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<document id="95B6A1E91BEF3F7858E5C6DC183D4AD8" ID-CLB-Dataset="31378" ID-DOI="10.11646/zootaxa.4350.3.1" ID-GBIF-Dataset="25b07a0d-7a27-460e-8813-713dbfb1888c" ID-ISSN="1175-5326" ID-Zenodo-Dep="1059179" ID-ZooBank="A3F42509-91EA-48BC-8C9D-253F5C57F20D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511167096685" checkinUser="plazi" docAuthor="Kluge, Nikita J., Gattolliat, Jean-Luc &amp; Salles, Frederico F." docDate="2017" docId="7C3A878E450E5A2FFF74FAB0FA7436B9" docLanguage="en" docName="zootaxa.4350.3.1.pdf" docOrigin="Zootaxa 4350 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crassabwa Lugo-Ortiz &amp; McCafferty 1996" docType="key" docVersion="7" lastPageNumber="407" masterDocId="8003FFF645085A29FFE3FF8CFFE83041" masterDocTitle="Redescription of the Afrotropical genus Crassabwa Lugo-Ortiz &amp; McCafferty 1996 (Ephemeroptera: Baetidae: Protopatellata)" masterLastPageNumber="435" masterPageNumber="401" pageNumber="407" updateTime="1698501443309" updateUser="ExternalLinkService">
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<mods:title id="CA37CCC90F2BB4A0D68D88F303DCF253">Redescription of the Afrotropical genus Crassabwa Lugo-Ortiz &amp; McCafferty 1996 (Ephemeroptera: Baetidae: Protopatellata)</mods:title>
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<mods:namePart id="B75E2BB9C8E11E9AF272F1051EEAC2BD">Kluge, Nikita J.</mods:namePart>
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<mods:namePart id="86ECF742395A83882E98B0C7A0AF5B1E">Gattolliat, Jean-Luc</mods:namePart>
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<mods:namePart id="FC8A912EAFAC20B37BE237269639EE58">Salles, Frederico F.</mods:namePart>
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<treatment id="7C3A878E450E5A2FFF74FAB0FA7436B9" ID-DOI="http://doi.org/10.5281/zenodo.6045783" ID-GBIF-Taxon="137453635" ID-Zenodo-Dep="6045783" LSID="urn:lsid:plazi:treatment:7C3A878E450E5A2FFF74FAB0FA7436B9" httpUri="http://treatment.plazi.org/id/7C3A878E450E5A2FFF74FAB0FA7436B9" lastPageNumber="407" pageId="6" pageNumber="407">
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<paragraph id="F42C3698450E5A2FFF74FAB0FE013517" blockId="6.[151,489,1340,1366]" box="[151,489,1340,1366]" pageId="6" pageNumber="407">
<heading id="AF6481F4450E5A2FFF74FAB0FE013517" bold="true" box="[151,489,1340,1366]" fontSize="11" level="1" pageId="6" pageNumber="407" reason="1">
<emphasis id="C6E7EA8A450E5A2FFF74FAB0FE013517" bold="true" box="[151,489,1340,1366]" pageId="6" pageNumber="407">
Key to larvae of
<taxonomicName id="33934D1B450E5A2FFE86FAB0FE013517" ID-CoL="3V95" ID-ENA="328958" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[357,489,1340,1366]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A450E5A2FFE86FAB0FE013517" bold="true" box="[357,489,1340,1366]" italics="true" pageId="6" pageNumber="407">Crassabwa</emphasis>
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</emphasis>
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</paragraph>
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<subSubSection id="BC896513450E5A2FFF74FA0DFA7436B9" pageId="6" pageNumber="407" type="key">
<keyStep id="4F672E3D450E5A2FFF74FA0DFA743593" pageId="6" pageNumber="407">
<paragraph id="F42C3698450E5A2FFF74FA0DFA7435F4" blockId="6.[151,1436,1409,1784]" pageId="6" pageNumber="407">
<keyLead id="4F6295AD450E5A2FFF74FA0DFA7435F4" pageId="6" pageNumber="407">
1. In last larval instar, tergalius I in proximal half with anal portion usually bent ventrally (
<figureCitation id="6CA82A1D450E5A2FFBC5FA0DFB9835D6" box="[1062,1136,1409,1431]" captionStart="FIGURES 23 30" captionStartId="13.[151,264,1589,1612]" captionTargetBox="[151,1436,203,1568]" captionTargetId="figure@13.[151,1436,193,1568]" captionTargetPageId="13" captionText="FIGURES 23 30. Crassabwa flava. 23, tergalius I of last instar larva in natural condition; 24, tergalius I spread on slide; 25, tergalius II; 26, costal rib of tergalius of penultimolarva (developing compressed ultimolarval denticles shown by interrupted line); 27 and 28, tergalii VI and VII; 29, tergalii II (in normal condition) and III (with dorsal cuticle partly separated from tissues); 30, enlarged fragment of this separated cuticle." httpUri="https://zenodo.org/record/1059191/files/figure.png" pageId="6" pageNumber="407">Figs 23</figureCitation>
,
<figureCitation id="6CA82A1D450E5A2FFB99FA0DFB7D35D6" box="[1146,1173,1409,1431]" captionStart="FIGURES 84 95" captionStartId="23.[151,264,1927,1950]" captionTargetBox="[155,1436,193,1897]" captionTargetId="figure@23.[151,1436,193,1907]" captionTargetPageId="23" captionText="FIGURES 84 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 91, tergalii II VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 93 and 95, holotype)." httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="6" pageNumber="407">84</figureCitation>
); other tergalii IIVII without such overturned portion (
<taxonomicName id="33934D1B450E5A2FFE1BFA13FD8C35F4" ID-CoL="3V95" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1996" box="[504,612,1439,1461]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="C6E7EA8A450E5A2FFE1BFA13FD8C35F4" box="[504,612,1439,1461]" italics="true" pageId="6" pageNumber="407">Crassabwa</emphasis>
</taxonomicName>
)...................................................................2
</keyLead>
</paragraph>
<paragraph id="F42C3698450E5A2FFF74FA31FA743593" blockId="6.[151,1436,1409,1784]" box="[151,1436,1468,1490]" pageId="6" pageNumber="407">
<keyLead id="4F6295AD450E5A2FFF74FA31FA743593" box="[151,1436,1468,1490]" pageId="6" pageNumber="407">- In last larval instar, tergalii not as above............................................................ other taxa.</keyLead>
</paragraph>
</keyStep>
<keyStep id="4F672E3D450E5A2FFF74FA55FA7D36C3" pageId="6" pageNumber="407">
<paragraph id="F42C3698450E5A2FFF74FA55FA74366B" blockId="6.[151,1436,1409,1784]" pageId="6" pageNumber="407">
<keyLead id="4F6295AD450E5A2FFF74FA55FA74366B" pageId="6" pageNumber="407">
2. In last instar larva, cuticle of fore protopteron with dark lines corresponding to convex longitudinal veins only (
<figureCitation id="6CA82A1D450E5A2FFAE7FA55FAD635AE" box="[1284,1342,1497,1519]" captionStart="FIGURES 1 4" captionStartId="8.[151,264,2012,2035]" captionTargetBox="[151,1425,539,1991]" captionTargetId="figure@8.[151,1436,539,1991]" captionTargetPageId="8" captionText="FIGURES 1 4. Crassabwa flava, last instar larvae. 1 3, dorsal view; 4, exuviae of half of pronotum and mesonotum." httpUri="https://zenodo.org/record/1059181/files/figure.png" pageId="6" pageNumber="407">Fig. 4</figureCitation>
). Mola of left mandible with even row of denticles (
<figureCitation id="6CA82A1D450E5A2FFD92FA7BFD43364C" box="[625,683,1527,1549]" captionStart="FIGURES 5 9" captionStartId="9.[151,264,1919,1942]" captionTargetBox="[151,1436,215,1890]" captionTargetId="figure@9.[151,1436,193,1899]" captionTargetPageId="9" captionText="FIGURES 5 9. Crassabwa flava. 5, labium (at left ventral view, with muscles of glossa, paraglossa and palp shown by interrupted lines; at right dorsal view); 6, apex of maxilla, dorsal view; 7, dentisetae and setae of inner-dorsal row of another individual; 8 and 9, left and right mandible of penultimolarva with developing ultimolarval mandibles inside. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/1059183/files/figure.png" pageId="6" pageNumber="407">Fig. 8</figureCitation>
). First tergalius relatively narrow (in spread condition) (
<figureCitation id="6CA82A1D450E5A2FFB24FA7BFAE7364C" box="[1223,1295,1527,1549]" captionStart="FIGURES 23 30" captionStartId="13.[151,264,1589,1612]" captionTargetBox="[151,1436,203,1568]" captionTargetId="figure@13.[151,1436,193,1568]" captionTargetPageId="13" captionText="FIGURES 23 30. Crassabwa flava. 23, tergalius I of last instar larva in natural condition; 24, tergalius I spread on slide; 25, tergalius II; 26, costal rib of tergalius of penultimolarva (developing compressed ultimolarval denticles shown by interrupted line); 27 and 28, tergalii VI and VII; 29, tergalii II (in normal condition) and III (with dorsal cuticle partly separated from tissues); 30, enlarged fragment of this separated cuticle." httpUri="https://zenodo.org/record/1059191/files/figure.png" pageId="6" pageNumber="407">Fig. 24</figureCitation>
)..................................................................................................
<taxonomicName id="33934D1B450E5A2FFB19F998FA74366B" ID-CoL="Z6BY" baseAuthorityName="Crass" baseAuthorityYear="1947" box="[1274,1436,1556,1578]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="C6E7EA8A450E5A2FFB19F998FA74366B" box="[1274,1436,1556,1578]" italics="true" pageId="6" pageNumber="407">Crassabwa flava</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F42C3698450E5A2FFF74F9BEFA7D36C3" blockId="6.[151,1436,1409,1784]" pageId="6" pageNumber="407">
<keyLead id="4F6295AD450E5A2FFF74F9BEFA7D36C3" pageId="6" pageNumber="407">
- In last instar larva, cuticle of fore protopteron with dark lines corresponding to convex, concave and cross veins (
<figureCitation id="6CA82A1D450E5A2FFAA5F9BDFA793606" box="[1350,1425,1585,1607]" captionStart="FIGURES 60 65" captionStartId="20.[151,264,1865,1888]" captionTargetBox="[157,1426,193,1824]" captionTargetId="figure@20.[151,1436,193,1844]" captionTargetPageId="20" captionText="FIGURES 60 65. Crassabwa ludmilae sp. n., holotype, ultimolarval exuviae. 60, abdominal terga and caudalii; 61, head with labrum and right mandible; 62, exuviae of half of pronotum and mesonotum; 63, fragment of cercus; 64, hind leg (triangle shows point where patella-tibial suture crosses inner margin); 65, abdominal terga IX and X." httpUri="https://zenodo.org/record/1059205/files/figure.png" pageId="6" pageNumber="407">Fig. 62</figureCitation>
). Mola of left mandible with tubercles bearing denticles (
<figureCitation id="6CA82A1D450E5A2FFD15F9C3FCD43624" box="[758,828,1615,1637]" captionStart="FIGURES 76 83" captionStartId="22.[151,264,1646,1669]" captionTargetBox="[154,1436,193,1625]" captionTargetId="figure@22.[151,1436,193,1625]" captionTargetPageId="22" captionText="FIGURES 76 83. Crassabwa ludmilae sp. n. 76 77, right and left mandibles; 78 79, biting edges of left and right mandibles of penultimolarva with developing ultimolarval mandibles inside; 80 82, biting edges of left and right mandibles of ultimolarva; 83, exuviae of labium with focus on ventral side." httpUri="https://zenodo.org/record/1059209/files/figure.png" pageId="6" pageNumber="407">Fig. 81</figureCitation>
). First tergalius relatively wide (in spread condition) (
<figureCitation id="6CA82A1D450E5A2FFAA1F9C3FA6F3624" box="[1346,1415,1615,1637]" captionStart="FIGURES 5 9" captionStartId="9.[151,264,1919,1942]" captionTargetBox="[151,1436,215,1890]" captionTargetId="figure@9.[151,1436,193,1899]" captionTargetPageId="9" captionText="FIGURES 5 9. Crassabwa flava. 5, labium (at left ventral view, with muscles of glossa, paraglossa and palp shown by interrupted lines; at right dorsal view); 6, apex of maxilla, dorsal view; 7, dentisetae and setae of inner-dorsal row of another individual; 8 and 9, left and right mandible of penultimolarva with developing ultimolarval mandibles inside. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/1059183/files/figure.png" pageId="6" pageNumber="407">Fig. 8 5</figureCitation>
)................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4F672E3D450E5A2FFF74F905FA7436B9" pageId="6" pageNumber="407">
<paragraph id="F42C3698450E5A2FFF74F905FA7436FC" blockId="6.[151,1436,1409,1784]" pageId="6" pageNumber="407">
<keyLead id="4F6295AD450E5A2FFF74F905FA7436FC" pageId="6" pageNumber="407">
3. Claw relatively short, with portion from distal denticle to apex shorter than portion from distal denticle to base (
<figureCitation id="6CA82A1D450E5A2FFAF2F905FAB336DE" box="[1297,1371,1673,1695]" captionStart="FIGURES 66 75" captionStartId="21.[151,264,1855,1878]" captionTargetBox="[186,1400,193,1812]" captionTargetId="figure@21.[170,1417,193,1835]" captionTargetPageId="21" captionText="FIGURES 66 75. Crassabwa ludmilae sp. n., larva. 66, claw; 67, fore leg (anterior view); 68, paraproct; 69 70, right and left mandibles (ventral view); 71, labrum (at left ventral view, at right dorsal view); 72, maxilla; 73, labial palp (ventral view); 74, labium (at left ventral view, at right dorsal view); 75, hypopharynx and superlinguae." httpUri="https://zenodo.org/record/1059207/files/figure.png" pageId="6" pageNumber="407">Figs 66</figureCitation>
,
<figureCitation id="6CA82A1D450E5A2FFA85F905FA9736DE" box="[1382,1407,1673,1695]" captionStart="FIGURES 84 95" captionStartId="23.[151,264,1927,1950]" captionTargetBox="[155,1436,193,1897]" captionTargetId="figure@23.[151,1436,193,1907]" captionTargetPageId="23" captionText="FIGURES 84 95. Crassabwa ludmilae sp. n. 84, tergalius I of last instar larva in natural condition; 85, tergalius I spread on slide; 86 91, tergalii II VII; 92, tergalius V with part of dorsal cuticle detached; 93, enlarged fragment of its detached cuticle; 94, part of cuticle from abdominal tergum III; 95, claw (84 93 and 95, holotype)." httpUri="https://zenodo.org/record/1059211/files/figure.png" pageId="6" pageNumber="407">95</figureCitation>
)................................................................................
<taxonomicName id="33934D1B450E5A2FFB79F92BFA8836FC" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[1178,1376,1703,1725]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="C6E7EA8A450E5A2FFB79F92BFA8836FC" box="[1178,1376,1703,1725]" italics="true" pageId="6" pageNumber="407">Crassabwa ludmilae</emphasis>
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<emphasis id="C6E7EA8A450E5A2FFA85F92BFA7436FC" bold="true" box="[1382,1436,1703,1725]" pageId="6" pageNumber="407">sp. n.</emphasis>
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</keyLead>
</paragraph>
<paragraph id="F42C3698450E5A2FFF74F949FA7436B9" blockId="6.[151,1436,1409,1784]" pageId="6" pageNumber="407">
<keyLead id="4F6295AD450E5A2FFF74F949FA7436B9" pageId="6" pageNumber="407">
- Claw relatively long, with portion from distal denticle to apex longer than portion from distal denticle to base (
<figureCitation id="6CA82A1D450E5A2FFAE4F948FAB0369B" box="[1287,1368,1732,1754]" captionStart="FIGURES 125 132" captionStartId="30.[151,264,1740,1763]" captionTargetBox="[180,1396,184,1698]" captionTargetId="figure@30.[164,1411,171,1705]" captionTargetPageId="30" captionText="FIGURES 125 132. Crassabwa ameliae sp. n., larva. 125, fore leg (anterior view); 126, claw; 127 128, right and left mandibles, ventral view; 129, labrum (at left ventral view, at right dorsal view); 130, maxilla; 131, labium (at left ventral view, at right dorsal view); 132, hypopharynx and superlinguae." httpUri="https://zenodo.org/record/1059223/files/figure.png" pageId="6" pageNumber="407">Fig. 126</figureCitation>
)....................................................................................
<taxonomicName id="33934D1B450E5A2FFB41F96DFA8836B6" authority="Kluge &amp; Gattolliat &amp; Salles, 2017" authorityName="Kluge &amp; Gattolliat &amp; Salles" authorityYear="2017" box="[1186,1376,1761,1783]" class="Insecta" family="Baetidae" genus="Crassabwa" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="ameliae" status="sp. nov.">
<emphasis id="C6E7EA8A450E5A2FFB41F96DFA8836B6" box="[1186,1376,1761,1783]" italics="true" pageId="6" pageNumber="407">Crassabwa ameliae</emphasis>
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<emphasis id="C6E7EA8A450E5A2FFA85F96EFA7436B9" bold="true" box="[1382,1436,1762,1784]" pageId="6" pageNumber="407">sp. n.</emphasis>
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