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<document id="5D12841AA3222433B996DEEB2C2C228D" ID-CLB-Dataset="27774" ID-DOI="10.11646/zootaxa.4531.3.7" ID-GBIF-Dataset="5aa5a37c-ff28-4de4-be53-9649408c8418" ID-ISSN="1175-5326" ID-Zenodo-Dep="2614772" ID-ZooBank="E61435D6-46FA-4B9E-A8ED-977B586C6406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553787154419" checkinUser="plazi" docAuthor="Sigwart, Julia D. &amp; Chen, Chong" docDate="2018" docId="6C070220D073FFCEADA4FBE78ACDFF2C" docLanguage="en" docName="zootaxa.4531.3.7.pdf" docOrigin="Zootaxa 4531 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hanleyella henrici Sigwart &amp; Chen 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="433" masterDocId="903E7A58D072FFCDAD33FFE88A04FFD4" masterDocTitle="A new deep water chiton (Mollusca: Polyplacophora) from hydrothermal vent ecosystems in the Okinawa Trough, Japan" masterLastPageNumber="436" masterPageNumber="430" pageNumber="431" updateTime="1698714902394" updateUser="ExternalLinkService">
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<mods:title id="6226E25A51FB14F2209F62C3B90E3EFB">A new deep water chiton (Mollusca: Polyplacophora) from hydrothermal vent ecosystems in the Okinawa Trough, Japan</mods:title>
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<mods:namePart id="2FF156B4C33897670A0B2092AABB3244">Sigwart, Julia D.</mods:namePart>
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<mods:namePart id="CFD4235981D4008CD2C8F87895FC7B75">Chen, Chong</mods:namePart>
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<emphasis id="D6DA6F24D073FFCCADA4FBE78BC6FBFE" bold="true" box="[151,450,1039,1066]" pageId="1" pageNumber="431">
<taxonomicName id="23AEC8B5D073FFCCADA4FBE78B7CFBFD" ID-CoL="6LBVV" authority="Sigwart &amp; Chen, 2018" authorityName="Sigwart &amp; Chen" authorityYear="2018" box="[151,376,1039,1065]" class="Polyplacophora" family="Leptochitonidae" genus="Hanleyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="431" phylum="Mollusca" rank="species" species="henrici" status="sp. nov.">
<emphasis id="D6DA6F24D073FFCCADA4FBE78B7CFBFD" bold="true" box="[151,376,1039,1065]" italics="true" pageId="1" pageNumber="431">Hanleyella henrici</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDE9D25FD073FFCCAC4CFBF88BC6FBFE" box="[383,450,1040,1066]" pageId="1" pageNumber="431" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="ACB4E0BDD073FFCCADA4FBDA8B04FB9F" box="[151,256,1074,1099]" pageId="1" pageNumber="431" type="description">
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<figureCitation id="7C95AFB3D073FFCCADA4FBDA8B04FB9F" box="[151,256,1074,1099]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1101,1123]" captionStartId-1="3.[151,250,1508,1530]" captionStartId-2="4.[151,250,1051,1073]" captionTargetBox-0="[168,1436,193,1079]" captionTargetBox-1="[150,1436,303,1486]" captionTargetBox-2="[150,1436,337,1030]" captionTargetId-0="figure@2.[151,1436,193,1079]" captionTargetId-1="figure@3.[150,1436,303,1486]" captionTargetId-2="figure@4.[150,1436,337,1030]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Hanleyella henrici n. sp., live images of the holotype (NSMT-Mo 79005), body length 8.9 mm. A. Underside of the head showing Schwabe organ. B. Side view. C. In natural position, live animal collected on shell material and re-attached to a rock post-sampling." captionText-1="FIGURE 2. Hanleyella henrici n. sp., SEM of the valves of the holotype (NSMT-Mo 79005). A. From top to bottom, head valve, ventral side of valve II, dorsal view of intermediate valve, and tail valve. B. Close of up granules in intermediate valve showing aesthete pores and the coalescing chains of granules on distal part of the central area. C. Higher magnification view of round, raised granules on lateral areas in intermediate valve shown in A." captionText-2="FIGURE 3. Hanleyella henrici n. sp., SEM of elements from the holotype (NSMT-Mo 79005). A. Perinotum section, showing dorsal (top) and ventral (bottom) scales. B. Higher magnification view of ribbed dorsal scales. C. Anterior portion of the radula." httpUri-0="https://zenodo.org/record/2614774/files/figure.png" httpUri-1="https://zenodo.org/record/2614776/files/figure.png" httpUri-2="https://zenodo.org/record/2614778/files/figure.png" pageId="1" pageNumber="431">Figs. 13</figureCitation>
</paragraph>
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Type material.
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:
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</emphasis>
NSMT-Mo 79005,
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site,
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hydrothermal vent field (
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),
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<location id="E171E5EDD073FFCCAC88FB76880BFB63" LSID="urn:lsid:plazi:treatment:6C070220D073FFCEADA4FBE78ACDFF2C:E171E5EDD073FFCCAC88FB76880BFB63" box="[443,527,1182,1207]" county="Aki" latitude="27.7687" longLatPrecision="1" longitude="126.90297" municipality="Iheya North" name="Trough" pageId="1" pageNumber="431" stateProvince="Okinawa">Trough</location>
,
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deep, taken by ROV
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<collectorName id="495BD6E0D073FFCCAE42FB768E1FFB63" box="[881,1051,1182,1207]" pageId="1" pageNumber="431">Hyper-Dolphin</collectorName>
</emphasis>
during dive #1616 using a scoop on-board R/
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V
<emphasis id="D6DA6F24D073FFCCAC0BFB2C8B7FFB0F" box="[312,379,1220,1243]" italics="true" pageId="1" pageNumber="431">Kaiyo</emphasis>
</collectorName>
cruise KY14-01,
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<collectingDate id="80546C1ED073FFCCAF70FB2A88B5FB0F" box="[579,689,1218,1243]" pageId="1" pageNumber="431" value="2014-01-25">2014/i/25</collectingDate>
</date>
, fixed and stored in 99% ethanol.
<materialsCitation id="54C6B96BD073FFCCA91EFB2A8F27FA93" ID-GBIF-Occurrence="2116190065" collectingDate="2015-08-03" collectionCode="NCBI, MH" county="Iheya North Original" elevation="950" latitude="27.787333" location="Iheya North" longLatPrecision="1" longitude="126.89843" municipality="North Original" pageId="1" pageNumber="431" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
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<typeStatus id="3B150D94D073FFCCA91EFB2A8E9EFB0F" box="[1069,1178,1218,1243]" pageId="1" pageNumber="431" type="paratype">Paratype</typeStatus>
#1:
</emphasis>
NSMT-Mo 79006,
<collectingCounty id="0D70CBBAD073FFCCADA4FB0E8B8CFB2B" box="[151,392,1254,1279]" pageId="1" pageNumber="431">
Iheya
<collectingMunicipality id="0475294CD073FFCCADECFB0E8B8CFB2B" box="[223,392,1254,1279]" pageId="1" pageNumber="431">North Original</collectingMunicipality>
</collectingCounty>
site,
<location id="E171E5EDD073FFCCACF8FB0E8850FB2B" LSID="urn:lsid:plazi:treatment:6C070220D073FFCEADA4FBE78ACDFF2C:E171E5EDD073FFCCACF8FB0E8850FB2B" box="[459,596,1254,1279]" county="Iheya North Original" latitude="27.787333" longLatPrecision="1" longitude="126.89843" municipality="North Original" name="Iheya North" pageId="1" pageNumber="431" stateProvince="Okinawa">Iheya North</location>
hydrothermal vent field (
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),
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<location id="E171E5EDD073FFCCA877FB0E8F9CFB2B" LSID="urn:lsid:plazi:treatment:6C070220D073FFCEADA4FBE78ACDFF2C:E171E5EDD073FFCCA877FB0E8F9CFB2B" box="[1348,1432,1254,1279]" county="Iheya North Original" latitude="27.787333" longLatPrecision="1" longitude="126.89843" municipality="North Original" name="Trough" pageId="1" pageNumber="431" stateProvince="Okinawa">Trough</location>
,
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deep, taken by ROV
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<location id="E171E5EDD073FFCCACE4FAE28886FAF7" LSID="urn:lsid:plazi:treatment:6C070220D073FFCEADA4FBE78ACDFF2C:E171E5EDD073FFCCACE4FAE28886FAF7" box="[471,642,1290,1315]" county="Iheya North Original" latitude="27.787333" longLatPrecision="1" longitude="126.89843" municipality="North Original" name="Hyper-Dolphin" pageId="1" pageNumber="431" stateProvince="Okinawa">Hyper-Dolphin</location>
</emphasis>
during dive #1861 using a suction sampler on-board R/V
<taxonomicName id="23AEC8B5D073FFCCA82BFAE28F98FAF7" authorityName="Miura &amp; Laubier" authorityYear="1990" box="[1304,1436,1290,1315]" class="Polychaeta" family="Chrysopetalidae" genus="Natsushima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="431" phylum="Annelida" rank="genus">
<emphasis id="D6DA6F24D073FFCCA82BFAE28F98FAF7" box="[1304,1436,1290,1315]" italics="true" pageId="1" pageNumber="431">Natsushima</emphasis>
</taxonomicName>
cruise NT15-13,
<date id="901095F6D073FFCCAC6AFAC68BD3FA93" box="[345,471,1326,1351]" pageId="1" pageNumber="431" value="2015-08-03">
<collectingDate id="80546C1ED073FFCCAC6AFAC68BD3FA93" box="[345,471,1326,1351]" pageId="1" pageNumber="431" value="2015-08-03">2015/viii/3</collectingDate>
</date>
, fixed and stored in 99% ethanol.
<collectionCode id="82BF2BF3D073FFCCAE50FAC789A2FA93" box="[867,934,1327,1351]" pageId="1" pageNumber="431">NCBI</collectionCode>
GenBank accession:
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<collectionCode id="82BF2BF3D073FFCCA9ADFAC68EC8FA93" box="[1182,1228,1326,1351]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="1" pageNumber="431">MH</collectionCode>
349538
</accessionNumber>
</materialsCitation>
.
<materialsCitation id="54C6B96BD073FFCCA81CFAC68F00FA67" ID-GBIF-Occurrence="2116189669" collectingDate="2016-12-04" collectionCode="KAIKO, NCBI, MH" collectorName="V Kairei &amp; GenBank" county="Fukai" elevation="1178" latitude="28.436995" location="Trough" longLatPrecision="1" longitude="128.18855" municipality="Higashi-Ensei" pageId="1" pageNumber="431" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
<emphasis id="D6DA6F24D073FFCCA81CFAC68AB8FABF" bold="true" pageId="1" pageNumber="431">
<typeStatus id="3B150D94D073FFCCA81CFAC68F98FA93" box="[1327,1436,1326,1351]" pageId="1" pageNumber="431" type="paratype">Paratype</typeStatus>
#2:
</emphasis>
NSMT-Mo 79007,
<collectingCounty id="0D70CBBAD073FFCCAC9EFABA8BE9FABF" box="[429,493,1362,1387]" pageId="1" pageNumber="431">Fukai</collectingCounty>
site (
<bibRefCitation id="803FCEC7D073FFCCAF04FABA890DFABF" author="Chen, C. &amp; Watanabe, H. K. &amp; Miyazaki, J. &amp; Kawagucci, S." box="[567,777,1362,1387]" pageId="1" pageNumber="431" pagination="4121" refId="ref2908" refString="Chen, C., Watanabe, H. K., Miyazaki, J. &amp; Kawagucci, S. (2017) Unanticipated discovery of two rare gastropod molluscs from recently located hydrothermally influenced areas in the Okinawa Trough. PeerJ, 5, e 4121. https: // doi. org / 10.7717 / peerj. 4121" type="journal article" year="2017">
Chen
<emphasis id="D6DA6F24D073FFCCAFB2FABC88C4FABF" box="[641,704,1362,1387]" italics="true" pageId="1" pageNumber="431">et al.</emphasis>
2017
</bibRefCitation>
),
<collectingMunicipality id="0475294CD073FFCCAE10FABA89C5FABF" box="[803,961,1362,1387]" pageId="1" pageNumber="431">Higashi-Ensei</collectingMunicipality>
hydrothermal vent field (
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<geoCoordinate id="819AD5F1D073FFCCADA4FA9F8B40FA5B" box="[151,324,1399,1423]" direction="east" orientation="longitude" pageId="1" pageNumber="431" precision="1" value="128.18855">128°11.3131E</geoCoordinate>
),
<collectingRegion id="266A7DD4D073FFCCAC68FA9E8BC4FA5B" box="[347,448,1398,1423]" country="Japan" name="Okinawa" pageId="1" pageNumber="431">Okinawa</collectingRegion>
<location id="E171E5EDD073FFCCACFAFA9E8819FA5B" LSID="urn:lsid:plazi:treatment:6C070220D073FFCEADA4FBE78ACDFF2C:E171E5EDD073FFCCACFAFA9E8819FA5B" box="[457,541,1398,1423]" county="Fukai" latitude="28.436995" longLatPrecision="1" longitude="128.18855" municipality="Higashi-Ensei" name="Trough" pageId="1" pageNumber="431" stateProvince="Okinawa">Trough</location>
,
<quantity id="23561ED3D073FFCCAF18FA9F8886FA5B" box="[555,642,1399,1423]" metricMagnitude="3" metricUnit="m" metricValue="1.178" pageId="1" pageNumber="431" unit="m" value="1178.0">
<elevation id="6F835405D073FFCCAF18FA9F8886FA5B" box="[555,642,1399,1423]" metricMagnitude="3" metricUnit="m" metricValue="1.178" pageId="1" pageNumber="431" unit="m" value="1178.0">1178 m</elevation>
</quantity>
deep, taken by ROV
<emphasis id="D6DA6F24D073FFCCAEB1FA9F89D2FA5A" box="[898,982,1399,1422]" italics="true" pageId="1" pageNumber="431">
<collectionCode id="82BF2BF3D073FFCCAEB1FA9F89D2FA5A" box="[898,982,1399,1422]" pageId="1" pageNumber="431">KAIKO</collectionCode>
</emphasis>
during dive #724 on-board R/
<collectorName id="495BD6E0D073FFCCA80BFA9F8F9FFA5B" box="[1336,1435,1399,1423]" pageId="1" pageNumber="431">
V
<emphasis id="D6DA6F24D073FFCCA866FA908F9FFA5B" box="[1365,1435,1400,1423]" italics="true" pageId="1" pageNumber="431">Kairei</emphasis>
</collectorName>
cruise KR16-16,
<date id="901095F6D073FFCCAC64FA728BC9FA67" box="[343,461,1434,1459]" pageId="1" pageNumber="431" value="2016-12-04">
<collectingDate id="80546C1ED073FFCCAC64FA728BC9FA67" box="[343,461,1434,1459]" pageId="1" pageNumber="431" value="2016-12-04">2016/xii/4</collectingDate>
</date>
, fixed and stored in 99% ethanol.
<collectionCode id="82BF2BF3D073FFCCAE78FA73898AFA67" box="[843,910,1435,1459]" pageId="1" pageNumber="431">NCBI</collectionCode>
<collectorName id="495BD6E0D073FFCCAEA6FA7289FBFA67" box="[917,1023,1434,1459]" pageId="1" pageNumber="431">GenBank</collectorName>
accession:
<accessionNumber id="FBFD2ED5D073FFCCA9B3FA728F00FA67" box="[1152,1284,1434,1459]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH349539" pageId="1" pageNumber="431">
<collectionCode id="82BF2BF3D073FFCCA9B3FA728EAAFA67" box="[1152,1198,1434,1459]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="1" pageNumber="431">MH</collectionCode>
349539
</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ACB4E0BDD073FFCCADF4FA568AEDFA2F" pageId="1" pageNumber="431" type="etymology">
<paragraph id="E411B336D073FFCCADF4FA568AEDFA2F" blockId="1.[151,1437,1146,1963]" pageId="1" pageNumber="431">
<emphasis id="D6DA6F24D073FFCCADF4FA568B4AFA03" bold="true" box="[199,334,1470,1495]" pageId="1" pageNumber="431">Etymology.</emphasis>
Named in honour of Henry Schwabe, who shares the Latin form of his name with his father Enrico.
</paragraph>
</subSubSection>
<subSubSection id="ACB4E0BDD073FFCCADF4F9EE892AF9CB" box="[199,814,1542,1568]" pageId="1" pageNumber="431" type="materials_examined">
<paragraph id="E411B336D073FFCCADF4F9EE892AF9CB" blockId="1.[151,1437,1146,1963]" box="[199,814,1542,1568]" pageId="1" pageNumber="431">
<emphasis id="D6DA6F24D073FFCCADF4F9EE8BB4F9CB" bold="true" box="[199,432,1542,1567]" pageId="1" pageNumber="431">Material examined.</emphasis>
Known only from the
<typeStatus id="3B150D94D073FFCCAF83F9E088E4F9F4" box="[688,736,1544,1568]" pageId="1" pageNumber="431">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="ACB4E0BDD073FFCCADF4F9C28AEBF9B3" pageId="1" pageNumber="431" type="distribution">
<paragraph id="E411B336D073FFCCADF4F9C28AEBF9B3" blockId="1.[151,1437,1146,1963]" pageId="1" pageNumber="431">
<emphasis id="D6DA6F24D073FFCCADF4F9C28B64F997" bold="true" box="[199,352,1578,1603]" pageId="1" pageNumber="431">Distribution.</emphasis>
Known only from the localities of the
<typeStatus id="3B150D94D073FFCCAE28F9C3894FF997" box="[795,843,1579,1603]" pageId="1" pageNumber="431">type</typeStatus>
series; deep-sea hydrothermal vents of the
<collectingRegion id="266A7DD4D073FFCCA804F9C28F98F997" box="[1335,1436,1578,1603]" country="Japan" name="Okinawa" pageId="1" pageNumber="431">Okinawa</collectingRegion>
Trough.
</paragraph>
</subSubSection>
<subSubSection id="ACB4E0BDD073FFCCADF4F99A8E74F8EB" pageId="1" pageNumber="431" type="diagnosis">
<paragraph id="E411B336D073FFCCADF4F99A8E74F8EB" blockId="1.[151,1437,1146,1963]" pageId="1" pageNumber="431">
<emphasis id="D6DA6F24D073FFCCADF4F99A8B45F95F" bold="true" box="[199,321,1650,1675]" pageId="1" pageNumber="431">Diagnosis.</emphasis>
Valves moderately elevated, not beaked. Intermediate valves side margins rounded, anterior margins straight, posterior margins slightly convex, lateral areas slightly inflated. Tail valve mucro anterior of central, postmucronal slope slightly concave. Tegmentum with pronounced longitudinal ridges formed of fused flattened round granules in central areas, lateral areas with occasional strongly pronounced raised spherical granules. Aesthete caps not projecting. Girdle narrow, dorsally with ribbed bluntly pointed scales, ventrally with smooth flat scales. Gills seven per side. Radula major lateral teeth with tridentate cusps.
</paragraph>
</subSubSection>
<subSubSection id="ACB4E0BDD073FFCEADF4F8A28ACDFF2C" lastPageId="3" lastPageNumber="433" pageId="1" pageNumber="431" type="description">
<paragraph id="E411B336D073FFCCADF4F8A289EBF87F" blockId="1.[151,1437,1146,1963]" pageId="1" pageNumber="431">
<emphasis id="D6DA6F24D073FFCCADF4F8A28B5DF8B7" bold="true" box="[199,345,1866,1891]" pageId="1" pageNumber="431">Description.</emphasis>
<typeStatus id="3B150D94D073FFCCAC53F8A28BCCF8B7" box="[352,456,1866,1891]" pageId="1" pageNumber="431" type="holotype">Holotype</typeStatus>
8.9 x
<quantity id="23561ED3D073FFCCAF21F8A38868F8B6" box="[530,620,1867,1891]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="431" unit="mm" value="4.3">4.3 mm</quantity>
. Shell carinated, moderately elevated (elevation ratio
<quantity id="23561ED3D073FFCCA9FDF8A38F19F8B7" box="[1230,1309,1866,1891]" metricMagnitude="-2" metricUnit="m" metricValue="1.0922" pageId="1" pageNumber="431" unit="in" value="0.43">0.43 in</quantity>
valve II of
<typeStatus id="3B150D94D073FFCCADA4F8868AF9F853" box="[151,253,1902,1927]" pageId="1" pageNumber="431" type="holotype">holotype</typeStatus>
), valves not beaked. Colour of tegmentum white, in
<typeStatus id="3B150D94D073FFCCAE55F88689CCF853" box="[870,968,1902,1927]" pageId="1" pageNumber="431" type="holotype">holotype</typeStatus>
valves and girdle covered dorsally with brown mineral deposits, pale newer growth visible on distal edges of valves.
</paragraph>
<caption id="B0D1E3BED070FFCFADA4FBA58B76FB74" httpUri="https://zenodo.org/record/2614774/files/figure.png" pageId="2" pageNumber="432" startId="2.[151,250,1101,1123]" targetBox="[168,1436,193,1079]" targetPageId="2">
<paragraph id="E411B336D070FFCFADA4FBA58B76FB74" blockId="2.[151,1436,1100,1184]" pageId="2" pageNumber="432">
<emphasis id="D6DA6F24D070FFCFADA4FBA58B10FBB6" bold="true" box="[151,276,1101,1123]" pageId="2" pageNumber="432">FIGURE 1.</emphasis>
<taxonomicName id="23AEC8B5D070FFCFAC28FBA48BDDFBB6" authority="Sigwart &amp; Chen, 2018" authorityName="Sigwart &amp; Chen" authorityYear="2018" box="[283,473,1100,1122]" class="Polyplacophora" family="Leptochitonidae" genus="Hanleyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidopleurida" pageId="2" pageNumber="432" phylum="Mollusca" rank="species" species="henrici" status="sp. nov.">
<emphasis id="D6DA6F24D070FFCFAC28FBA48BDDFBB6" box="[283,473,1100,1122]" italics="true" pageId="2" pageNumber="432">Hanleyella henrici</emphasis>
</taxonomicName>
<emphasis id="D6DA6F24D070FFCFACD3FBA5881EFBB6" bold="true" box="[480,538,1101,1122]" pageId="2" pageNumber="432">
<taxonomicNameLabel id="CDE9D25FD070FFCFACD3FBA5881EFBB6" box="[480,538,1101,1122]" pageId="2" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, live images of the holotype (NSMT-Mo 79005), body length 8.9 mm.
<emphasis id="D6DA6F24D070FFCFA9C4FBA58F14FBB6" bold="true" box="[1271,1296,1101,1122]" pageId="2" pageNumber="432">A.</emphasis>
Underside of the head showing Schwabe organ.
<emphasis id="D6DA6F24D070FFCFACC0FB84880EFB55" bold="true" box="[499,522,1132,1153]" pageId="2" pageNumber="432">B.</emphasis>
Side view.
<emphasis id="D6DA6F24D070FFCFAF4CFB83889CFB55" bold="true" box="[639,664,1131,1153]" pageId="2" pageNumber="432">C.</emphasis>
In natural position, live animal collected on shell material and re-attached to a rock post-sampling.
</paragraph>
</caption>
<paragraph id="E411B336D070FFCFADF4FB2F890CFA9F" blockId="2.[151,1437,1223,2004]" pageId="2" pageNumber="432">Head valve slightly more than semicircular, slightly more than 10% wider than tail valve. Intermediate valves rectangular, lateral areas slightly inflated, anterior margins nearly straight, except margin of valve II convex, posterior margins slightly convex. Tail valve with mucro anterior of centre, anterior margin of jugal area straight except in valve II, postmucronal slope slightly concave.</paragraph>
<paragraph id="E411B336D070FFCFADF4FAB08E24F9F0" blockId="2.[151,1437,1223,2004]" pageId="2" pageNumber="432">Tegmentum of central areas and antemucronal area with elongate round granules (approx. 100 × 6575 µm) fused to form pronounced longitunidal ribs, about 15 ribs per side in intermediate valves. Ribs becoming more irregular and closer set toward jugum; on distal pleural areas, ribs curving inward as they join lateral area. Lateral areas, head valve, and postmucronal area of tail valve sculpted with distinctive, highly raised irregularly arranged spherical granules (approx. diameter 100 µm). Some growth marks visible especially on the postmucronal area and head valve, and lateral areas continuing into central areas of intermediate valves.</paragraph>
<paragraph id="E411B336D070FFCFADF4F9C78B21F95B" blockId="2.[151,1437,1223,2004]" pageId="2" pageNumber="432">Each granule with one megalaesthete (7 µm diameter) surrounded by 46 slightly smaller micraesthetes (45 µm), irregularly arranged. Single aesthete pores (7 µm diameter) arranged in spaces between spherical granules on lateral areas.</paragraph>
<paragraph id="E411B336D070FFCFADF4F97388ECF928" blockId="2.[151,1437,1223,2004]" pageId="2" pageNumber="432">Articulamentum well developed, eaves of intermediate valves spongy and porous texture to jugal area and lateral area where tegmentum overhangs the primordium of an insertion plate; apophyses small, widely separated, anteriorly rounded with long straight slope to jugum.</paragraph>
<paragraph id="E411B336D070FFCFADF4F8EF8F04F8BC" blockId="2.[151,1437,1223,2004]" pageId="2" pageNumber="432">Girdle narrow, dorsally covered in blunt scales with 45 ribs, scales 60 × 30 µm, longer toward valve interstices. Occasional marginal spicules, cylindrical, smooth, over 130 µm long. Ventrally, girdle covered in elongate, smooth, flat, bluntly pointed scales 60 × 24 µm, with four short ribs at tip on one side only.</paragraph>
<paragraph id="E411B336D070FFCFADF4F89B89E5F800" blockId="2.[151,1437,1223,2004]" pageId="2" pageNumber="432">Radula major lateral teeth flat and wide with tridentate heads, interior denticle slightly longer than the exterior denticle; outer denticle well formed but worn. Central tooth flat, with a short blade that forms a curled edge extending down both lateral edges of the tooth. First laterals short, pointed.</paragraph>
<paragraph id="E411B336D071FFCEADF4FF7F8ACDFF2C" blockId="3.[151,1436,151,248]" pageId="3" pageNumber="433">
Gills seven per side, without interspace, extending posteriorly to the anus. Schwabe organ (
<bibRefCitation id="803FCEC7D071FFCEACB7FF548867FF00" author="Sigwart, J. D. &amp; Sumner-Rooney, L. H. &amp; Schwabe, E. &amp; Hess, M. &amp; Brennan, G. P. &amp; Schrodl, M." box="[388,611,188,213]" pageId="3" pageNumber="433" pagination="7" refId="ref3970" refString="Sigwart, J. D., Sumner-Rooney, L. H., Schwabe, E., Hess, M., Brennan, G. P. &amp; Schrodl, M. (2014) A new sensory organ in &quot; primitive &quot; molluscs (Polyplacophora: Lepidopleurida), and its context in the nervous system of chitons. Frontiers in Zoology, 11, 7. https: // doi. org / 10.1186 / 1742 - 9994 - 11 - 7" type="journal article" year="2014">
Sigwart
<emphasis id="D6DA6F24D071FFCEACD5FF558824FF01" box="[486,544,188,213]" italics="true" pageId="3" pageNumber="433">et al.</emphasis>
2014
</bibRefCitation>
) present under the corner of the mouth lappets, dark and triangular (
<figureCitation id="7C95AFB3D071FFCEA846FF548ABFFF2C" captionStart="FIGURE 1" captionStartId="2.[151,250,1101,1123]" captionTargetBox="[168,1436,193,1079]" captionTargetId="figure@2.[151,1436,193,1079]" captionTargetPageId="2" captionText="FIGURE 1. Hanleyella henrici n. sp., live images of the holotype (NSMT-Mo 79005), body length 8.9 mm. A. Underside of the head showing Schwabe organ. B. Side view. C. In natural position, live animal collected on shell material and re-attached to a rock post-sampling." httpUri="https://zenodo.org/record/2614774/files/figure.png" pageId="3" pageNumber="433">Fig 1A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>