263 lines
21 KiB
XML
263 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/BDJ.3.e4596" ID-PMC="PMC4698461" ID-Pensoft-Pub="1314-2828--4596" ID-PubMed="26752964" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1314-2828--e4596" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Nine new species of Itaplectops (Diptera: Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, northwestern Costa Rica, with a key to Itaplectops species" checkinTime="1585357573225" checkinUser="pensoft" docAuthor="Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Janzen, Daniel H. & Hallwachs, Winnie" docDate="2015" docId="6C2DBFF4D4F859C45498DC31BB50D21B" docLanguage="en" docName="BiodivDatJour 3: e4596" docOrigin="Biodiversity Data Journal 3" docSource="http://dx.doi.org/10.3897/BDJ.3.e4596" docTitle="Itaplectops griseobasis Fleming, Wood, Smith, Janzen & Hallwachs, 2015, sp. n." docType="treatment" docVersion="3" lastPageNumber="4596" masterDocId="A4229B60FFCCFFBA4464FFC3AE4D9158" masterDocTitle="Nine new species of Itaplectops (Diptera: Tachinidae) reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica, with a key to Itaplectops species" masterLastPageNumber="4596" masterPageNumber="4596" pageNumber="4596" updateTime="1668122940553" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Nine new species of Itaplectops (Diptera: Tachinidae) reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica, with a key to Itaplectops species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fleming, AJ</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wood, D. Monty</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Smith, M. Alex</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Janzen, Daniel H.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hallwachs, Winnie</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Biodiversity Data Journal</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>4596</mods:start>
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<mods:end>4596</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/BDJ.3.e4596</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.3.e4596</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2828--4596</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:6C2DBFF4D4F859C45498DC31BB50D21B" httpUri="http://treatment.plazi.org/id/6C2DBFF4D4F859C45498DC31BB50D21B" lastPageNumber="4596" pageId="0" pageNumber="4596">
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<subSubSection pageId="0" pageNumber="4596" type="nomenclature">
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<paragraph pageId="0" pageNumber="4596">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:F334625E-1950-4156-B60E-CA3658F8D3F9" class="Insecta" family="Tachinidae" genus="Itaplectops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Itaplectops griseobasis" order="Diptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="griseobasis">Itaplectops griseobasis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="4596">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="materials_examined">
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<paragraph pageId="0" pageNumber="4596">Materials</paragraph>
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<paragraph pageId="0" pageNumber="4596">
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<materialsCitation collectingDate="2005-04-04" collectingMethod="reared from caterpillar of Euprosternawemilleri (Limacodidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Gloria Sihezar" country="Costa Rica" latitude="10.9" location="Sector San Cristobal" longitude="-85.373" pageId="0" pageNumber="4596" specimenCode="DHJPAR 0007073" specimenCount="1" typeStatus="Holotype">
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Type status:
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<typeStatus pageId="0" pageNumber="4596">Holotype</typeStatus>
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. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
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<specimenCode pageId="0" pageNumber="4596">DHJPAR0007073</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="4596">D.H. Janzen & W. Hallwachs, Gloria Sihezar</collectorName>
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; individualID: DHJPAR0007073; individualCount:
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<specimenCount pageId="0" pageNumber="4596">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="4596">F</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="4596">adult</specimenType>
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; preparations: pinned; otherCatalogNumbers: ASTAV315-06, 06-SRNP-2159; Taxon: scientificName: Itaplectopsgriseobasis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Itaplectops; specificEpithet: griseobasis; scientificNameAuthorship: Fleming & Wood; Location: continent: Central America; country:
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<collectingCountry pageId="0" pageNumber="4596">Costa Rica</collectingCountry>
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; countryCode: CR; stateProvince: Alajuela; county: Area de Conservacion Guanacaste; locality:
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<location LSID="urn:lsid:plazi:treatment:6C2DBFF4D4F859C45498DC31BB50D21B:278525448DA3D8BDE2A2434D86F0AD0C" country="Costa Rica" latitude="10.9" longitude="-85.373" name="Sector San Cristobal" pageId="0" pageNumber="4596">Sector San Cristobal</location>
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; verbatimLocality: Rio Blanco Abajo; verbatimElevation:
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<elevation pageId="0" pageNumber="4596">500</elevation>
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; verbatimLatitude: 10.9; verbatimLongitude: -85.373; verbatimCoordinateSystem: Decimal; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="4596" value="10.9">10.9</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="4596" value="-85.373">-85.373</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="4596">AJ Fleming</determinerName>
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; dateIdentified: 2014; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="4596">reared from caterpillar of Euprosternawemilleri (Limacodidae)</collectingMethod>
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; verbatimEventDate:
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<collectingDate pageId="0" pageNumber="4596" value="2005-04-04">04-Apr-2005</collectingDate>
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; Record Level: language: en; institutionCode:
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<collectionCode pageId="0" pageNumber="4596">CNC</collectionCode>
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; collectionCode:
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<collectionCode pageId="0" pageNumber="4596">Insects</collectionCode>
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; basisOfRecord: Pinned Specimen
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</materialsCitation>
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<materialsCitation collectingDate="2008-11-29" collectingMethod="reared from caterpillar of Natadamichorta (Limacodidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Petrona Rios" country="Costa Rica" latitude="11.019" location="Sector Pitilla" longitude="-85.41" pageId="0" pageNumber="4596" specimenCode="DHJPAR 0030159" specimenCount="1" typeStatus="Paratype">
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Type status:
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<typeStatus pageId="0" pageNumber="4596">Paratype</typeStatus>
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. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
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<specimenCode pageId="0" pageNumber="4596">DHJPAR0030159</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="4596">D.H. Janzen & W. Hallwachs, Petrona Rios</collectorName>
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; individualID: DHJPAR0030159; individualCount:
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<specimenCount pageId="0" pageNumber="4596">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="4596">F</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="4596">adult</specimenType>
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; preparations: pinned; otherCatalogNumbers: ASHYB903-09, 08-SRNP-32765; Taxon: scientificName: Itaplectopsgriseobasis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Itaplectops; specificEpithet: griseobasis; scientificNameAuthorship: Fleming & Wood; Location: continent: Central America; country:
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<collectingCountry pageId="0" pageNumber="4596">Costa Rica</collectingCountry>
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; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservacion Guanacaste; locality:
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<location LSID="urn:lsid:plazi:treatment:6C2DBFF4D4F859C45498DC31BB50D21B:07C4AA3CEBA56593767B1B4755B318BD" country="Costa Rica" latitude="11.019" longitude="-85.41" name="Sector Pitilla" pageId="0" pageNumber="4596">Sector Pitilla</location>
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; verbatimLocality: Pasmompa; verbatimElevation:
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<elevation pageId="0" pageNumber="4596">440</elevation>
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; verbatimLatitude: 11.019; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="4596" value="11.019">11.019</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="4596" value="-85.41">-85.41</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="4596">AJ Fleming</determinerName>
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; dateIdentified: 2014; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="4596">reared from caterpillar of Natadamichorta (Limacodidae)</collectingMethod>
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; verbatimEventDate:
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<collectingDate pageId="0" pageNumber="4596" value="2008-11-29">29-Nov-2008</collectingDate>
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; Record Level: language: en; institutionCode:
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<collectionCode pageId="0" pageNumber="4596">CNC</collectionCode>
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; collectionCode:
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<collectionCode pageId="0" pageNumber="4596">Insects</collectionCode>
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; basisOfRecord: Pinned Specimen
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</materialsCitation>
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<materialsCitation collectingDate="2007-08-10" collectingMethod="reared from caterpillar of Natadamichorta (Limacodidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Jose Perez" country="Costa Rica" latitude="10.933" location="Sector Rincon Rain Forest" longitude="-85.253" pageId="0" pageNumber="4596" specimenCode="DHJPAR 0020994" specimenCount="1" typeStatus="Paratype">
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Type status:
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<typeStatus pageId="0" pageNumber="4596">Paratype</typeStatus>
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. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
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<specimenCode pageId="0" pageNumber="4596">DHJPAR0020994</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="4596">D.H. Janzen & W. Hallwachs, Jose Perez</collectorName>
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; individualID: DHJPAR0020994; individualCount:
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<specimenCount pageId="0" pageNumber="4596">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="4596">F</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="4596">adult</specimenType>
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; preparations: pinned; otherCatalogNumbers: ASTA1337-07, 07-SRNP-41902; Taxon: scientificName: Itaplectopsgriseobasis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Itaplectops; specificEpithet: griseobasis; scientificNameAuthorship: Fleming & Wood; Location: continent: Central America; country:
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<collectingCountry pageId="0" pageNumber="4596">Costa Rica</collectingCountry>
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; countryCode: CR; stateProvince: Alajuela; county: Area de Conservacion Guanacaste; locality:
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<location LSID="urn:lsid:plazi:treatment:6C2DBFF4D4F859C45498DC31BB50D21B:22838C200C277217EE8C76DD0FABA2F4" country="Costa Rica" latitude="10.933" longitude="-85.253" name="Sector Rincon Rain Forest" pageId="0" pageNumber="4596">Sector Rincon Rain Forest</location>
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; verbatimLocality: Laureles; verbatimElevation:
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<elevation pageId="0" pageNumber="4596">95</elevation>
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; verbatimLatitude: 10.933; verbatimLongitude: -85.253; verbatimCoordinateSystem: Decimal; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="4596" value="10.933">10.933</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="4596" value="-85.253">-85.253</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="4596">AJ Fleming</determinerName>
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; dateIdentified: 2014; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="4596">reared from caterpillar of Natadamichorta (Limacodidae)</collectingMethod>
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; verbatimEventDate:
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<collectingDate pageId="0" pageNumber="4596" value="2007-08-10">10-Aug-2007</collectingDate>
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; Record Level: language: en; institutionCode:
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<collectionCode pageId="0" pageNumber="4596">CNC</collectionCode>
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; collectionCode:
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<collectionCode pageId="0" pageNumber="4596">Insects</collectionCode>
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; basisOfRecord: Pinned Specimen
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</materialsCitation>
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<materialsCitation collectingDate="2008-01-13" collectingMethod="reared from caterpillar of Natadalalogamezi (Limacodidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Anabelle Cordoba" country="Costa Rica" latitude="10.916" location="Sector San Cristobal" longitude="-85.379" pageId="0" pageNumber="4596" specimenCode="DHJPAR 0023604" specimenCount="1" typeStatus="Paratype">
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Type status:
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<typeStatus pageId="0" pageNumber="4596">Paratype</typeStatus>
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. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
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<specimenCode pageId="0" pageNumber="4596">DHJPAR0023604</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="4596">D.H. Janzen & W. Hallwachs, Anabelle Cordoba</collectorName>
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; individualID: DHJPAR0023604; individualCount:
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<specimenCount pageId="0" pageNumber="4596">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="4596">F</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="4596">adult</specimenType>
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; preparations: pinned; otherCatalogNumbers: ASTAW461-08, 07-SRNP-5030; Taxon: scientificName: Itaplectopsgriseobasis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Itaplectops; specificEpithet: griseobasis; scientificNameAuthorship: Fleming & Wood; Location: continent: Central America; country:
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<collectingCountry pageId="0" pageNumber="4596">Costa Rica</collectingCountry>
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; countryCode: CR; stateProvince: Alajuela; county: Area de Conservacion Guanacaste; locality:
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<location LSID="urn:lsid:plazi:treatment:6C2DBFF4D4F859C45498DC31BB50D21B:D4EB7D5A39D3198558E22680A41FA3CA" country="Costa Rica" latitude="10.916" longitude="-85.379" name="Sector San Cristobal" pageId="0" pageNumber="4596">Sector San Cristobal</location>
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; verbatimLocality: Puente Palma; verbatimElevation:
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<elevation pageId="0" pageNumber="4596">460</elevation>
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; verbatimLatitude: 10.916; verbatimLongitude: -85.379; verbatimCoordinateSystem: Decimal; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="4596" value="10.916">10.916</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="4596" value="-85.379">-85.379</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="4596">AJ Fleming</determinerName>
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; dateIdentified: 2014; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="4596">reared from caterpillar of Natadalalogamezi (Limacodidae)</collectingMethod>
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; verbatimEventDate:
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<collectingDate pageId="0" pageNumber="4596" value="2008-01-13">13-Jan-2008</collectingDate>
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; Record Level: language: en; institutionCode:
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<collectionCode pageId="0" pageNumber="4596">CNC</collectionCode>
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; collectionCode:
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<collectionCode pageId="0" pageNumber="4596">Insects</collectionCode>
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; basisOfRecord: Pinned Specimen
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="description">
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<paragraph pageId="0" pageNumber="4596">Description</paragraph>
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<paragraph pageId="0" pageNumber="4596">Male: unknown at this time.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Female</paragraph>
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<paragraph pageId="0" pageNumber="4596">Length: 5-7mm.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Head (Fig. 8c): proclinate orbital bristles present; first flagellomere entirely dark or brownish orange over at least 1/2 of its surface; arista dark brown over 4/5 of its length, bright orange basally, with gradual taper; first flagellomere reaching facial margin; ocellar bristles reduced, almost hair-like, less than the length of pedicel, arising behind anterior ocellus; ocellar triangle covered in small proclinate hairs; frontal vitta approximately 2x as wide as fronto-orbital plate; facial ridge bare, 4-5 supravibrissal bristles; fronto-orbital plate silver; parafacial entirely silver and bare; fronto-orbital plate with fine bristles not confined to a row and interspersed with frontal bristles, these not extending past lowest frontal bristle.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Thorax (Fig. 8a, b): three postsutural supra-alar bristles, anteriormost extremely reduced in size so as to appear hair-like; katepisternum with 2 bristles of equal size, anteriormost arising directly below or slightly in front of suture; apical scutellar bristles short, up to 1/2 length of subapical scutellars; subapical scutellar bristles divergent; scutellum lacking discal bristles.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Wings (Fig. 8a): smoky yellow.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Legs (Fig. 8b): Appearing light colored overall, femur and tibia yellow, and tarsi haired, appearing black but with yellow ground color; dorso-ventral margin of hind tarsi with yellow tufts of bristles at tarsal apices.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Abdomen (Fig. 8a, b): T1+2 with mid-dorsal depression extending halfway along its length, but not reaching tergal margin; median marginal bristles present on all tergites. Discal bristles present on T3, these are shorter than marginal bristles but still obviously bristles. Silver tomentosity entirely covering T1+2 and on anterior margins of abdominal segments T3 and T4, covering up to 1/2 of tergal surface.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="diagnosis">
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<paragraph pageId="0" pageNumber="4596">Diagnosis</paragraph>
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<paragraph pageId="0" pageNumber="4596">
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<taxonomicName class="Insecta" family="Tachinidae" genus="Itaplectops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Itaplectops griseobasis" order="Diptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="griseobasis">Itaplectops griseobasis</taxonomicName>
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is to date only known from females. It is distinguished by the following combination of traits: first flagellomere brown/black over at least 1/2 of surface; legs yellow with black tarsi; median marginal bristles present on all tergites; discal bristles present at least on T3 (in some cases these can appear reduced but still present); silver tomentosity present on all of T1+2 and on margins of abdominal segments T3, T4, and the underside of T5. It can be distinguished from its most similar congener,
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<taxonomicName class="Insecta" family="Tachinidae" genus="Itaplectops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Itaplectops akselpalolai" order="Diptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="akselpalolai">Itaplectops akselpalolai</taxonomicName>
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, following couplet 9 in the key to
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<taxonomicName class="Insecta" family="Tachinidae" genus="Itaplectops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Itaplectops" order="Diptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="genus">Itaplectops</taxonomicName>
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(below).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="etymology">
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<paragraph pageId="0" pageNumber="4596">Etymology</paragraph>
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<paragraph pageId="0" pageNumber="4596">
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From the Latin adjective grisea, meaning
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<normalizedToken originalValue="“gray”">"gray"</normalizedToken>
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, in reference to the silver tomentosity on the entirety of T1+2, and the Latin nound basis meaning "foundation".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="distribution">
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<paragraph pageId="0" pageNumber="4596">Distribution</paragraph>
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<paragraph pageId="0" pageNumber="4596">Costa Rica, ACG, Prov. Alajuela, rain forest.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="ecology">
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<paragraph pageId="0" pageNumber="4596">Ecology</paragraph>
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<subSection pageId="0" pageNumber="4596" type="hosts">
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<paragraph pageId="0" pageNumber="4596">Hosts</paragraph>
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<paragraph pageId="0" pageNumber="4596">
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Reared from caterpillars of 3 species of
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<taxonomicName family="Limacodidae" lsidName="" pageId="0" pageNumber="4596" rank="family">Limacodidae</taxonomicName>
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:
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<taxonomicName class="Insecta" family="Limacodidae" genus="Euprosterna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euprosterna wemilleri" order="Lepidoptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="wemilleri">Euprosterna wemilleri</taxonomicName>
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Corrales & Epstein, 2000;
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<taxonomicName class="Insecta" family="Limacodidae" genus="Natada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Natada lalogamezi" order="Lepidoptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="lalogamezi">Natada lalogamezi</taxonomicName>
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Corrales, 2000; and
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<taxonomicName class="Insecta" family="Limacodidae" genus="Natada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Natada michorta" order="Lepidoptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="michorta">Natada michorta</taxonomicName>
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Dyar, 1912.
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</paragraph>
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</subSection>
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</subSubSection>
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</treatment>
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</document> |