIGURES 96 – 103. Crassabwa ludmilae sp. n. 96 – 97, female imagoes; 98, postsubalar sclerite of subimaginal exuviae (arrow shows concavity of dorsal margin of posterior-dorsal process); 99, half of exuviae of subimaginal mesonotum; 100 – 101, male imago; 102, fragment of fore leg of male imago with apex of tibia and 1 st and 2 nd tarsomeres; 103, apex of tibia and tarsus of hind leg." httpUri="https://zenodo.org/record/1059213/files/figure.png" pageId="27" pageNumber="428">Figs 96–97</figureCitation>
). Head ocher. Thorax ventrally lighter than dorsally. Coloration of legs, abdomen and cerci as in male.
</paragraph>
<paragraph id="F42C369845135A32FF24FB5CFEB3354D" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="428">
<emphasis id="C6E7EA8A45135A32FF24FB5CFEE934A6" bold="true" box="[199,257,1232,1255]" italics="true" pageId="27" pageNumber="428">Eggs</emphasis>
(
<figureCitation id="6CA82A1D45135A32FEFBFB43FE5634A9" box="[280,446,1231,1256]" captionStart="FIGURES 108 – 113" captionStartId="26.[151,264,1682,1705]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@26.[151,1436,193,1653]" captionTargetPageId="26" captionText="FIGURES 108 – 113. Crassabwa ludmilae sp. n., eggs extracted from holotype." httpUri="https://zenodo.org/record/1059217/files/figure.png" pageId="27" pageNumber="428">Figs 108–113</figureCitation>
). Nearly ellipsoid, 0.15 mm length; chorion with relief characterized above [see 
<taxonomicName id="33934D1B45135A32FF74FB78FEFD354D" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[151,277,1268,1292]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A45135A32FF74FB78FEFD354D" box="[151,277,1268,1292]" italics="true" pageId="27" pageNumber="428">Crassabwa</emphasis>
</taxonomicName>
(33)].
</paragraph>
<paragraph id="F42C369845135A32FF24FA9BFD573571" blockId="27.[151,1437,151,1581]" box="[199,703,1303,1328]" pageId="27" pageNumber="428">
<emphasis id="C6E7EA8A45135A32FF24FA9BFEA63571" bold="true" box="[199,334,1303,1328]" pageId="27" pageNumber="428">Dimension.</emphasis>
Fore wing length of male 
<quantity id="336B9B7D45135A32FD94FA95FD533571" box="[631,699,1305,1328]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="27" pageNumber="428" unit="mm" value="7.0">7 mm</quantity>
.
</paragraph>
<paragraph id="F42C369845135A32FF24FAB0FACC366C" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="428">
<emphasis id="C6E7EA8A45135A32FF24FAB0FE893514" bold="true" box="[199,353,1340,1365]" pageId="27" pageNumber="428">Comparison.</emphasis>
Larva of the new species 
<taxonomicName id="33934D1B45135A32FD42FAB0FCC13515" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[673,809,1340,1364]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A45135A32FD42FAB0FCC13515" box="[673,809,1340,1364]" italics="true" pageId="27" pageNumber="428">C. ludmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDD457F145135A32FCD7FAB0FC903514" box="[820,888,1340,1365]" pageId="27" pageNumber="428" rank="species">
<emphasis id="C6E7EA8A45135A32FCD7FAB0FC903514" bold="true" box="[820,888,1340,1365]" pageId="27" pageNumber="428">sp. n.</emphasis>
</taxonomicNameLabel>
can be distinguished from that of 
<taxonomicName id="33934D1B45135A32FAF4FAB0FA9D3515" authorityName=": Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[1303,1397,1340,1364]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="C6E7EA8A45135A32FAF4FAB0FA9D3515" box="[1303,1397,1340,1364]" italics="true" pageId="27" pageNumber="428">C. flava</emphasis>
</taxonomicName>
by structure of mola of left and right mandibles, presence of setae on left mandible, cuticular coloration of fore protoptera in last larval instar, non-bifurcate setae on tibiae, and wider tergalii of first pair. Imagoes of both sexes of the new species 
<taxonomicName id="33934D1B45135A32FEAEFA24FE393581" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[333,465,1448,1472]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A45135A32FEAEFA24FE393581" box="[333,465,1448,1472]" italics="true" pageId="27" pageNumber="428">C. ludmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDD457F145135A32FE34FA2BFDFE3581" box="[471,534,1447,1472]" pageId="27" pageNumber="428" rank="species">
<emphasis id="C6E7EA8A45135A32FE34FA2BFDFE3581" bold="true" box="[471,534,1447,1472]" pageId="27" pageNumber="428">sp. n.</emphasis>
</taxonomicNameLabel>
differ from 
<taxonomicName id="33934D1B45135A32FD7CFA24FD113581" authorityName=": Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[671,761,1448,1472]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="C6E7EA8A45135A32FD7CFA24FD5A35FE" box="[671,690,1448,1471]" italics="true" pageId="27" pageNumber="428">C</emphasis>
. 
<emphasis id="C6E7EA8A45135A32FD23FA24FD113581" box="[704,761,1448,1472]" italics="true" pageId="27" pageNumber="428">flava</emphasis>
</taxonomicName>
by absence of dark coloration at bases of tibiae and absence of dark sigmoid marks on abdominal sterna. Male imaginal genitals of 
<taxonomicName id="33934D1B45135A32FC2EFA40FBBA35A5" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[973,1106,1484,1508]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A45135A32FC2EFA40FBBA35A5" box="[973,1106,1484,1508]" italics="true" pageId="27" pageNumber="428">C. ludmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDD457F145135A32FBBFFA40FB7635A4" box="[1116,1182,1484,1509]" pageId="27" pageNumber="428" rank="species">
<emphasis id="C6E7EA8A45135A32FBBFFA40FB7635A4" bold="true" box="[1116,1182,1484,1509]" pageId="27" pageNumber="428">sp. n.</emphasis>
</taxonomicNameLabel>
differ from that of 
<taxonomicName id="33934D1B45135A32FA61FA40FF273649" authorityName=": Lugo-Ortiz &amp; McCafferty" authorityYear="1996" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="C6E7EA8A45135A32FA61FA40FF273649" italics="true" pageId="27" pageNumber="428">C. flava</emphasis>
</taxonomicName>
by wider gonovectes and shape of first segment of gonostylus. Eggs of 
<taxonomicName id="33934D1B45135A32FC0EFA7CFB983649" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[1005,1136,1520,1544]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A45135A32FC0EFA7CFB983649" box="[1005,1136,1520,1544]" italics="true" pageId="27" pageNumber="428">C. ludmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDD457F145135A32FB95FA63FB5D3649" box="[1142,1205,1519,1544]" pageId="27" pageNumber="428" rank="species">
<emphasis id="C6E7EA8A45135A32FB95FA63FB5D3649" bold="true" box="[1142,1205,1519,1544]" pageId="27" pageNumber="428">sp. n.</emphasis>
</taxonomicNameLabel>
are similar to that of 
<taxonomicName id="33934D1B45135A32FF74F998FF07366D" authorityName=": Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[151,239,1556,1580]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="C6E7EA8A45135A32FF74F998FF07366D" box="[151,239,1556,1580]" italics="true" pageId="27" pageNumber="428">C. flava</emphasis>
</taxonomicName>
, but shorter and more gradually narrowing toward poles (i.e. ellipsoid rater than elongate-oval).
</paragraph>
</subSubSection>
</treatment>
</document>