213 lines
31 KiB
XML
213 lines
31 KiB
XML
<document id="40E12D8F1453602ACB297DCB17A6C279" ID-DOI="10.1649/072.066.0202" ID-ISSN="1938-4394" ID-PMC="PMC4148458" ID-PubMed="25177045" ID-Zenodo-Dep="7311791" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630187931888" checkinUser="diego" docAuthor="Jean, April, Telles, Nicole D., Gibson, J. Randy, Foley, Dan & Miller, Kelly B." docDate="2012" docId="2A4C87D1E205FF82FF595382C3D4F992" docLanguage="en" docName="072.066.0202.pdf.imf" docOrigin="The Coleopterists Bulletin 66 (2)" docSource="http://dx.doi.org/10.1649/072.066.0202" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Psychopomporus Jean, Telles, and Miller, new genus" docType="treatment" docVersion="4" lastPageNumber="106" masterDocId="D675FFA9E204FF83FFE15616C747FF95" masterDocTitle="Description of a New Genus and Species of Stygobiontic Diving Beetle, Psychopomporus felipiJean, Telles, and Miller (Coleoptera: Dytiscidae: Hydroporinae), from the Edwards-Trinity Aquifer System of Texas, USA" masterLastPageNumber="110" masterPageNumber="105" pageNumber="106" updateTime="1699180466305" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="BB284F99E607D8EF6CBFA94F642512B5">Description of a New Genus and Species of Stygobiontic Diving Beetle, Psychopomporus felipiJean, Telles, and Miller (Coleoptera: Dytiscidae: Hydroporinae), from the Edwards-Trinity Aquifer System of Texas, USA</mods:title>
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<mods:namePart id="A045B554D8B4B6503F2493599F2DCD3C">Jean, April</mods:namePart>
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<mods:namePart id="744B17A18204B2FE1D016BC04E28334C">Telles, Nicole D.</mods:namePart>
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<mods:namePart id="B67AF473ED04E7DF4EEA392DEA01219F">Gibson, J. Randy</mods:namePart>
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<mods:namePart id="FB5A9EA84252495B5D09A7DF3DC2CB3C">Foley, Dan</mods:namePart>
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<mods:namePart id="9A37B6F6B324B6A1DA35DFC2B8383ADC">Miller, Kelly B.</mods:namePart>
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<mods:title id="9F9C05DDB155D1E5C40BCB78DA9F2D45">The Coleopterists Bulletin</mods:title>
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<mods:date id="970998689A6C1B2AF53C5EA2E8920CD6">2012</mods:date>
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<mods:number id="84CB369BF46E21A5F8085077444FC4B3">2012-06-30</mods:number>
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<mods:identifier id="EB6FDA9112923DB3B78D4327CC2A1EA7" type="DOI">10.1649/072.066.0202</mods:identifier>
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<mods:identifier id="276FFF467182A365FD32151E4AD32433" type="ISSN">1938-4394</mods:identifier>
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<treatment id="2A4C87D1E205FF82FF595382C3D4F992" LSID="urn:lsid:plazi:treatment:2A4C87D1E205FF82FF595382C3D4F992" httpUri="http://treatment.plazi.org/id/2A4C87D1E205FF82FF595382C3D4F992" lastPageNumber="106" pageId="1" pageNumber="106">
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<subSubSection id="EAFF654CE205FF82FF595382C687FA51" pageId="1" pageNumber="106" type="nomenclature">
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<paragraph id="A25A36C7E205FF82FF595382C687FA51" blockId="1.[184,605,1428,1503]" pageId="1" pageNumber="106">
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<heading id="F91281ABE205FF82FF595382C687FA51" centered="true" fontSize="10" level="2" pageId="1" pageNumber="106" reason="2">
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<emphasis id="9091EAD5E205FF82FF595382C687FA51" bold="true" pageId="1" pageNumber="106">
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<taxonomicName id="65E54D44E205FF82FF595382C51EFA3C" ID-CoL="8LSTN" authority="Jean, Telles, and Miller" authorityName="Jean, Telles, and Miller" box="[184,601,1428,1450]" class="Insecta" family="Dytiscidae" genus="Psychopomporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="9091EAD5E205FF82FF595382C627FA3F" bold="true" box="[184,352,1428,1450]" italics="true" pageId="1" pageNumber="106">Psychopomporus</emphasis>
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Jean, Telles, and Miller
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</taxonomicName>
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,
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<taxonomicNameLabel id="8BA257AEE205FF82FEB453B9C687FA51" box="[341,448,1455,1476]" pageId="1" pageNumber="106" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="EAFF654CE205FF82FEDA53DFC69EFA4B" box="[315,473,1481,1503]" pageId="1" pageNumber="106" type="description">
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<paragraph id="A25A36C7E205FF82FEDA53DFC69EFA4B" blockId="1.[184,605,1428,1503]" box="[315,473,1481,1503]" pageId="1" pageNumber="106">
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(
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<figureCitation id="3ADE2A42E205FF82FEA353DFC695FA4B" box="[322,466,1481,1503]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="2.[197,237,1657,1676]" captionStartId-1="3.[175,207,1151,1170]" captionTargetBox-0="[173,1193,183,1633]" captionTargetBox-1="[176,1121,218,1100]" captionTargetId-0="figure-0@2.[173,1193,183,1633]" captionTargetId-1="figure-266@3.[151,1171,183,1128]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." captionText-1="Fig. 7. Distribution of Psychopomporus felipi and other Texas subterranean diving beetles, showing extent of Edwards-Trinity Aquifer." figureDoi-0="http://doi.org/10.5281/zenodo.7311793" figureDoi-1="http://doi.org/10.5281/zenodo.7311795" httpUri-0="https://zenodo.org/record/7311793/files/figure.png" httpUri-1="https://zenodo.org/record/7311795/files/figure.png" pageId="1" pageNumber="106">Figs. 1–4, 6–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="EAFF654CE205FF82FF4E53E2C538F99C" box="[175,639,1524,1545]" pageId="1" pageNumber="106" type="type_taxon">
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<paragraph id="A25A36C7E205FF82FF4E53E2C538F99C" blockId="1.[151,639,1524,1729]" box="[175,639,1524,1545]" pageId="1" pageNumber="106">
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<emphasis id="9091EAD5E205FF82FF4E53E2C677F99C" bold="true" box="[175,304,1524,1545]" pageId="1" pageNumber="106">
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<typeStatus id="7D5E8865E205FF82FF4E53E2C799F99C" box="[175,222,1524,1545]" pageId="1" pageNumber="106">Type</typeStatus>
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Species.
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</emphasis>
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<taxonomicName id="65E54D44E205FF82FEDC53E2C54EF99C" ID-CoL="8LVH5" authorityName="JEAN, TELLES, AND MILLER" authorityYear="2011" box="[317,521,1524,1545]" class="Insecta" family="Dytiscidae" genus="Psychopomporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="species" species="felipi" status="sp. nov.">
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<emphasis id="9091EAD5E205FF82FEDC53E2C54EF99C" box="[317,521,1524,1545]" italics="true" pageId="1" pageNumber="106">Psychopomporus felipi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8BA257AEE205FF82FDEF53E2C53DF99C" box="[526,634,1524,1545]" pageId="1" pageNumber="106" rank="species">new species</taxonomicNameLabel>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="EAFF654CE205FF82FF4E5018C492FDDE" pageId="1" pageNumber="106" type="diagnosis">
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<paragraph id="A25A36C7E205FF82FF4E5018C492FDDE" blockId="1.[151,639,1524,1729]" lastBlockId="1.[685,1173,194,1543]" pageId="1" pageNumber="106">
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<emphasis id="9091EAD5E205FF82FF4E5018C65FF9B6" bold="true" box="[175,280,1550,1571]" pageId="1" pageNumber="106">Diagnosis.</emphasis>
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Adults of
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<taxonomicName id="65E54D44E205FF82FE685019C56AF9B1" box="[393,557,1551,1572]" class="Insecta" family="Dytiscidae" genus="Psychopomporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">
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<emphasis id="9091EAD5E205FF82FE685019C56AF9B1" box="[393,557,1551,1572]" italics="true" pageId="1" pageNumber="106">Psychopomporus</emphasis>
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</taxonomicName>
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are distinct among
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<taxonomicName id="65E54D44E205FF82FEEA503FC6CBF9AB" authorityName="Aube" authorityYear="1836" box="[267,396,1577,1598]" pageId="1" pageNumber="106" rank="subFamily" subFamily="Hydroporinae">Hydroporinae</taxonomicName>
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based on the combination of the following character states: 1) adaptations to subterranean environment including light pigmentation, fused elytra, and metathoracic wings and compound eyes reduced (
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<figureCitation id="3ADE2A42E205FF82FE875084C6D9F932" box="[358,414,1682,1703]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 1</figureCitation>
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); 2) prosternal process strongly declivous, apically reaching the anterio- medial projection of the metasternum (
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<figureCitation id="3ADE2A42E205FF82FBF156D4C322FF42" box="[1040,1125,194,215]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Figs. 2, 3</figureCitation>
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) and with a prominent projection anteriorly (
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<figureCitation id="3ADE2A42E205FF82FBA656CBC3C1FF67" box="[1095,1158,221,242]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 3</figureCitation>
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); 3) mesotibia apically expanded and strongly arcuate medially (
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<figureCitation id="3ADE2A42E205FF82FCF15704C40AFEB2" box="[784,845,274,295]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 2</figureCitation>
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); and 4) elytral epipleuron broad and flat anteriorly (
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<figureCitation id="3ADE2A42E205FF82FC86573AC4E4FED4" box="[871,931,300,321]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 2</figureCitation>
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). The general body form and obvious cave adaptations are similar to other stygobiontic members of
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<taxonomicName id="65E54D44E205FF82FC425774C34EFEE2" authorityName="Leach" authorityYear="1815" box="[931,1033,354,375]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
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, suggestive of convergent adaption to the subterranean environment. Many stygobiontic diving beetles have a shortened prosternal process that does not reach the anterior margin of the metasternum and a distinct habitus with the pronotum cordate and a strong discontinuity in the curvature between the pronotum and elytron. These features are not present in
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<taxonomicName id="65E54D44E205FF82FD4C5420C413FDDE" box="[685,852,566,587]" class="Insecta" family="Dytiscidae" genus="Psychopomporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">
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<emphasis id="9091EAD5E205FF82FD4C5420C413FDDE" box="[685,852,566,587]" italics="true" pageId="1" pageNumber="106">Psychopomporus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="3ADE2A42E205FF82FC865420C48FFDDE" box="[871,968,565,587]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Figs. 1–3</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="EAFF654CE205FF82FD245447C33FFD20" pageId="1" pageNumber="106" type="etymology">
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<paragraph id="A25A36C7E205FF82FD245447C33FFD20" blockId="1.[685,1173,194,1543]" pageId="1" pageNumber="106">
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<emphasis id="9091EAD5E205FF82FD245447C473FDF3" bold="true" box="[709,820,593,614]" pageId="1" pageNumber="106">Etymology.</emphasis>
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The genus is named
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<taxonomicName id="65E54D44E205FF82FC175447C3D3FDF3" box="[1014,1172,593,614]" class="Insecta" family="Dytiscidae" genus="Psychopomporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">
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<emphasis id="9091EAD5E205FF82FC175447C3D3FDF3" box="[1014,1172,593,614]" italics="true" pageId="1" pageNumber="106">Psychopomporus</emphasis>
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</taxonomicName>
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(masculine), from the Greek
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<emphasis id="9091EAD5E205FF82FC23547DC30BFD15" box="[962,1100,619,640]" italics="true" pageId="1" pageNumber="106">psychopompos</emphasis>
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, meaning “guide of souls” and
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<emphasis id="9091EAD5E205FF82FC4A5493C4A9FD0E" box="[939,1006,645,667]" italics="true" pageId="1" pageNumber="106">–porus</emphasis>
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||
, a common root for genus names in the subfamily
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<taxonomicName id="65E54D44E205FF82FC0E54B6C334FD20" authorityName="Aube" authorityYear="1836" box="[1007,1139,672,693]" pageId="1" pageNumber="106" rank="subFamily" subFamily="Hydroporinae">Hydroporinae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="EAFF654CE205FF82FD2454ADC4F2FC31" pageId="1" pageNumber="106" type="distribution">
|
||
<paragraph id="A25A36C7E205FF82FD2454ADC4F2FC31" blockId="1.[685,1173,194,1543]" pageId="1" pageNumber="106">
|
||
<emphasis id="9091EAD5E205FF82FD2454ADC40FFD45" bold="true" box="[709,840,699,720]" pageId="1" pageNumber="106">Distribution.</emphasis>
|
||
Specimens of the single species in this genus were recovered in the flow from two springs discharging into the western side of San Felipe Creek (
|
||
<geoCoordinate id="C7D15000E205FF82FCAB551CC48CFC8A" box="[842,971,778,799]" degrees="29.37234" direction="north" orientation="latitude" pageId="1" pageNumber="106" precision="1" value="29.37234">29.37234°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C7D15000E205FF82FC3D551CC32BFC8A" box="[988,1132,778,799]" degrees="100.88602" direction="west" orientation="longitude" pageId="1" pageNumber="106" precision="1" value="-100.88602">100.88602°W</geoCoordinate>
|
||
) in Del Rio, Val Verde County,
|
||
<collectingRegion id="6021F825E205FF82FC245533C347FCAF" box="[965,1024,805,826]" country="United States of America" name="Texas" pageId="1" pageNumber="106">Texas</collectingRegion>
|
||
,
|
||
<collectingCountry id="DAF27657E205FF82FBEA5533C37CFCAF" box="[1035,1083,805,826]" name="United States of America" pageId="1" pageNumber="106">USA</collectingCountry>
|
||
(
|
||
<figureCitation id="3ADE2A42E205FF82FBAB5533C3CFFCAF" box="[1098,1160,805,826]" captionStart="Fig" captionStartId="3.[175,207,1151,1170]" captionTargetBox="[176,1121,218,1100]" captionTargetId="figure-266@3.[151,1171,183,1128]" captionTargetPageId="3" captionText="Fig. 7. Distribution of Psychopomporus felipi and other Texas subterranean diving beetles, showing extent of Edwards-Trinity Aquifer." figureDoi="http://doi.org/10.5281/zenodo.7311795" httpUri="https://zenodo.org/record/7311795/files/figure.png" pageId="1" pageNumber="106">Fig. 7</figureCitation>
|
||
). These springs discharge from the Edwards-Trinity aquifer system. The distribution of
|
||
<taxonomicName id="65E54D44E205FF82FC18554DC37AFCE5" authorityName="JEAN, TELLES, AND MILLER" authorityYear="2011" box="[1017,1085,859,880]" class="Insecta" family="Dytiscidae" genus="Psychopomporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="species" species="felipi">
|
||
<emphasis id="9091EAD5E205FF82FC18554DC37AFCE5" box="[1017,1085,859,880]" italics="true" pageId="1" pageNumber="106">P. felipi</emphasis>
|
||
</taxonomicName>
|
||
throughout this aquifer, as well as possible occurrence in other aquifers, is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EAFF654CE205FF82FD2455BCC3D4F992" pageId="1" pageNumber="106" type="discussion">
|
||
<paragraph id="A25A36C7E205FF82FD2455BCC3D4F992" blockId="1.[685,1173,194,1543]" pageId="1" pageNumber="106">
|
||
<emphasis id="9091EAD5E205FF82FD2455BCC470FC2A" bold="true" box="[709,823,938,959]" pageId="1" pageNumber="106">Discussion.</emphasis>
|
||
<taxonomicName id="65E54D44E205FF82FCA455BCC4B5FC2A" box="[837,1010,938,959]" class="Insecta" family="Dytiscidae" genus="Psychopomporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9091EAD5E205FF82FCA455BCC4B5FC2A" box="[837,1010,938,959]" italics="true" pageId="1" pageNumber="106">Psychopomporus</emphasis>
|
||
</taxonomicName>
|
||
belongs within the tribe
|
||
<taxonomicName id="65E54D44E205FF82FCE455D2C356FC4C" authority="(Hydroporinae)" baseAuthorityName="Hydroporinae" box="[773,1041,964,985]" pageId="1" pageNumber="106" rank="tribe" tribe="Hydroporini">Hydroporini (Hydroporinae)</taxonomicName>
|
||
based on the following characters: 1) pronotum lacking bilateral basal plicae (
|
||
<figureCitation id="3ADE2A42E205FF82FCD355ECC429FB9A" box="[818,878,1018,1039]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 1</figureCitation>
|
||
); 2) scutellum not externally visible (
|
||
<figureCitation id="3ADE2A42E205FF82FD1D5202C471FBBC" box="[764,822,1044,1065]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 1</figureCitation>
|
||
); 3) pro- and mesotarsi moderately pseudotetramerous, though not strongly so (
|
||
<figureCitation id="3ADE2A42E205FF82FBAC5239C3CFFBD1" box="[1101,1160,1071,1092]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 2</figureCitation>
|
||
); 4) prosternal process strongly declivous (
|
||
<figureCitation id="3ADE2A42E205FF82FBCC525FC3C0FBCB" box="[1069,1159,1097,1118]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Figs. 2, 3</figureCitation>
|
||
); 5) apices of elytra evenly rounded (
|
||
<figureCitation id="3ADE2A42E205FF82FC0C5272C303FBEC" box="[1005,1092,1124,1145]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Figs. 1, 2</figureCitation>
|
||
); 6) posterior margin of metacoxal processes in similar plane as abdominal sternum (
|
||
<figureCitation id="3ADE2A42E205FF82FC67528FC4F8FB3B" box="[902,959,1177,1198]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 2</figureCitation>
|
||
); 7) metacoxal process with lobes small but distinct, covering the base of the metatrochanter (
|
||
<figureCitation id="3ADE2A42E205FF82FC8C52D8C4EDFB76" box="[877,938,1230,1251]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 2</figureCitation>
|
||
); 8) metatarsal claws of equal length (
|
||
<figureCitation id="3ADE2A42E205FF82FCD452FFC435FB6B" box="[821,882,1257,1278]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 2</figureCitation>
|
||
); 9) male genitalia bilaterally symmetrical (
|
||
<figureCitation id="3ADE2A42E205FF82FCCC5315C420FA8D" box="[813,871,1283,1304]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 4</figureCitation>
|
||
); and 10) male lateral lobe with a single segment (
|
||
<figureCitation id="3ADE2A42E205FF82FC845308C4E4FAA6" box="[869,931,1310,1331]" captionStart="Figs" captionStartId="2.[197,237,1657,1676]" captionTargetBox="[173,1193,183,1633]" captionTargetId="figure-0@2.[173,1193,183,1633]" captionTargetPageId="2" captionText="Figs. 1–6. Psychopomporus felipi. 1) Dorsal habitus; 2) Ventral habitus; 3) Prosternal process, left lateral aspect, arrow points at medial prosternal prominences; 4) Male median lobe, ventral aspect; 5) Male median lobe, right lateral aspect; 6) Male right lateral lobe, right lateral aspect. Scale bar = 1 mm for Figs. 1–3 only." figureDoi="http://doi.org/10.5281/zenodo.7311793" httpUri="https://zenodo.org/record/7311793/files/figure.png" pageId="1" pageNumber="106">Fig. 6</figureCitation>
|
||
). Based on morphology alone, relationships of
|
||
<taxonomicName id="65E54D44E205FF82FC74532FC37CFADB" box="[917,1083,1337,1358]" class="Insecta" family="Dytiscidae" genus="Psychopomporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9091EAD5E205FF82FC74532FC37CFADB" box="[917,1083,1337,1358]" italics="true" pageId="1" pageNumber="106">Psychopomporus</emphasis>
|
||
</taxonomicName>
|
||
to other
|
||
<taxonomicName id="65E54D44E205FF82FD4C5345C467FAFD" box="[685,800,1363,1384]" pageId="1" pageNumber="106" rank="tribe" tribe="Hydroporini">Hydroporini</taxonomicName>
|
||
(and relationships among
|
||
<taxonomicName id="65E54D44E205FF82FBC05345C3D3FAFD" box="[1057,1172,1363,1384]" pageId="1" pageNumber="106" rank="tribe" tribe="Hydroporini">Hydroporini</taxonomicName>
|
||
taxa in general) are not clear and a comprehensive analysis of this group using morphology and molecular data are needed for placing this unusual taxon. Such an analysis is currently under way with preliminary results suggesting that
|
||
<taxonomicName id="65E54D44E205FF82FC3653CEC33FFA78" box="[983,1144,1496,1517]" class="Insecta" family="Dytiscidae" genus="Psychopomporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9091EAD5E205FF82FC3653CEC33FFA78" box="[983,1144,1496,1517]" italics="true" pageId="1" pageNumber="106">Psychopomporus</emphasis>
|
||
</taxonomicName>
|
||
is, indeed, a valid genus (K. Miller, personal observation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |