440 lines
47 KiB
XML
440 lines
47 KiB
XML
<document id="5FA04A612C4E15211F382FF59493F301" ID="10.11646/zootaxa.4196.4.1" ID-DOI="10.11646/zootaxa.4196.4.1" ID-GBIF-Dataset="921bb78e-4a75-4fa5-82b6-5c7a75c7ad1e" ID-ISSN="1175-5326" ID-Zenodo-Dep="168273" ID-ZooBank="80863DAE-C9E4-466D-9E27-AD938E826D4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479971428066" checkinUser="plazi" docAuthor="Samyn, Yves & Vandenspiegel, Didier" docDate="2016" docId="6C2D24149066A02FFF79FBAEB3EBF9DA" docLanguage="en" docName="zootaxa.4196.4.1.pdf" docOrigin="Zootaxa 4196 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holothuria (Metriatyla) alex Samyn, sp.nov." docType="treatment" docVersion="14" lastPageNumber="462" masterDocId="90145C6C906FA024FFEEFFFBB052FFF6" masterDocTitle="Sublittoral and bathyal sea cucumbers (Echinodermata: Holothuroidea) from the Northern Mozambique Channel with description of six new species" masterLastPageNumber="497" masterPageNumber="451" pageNumber="460" updateTime="1698394447745" updateUser="plazi">
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<mods:titleInfo id="D721D851FF472283D021082E8AF546E4">
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<mods:title id="CA411D12BBD9739A4DCFF6012881CC2B">Sublittoral and bathyal sea cucumbers (Echinodermata: Holothuroidea) from the Northern Mozambique Channel with description of six new species</mods:title>
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<mods:name id="176FE0F2973E5CB8DE32C3CEF6417A53" type="personal">
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<mods:roleTerm id="74A76AA4B32193877B8A4E339DD4C030">Author</mods:roleTerm>
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<mods:namePart id="26F3D7289C57AD560AB4AA9E1AF3CA56">Samyn, Yves</mods:namePart>
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<mods:name id="2B94EBF1391500AED7E40F1DE7851A64" type="personal">
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<mods:roleTerm id="878C40C07D658A6372A51D00FF014C2D">Author</mods:roleTerm>
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<mods:namePart id="5517C2FC32B7D13FE4DE75A1428D1FBD">Vandenspiegel, Didier</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7FE5817F5D1EE17149BDED9BD8283E50">text</mods:typeOfResource>
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<mods:title id="C3E97053BD5C1B3F4B578D45200BB002">Zootaxa</mods:title>
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<mods:part id="1CDD187360902E10687329DBFA8180D2">
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<mods:date id="60C509E474E14ACE58F98AE375F35328">2016</mods:date>
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<mods:detail id="E3572F1BD1B7205BE2232E8FC6B0F2A8" type="volume">
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<mods:number id="E886F795EDB1BD46B55BDFEC1A4BD2E9">4196</mods:number>
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<mods:number id="0BFC00F0729752C8BC9C02ECDBB0849E">4</mods:number>
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<mods:identifier id="4B1FF482E3E845E6AFEE818774A46CAD" type="DOI">10.11646/zootaxa.4196.4.1</mods:identifier>
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<mods:identifier id="9C215882DCB3AB0D71FF7D52E517EC64" type="ISSN">1175-5326</mods:identifier>
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<treatment id="6C2D24149066A02FFF79FBAEB3EBF9DA" ID-DOI="http://doi.org/10.5281/zenodo.5107176" ID-GBIF-Taxon="124831669" ID-Zenodo-Dep="5107176" LSID="urn:lsid:plazi:treatment:6C2D24149066A02FFF79FBAEB3EBF9DA" httpUri="http://treatment.plazi.org/id/6C2D24149066A02FFF79FBAEB3EBF9DA" lastPageId="11" lastPageNumber="462" pageId="9" pageNumber="460">
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<subSubSection id="AC9EC6899066A02DFF79FBAEB176FB66" pageId="9" pageNumber="460" type="nomenclature">
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<paragraph id="E43B95029066A02DFF79FBAEB2F8FB99" blockId="9.[151,682,1109,1168]" box="[151,682,1109,1135]" pageId="9" pageNumber="460">
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<heading id="BF73226E9066A02DFF79FBAEB2F8FB99" bold="true" box="[151,682,1109,1135]" fontSize="11" level="1" pageId="9" pageNumber="460" reason="1">
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<emphasis id="D6F049109066A02DFF79FBAEB2F8FB99" bold="true" box="[151,682,1109,1135]" pageId="9" pageNumber="460">
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<taxonomicName id="2384EE819066A02DFF79FBAEB21FFB99" authority="Samyn" authorityName="Samyn" box="[151,589,1109,1135]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="9" pageNumber="460" phylum="Echinodermata" rank="species" species="alex" status="sp. nov." subGenus="Metriatyla">
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||
<emphasis id="D6F049109066A02DFF79FBAEB1A3FB99" bold="true" box="[151,497,1109,1135]" italics="true" pageId="9" pageNumber="460">Holothuria (Metriatyla) alex</emphasis>
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Samyn
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</taxonomicName>
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<taxonomicNameLabel id="CDC3F46B9066A02DFDBBFBAEB2F8FB99" box="[597,682,1109,1135]" pageId="9" pageNumber="460" rank="species">sp.nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="E43B95029066A02DFF79FB8CB176FB66" blockId="9.[151,682,1109,1168]" box="[151,292,1143,1168]" pageId="9" pageNumber="460">
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||
(
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<figureCitation id="7CBF89879066A02DFF71FB8CB0B0FB66" box="[159,226,1143,1168]" captionStart="FIGURE 4" captionStartId="10.[151,250,1904,1926]" captionTargetBox="[151,1436,193,1876]" captionTargetId="figure@10.[151,1436,193,1883]" captionTargetPageId="10" captionText="FIGURE 4. Holothuria (Metriatyla) alex sp. nov. A – B: Dorsal (A) and ventral (B) views of holotype; C – F: SEM photos of ossicles from tentacles (C), dorsal body wall (D), lateral papillae (E) and dorsal papillae (F). Scale bars: A, B = 1 cm; C, E, F = 20 µm; D = 10 µm." httpUri="https://zenodo.org/record/258664/files/figure.png" pageId="9" pageNumber="460">Fig. 4</figureCitation>
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A
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<emphasis id="D6F049109066A02DFF12FB8CB158FB66" bold="true" box="[252,266,1143,1168]" pageId="9" pageNumber="460">–</emphasis>
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F)
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</paragraph>
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</subSubSection>
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<subSubSection id="AC9EC6899066A02DFF79FB44B573FA0E" pageId="9" pageNumber="460" type="materials_examined">
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||
<paragraph id="E43B95029066A02DFF79FB44B491FB2E" blockId="9.[151,1437,1215,2032]" box="[151,1219,1215,1240]" pageId="9" pageNumber="460">
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<emphasis id="D6F049109066A02DFF79FB44B1D2FB21" bold="true" box="[151,384,1215,1240]" pageId="9" pageNumber="460">Material examined.</emphasis>
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<emphasis id="D6F049109066A02DFE69FB3AB273FB2E" box="[391,545,1215,1240]" italics="true" pageId="9" pageNumber="460">Type material</emphasis>
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:
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<emphasis id="D6F049109066A02DFDDEFB44B291FB2E" bold="true" box="[560,707,1215,1240]" pageId="9" pageNumber="460">IE-2007-779</emphasis>
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(holotype, sampled in front Mahajamba Bay)
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</paragraph>
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<paragraph id="E43B95029066A02DFF29FB18B4CBFA22" blockId="9.[151,1437,1215,2032]" pageId="9" pageNumber="460">
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<materialsCitation id="54EC9F5F9066A02DFF29FB18B3D6FAD6" ID-GBIF-Occurrence="1343648481" collectionCode="ZMB" collectorName="V. Martens" country="Indonesia" location="Ech" municipality="Semper" pageId="9" pageNumber="460" specimenCount="1" typeStatus="holotype">
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<emphasis id="D6F049109066A02DFF29FB18B341FB0A" bold="true" box="[199,787,1251,1276]" pageId="9" pageNumber="460">
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Comparative material examined.
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<collectionCode id="82950DC79066A02DFDB2FB18B2CEFB0A" box="[604,668,1251,1276]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="9" pageNumber="460">ZMB</collectionCode>
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<location id="E15BC3D99066A02DFD4AFB18B281FB0A" LSID="urn:lsid:plazi:treatment:6C2D24149066A02FFF79FBAEB3EBF9DA:E15BC3D99066A02DFD4AFB18B281FB0A" box="[676,723,1251,1276]" country="Indonesia" municipality="Semper" name="Ech" pageId="9" pageNumber="460">Ech</location>
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1681
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</emphasis>
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||
(
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<typeStatus id="3B3F2BA09066A02DFCCDFB18B3D5FB0A" box="[803,903,1251,1276]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
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of
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<emphasis id="D6F049109066A02DFC41FB18B56AFB0A" box="[943,1336,1251,1276]" italics="true" pageId="9" pageNumber="460">
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<taxonomicName id="2384EE819066A02DFC41FB18B563FB0A" box="[943,1329,1251,1276]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="9" pageNumber="460" phylum="Echinodermata" rank="species" species="martensii" subGenus="Metriatyla">Holothuria (Metriatyla) martensii</taxonomicName>
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;
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</emphasis>
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<collectingMunicipality id="045F0F789066A02DFAAEFB1FB5C5FB0A" box="[1344,1431,1252,1276]" pageId="9" pageNumber="460">Semper</collectingMunicipality>
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, 1868 (
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<figureCitation id="7CBF89879066A02DFF30FAFCB170FAD6" box="[222,290,1287,1312]" captionStart="FIGURE 5" captionStartId="11.[151,250,517,539]" captionTargetBox="[205,1381,193,494]" captionTargetId="figure@11.[205,1381,193,495]" captionTargetPageId="11" captionText="FIGURE 5. Holothuria (Metriatyla) martensii Semper, 1868. Label and dorsal view of the holotype." httpUri="https://zenodo.org/record/168278/files/figure.png" pageId="9" pageNumber="460">Fig. 5</figureCitation>
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);
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<collectingCountry id="9C93D5929066A02DFED9FAFCB1DCFAD6" box="[311,398,1287,1312]" name="Indonesia" pageId="9" pageNumber="460">Ambon</collectingCountry>
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,
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<collectingCountry id="9C93D5929066A02DFE77FAFCB25BFAD6" box="[409,521,1287,1312]" name="Indonesia" pageId="9" pageNumber="460">Indonesia</collectingCountry>
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||
, leg.
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<collectorName id="4971F0D49066A02DFDADFAF3B292FAE9" box="[579,704,1288,1311]" pageId="9" pageNumber="460">V. Martens</collectorName>
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||
, unknown depth)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="54EC9F5F9066A02DFC60FAFCB23CFA9E" ID-GBIF-Occurrence="1343648496" collectingDate="1962-09-15" collectionCode="MNHN" collectorName="Thomassin" country="Madagascar" county="Mahavatsy" location="EchHh" municipality="Tulear" pageId="9" pageNumber="460" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="D6F049109066A02DFC60FAFCB4D2FAD6" bold="true" box="[910,1152,1287,1312]" pageId="9" pageNumber="460">
|
||
<collectionCode id="82950DC79066A02DFC60FAFCB3B5FAD6" box="[910,999,1287,1312]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="460">MNHN</collectionCode>
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||
<location id="E15BC3D99066A02DFC03FAFCB413FAD6" LSID="urn:lsid:plazi:treatment:6C2D24149066A02FFF79FBAEB3EBF9DA:E15BC3D99066A02DFC03FAFCB413FAD6" box="[1005,1089,1287,1312]" country="Madagascar" county="Mahavatsy" municipality="Tulear" name="EchHh" pageId="9" pageNumber="460">EchHh</location>
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||
3894
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||
</emphasis>
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||
(
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||
<taxonomicName id="2384EE819066A02DFB7EFAFCB1DDFAB2" authority="Massin, 1987" authorityName="Massin" authorityYear="1987" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="9" pageNumber="460" phylum="Echinodermata" rank="species" species="horrida" subGenus="Metriatyla">
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||
<emphasis id="D6F049109066A02DFB7EFAFCB0BFFAB2" italics="true" pageId="9" pageNumber="460">Holothuria (Metriatyla) horrida</emphasis>
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||
<bibRefCitation id="8015E8F39066A02DFF1AFAD0B1DDFAB2" author="Massin" box="[244,399,1323,1348]" pageId="9" pageNumber="460" refString="Massin, C. (1987) Holothuries nouvelles et peu connues recoltees en Indonesie au cours de la Snellius II Expedition. Bulletin de l'Institut Royal des Sciences naturelles de Belgique, Biologie, 57, 97 - 121." type="journal article" year="1987">Massin, 1987</bibRefCitation>
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||
</taxonomicName>
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||
, erroneously identified as
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||
<taxonomicName id="2384EE819066A02DFD57FAD6B3BEFAB2" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" box="[697,1004,1323,1348]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="9" pageNumber="460" phylum="Echinodermata" rank="species" species="martensii">
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||
<emphasis id="D6F049109066A02DFD57FAD6B314FAB2" box="[697,838,1325,1348]" italics="true" pageId="9" pageNumber="460">H. martensii</emphasis>
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||
Semper, 1868
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</taxonomicName>
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,
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<collectingMunicipality id="045F0F789066A02DFC16FAD0B410FAB2" box="[1016,1090,1323,1348]" pageId="9" pageNumber="460">Tuléar</collectingMunicipality>
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||
,
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||
<collectingCounty id="0D5AED8E9066A02DFBA0FAD0B498FAB2" box="[1102,1226,1323,1348]" pageId="9" pageNumber="460">Mahavatsy</collectingCounty>
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||
,
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||
<collectingCountry id="9C93D5929066A02DFB38FAD0B50DFAB2" box="[1238,1375,1323,1348]" name="Madagascar" pageId="9" pageNumber="460">Madagascar</collectingCountry>
|
||
, coll.
|
||
<collectorName id="4971F0D49066A02DFF79FAB4B147FA9E" box="[151,277,1359,1384]" pageId="9" pageNumber="460">Thomassin</collectorName>
|
||
,
|
||
<date id="903AB3C29066A02DFECCFAB4B1F6FA9E" box="[290,420,1359,1384]" pageId="9" pageNumber="460" value="1962-09-15">
|
||
<collectingDate id="807E4A2A9066A02DFECCFAB4B1F6FA9E" box="[290,420,1359,1384]" pageId="9" pageNumber="460" value="1962-09-15">15.IX.1962</collectingDate>
|
||
</date>
|
||
, unknown depth)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="54EC9F5F9066A02DFD92FAB4B1B0FA46" ID-GBIF-Occurrence="1343648489" collectingDate="1962-09-23" collectionCode="MNHN" collectorName="Thomassin" country="Madagascar" location="EchHh" municipality="Tulear" pageId="9" pageNumber="460" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="D6F049109066A02DFD92FAB4B321FA9E" bold="true" box="[636,883,1359,1384]" pageId="9" pageNumber="460">
|
||
<collectionCode id="82950DC79066A02DFD92FAB4B287FA9E" box="[636,725,1359,1384]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="460">MNHN</collectionCode>
|
||
<location id="E15BC3D99066A02DFD33FAB4B363FA9E" LSID="urn:lsid:plazi:treatment:6C2D24149066A02FFF79FBAEB3EBF9DA:E15BC3D99066A02DFD33FAB4B363FA9E" box="[733,817,1359,1384]" country="Madagascar" municipality="Tulear" name="EchHh" pageId="9" pageNumber="460">EchHh</location>
|
||
3906
|
||
</emphasis>
|
||
(
|
||
<taxonomicName id="2384EE819066A02DFC6BFAB4B5CAFA9E" authority="Massin, 1997" authorityName="Massin" authorityYear="1997" box="[901,1432,1359,1384]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="9" pageNumber="460" phylum="Echinodermata" rank="species" species="horrida" subGenus="Metriatyla">
|
||
<emphasis id="D6F049109066A02DFC6BFAB4B4A0FA9E" box="[901,1266,1359,1384]" italics="true" pageId="9" pageNumber="460">Holothuria (Metriatyla) horrida</emphasis>
|
||
Massin, 1997
|
||
</taxonomicName>
|
||
, erroneously identified as
|
||
<taxonomicName id="2384EE819066A02DFE54FA8EB2A0FA7A" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" box="[442,754,1395,1420]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="9" pageNumber="460" phylum="Echinodermata" rank="species" species="martensii">
|
||
<emphasis id="D6F049109066A02DFE54FA8EB21BFA7A" box="[442,585,1397,1420]" italics="true" pageId="9" pageNumber="460">H. martensii</emphasis>
|
||
Semper, 1868
|
||
</taxonomicName>
|
||
,
|
||
<collectingMunicipality id="045F0F789066A02DFD11FA88B31BFA7A" box="[767,841,1395,1420]" pageId="9" pageNumber="460">Tuléar</collectingMunicipality>
|
||
, devant labo, st. 450,
|
||
<collectingCountry id="9C93D5929066A02DFBA7FA88B480FA7A" box="[1097,1234,1395,1420]" name="Madagascar" pageId="9" pageNumber="460">Madagascar</collectingCountry>
|
||
, coll.
|
||
<collectorName id="4971F0D49066A02DFAF4FA88B5CAFA7A" box="[1306,1432,1395,1420]" pageId="9" pageNumber="460">Thomassin</collectorName>
|
||
,
|
||
<date id="903AB3C29066A02DFF79FA6CB14BFA46" box="[151,281,1431,1456]" pageId="9" pageNumber="460" value="1962-09-23">
|
||
<collectingDate id="807E4A2A9066A02DFF79FA6CB14BFA46" box="[151,281,1431,1456]" pageId="9" pageNumber="460" value="1962-09-23">23.IX.1962</collectingDate>
|
||
</date>
|
||
, unknown depth)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="54EC9F5F9066A02DFE00FA6CB4CBFA22" ID-GBIF-Occurrence="1343648491" collectionCode="MNHN" collectorName="Decary" country="Madagascar" location="EchHh" municipality="Antsirane" pageId="9" pageNumber="460" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="D6F049109066A02DFE00FA6CB2B6FA46" bold="true" box="[494,740,1431,1456]" pageId="9" pageNumber="460">
|
||
<collectionCode id="82950DC79066A02DFE00FA6CB215FA46" box="[494,583,1431,1456]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="460">MNHN</collectionCode>
|
||
<location id="E15BC3D99066A02DFDA1FA6CB2F1FA46" LSID="urn:lsid:plazi:treatment:6C2D24149066A02FFF79FBAEB3EBF9DA:E15BC3D99066A02DFDA1FA6CB2F1FA46" box="[591,675,1431,1456]" country="Madagascar" municipality="Antsirane" name="EchHh" pageId="9" pageNumber="460">EchHh</location>
|
||
7030
|
||
</emphasis>
|
||
(
|
||
<taxonomicName id="2384EE819066A02DFD1BFA6CB4C5FA46" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" box="[757,1175,1431,1456]" class="Holothuroidea" family="Holothuriidae" genus="Bohadschia" kingdom="Animalia" order="Aspidochirotida" pageId="9" pageNumber="460" phylum="Echinodermata" rank="species" species="vitiensis">
|
||
<emphasis id="D6F049109066A02DFD1BFA6CB38CFA46" box="[757,990,1431,1456]" italics="true" pageId="9" pageNumber="460">Bohadschia vitiensis</emphasis>
|
||
(Semper, 1868)
|
||
</taxonomicName>
|
||
, erroneously identified as
|
||
<taxonomicName id="2384EE819066A02DFF5BFA46B1B4FA22" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" box="[181,486,1467,1492]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="9" pageNumber="460" phylum="Echinodermata" rank="species" species="martensii">
|
||
<emphasis id="D6F049109066A02DFF5BFA46B113FA22" box="[181,321,1469,1492]" italics="true" pageId="9" pageNumber="460">H. martensii</emphasis>
|
||
Semper, 1868
|
||
</taxonomicName>
|
||
,
|
||
<collectingMunicipality id="045F0F789066A02DFE1FFA40B233FA22" box="[497,609,1467,1492]" pageId="9" pageNumber="460">Antsirane</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="9C93D5929066A02DFD85FA40B2A6FA22" box="[619,756,1467,1492]" name="Madagascar" pageId="9" pageNumber="460">Madagascar</collectingCountry>
|
||
, coll.
|
||
<collectorName id="4971F0D49066A02DFCD9FA47B3DBFA22" box="[823,905,1468,1492]" pageId="9" pageNumber="460">Decary</collectorName>
|
||
, 1919, unknown depth).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="E43B95029066A02DFF29FA24B573FA0E" blockId="9.[151,1437,1215,2032]" box="[199,1313,1503,1528]" pageId="9" pageNumber="460">
|
||
<materialsCitation id="54EC9F5F9066A02DFF29FA24B573FA0E" ID-GBIF-Occurrence="1343648480" box="[199,1313,1503,1528]" collectorName="Station CP" country="Madagascar" location="Mahajamba Bay" pageId="9" pageNumber="460" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="D6F049109066A02DFF29FA24B10FFA0E" bold="true" box="[199,349,1503,1528]" pageId="9" pageNumber="460">
|
||
<typeStatus id="3B3F2BA09066A02DFF29FA24B150FA0E" box="[199,258,1503,1528]" pageId="9" pageNumber="460">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="9C93D5929066A02DFE82FA24B1A7FA0E" box="[364,501,1503,1528]" name="Madagascar" pageId="9" pageNumber="460">Madagascar</collectingCountry>
|
||
,
|
||
<location id="E15BC3D99066A02DFDEEFA24B2EBFA0E" LSID="urn:lsid:plazi:treatment:6C2D24149066A02FFF79FBAEB3EBF9DA:E15BC3D99066A02DFDEEFA24B2EBFA0E" box="[512,697,1503,1528]" country="Madagascar" name="Mahajamba Bay" pageId="9" pageNumber="460">Mahajamba Bay</location>
|
||
,
|
||
<collectorName id="4971F0D49066A02DFD2AFA24B36EFA0E" box="[708,828,1503,1528]" pageId="9" pageNumber="460">Station CP</collectorName>
|
||
3218 (Decimal coordinates: -14.97; 46.94)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC9EC6899066A02DFF29F9F8B418F97E" pageId="9" pageNumber="460" type="etymology">
|
||
<paragraph id="E43B95029066A02DFF29F9F8B191F992" blockId="9.[151,1437,1215,2032]" pageId="9" pageNumber="460">
|
||
<emphasis id="D6F049109066A02DFF29F9F8B114F9EA" bold="true" box="[199,326,1539,1564]" pageId="9" pageNumber="460">Etymology</emphasis>
|
||
. This very handsome species is named after
|
||
<taxonomicName id="2384EE819066A02DFCA9F9F8B385F9EA" authority="Samyn" authorityName="Samyn" box="[839,983,1539,1564]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="9" pageNumber="460" phylum="Echinodermata" rank="species" species="alex" status="sp. nov.">Alex Samyn</taxonomicName>
|
||
, first daughter of Reen Tallon and Yves Samyn, in recognition of her support, when her dad is writing up his taxonomic work at home. The species epithet has been put in apposition.
|
||
</paragraph>
|
||
<paragraph id="E43B95029066A02DFF29F994B418F97E" blockId="9.[151,1437,1215,2032]" box="[199,1098,1647,1672]" pageId="9" pageNumber="460">
|
||
<emphasis id="D6F049109066A02DFF29F994B201F97E" bold="true" box="[199,595,1647,1672]" pageId="9" pageNumber="460">Known geographical distribution</emphasis>
|
||
. For now only known from the type locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC9EC6899066A02FFF29F968B53BFC76" lastPageId="11" lastPageNumber="462" pageId="9" pageNumber="460" type="description">
|
||
<paragraph id="E43B95029066A02DFF29F968B2B5F896" blockId="9.[151,1437,1215,2032]" pageId="9" pageNumber="460">
|
||
<emphasis id="D6F049109066A02DFF29F968B208F95A" bold="true" box="[199,602,1683,1708]" pageId="9" pageNumber="460">
|
||
Taxonomic description (
|
||
<typeStatus id="3B3F2BA09066A02DFE06F968B200F95A" box="[488,594,1683,1708]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
|
||
)
|
||
</emphasis>
|
||
.
|
||
<emphasis id="D6F049109066A02DFD84F968B360F95A" box="[618,818,1683,1708]" italics="true" pageId="9" pageNumber="460">External anatomy</emphasis>
|
||
—
|
||
<quantity id="237C38E79066A02DFCBEF968B3CDF95D" box="[848,927,1683,1708]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="460" unit="mm" value="55.0">55 mm</quantity>
|
||
long and
|
||
<quantity id="237C38E79066A02DFBFCF96FB430F95D" box="[1042,1122,1684,1708]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="460" unit="mm" value="24.0">24 mm</quantity>
|
||
wide at mid body, tapering acutely posteriorly. Trivium flat, separated from more convex bivium by fringe of well-developed lateral papillae (fig. 4A,B). Mouth ventral, surrounded by collar of reduced papillae, especially ventrally. Number of tentacles could not be determined due to contraction. Anus terminal, unguarded by papillae. Color in alcohol: bivium dark beige with some darker spots forming more or less two transversal lines with beige, evenly spread, papillae; trivium brown, with evenly scattered beige tube feet.
|
||
</paragraph>
|
||
<paragraph id="E43B95029066A02DFF29F890B351F806" blockId="9.[151,1437,1215,2032]" pageId="9" pageNumber="460">
|
||
<emphasis id="D6F049109066A02DFF29F890B1D8F872" box="[199,394,1899,1924]" italics="true" pageId="9" pageNumber="460">Internal anatomy</emphasis>
|
||
: specimen partly eviscerated, leaving only part of the gut, the right respiratory tree and the oral structures. Calcareous ring narrow. Single short stone canal,
|
||
<quantity id="237C38E79066A02DFC7DF86BB38AF851" box="[915,984,1936,1960]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="460" unit="mm" value="4.0">4 mm</quantity>
|
||
long, ending in spherical madreporite. Polian vesicles not observed. Gonad not observed. Gut filled with very fine mud. Longitudinal muscles very flat, bifid, only
|
||
<quantity id="237C38E79066A02DFEFDF823B101F819" box="[275,339,2008,2032]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="460" unit="mm" value="3.0">3 mm</quantity>
|
||
wide. Cuvierian tubules not observed.
|
||
</paragraph>
|
||
<caption id="B0FBC58A9065A02EFF79F88BB102F835" httpUri="https://zenodo.org/record/258664/files/figure.png" pageId="10" pageNumber="461" targetBox="[151,1436,193,1876]" targetPageId="10">
|
||
<paragraph id="E43B95029065A02EFF79F88BB102F835" blockId="10.[151,1436,1904,1987]" pageId="10" pageNumber="461">
|
||
<emphasis id="D6F049109065A02EFF79F88BB146F873" bold="true" box="[151,276,1904,1926]" pageId="10" pageNumber="461">FIGURE 4.</emphasis>
|
||
<taxonomicName id="2384EE819065A02EFEF2F88BB213F870" box="[284,577,1904,1926]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="10" pageNumber="461" phylum="Echinodermata" rank="species" species="alex" status="sp. nov." subGenus="Metriatyla">
|
||
<emphasis id="D6F049109065A02EFEF2F88BB213F870" box="[284,577,1904,1926]" italics="true" pageId="10" pageNumber="461">Holothuria (Metriatyla) alex</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D6F049109065A02EFDA7F88BB281F873" bold="true" box="[585,723,1904,1925]" pageId="10" pageNumber="461">
|
||
<taxonomicNameLabel id="CDC3F46B9065A02EFDA7F88BB2C9F873" box="[585,667,1904,1925]" pageId="10" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A–B
|
||
</emphasis>
|
||
: Dorsal (A) and ventral (B) views of holotype;
|
||
<emphasis id="D6F049109065A02EFB2DF88BB4A3F870" bold="true" box="[1219,1265,1904,1926]" pageId="10" pageNumber="461">C–F</emphasis>
|
||
: SEM photos of ossicles from tentacles (C), dorsal body wall (D), lateral papillae (E) and dorsal papillae (F). Scale bars: A,B = 1cm; C,E,F = 20µm; D = 10µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B0FBC58A9064A02FFF79FDFEB4C5FDEC" box="[151,1175,516,539]" httpUri="https://zenodo.org/record/168278/files/figure.png" pageId="11" pageNumber="462" targetBox="[205,1381,193,494]" targetPageId="11">
|
||
<paragraph id="E43B95029064A02FFF79FDFEB4C5FDEC" blockId="11.[151,1175,516,539]" box="[151,1175,516,539]" pageId="11" pageNumber="462">
|
||
<emphasis id="D6F049109064A02FFF79FDFEB140FDEC" bold="true" box="[151,274,517,539]" pageId="11" pageNumber="462">FIGURE 5.</emphasis>
|
||
<taxonomicName id="2384EE819064A02FFEF7FDFFB355FDEC" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" box="[281,775,516,539]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="martensii" subGenus="Metriatyla">
|
||
<emphasis id="D6F049109064A02FFEF7FDFFB220FDED" box="[281,626,516,539]" italics="true" pageId="11" pageNumber="462">Holothuria (Metriatyla) martensii</emphasis>
|
||
Semper, 1868
|
||
</taxonomicName>
|
||
. Label and dorsal view of the holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E43B95029064A02FFF29FDBCB53BFC76" blockId="11.[151,1437,583,1580]" pageId="11" pageNumber="462">
|
||
<emphasis id="D6F049109064A02FFF29FDBCB176FD96" box="[199,292,583,608]" italics="true" pageId="11" pageNumber="462">Ossicles</emphasis>
|
||
: Tentacles with spiny rods, occasionally with some perforations, 110
|
||
<emphasis id="D6F049109064A02FFBDDFDBCB413FD96" bold="true" box="[1075,1089,583,608]" pageId="11" pageNumber="462">–</emphasis>
|
||
480 µm long (fig. 4C). Ventral and dorsal body wall with tables, buttons and few fenestrated ellipsoids. Tables with disc on average 60 µm across, perforated by irregularly distributed central holes and 14 to 16 peripheral holes, rim of the disc undulating, edge smooth to spiny, spire low (1
|
||
<emphasis id="D6F049109064A02FFE30FD48B1BEFD3A" bold="true" box="[478,492,691,716]" pageId="11" pageNumber="462">–</emphasis>
|
||
2 cross beams) with four pillars (30
|
||
<emphasis id="D6F049109064A02FFC93FD48B3D9FD3A" bold="true" box="[893,907,691,716]" pageId="11" pageNumber="462">–</emphasis>
|
||
38µm high) ending in simple spiny crown (fig. 4D). Buttons very wide (31
|
||
<emphasis id="D6F049109064A02FFE23FD2CB189FD06" bold="true" box="[461,475,727,752]" pageId="11" pageNumber="462">–</emphasis>
|
||
48 µm wide and µm 50
|
||
<emphasis id="D6F049109064A02FFD0BFD2CB2A1FD06" bold="true" box="[741,755,727,752]" pageId="11" pageNumber="462">–</emphasis>
|
||
68 long), smooth to knobbed with a central bridge (fig. 4D). Dorsal papillae with tables, plates and numerous ellipsoids (36
|
||
<emphasis id="D6F049109064A02FFC9AFD00B3D0FCE2" bold="true" box="[884,898,763,788]" pageId="11" pageNumber="462">–</emphasis>
|
||
50µm wide, 45
|
||
<emphasis id="D6F049109064A02FFBDDFD00B413FCE2" bold="true" box="[1075,1089,763,788]" pageId="11" pageNumber="462">–</emphasis>
|
||
53µm long); plates flat or tridimensional (123
|
||
<emphasis id="D6F049109064A02FFEB4FCE4B13AFCCE" bold="true" box="[346,360,799,824]" pageId="11" pageNumber="462">–</emphasis>
|
||
185µm long,), tables similar to those of the body wall with a few tables with a higher spire (43
|
||
<emphasis id="D6F049109064A02FFA60FCE4B5CEFCCE" bold="true" box="[1422,1436,799,824]" pageId="11" pageNumber="462">–</emphasis>
|
||
72µm, up to five cross beams, fig. 4F). Lateral papillae with numerous fenestrated ellipsoids in addition to plates and tables similar to those of the dorsal papillae (fig. 4E). Longitudinal muscles and cloaca devoid of ossicles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC9EC6899064A02FFF29FC70B481FAA2" pageId="11" pageNumber="462" type="discussion">
|
||
<paragraph id="E43B95029064A02FFF29FC70B4C1FBF9" blockId="11.[151,1437,583,1580]" pageId="11" pageNumber="462">
|
||
<emphasis id="D6F049109064A02FFF29FC70B169FC52" bold="true" box="[199,315,907,932]" pageId="11" pageNumber="462">Remarks.</emphasis>
|
||
It is with some doubt that we assigned this species to the subgenus
|
||
<taxonomicName id="2384EE819064A02FFB6CFC70B4AAFC52" box="[1154,1272,907,932]" class="Holothuroidea" family="Holothuriidae" genus="Metriatyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="D6F049109064A02FFB6CFC70B4AAFC52" box="[1154,1272,907,932]" italics="true" pageId="11" pageNumber="462">Metriatyla</emphasis>
|
||
</taxonomicName>
|
||
as its ossicle assemblage does not completely fit with the diagnosis of the subgenus.
|
||
<taxonomicName id="2384EE819064A02FFC32FC4BB463FC3E" box="[988,1073,943,968]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex" status="sp. nov.">
|
||
<emphasis id="D6F049109064A02FFC32FC4BB463FC3E" box="[988,1073,943,968]" italics="true" pageId="11" pageNumber="462">H. alex</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D6F049109064A02FFBD5FC54B4CAFC3E" bold="true" box="[1083,1176,943,968]" pageId="11" pageNumber="462">
|
||
<taxonomicNameLabel id="CDC3F46B9064A02FFBD5FC54B4CAFC3E" box="[1083,1176,943,968]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from the other species in
|
||
<taxonomicName id="2384EE819064A02FFEE3FC28B1D1FC1A" box="[269,387,979,1004]" class="Holothuroidea" family="Holothuriidae" genus="Metriatyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="D6F049109064A02FFEE3FC28B1D1FC1A" box="[269,387,979,1004]" italics="true" pageId="11" pageNumber="462">Metriatyla</emphasis>
|
||
</taxonomicName>
|
||
in having tables with discs that, when the spire is low, are turned upwards and are nodulous.
|
||
<taxonomicName id="2384EE819064A02FFF79FC03B0BAFBE6" box="[151,232,1015,1040]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex">
|
||
<emphasis id="D6F049109064A02FFF79FC03B0BAFBE6" box="[151,232,1015,1040]" italics="true" pageId="11" pageNumber="462">H. alex</emphasis>
|
||
</taxonomicName>
|
||
also presents fenestrated ellipsoids, whereas no other species in
|
||
<taxonomicName id="2384EE819064A02FFC52FC0CB460FBE6" box="[956,1074,1015,1040]" class="Holothuroidea" family="Holothuriidae" genus="Metriatyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="D6F049109064A02FFC52FC0CB460FBE6" box="[956,1074,1015,1040]" italics="true" pageId="11" pageNumber="462">Metriatyla</emphasis>
|
||
</taxonomicName>
|
||
does so.
|
||
</paragraph>
|
||
<paragraph id="E43B95029064A02FFF29FBE7B25CFB69" blockId="11.[151,1437,583,1580]" pageId="11" pageNumber="462">
|
||
In terms of external morphology
|
||
<taxonomicName id="2384EE819064A02FFDACFBE7B29CFBC2" box="[578,718,1051,1076]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex" status="sp. nov." subGenus="Metriatyla">
|
||
<emphasis id="D6F049109064A02FFDACFBE7B29CFBC2" box="[578,718,1051,1076]" italics="true" pageId="11" pageNumber="462">H. (M.) alex</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D6F049109064A02FFD38FBE0B360FBC2" bold="true" box="[726,818,1051,1076]" pageId="11" pageNumber="462">
|
||
<taxonomicNameLabel id="CDC3F46B9064A02FFD38FBE0B360FBC2" box="[726,818,1051,1076]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is close to
|
||
<taxonomicName id="2384EE819064A02FFC58FBE7B4A2FBC2" authority="Ludwig, 1891" authorityName="Ludwig" authorityYear="1891" box="[950,1264,1051,1076]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="kurti" subGenus="Metriatyla">
|
||
<emphasis id="D6F049109064A02FFC58FBE7B41BFBC2" box="[950,1097,1051,1076]" italics="true" pageId="11" pageNumber="462">H. (M.) kurti</emphasis>
|
||
<bibRefCitation id="8015E8F39064A02FFBBFFBE0B4A2FBC2" author="Ludwig" box="[1105,1264,1051,1076]" pageId="11" pageNumber="462" refString="Ludwig, H. (1891; 1889 - 92) Echinodermen: Die Seewalzen. In: H. G. Bronn, ed. Broon's Klassen und Ordnungen des Thierreichs. Bd. 2, Abteilung 3, Buch 1, i - iv, 1 - 460, 17 plates. CF. Winter'sche, Leipzig." type="book chapter" year="1891">Ludwig, 1891</bibRefCitation>
|
||
</taxonomicName>
|
||
(see below for description).
|
||
<taxonomicName id="2384EE819064A02FFEC2FBBBB12CFBAE" box="[300,382,1087,1112]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex">
|
||
<emphasis id="D6F049109064A02FFEC2FBBBB12CFBAE" box="[300,382,1087,1112]" italics="true" pageId="11" pageNumber="462">H. alex</emphasis>
|
||
</taxonomicName>
|
||
can however easily be separated from that species because it lacks the ‘spatulate’ tables that characterize
|
||
<taxonomicName id="2384EE819064A02FFECBFB9FB12EFB8A" box="[293,380,1123,1148]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="kurti">
|
||
<emphasis id="D6F049109064A02FFECBFB9FB12EFB8A" box="[293,380,1123,1148]" italics="true" pageId="11" pageNumber="462">H. kurti</emphasis>
|
||
</taxonomicName>
|
||
(cf fig. 7C) and because of its very characteristic ellipsoids (fig. 4 E,F) and wide buttons (fig. 4D) which are absent in
|
||
<taxonomicName id="2384EE819064A02FFE47FB73B253FB56" box="[425,513,1159,1184]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="kurti">
|
||
<emphasis id="D6F049109064A02FFE47FB73B253FB56" box="[425,513,1159,1184]" italics="true" pageId="11" pageNumber="462">H. kurti</emphasis>
|
||
</taxonomicName>
|
||
..
|
||
</paragraph>
|
||
<paragraph id="E43B95029064A02FFF29FB57B481FAA2" blockId="11.[151,1437,583,1580]" pageId="11" pageNumber="462">
|
||
At a first glance one could also think that
|
||
<taxonomicName id="2384EE819064A02FFD72FB57B2BFFB32" box="[668,749,1195,1220]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex" status="sp. nov.">
|
||
<emphasis id="D6F049109064A02FFD72FB57B2BFFB32" box="[668,749,1195,1220]" italics="true" pageId="11" pageNumber="462">H. alex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CDC3F46B9064A02FFD1AFB50B31CFB32" box="[756,846,1195,1220]" pageId="11" pageNumber="462" rank="species">
|
||
<emphasis id="D6F049109064A02FFD1AFB50B31CFB32" bold="true" box="[756,846,1195,1220]" pageId="11" pageNumber="462">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is just a juvenile form of the species described as
|
||
<taxonomicName id="2384EE819064A02FFA6FFB57B1C4FB1E" authority="Jaeger, 1833" authorityName="Jaeger" authorityYear="1833" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="ocellata">
|
||
<emphasis id="D6F049109064A02FFA6FFB57B0A3FB1E" italics="true" pageId="11" pageNumber="462">H. ocellata</emphasis>
|
||
(
|
||
<bibRefCitation id="8015E8F39064A02FFF11FB2BB1DCFB1E" author="Jaeger" box="[255,398,1232,1256]" pageId="11" pageNumber="462" refString="Jaeger, G. F. (1833) De Holothuriis. Dissertatio Inauguralis, Turici. laguerda, 40 pps., 3 pls." type="journal article" year="1833">Jaeger, 1833</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="8015E8F39064A02FFE2EFB2BB2D1FB1E" author="Teo" box="[448,643,1232,1256]" pageId="11" pageNumber="462" refString="Teo, S. & Ng, C. S. L. (2009) New record of a sea cucumber, Holothuria ocellata (Jaeger, 1833) (Holothuroidea: Aspidochirotida: Holothuriidae) in Singapore. Nature in Singapore, 2, 411 - 414." type="journal article" year="2009">Teo & Ng (2009)</bibRefCitation>
|
||
. However, the coloration pattern (papillae surrounded by a dark ring in
|
||
<taxonomicName id="2384EE819064A02FFF79FB0FB147FAFA" box="[151,277,1267,1292]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="ocellata">
|
||
<emphasis id="D6F049109064A02FFF79FB0FB147FAFA" box="[151,277,1267,1292]" italics="true" pageId="11" pageNumber="462">H. ocellata</emphasis>
|
||
</taxonomicName>
|
||
(sensu
|
||
<bibRefCitation id="8015E8F39064A02FFE85FB0FB24CFAFA" author="Teo" box="[363,542,1268,1292]" pageId="11" pageNumber="462" refString="Teo, S. & Ng, C. S. L. (2009) New record of a sea cucumber, Holothuria ocellata (Jaeger, 1833) (Holothuroidea: Aspidochirotida: Holothuriidae) in Singapore. Nature in Singapore, 2, 411 - 414." type="journal article" year="2009">Teo & Ng 2009</bibRefCitation>
|
||
) which is not the case in
|
||
<taxonomicName id="2384EE819064A02FFCD1FB0FB3C3FAFA" box="[831,913,1267,1292]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex">
|
||
<emphasis id="D6F049109064A02FFCD1FB0FB3C3FAFA" box="[831,913,1267,1292]" italics="true" pageId="11" pageNumber="462">H. alex</emphasis>
|
||
</taxonomicName>
|
||
), the form of the lateral rim of papillae (fused papillae in
|
||
<taxonomicName id="2384EE819064A02FFEFAFAE3B1C3FAC6" box="[276,401,1303,1328]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="ocellata">
|
||
<emphasis id="D6F049109064A02FFEFAFAE3B1C3FAC6" box="[276,401,1303,1328]" italics="true" pageId="11" pageNumber="462">H. ocellata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="D6F049109064A02FFE4CFAE3B1B2FAD9" box="[418,480,1304,1327]" italics="true" pageId="11" pageNumber="462">sensu</emphasis>
|
||
<bibRefCitation id="8015E8F39064A02FFE06FAE3B2CEFAC6" author="Teo" box="[488,668,1304,1328]" pageId="11" pageNumber="462" refString="Teo, S. & Ng, C. S. L. (2009) New record of a sea cucumber, Holothuria ocellata (Jaeger, 1833) (Holothuroidea: Aspidochirotida: Holothuriidae) in Singapore. Nature in Singapore, 2, 411 - 414." type="journal article" year="2009">Teo & Ng 2009</bibRefCitation>
|
||
), not so in
|
||
<taxonomicName id="2384EE819064A02FFCF7FAE3B339FAC6" box="[793,875,1303,1328]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex">
|
||
<emphasis id="D6F049109064A02FFCF7FAE3B339FAC6" box="[793,875,1303,1328]" italics="true" pageId="11" pageNumber="462">H. alex</emphasis>
|
||
</taxonomicName>
|
||
), and especially the ossicle assemblage (tack-like tables present in
|
||
<taxonomicName id="2384EE819064A02FFEBAFAC7B182FAA2" box="[340,464,1339,1364]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="ocellata">
|
||
<emphasis id="D6F049109064A02FFEBAFAC7B182FAA2" box="[340,464,1339,1364]" italics="true" pageId="11" pageNumber="462">H. ocellata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="D6F049109064A02FFE0EFAC7B24CFAA5" box="[480,542,1340,1363]" italics="true" pageId="11" pageNumber="462">sensu</emphasis>
|
||
<bibRefCitation id="8015E8F39064A02FFDC8FAC7B28DFAA2" author="Teo" box="[550,735,1340,1364]" pageId="11" pageNumber="462" refString="Teo, S. & Ng, C. S. L. (2009) New record of a sea cucumber, Holothuria ocellata (Jaeger, 1833) (Holothuroidea: Aspidochirotida: Holothuriidae) in Singapore. Nature in Singapore, 2, 411 - 414." type="journal article" year="2009">Teo & Ng, 2009</bibRefCitation>
|
||
), absent in
|
||
<taxonomicName id="2384EE819064A02FFCB0FAC7B3FDFAA2" box="[862,943,1339,1364]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex">
|
||
<emphasis id="D6F049109064A02FFCB0FAC7B3FDFAA2" box="[862,943,1339,1364]" italics="true" pageId="11" pageNumber="462">H. alex</emphasis>
|
||
</taxonomicName>
|
||
) is significantly different.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC9EC6899064A02FFF29FA9BB3EBF9DA" pageId="11" pageNumber="462" type="reference_group">
|
||
<paragraph id="E43B95029064A02FFF29FA9BB3EBF9DA" blockId="11.[151,1437,583,1580]" pageId="11" pageNumber="462">
|
||
<taxonomicName id="2384EE819064A02FFF29FA9BB14AFA8E" box="[199,280,1375,1400]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex" status="sp. nov.">
|
||
<emphasis id="D6F049109064A02FFF29FA9BB14AFA8E" box="[199,280,1375,1400]" italics="true" pageId="11" pageNumber="462">H. alex</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D6F049109064A02FFEF0FAA4B12AFA8E" bold="true" box="[286,376,1375,1400]" pageId="11" pageNumber="462">
|
||
<taxonomicNameLabel id="CDC3F46B9064A02FFEF0FAA4B12AFA8E" box="[286,376,1375,1400]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also resembles
|
||
<taxonomicName id="2384EE819064A02FFDC7FA9BB304FA8E" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" box="[553,854,1375,1400]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="martensii">
|
||
<emphasis id="D6F049109064A02FFDC7FA9BB2E7FA81" box="[553,693,1376,1399]" italics="true" pageId="11" pageNumber="462">H. martensii</emphasis>
|
||
Semper,
|
||
<quantity id="237C38E79064A02FFCF0FAA4B304FA8E" box="[798,854,1375,1400]" metricMagnitude="1" metricUnit="m" metricValue="4.74472" pageId="11" pageNumber="462" unit="in" value="1868.0">1868</quantity>
|
||
</taxonomicName>
|
||
in presenting tables with reduced disc and with tall, smooth spires. We studied the ossicle assemblage of the holotype of
|
||
<taxonomicName id="2384EE819064A02FFC43FA7FB469FA6D" box="[941,1083,1412,1435]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="martensii">
|
||
<emphasis id="D6F049109064A02FFC43FA7FB469FA6D" box="[941,1083,1412,1435]" italics="true" pageId="11" pageNumber="462">H. martensii</emphasis>
|
||
</taxonomicName>
|
||
(fig.5) and of three specimens from Madagascar which
|
||
<bibRefCitation id="8015E8F39064A02FFE5FFA5CB2DDFA36" author="Cherbonnier" box="[433,655,1447,1472]" pageId="11" pageNumber="462" refString="Cherbonnier, G. (1988) Echinodermes: Holothurides. Faune de Madagascar, 70, 1 - 292." type="journal article" year="1988">Cherbonnier (1988)</bibRefCitation>
|
||
identified as
|
||
<emphasis id="D6F049109064A02FFCC7FA53B3ECFA49" box="[809,958,1448,1471]" italics="true" pageId="11" pageNumber="462">
|
||
<taxonomicName id="2384EE819064A02FFCC7FA53B3E9FA49" box="[809,955,1448,1471]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="martensii">H. martensii</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
We came to the conclusion that
|
||
<taxonomicName id="2384EE819064A02FFAC1FA53B5D3FA36" box="[1327,1409,1447,1472]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="alex">
|
||
<emphasis id="D6F049109064A02FFAC1FA53B5D3FA36" box="[1327,1409,1447,1472]" italics="true" pageId="11" pageNumber="462">H. alex</emphasis>
|
||
</taxonomicName>
|
||
is markedly different because it presents fenestrated spheres, tables with an upward turned disc and wide buttons. Of the three specimens
|
||
<bibRefCitation id="8015E8F39064A02FFE49FA14B2D5F9FE" author="Cherbonnier" box="[423,647,1519,1544]" pageId="11" pageNumber="462" refString="Cherbonnier, G. (1988) Echinodermes: Holothurides. Faune de Madagascar, 70, 1 - 292." type="journal article" year="1988">Cherbonnier (1988)</bibRefCitation>
|
||
identified as
|
||
<taxonomicName id="2384EE819064A02FFCCAFA0BB3E0F9F1" box="[804,946,1520,1543]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="martensii">
|
||
<emphasis id="D6F049109064A02FFCCAFA0BB3E0F9F1" box="[804,946,1520,1543]" italics="true" pageId="11" pageNumber="462">H. martensii</emphasis>
|
||
</taxonomicName>
|
||
, one turned out to be
|
||
<taxonomicName id="2384EE819064A02FFB5DFA14B116F9DA" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" class="Holothuroidea" family="Holothuriidae" genus="Bohadschia" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="vitiensis">
|
||
<emphasis id="D6F049109064A02FFB5DFA14B5CEF9F1" box="[1203,1436,1519,1544]" italics="true" pageId="11" pageNumber="462">Bohadschia vitiensis</emphasis>
|
||
(Semper, 1868)
|
||
</taxonomicName>
|
||
and two
|
||
<taxonomicName id="2384EE819064A02FFE45F9E8B3E7F9DA" authority="Massin, 1987" authorityName="Massin" authorityYear="1987" box="[427,949,1555,1580]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="horrida" subGenus="Metriatyla">
|
||
<emphasis id="D6F049109064A02FFE45F9E8B346F9DA" box="[427,788,1555,1580]" italics="true" pageId="11" pageNumber="462">Holothuria (Metriatyla) horrida</emphasis>
|
||
<bibRefCitation id="8015E8F39064A02FFCF5F9E8B3E7F9DA" author="Massin" box="[795,949,1555,1580]" pageId="11" pageNumber="462" refString="Massin, C. (1987) Holothuries nouvelles et peu connues recoltees en Indonesie au cours de la Snellius II Expedition. Bulletin de l'Institut Royal des Sciences naturelles de Belgique, Biologie, 57, 97 - 121." type="journal article" year="1987">Massin, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |