451 lines
54 KiB
XML
451 lines
54 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4763990" ID-GBIF-Dataset="8b5fc63e-4bcb-48d0-8015-bda6684e7590" ID-GBIF-Taxon="182075258" ID-ISSN="1854-0392" ID-Zenodo-Dep="4763990" ID-ZooBank="2DCA0E5D-15BE-4F6B-AF64-3AC7AA19B1F4" LSID="urn:lsid:zoobank.org:act:8CE031D8-2F45-4F1F-8750-61FA66B13E3B" checkinTime="1620964410999" checkinUser="felipe" docAuthor="Szczytko, Stanley W. & Stewart, Kenneth W." docDate="2013" docId="6C2B87F1FFEFFF9F44ACDE349067FEF4" docLanguage="en" docName="Illiesia.9.04.28-33.pdf" docOrigin="Illiesia 9 (4)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Isoperla umpqua Szczytko & Stewart 2013, sp. n." docType="treatment" docUuid="8CE031D8-2F45-4F1F-8750-61FA66B13E3B" docUuidSource="ZooBank" docVersion="6" lastPageId="5" lastPageNumber="31" masterDocId="9012FF89FFEFFF9A4740D97A925CFFB4" masterDocTitle="Isoperla Umpqua A New Species Of Western Nearctic Stonefly (Plecoptera: Isoperlinae)" masterLastPageNumber="33" masterPageNumber="28" pageId="0" pageNumber="28" updateTime="1643536171962" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Isoperla Umpqua A New Species Of Western Nearctic Stonefly (Plecoptera: Isoperlinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Szczytko, Stanley W.</mods:namePart>
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<mods:affiliation>College of Natural Resources, University of Wisconsin, Stevens Point, WI 54481, U. S. A. E-mail: sszczytk @ uwsp. edu</mods:affiliation>
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<mods:nameIdentifier type="email">sszczytk@uwsp.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stewart, Kenneth W.</mods:namePart>
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<mods:affiliation>Deceased Formerly, Dept. of Biological Sciences, University of North Texas, Denton, TX 76203, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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||
<mods:title>Illiesia</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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||
<mods:date>2013</mods:date>
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||
<mods:detail type="volume">
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<mods:number>9</mods:number>
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||
</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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||
</mods:detail>
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<mods:extent unit="page">
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<mods:start>28</mods:start>
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<mods:end>33</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760858</mods:identifier>
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<mods:identifier type="GBIF-Dataset">8b5fc63e-4bcb-48d0-8015-bda6684e7590</mods:identifier>
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<mods:identifier type="ISSN">1854-0392</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4760858</mods:identifier>
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<mods:identifier type="ZooBank">2DCA0E5D-15BE-4F6B-AF64-3AC7AA19B1F4</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4763990" ID-GBIF-Taxon="182075258" ID-Zenodo-Dep="4763990" LSID="urn:lsid:zoobank.org:act:8CE031D8-2F45-4F1F-8750-61FA66B13E3B" httpUri="http://treatment.plazi.org/id/6C2B87F1FFEFFF9F44ACDE349067FEF4" lastPageId="5" lastPageNumber="31" pageId="0" pageNumber="28">
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<subSubSection box="[1004,1275,1870,1894]" pageId="0" pageNumber="28" type="nomenclature">
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||
<paragraph blockId="0.[1004,1275,1870,1894]" box="[1004,1275,1870,1894]" pageId="0" pageNumber="28">
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||
<heading bold="true" box="[1004,1275,1870,1894]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="28" reason="2">
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||
<emphasis bold="true" box="[1004,1275,1870,1894]" pageId="0" pageNumber="28">
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<taxonomicName authority="Szczytko & Stewart, 2013" authorityName="Szczytko & Stewart" authorityYear="2013" box="[1004,1203,1870,1894]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="28" phylum="Arthropoda" rank="species" species="umpqua" status="sp. n.">
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<emphasis bold="true" box="[1004,1203,1870,1894]" italics="true" pageId="0" pageNumber="28">Isoperla umpqua</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1210,1275,1870,1894]" pageId="0" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[1074,1204,1906,1930]" pageId="0" pageNumber="28" type="description">
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||
<paragraph blockId="0.[1074,1204,1906,1930]" box="[1074,1204,1906,1930]" pageId="0" pageNumber="28">
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||
(
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<figureCitation box="[1083,1195,1906,1930]" captionStart-0="Figs" captionStart-1="Fig" captionStart-2="Figs" captionStartId-0="1.[189,241,1752,1776]" captionStartId-1="3.[189,230,1504,1528]" captionStartId-2="4.[189,241,1823,1847]" captionTargetBox-0="[280,1383,215,1698]" captionTargetBox-1="[229,744,225,1455]" captionTargetBox-2="[250,1383,251,1752]" captionTargetId-0="figure-124@1.[249,1383,215,1681]" captionTargetId-1="figure-511@3.[229,758,167,1455]" captionTargetId-2="figure-45@4.[249,1383,251,1752]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." captionText-1="Fig. 8. Isoperla umpqua larval habitus - line = 1.0 mm" captionText-2="Figs. 9-15. Isoperla umpqua. 9. Left lacinia. 10. Left mandible. 11. Right mandible. 12. Ova showing concave profile. 13. Ova. 14. Detail of ova chorion and micropyles. 15. Detail of right mandible." figureDoi-0="http://doi.org/10.5281/zenodo.4760860" figureDoi-1="http://doi.org/10.5281/zenodo.4760864" figureDoi-2="http://doi.org/10.5281/zenodo.4760868" httpUri-0="https://zenodo.org/record/4760860/files/figure.png" httpUri-1="https://zenodo.org/record/4760864/files/figure.png" httpUri-2="https://zenodo.org/record/4760868/files/figure.png" pageId="0" pageNumber="28">Figs. 1-15</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4760860" ID-Zenodo-Dep="4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="1" pageNumber="29" startId="1.[189,241,1752,1776]" targetBox="[280,1383,215,1698]" targetPageId="1">
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<paragraph blockId="1.[189,1443,1752,1918]" pageId="1" pageNumber="29">
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Figs. 1-7.
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<taxonomicName authorityName="Szczytko & Stewart" authorityYear="2013" box="[299,477,1752,1776]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="29" phylum="Arthropoda" rank="species" species="umpqua">
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<emphasis box="[299,477,1752,1776]" italics="true" pageId="1" pageNumber="29">Isoperla umpqua</emphasis>
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</taxonomicName>
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. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern.
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||
</paragraph>
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||
</caption>
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||
<subSubSection pageId="2" pageNumber="30" type="reference_group">
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||
<paragraph blockId="2.[227,797,225,279]" pageId="2" pageNumber="30">
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||
<taxonomicName authority="Szczytko and Stewart 2002" authorityName="Szczytko and Stewart" authorityYear="2002" box="[227,644,225,247]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="30" phylum="Arthropoda" rank="species" species="muir">
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<emphasis box="[227,356,225,246]" italics="true" pageId="2" pageNumber="30">Isoperla muir</emphasis>
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Szczytko and Stewart 2002
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</taxonomicName>
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, 130:233. Muir Creek, Douglas Co.,
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<collectingRegion box="[445,528,257,278]" country="United States of America" name="Oregon" pageId="2" pageNumber="30">Oregon</collectingRegion>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="2" pageNumber="30" type="materials_examined">
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||
<paragraph blockId="2.[189,798,325,1914]" pageId="2" pageNumber="30">
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||
<emphasis bold="true" box="[189,439,326,350]" pageId="2" pageNumber="30">Material examined.</emphasis>
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||
<materialsCitation ID-GBIF-Occurrence="3111753310" collectingDate="2002-05-12" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Muir Crk." pageId="2" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Oregon" typeStatus="holotype">
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<emphasis bold="true" box="[459,573,326,350]" pageId="2" pageNumber="30">
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||
<typeStatus box="[459,573,326,350]" pageId="2" pageNumber="30">Holotype</typeStatus>
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||
</emphasis>
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||
<specimenCount box="[592,612,325,350]" pageId="2" pageNumber="30" type="male">♂</specimenCount>
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||
,
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||
<emphasis bold="true" box="[639,791,326,350]" pageId="2" pageNumber="30">
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||
<collectingCountry box="[639,697,326,350]" name="United States of America" pageId="2" pageNumber="30">USA</collectingCountry>
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||
<emphasis bold="true" box="[717,730,329,350]" italics="true" pageId="2" pageNumber="30">–</emphasis>
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||
<collectingRegion box="[750,791,326,350]" pageId="2" pageNumber="30">OR</collectingRegion>
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||
</emphasis>
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||
:
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||
<collectingCounty box="[189,335,362,386]" pageId="2" pageNumber="30">Douglas Co.</collectingCounty>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:6C2B87F1FFEFFF9F44ACDE349067FEF4:E15D603CFFEDFF984619D8109393FE36" box="[345,463,362,386]" country="United States of America" county="Douglas Co." name="Muir Crk." pageId="2" pageNumber="30" stateProvince="OR">Muir Crk.</location>
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, Hwy 230,
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<locationDeviation pageId="2" pageNumber="30">downstream from bridge</locationDeviation>
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||
,
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||
<date box="[280,399,397,421]" pageId="2" pageNumber="30" value="2002-05-12">
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||
<collectingDate box="[280,399,397,421]" pageId="2" pageNumber="30" value="2002-05-12">12/V/2002</collectingDate>
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||
</date>
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||
,
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||
<collectorName box="[410,576,397,421]" pageId="2" pageNumber="30">S.W. Szczytko</collectorName>
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and
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||
<collectorName box="[634,792,397,421]" pageId="2" pageNumber="30">K.W. Stewart</collectorName>
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||
, (emerged in laboratory) (
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||
<collectionCode box="[493,575,433,457]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="30" type="Museum">USNM</collectionCode>
|
||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation ID-GBIF-Occurrence="3111753309" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
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||
<emphasis bold="true" box="[600,719,433,457]" pageId="2" pageNumber="30">
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||
<typeStatus box="[600,719,433,457]" pageId="2" pageNumber="30">Paratypes</typeStatus>
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||
</emphasis>
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||
(same locality),
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||
<date box="[304,425,468,492]" pageId="2" pageNumber="30" value="2002-05-25">
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||
25/
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||
<collectionCode box="[339,359,468,492]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="30" type="Museum">V</collectionCode>
|
||
/2002
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||
</date>
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||
),
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||
<collectorName box="[449,568,469,493]" pageId="2" pageNumber="30">B.P. Stark</collectorName>
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||
& D.D.
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||
<location LSID="urn:lsid:plazi:treatment:6C2B87F1FFEFFF9F44ACDE349067FEF4:E15D603CFFEDFF9845E8D8AF90A3FE59" box="[680,767,469,493]" country="United States of America" name="Ziegler" pageId="2" pageNumber="30">Ziegler</location>
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||
,
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||
<specimenCount pageId="2" pageNumber="30" type="larva">8 larvae</specimenCount>
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||
(
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||
<collectionCode box="[281,358,504,528]" pageId="2" pageNumber="30">BYUC</collectionCode>
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||
);
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||
</materialsCitation>
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||
<materialsCitation ID-GBIF-Occurrence="3111753305" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
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||
upstream from bridge,
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||
<date box="[673,793,504,528]" pageId="2" pageNumber="30" value="2001-05-19">
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||
<collectingDate box="[673,793,504,528]" pageId="2" pageNumber="30" value="2001-05-19">19/V/2001</collectingDate>
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||
</date>
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||
, B.P.
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||
<collectorName box="[241,302,540,564]" pageId="2" pageNumber="30">Stark</collectorName>
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and
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||
<collectorName box="[359,517,540,564]" pageId="2" pageNumber="30">K.W. Stewart</collectorName>
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||
,
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||
<specimenCount box="[527,561,538,564]" pageId="2" pageNumber="30" type="male">3♂</specimenCount>
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||
,
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||
<specimenCount box="[574,678,540,564]" pageId="2" pageNumber="30" type="larva">21 larvae</specimenCount>
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||
(emerged in laboratory
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||
<date box="[359,480,575,599]" pageId="2" pageNumber="30" value="2001-05-25">
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||
<collectingDate box="[359,480,575,599]" pageId="2" pageNumber="30" value="2001-05-25">
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||
25/
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||
<collectionCode box="[395,415,575,599]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="30" type="Museum">V</collectionCode>
|
||
/2001
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||
</collectingDate>
|
||
</date>
|
||
), (
|
||
<collectionCode box="[511,590,575,599]" pageId="2" pageNumber="30">BYUC</collectionCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3111753302" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
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||
downstream of bridge,
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||
<date box="[280,399,611,635]" pageId="2" pageNumber="30" value="2002-05-12">
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||
<collectingDate box="[280,399,611,635]" pageId="2" pageNumber="30" value="2002-05-12">12/V/2002</collectingDate>
|
||
</date>
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||
, S.W.
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||
<collectorName box="[471,576,611,635]" pageId="2" pageNumber="30">Szczytko</collectorName>
|
||
and
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||
<collectorName box="[634,792,611,635]" pageId="2" pageNumber="30">K.W. Stewart</collectorName>
|
||
,
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||
<specimenCount box="[189,223,645,670]" pageId="2" pageNumber="30" type="male">2♂</specimenCount>
|
||
(emerged in laboratory
|
||
<date box="[520,639,646,670]" pageId="2" pageNumber="30" value="2002-05-25">
|
||
<collectingDate box="[520,639,646,670]" pageId="2" pageNumber="30" value="2002-05-25">
|
||
25/
|
||
<collectionCode box="[555,575,646,670]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="30" type="Museum">V</collectionCode>
|
||
/2002
|
||
</collectingDate>
|
||
</date>
|
||
) (
|
||
<collectionCode box="[663,740,646,670]" pageId="2" pageNumber="30">BYUC</collectionCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3111753311" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount box="[764,797,645,670]" pageId="2" pageNumber="30" type="male">1♂</specimenCount>
|
||
(emerged in laboratory),
|
||
<date box="[481,600,682,706]" pageId="2" pageNumber="30" value="2002-05-27">
|
||
27/
|
||
<collectionCode box="[516,536,682,706]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="30" type="Museum">V</collectionCode>
|
||
/2002
|
||
</date>
|
||
) (
|
||
<collectionCode box="[621,698,682,706]" pageId="2" pageNumber="30">BYUC</collectionCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3111753306" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount box="[718,751,681,706]" pageId="2" pageNumber="30" type="male">2♂</specimenCount>
|
||
,
|
||
<specimenCount box="[764,797,681,706]" pageId="2" pageNumber="30" type="female">1♀</specimenCount>
|
||
(emerged in laboratory
|
||
<date box="[503,622,717,741]" pageId="2" pageNumber="30" value="2002-05-30">
|
||
<collectingDate box="[503,622,717,741]" pageId="2" pageNumber="30" value="2002-05-30">
|
||
30/
|
||
<collectionCode box="[538,557,717,741]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="30" type="Museum">V</collectionCode>
|
||
/2002
|
||
</collectingDate>
|
||
</date>
|
||
) (
|
||
<collectionCode box="[655,732,717,741]" pageId="2" pageNumber="30">BYUC</collectionCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3111753312" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount box="[764,797,716,741]" pageId="2" pageNumber="30" type="male">1♂</specimenCount>
|
||
(emerged in laboratory
|
||
<date box="[465,595,753,777]" pageId="2" pageNumber="30" value="2002-06-01">
|
||
<collectingDate box="[465,595,753,777]" pageId="2" pageNumber="30" value="2002-06-01">
|
||
01/
|
||
<collectionCode box="[500,530,753,777]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="30">VI</collectionCode>
|
||
/2002
|
||
</collectingDate>
|
||
</date>
|
||
), (
|
||
<collectionCode box="[621,700,753,777]" pageId="2" pageNumber="30">BYUC</collectionCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3111753308" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount box="[718,751,752,777]" pageId="2" pageNumber="30" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount box="[764,797,752,777]" pageId="2" pageNumber="30" type="female">2♀</specimenCount>
|
||
(emerged in laboratory
|
||
<date box="[506,620,788,812]" pageId="2" pageNumber="30" value="2002-06-02">
|
||
<collectingDate box="[506,620,788,812]" pageId="2" pageNumber="30" value="2002-06-02">
|
||
2/
|
||
<collectionCode box="[527,556,788,812]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="30">VI</collectionCode>
|
||
/2002
|
||
</collectingDate>
|
||
</date>
|
||
) (
|
||
<collectionCode box="[654,731,788,812]" pageId="2" pageNumber="30">BYUC</collectionCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3111753307" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount box="[764,797,787,812]" pageId="2" pageNumber="30" type="female">2♀</specimenCount>
|
||
(emerged in laboratory
|
||
<date box="[501,617,824,848]" pageId="2" pageNumber="30" value="2002-06-01">
|
||
<collectingDate box="[501,617,824,848]" pageId="2" pageNumber="30" value="2002-06-01">
|
||
1/
|
||
<collectionCode box="[519,549,824,848]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="30">VI</collectionCode>
|
||
/2002
|
||
</collectingDate>
|
||
</date>
|
||
), (
|
||
<collectionCode box="[655,734,824,848]" pageId="2" pageNumber="30">BYUC</collectionCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3111753301" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount box="[764,797,823,848]" pageId="2" pageNumber="30" type="male">1♂</specimenCount>
|
||
(emerged in laboratory
|
||
<date box="[501,617,859,883]" pageId="2" pageNumber="30" value="2002-06-05">
|
||
<collectingDate box="[501,617,859,883]" pageId="2" pageNumber="30" value="2002-06-05">
|
||
5/
|
||
<collectionCode box="[522,552,859,883]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="30">VI</collectionCode>
|
||
/2002
|
||
</collectingDate>
|
||
</date>
|
||
), (
|
||
<collectionCode box="[655,734,860,884]" pageId="2" pageNumber="30">BYUC</collectionCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3111753303" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount box="[764,797,858,884]" pageId="2" pageNumber="30" type="female">1♀</specimenCount>
|
||
(emerged in laboratory
|
||
<date box="[512,642,895,919]" pageId="2" pageNumber="30" value="2002-06-11">
|
||
<collectingDate box="[512,642,895,919]" pageId="2" pageNumber="30" value="2002-06-11">
|
||
11/
|
||
<collectionCode box="[545,575,895,919]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="30">VI</collectionCode>
|
||
/2002
|
||
</collectingDate>
|
||
</date>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3111753304" collectingDate="2002-05-12" collectingDateMax="2001-05-25" collectingDateMin="2001-05-19" collectionCode="USNM" collectorName="S. W. Szczytko & K. W. Stewart" country="United States of America" county="Douglas Co." location="Hwy" pageId="2" pageNumber="30" specimenCount="1" specimenCount-female="1" specimenCount-larva="8" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount box="[676,796,895,919]" pageId="2" pageNumber="30" type="larva">19 larvae</specimenCount>
|
||
(
|
||
<collectionCode box="[197,274,931,955]" pageId="2" pageNumber="30">BYUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="2.[189,798,325,1914]" lastBlockId="2.[835,1443,224,1919]" pageId="2" pageNumber="30">
|
||
<emphasis bold="true" box="[189,258,966,990]" pageId="2" pageNumber="30">Male.</emphasis>
|
||
Macropterous. Body length 9.0 – 11.0 mm; forewing length 9.0 – 10.0 mm. General body color pale brown with dark brown markings. Dorsum of head with medium brown “M” bands anterior to median ocellus extending anterior and lateral to near base of antennae; medium brown bands connecting ocelli and extending laterally to near base of antennae with dark brown terminal spots; pale brown median area from anterior ocellus to posterior margin of head (
|
||
<figureCitation box="[381,445,1286,1310]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 1</figureCitation>
|
||
). Antennae pale brown; scape dark brown. Pronotum with wide median pale stripe, disks cream, rugosities, irregular shaped, raised, dark brown, anterior, posterior margins pale, lateral margins with a wide pale band (
|
||
<figureCitation box="[580,646,1428,1452]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 1</figureCitation>
|
||
). Meso- and metanota dark brown, with irregular pale spots. Wings pale, veins medium brown. Legs pale brown, dorsal surface of femora with dark, ovoid, proximal spot, outer surface with a thin incomplete median, longitudinal, brown band. Abdominal terga pale brown with 3 lateral, and 2 mesal longitudinal rows of small dark spots and faint median longitudinal band. Ninth and 10
|
||
<superScript attach="left" box="[415,430,1712,1726]" fontSize="6" pageId="2" pageNumber="30">th</superScript>
|
||
terga with mesal patch of short stout spinulae. Tenth tergum weakly sclerotized along midline; paraprocts sharply pointed, extending over ⅓ length of 10
|
||
<superScript attach="left" box="[435,450,1818,1832]" fontSize="6" pageId="2" pageNumber="30">th</superScript>
|
||
tergum (
|
||
<figureCitation box="[567,638,1819,1843]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 3</figureCitation>
|
||
). Abdominal sterna pale brown with paired dark anteormedian spots on segments 1-10 and smaller paired posteromedian dark spots. Eight and 9
|
||
<superScript attach="left" box="[1290,1305,224,238]" fontSize="6" pageId="2" pageNumber="30">th</superScript>
|
||
sterna with mesal patch of long setae; 8th sternum with a well developed vesicle, 1.25 times as long as wide, evenly rounded and expanded slightly apically; posterior margin fringed with medium length setae (
|
||
<figureCitation box="[1359,1429,367,391]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 2</figureCitation>
|
||
). Aedeagus mostly membranous with large paired anterodorsal (
|
||
<figureCitation box="[1001,1078,438,462]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 5f</figureCitation>
|
||
) and small paired dorsal (
|
||
<figureCitation captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 5e</figureCitation>
|
||
) lobes; small posterodorsal patch of dense medium stout spinulae (
|
||
<figureCitation box="[1121,1203,509,533]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 5d</figureCitation>
|
||
); wide median band of dense medium stout spinulae (
|
||
<figureCitation box="[1232,1314,544,568]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 5b</figureCitation>
|
||
); proximal area expanded and membranous (
|
||
<figureCitation box="[1242,1323,580,604]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 5a</figureCitation>
|
||
), exposed posterodorsal sclerotized rod, expanded slightly apically, base embedded and bifurcated with wide slender incurving arms (
|
||
<figureCitation box="[1123,1240,686,710]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Figs. 5c, 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="2.[835,1443,224,1919]" pageId="2" pageNumber="30">
|
||
<emphasis bold="true" box="[835,929,722,746]" pageId="2" pageNumber="30">Female.</emphasis>
|
||
Macropterous. Body length 11.0 mm; forewing length 12.0 mm. Body coloration and morphology similar to male. Eighth sternum with posteromedian dark brown patch and broad medium brown inverted triangular patch. Subgenital plate produced over ⅓ length of 9
|
||
<superScript attach="left" box="[1213,1228,899,913]" fontSize="6" pageId="2" pageNumber="30">th</superScript>
|
||
tergum, broadly rounded with slight posteromedial emargination (
|
||
<figureCitation box="[843,911,971,995]" captionStart="Figs" captionStartId="1.[189,241,1752,1776]" captionTargetBox="[280,1383,215,1698]" captionTargetId="figure-124@1.[249,1383,215,1681]" captionTargetPageId="1" captionText="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." figureDoi="http://doi.org/10.5281/zenodo.4760860" httpUri="https://zenodo.org/record/4760860/files/figure.png" pageId="2" pageNumber="30">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="3" lastPageNumber="31" pageId="2" pageNumber="30" type="description">
|
||
<paragraph blockId="2.[835,1443,224,1919]" lastBlockId="3.[835,1444,225,1920]" lastPageId="3" lastPageNumber="31" pageId="2" pageNumber="30">
|
||
<emphasis bold="true" box="[835,910,1007,1031]" pageId="2" pageNumber="30">Larva.</emphasis>
|
||
Body length of mature larvae 11.0 – 14.0 mm. General body color medium brown. Dorsum of head covered with short, blunt, stout spinulae, spinulae absent in pale patches; mostly brown with various pale areas (
|
||
<figureCitation box="[968,1068,1149,1173]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="1.[189,241,1752,1776]" captionStartId-1="3.[189,230,1504,1528]" captionTargetBox-0="[280,1383,215,1698]" captionTargetBox-1="[229,744,225,1455]" captionTargetId-0="figure-124@1.[249,1383,215,1681]" captionTargetId-1="figure-511@3.[229,758,167,1455]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." captionText-1="Fig. 8. Isoperla umpqua larval habitus - line = 1.0 mm" figureDoi-0="http://doi.org/10.5281/zenodo.4760860" figureDoi-1="http://doi.org/10.5281/zenodo.4760864" httpUri-0="https://zenodo.org/record/4760860/files/figure.png" httpUri-1="https://zenodo.org/record/4760864/files/figure.png" pageId="2" pageNumber="30">Figs. 7-8</figureCitation>
|
||
); thin, pale “M” shaped pattern anterior to median ocellus; thin, pale, slanting bars near antennal bases; interocellar area medium brown; ecdysal stem and “Y” arms thin pale lines; posterolateral areas with large pale spots infused with thin dark lines; occiput with sinuous row of stout, medium length setae interrupted medially; labrum mostly medium brown, lighter medially; antennae medium brown (
|
||
<figureCitation box="[1201,1319,1433,1457]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="1.[189,241,1752,1776]" captionStartId-1="3.[189,230,1504,1528]" captionTargetBox-0="[280,1383,215,1698]" captionTargetBox-1="[229,744,225,1455]" captionTargetId-0="figure-124@1.[249,1383,215,1681]" captionTargetId-1="figure-511@3.[229,758,167,1455]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." captionText-1="Fig. 8. Isoperla umpqua larval habitus - line = 1.0 mm" figureDoi-0="http://doi.org/10.5281/zenodo.4760860" figureDoi-1="http://doi.org/10.5281/zenodo.4760864" httpUri-0="https://zenodo.org/record/4760860/files/figure.png" httpUri-1="https://zenodo.org/record/4760864/files/figure.png" pageId="2" pageNumber="30">Figs. 7-8</figureCitation>
|
||
). Lacinia triangulate, bidentate; subapical tooth ca. ½ length of apical tooth; 1 axillary seta; 8-10 long, stout marginal setae below subapical tooth; sparse, irregularly spaced row of long, fine submarginal setae below row of stout setae proximal to ¾ length of lacinia (
|
||
<figureCitation box="[843,909,1646,1671]" captionStart="Figs" captionStartId="4.[189,241,1823,1847]" captionTargetBox="[250,1383,251,1752]" captionTargetId="figure-45@4.[249,1383,251,1752]" captionTargetPageId="4" captionText="Figs. 9-15. Isoperla umpqua. 9. Left lacinia. 10. Left mandible. 11. Right mandible. 12. Ova showing concave profile. 13. Ova. 14. Detail of ova chorion and micropyles. 15. Detail of right mandible." figureDoi="http://doi.org/10.5281/zenodo.4760868" httpUri="https://zenodo.org/record/4760868/files/figure.png" pageId="2" pageNumber="30">Fig. 9</figureCitation>
|
||
). Mandibles with 6 cusps; large median row of long medium stout setae below outer apical cusp to near mandibular base; inner marginal fringe of 25 - 30 long, stout setae; right mandible with row of short stout acanthae below inner 3rd apical cusp (
|
||
<figureCitation captionStart="Figs" captionStartId="4.[189,241,1823,1847]" captionTargetBox="[250,1383,251,1752]" captionTargetId="figure-45@4.[249,1383,251,1752]" captionTargetPageId="4" captionText="Figs. 9-15. Isoperla umpqua. 9. Left lacinia. 10. Left mandible. 11. Right mandible. 12. Ova showing concave profile. 13. Ova. 14. Detail of ova chorion and micropyles. 15. Detail of right mandible." figureDoi="http://doi.org/10.5281/zenodo.4760868" httpUri="https://zenodo.org/record/4760868/files/figure.png" pageId="2" pageNumber="30">Figs. 10- 15</figureCitation>
|
||
), right mandible with dense brush of medium length, stout setae below inner 3
|
||
<superScript attach="left" box="[1224,1240,1859,1873]" fontSize="6" pageId="2" pageNumber="30">rd</superScript>
|
||
apical cusp (
|
||
<figureCitation captionStart="Figs" captionStartId="4.[189,241,1823,1847]" captionTargetBox="[250,1383,251,1752]" captionTargetId="figure-45@4.[249,1383,251,1752]" captionTargetPageId="4" captionText="Figs. 9-15. Isoperla umpqua. 9. Left lacinia. 10. Left mandible. 11. Right mandible. 12. Ova showing concave profile. 13. Ova. 14. Detail of ova chorion and micropyles. 15. Detail of right mandible." figureDoi="http://doi.org/10.5281/zenodo.4760868" httpUri="https://zenodo.org/record/4760868/files/figure.png" pageId="2" pageNumber="30">Fig. 11</figureCitation>
|
||
). Pronotum mostly medium brown with pale, thin rugosities; discs with sparse short, black clothing hairs; margins fringed with dense, short stout setae (
|
||
<figureCitation box="[197,304,1681,1705]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="1.[189,241,1752,1776]" captionStartId-1="3.[189,230,1504,1528]" captionTargetBox-0="[280,1383,215,1698]" captionTargetBox-1="[229,744,225,1455]" captionTargetId-0="figure-124@1.[249,1383,215,1681]" captionTargetId-1="figure-511@3.[229,758,167,1455]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="Figs. 1-7. Isoperla umpqua. 1. Adult head pattern. 2. Terminal male sterna. 3. Terminal male terga. 4. Terminal female sterna. 5. Male aedeagus; a. expanded proximal area, b. wide median band of stout spinulae, c. posterodorsal sclerotized rod, d. posterodorsal dense band of medium stout spinulae, e. paired small dorsal lobes, f. paired anterodorsal lobes. 6. Detail of posterodorsal sclerotized aedeagal rod. 7. Larval head & pronotum with spinule pattern." captionText-1="Fig. 8. Isoperla umpqua larval habitus - line = 1.0 mm" figureDoi-0="http://doi.org/10.5281/zenodo.4760860" figureDoi-1="http://doi.org/10.5281/zenodo.4760864" httpUri-0="https://zenodo.org/record/4760860/files/figure.png" httpUri-1="https://zenodo.org/record/4760864/files/figure.png" pageId="3" pageNumber="31">Figs. 7-8</figureCitation>
|
||
). Anterior and posterior intersegmental membrane of pronotum with dense band of medium length stout hairs and chloride cells. Meso- metanota with sparse median patches of short stout spinulae and black clothing hairs. Mesonotum with dark brown lateral bands and 2 median dark brown bands; metanotum with wide, dark brown lateral bands and 2 wide, median dark brown bands (
|
||
<figureCitation box="[1289,1361,225,249]" captionStart="Fig" captionStartId="3.[189,230,1504,1528]" captionTargetBox="[229,744,225,1455]" captionTargetId="figure-511@3.[229,758,167,1455]" captionTargetPageId="3" captionText="Fig. 8. Isoperla umpqua larval habitus - line = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.4760864" httpUri="https://zenodo.org/record/4760864/files/figure.png" pageId="3" pageNumber="31">Fig. 8</figureCitation>
|
||
). Legs medium brown, outer surface of femora with scattered medium length, stout setae and black clothing hairs; median pale, thin longitudinal band with no setae or clothing hairs; proximal ovoid, pale spot on dorsal surface void of setae or clothing hairs; outer tibia surface with sparse medium length stout setae and clothing hairs. Abdominal terga with wide median pale brown patches that decrease in size posteriorly, forming a broad median longitudinal band, bordered laterally on each side by a dark longitudinal band; posterior margins of terga fringed with very short stout setae and occasional longer setae, lateral margins with more longer setae; dorsal surface with scattered heavy setae and clothing hairs; intersegmental membranes with concentrated chloride epithelia (
|
||
<figureCitation box="[1054,1119,793,817]" captionStart="Fig" captionStartId="3.[189,230,1504,1528]" captionTargetBox="[229,744,225,1455]" captionTargetId="figure-511@3.[229,758,167,1455]" captionTargetPageId="3" captionText="Fig. 8. Isoperla umpqua larval habitus - line = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.4760864" httpUri="https://zenodo.org/record/4760864/files/figure.png" pageId="3" pageNumber="31">Fig. 8</figureCitation>
|
||
). Cerci medium brown with posterior whorl of short setae.
|
||
</paragraph>
|
||
<paragraph blockId="3.[835,1444,225,1920]" pageId="3" pageNumber="31">
|
||
<emphasis bold="true" box="[835,918,864,888]" pageId="3" pageNumber="31">Ovum.</emphasis>
|
||
General shape oblong, cross section concave, collar absent (
|
||
<figureCitation box="[997,1123,900,924]" captionStart="Figs" captionStartId="4.[189,241,1823,1847]" captionTargetBox="[250,1383,251,1752]" captionTargetId="figure-45@4.[249,1383,251,1752]" captionTargetPageId="4" captionText="Figs. 9-15. Isoperla umpqua. 9. Left lacinia. 10. Left mandible. 11. Right mandible. 12. Ova showing concave profile. 13. Ova. 14. Detail of ova chorion and micropyles. 15. Detail of right mandible." figureDoi="http://doi.org/10.5281/zenodo.4760868" httpUri="https://zenodo.org/record/4760868/files/figure.png" pageId="3" pageNumber="31">Figs. 12-13</figureCitation>
|
||
). Color pale brown. Length
|
||
<quantity box="[835,988,935,960]" metricMagnitude="2" metricUnit="m" metricValue="2.875" metricValueMax="2.95" metricValueMin="2.8" pageId="3" pageNumber="31" unit="m" value="287.5" valueMax="295.0" valueMin="280.0">280 - 295 m</quantity>
|
||
; width
|
||
<quantity box="[1078,1231,936,960]" metricMagnitude="2" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="3" pageNumber="31" unit="m" value="215.0" valueMax="220.0" valueMin="210.0">
|
||
210 -
|
||
<specimenCount box="[1142,1231,936,960]" pageId="3" pageNumber="31" type="generic">220 m</specimenCount>
|
||
</quantity>
|
||
. Choronic surface covered with numerous shallow, round follicle cell impressions (FCI’s), FCI’s near posterior pole shallower; walls of hexagonal follicle cell impressions faintly visible (
|
||
<figureCitation box="[1168,1298,1078,1102]" captionStart="Figs" captionStartId="4.[189,241,1823,1847]" captionTargetBox="[250,1383,251,1752]" captionTargetId="figure-45@4.[249,1383,251,1752]" captionTargetPageId="4" captionText="Figs. 9-15. Isoperla umpqua. 9. Left lacinia. 10. Left mandible. 11. Right mandible. 12. Ova showing concave profile. 13. Ova. 14. Detail of ova chorion and micropyles. 15. Detail of right mandible." figureDoi="http://doi.org/10.5281/zenodo.4760868" httpUri="https://zenodo.org/record/4760868/files/figure.png" pageId="3" pageNumber="31">Figs. 12-13</figureCitation>
|
||
). Micropyle row subequatorial; orifices small and not elevated (
|
||
<figureCitation box="[843,925,1149,1173]" captionStart="Figs" captionStartId="4.[189,241,1823,1847]" captionTargetBox="[250,1383,251,1752]" captionTargetId="figure-45@4.[249,1383,251,1752]" captionTargetPageId="4" captionText="Figs. 9-15. Isoperla umpqua. 9. Left lacinia. 10. Left mandible. 11. Right mandible. 12. Ova showing concave profile. 13. Ova. 14. Detail of ova chorion and micropyles. 15. Detail of right mandible." figureDoi="http://doi.org/10.5281/zenodo.4760868" httpUri="https://zenodo.org/record/4760868/files/figure.png" pageId="3" pageNumber="31">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="31" type="distribution">
|
||
<paragraph blockId="3.[835,1444,225,1920]" pageId="3" pageNumber="31">
|
||
<emphasis bold="true" box="[835,994,1184,1208]" pageId="3" pageNumber="31">Distribution.</emphasis>
|
||
<taxonomicName authorityName="Szczytko & Stewart" authorityYear="2013" box="[1005,1188,1184,1208]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[1005,1188,1184,1208]" italics="true" pageId="3" pageNumber="31">Isoperla umpqua</emphasis>
|
||
</taxonomicName>
|
||
is known only from Muir Creek, Douglas County,
|
||
<collectingRegion box="[1181,1271,1220,1244]" country="United States of America" name="Oregon" pageId="3" pageNumber="31">Oregon</collectingRegion>
|
||
. It is surprising that this species has not been collected from other streams in the Cascade Mountain Range since the color pattern of the larvae is distinctive from other western Nearctic
|
||
<taxonomicName box="[1041,1223,1362,1386]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[1041,1125,1362,1386]" italics="true" pageId="3" pageNumber="31">Isoperla</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="31" type="etymology">
|
||
<paragraph blockId="3.[835,1444,225,1920]" pageId="3" pageNumber="31">
|
||
<emphasis bold="true" box="[835,973,1398,1422]" pageId="3" pageNumber="31">Etymology.</emphasis>
|
||
We honor the people of the Umpqua Native American Tribe which is federally recognized as a Cow Creek Band of the Umpqua Tribe. These native Americans live in south-central
|
||
<collectingRegion box="[1295,1384,1505,1529]" country="United States of America" name="Oregon" pageId="3" pageNumber="31">Oregon</collectingRegion>
|
||
near the
|
||
<typeStatus box="[879,931,1541,1565]" pageId="3" pageNumber="31">type</typeStatus>
|
||
locality of this stonefly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4760864" ID-Zenodo-Dep="4760864" box="[189,794,1504,1528]" httpUri="https://zenodo.org/record/4760864/files/figure.png" pageId="3" pageNumber="31" startId="3.[189,230,1504,1528]" targetBox="[229,744,225,1455]" targetPageId="3">
|
||
<paragraph blockId="3.[189,794,1504,1528]" box="[189,794,1504,1528]" pageId="3" pageNumber="31">
|
||
Fig. 8.
|
||
<taxonomicName authorityName="Szczytko & Stewart" authorityYear="2013" box="[267,446,1504,1528]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[267,446,1504,1528]" italics="true" pageId="3" pageNumber="31">Isoperla umpqua</emphasis>
|
||
</taxonomicName>
|
||
larval habitus - line = 1.0 mm
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="5" lastPageNumber="33" pageId="3" pageNumber="31" type="diagnosis">
|
||
<paragraph blockId="3.[835,1444,225,1920]" lastBlockId="5.[189,798,225,320]" lastPageId="5" lastPageNumber="33" pageId="3" pageNumber="31">
|
||
<emphasis bold="true" box="[835,964,1575,1599]" pageId="3" pageNumber="31">Diagnosis.</emphasis>
|
||
In the male, the clavate sclerotized posterior process of the aedeagus is most similar to
|
||
<taxonomicName authority="Jewett" authorityName="Jewett" authorityYear="1962" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="adunca">
|
||
<emphasis italics="true" pageId="3" pageNumber="31">I. adunca</emphasis>
|
||
Jewett
|
||
</taxonomicName>
|
||
, however it is larger and the shape and spinule patterns of the aedeagus easily distinguishes
|
||
<taxonomicName authorityName="Szczytko & Stewart" authorityYear="2013" box="[835,944,1719,1742]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[835,844,1719,1742]" italics="true" pageId="3" pageNumber="31">I</emphasis>
|
||
.
|
||
<emphasis box="[856,944,1719,1742]" italics="true" pageId="3" pageNumber="31">umpqua</emphasis>
|
||
</taxonomicName>
|
||
. The female subgenital plate with a shallow emargination and rounded lateral margins is similar to
|
||
<taxonomicName authority="Frison" authorityName="Frison" authorityYear="1937" box="[864,1023,1789,1813]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="pinta">
|
||
<emphasis box="[864,940,1790,1813]" italics="true" pageId="3" pageNumber="31">I. pinta</emphasis>
|
||
Frison
|
||
</taxonomicName>
|
||
, but lacks the pronotal checkerboard pattern of
|
||
<taxonomicName authorityName="Frison" authorityYear="1937" box="[978,1067,1825,1848]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="pinta">
|
||
<emphasis box="[978,1067,1825,1848]" italics="true" pageId="3" pageNumber="31">I. pinta</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Sandberg, J. B." box="[1092,1292,1825,1849]" pageId="3" pageNumber="31" pagination="202 - 258" refId="ref2400" refString="Sandberg, J. B. 2011. The Isoperla of California (Plecoptera: Perlodidae); larval descriptions and a key to 17 western Nearctic species. Illiesia, 7 (22): 202 - 258." type="journal article" year="2011">Sandberg (2011)</bibRefCitation>
|
||
provides a detailed description of the nymph and especially in mouthpart characters is also most similar to
|
||
<taxonomicName authorityName="Jewett" authorityYear="1962" box="[1341,1437,1896,1920]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="adunca">
|
||
<emphasis box="[1341,1437,1896,1920]" italics="true" pageId="3" pageNumber="31">I. adunca</emphasis>
|
||
</taxonomicName>
|
||
, but can be separated by the characters of the maxilla. Additionally,
|
||
<taxonomicName authorityName="Szczytko & Stewart" authorityYear="2013" box="[354,464,261,284]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[354,363,261,284]" italics="true" pageId="5" pageNumber="33">I</emphasis>
|
||
.
|
||
<emphasis box="[376,464,261,284]" italics="true" pageId="5" pageNumber="33">umpqua</emphasis>
|
||
</taxonomicName>
|
||
appears to restricted to
|
||
<typeStatus box="[745,797,261,285]" pageId="5" pageNumber="33">type</typeStatus>
|
||
locality in south-central
|
||
<collectingRegion box="[475,567,296,320]" country="United States of America" name="Oregon" pageId="5" pageNumber="33">Oregon</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |