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<document id="87607F7D01A5C20991343B2A05EF4A1E" ID-CLB-Dataset="31787" ID-DOI="10.11646/zootaxa.4337.4.3" ID-GBIF-Dataset="c9973f7f-1bbb-4f66-95b0-77913ab30c09" ID-ISSN="1175-5326" ID-Zenodo-Dep="1034252" ID-ZooBank="2D4807A5-EAF1-4B93-8833-B3C71BC53A9D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1508523566360" checkinUser="plazi" docAuthor="Gupta, Devanshu &amp; Chandra, Kailash" docDate="2017" docId="03D987F3ED49FFB3ADB7F9E6FC05F83B" docLanguage="en" docName="zootaxa.4337.4.3.pdf" docOrigin="Zootaxa 4337 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tropiorhynchus umbrinus Machatschke 1954" docType="treatment" docVersion="7" lastPageNumber="514" masterDocId="FFE0FF8BED4CFFB6AD20FFC2FF8BFFB3" masterDocTitle="Tropiorhynchus annandalei (Coleoptera: Scarabaeidae: Rutelinae), a new species from Maharashtra, India" masterLastPageNumber="522" masterPageNumber="509" pageNumber="514" updateTime="1698469998896" updateUser="ExternalLinkService">
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<mods:title id="F2E851ED1F168F1B1459B6F1356FFE01">Tropiorhynchus annandalei (Coleoptera: Scarabaeidae: Rutelinae), a new species from Maharashtra, India</mods:title>
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<mods:namePart id="247AD22D9FEACE21C75FFE7D73A5FA64">Gupta, Devanshu</mods:namePart>
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<mods:date id="0A4E07B9B0901C153690155787D436AD">2017</mods:date>
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<treatment id="03D987F3ED49FFB3ADB7F9E6FC05F83B" ID-DOI="http://doi.org/10.5281/zenodo.6042703" ID-GBIF-Taxon="135452496" ID-Zenodo-Dep="6042703" LSID="urn:lsid:plazi:treatment:03D987F3ED49FFB3ADB7F9E6FC05F83B" httpUri="http://treatment.plazi.org/id/03D987F3ED49FFB3ADB7F9E6FC05F83B" lastPageNumber="514" pageId="5" pageNumber="514">
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<paragraph id="8BCF36E5ED49FFB3ADB7F9E6FD47F98D" blockId="5.[151,716,1572,1631]" box="[151,716,1572,1598]" pageId="5" pageNumber="514">
<heading id="D0878189ED49FFB3ADB7F9E6FD47F98D" bold="true" box="[151,716,1572,1598]" fontSize="11" level="1" pageId="5" pageNumber="514" reason="1">
<taxonomicName id="4C704D66ED49FFB3ADB7F9E6FD47F98D" authority="Machatschke, 1954" authorityName="Machatschke" authorityYear="1954" box="[151,716,1572,1598]" class="Insecta" family="Rutelidae" genus="Tropiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="514" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B904EAF7ED49FFB3ADB7F9E6FD47F98D" bold="true" box="[151,716,1572,1598]" pageId="5" pageNumber="514">
<emphasis id="B904EAF7ED49FFB3ADB7F9E6FE5EF98D" bold="true" box="[151,469,1572,1598]" italics="true" pageId="5" pageNumber="514">Tropiorhynchus umbrinus</emphasis>
<bibRefCitation id="EFE14B14ED49FFB3ACFDF9E6FD47F98D" author="Machatschke" box="[477,716,1572,1598]" pageId="5" pageNumber="514" refString="Machatschke, J. W. (1954) Ein neuer Tropiorhynchus, gleichzeitig Revision der Gattung. Beitragezur Entomologie, 4, 57 - 70." type="journal article" year="1954">Machatschke, 1954</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCF36E5ED49FFB3ADB7F984FD23F9EC" blockId="5.[151,716,1572,1631]" box="[151,680,1606,1631]" pageId="5" pageNumber="514">
(
<figureCitation id="134B2A60ED49FFB3ADBFF984FF66F9EC" box="[159,237,1606,1631]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1909,1932]" captionTargetBox="[163,1436,460,1878]" captionTargetId="figure@2.[151,1436,445,1889]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus, male dorsal view. 1, Tropiorhynchus orientis; 2, T. podagricus; 3, T. umbrinus; 4, T. annandalei (holotype)." httpUri="https://zenodo.org/record/1034254/files/figure.png" pageId="5" pageNumber="514">Figs. 3</figureCitation>
,
<figureCitation id="134B2A60ED49FFB3ADDAF984FE81F9EC" box="[250,266,1606,1631]" captionStart="FIGURE 5" captionStartId="3.[151,250,1316,1338]" captionTargetBox="[157,1436,155,1294]" captionTargetId="figure@3.[151,1436,147,1294]" captionTargetPageId="3" captionText="FIGURE 5. Map of India and adjacent countries showing distribution of the species belonging to the genus Tropiorhynchus." httpUri="https://zenodo.org/record/1034256/files/figure.png" pageId="5" pageNumber="514">5</figureCitation>
,
<figureCitation id="134B2A60ED49FFB3AC36F985FEAEF9EC" box="[278,293,1607,1631]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1573,1596]" captionTargetBox="[166,1424,611,1552]" captionTargetId="figure@4.[151,1436,591,1552]" captionTargetPageId="4" captionText="FIGURES 6 9. Head. 6, Tropiorhynchus orientis; 7, T. podagricus; 8, T. umbrinus; 9, T. annandalei (holotype)." httpUri="https://zenodo.org/record/1034258/files/figure.png" pageId="5" pageNumber="514">8</figureCitation>
,
<figureCitation id="134B2A60ED49FFB3AC12F985FEDBF9EC" box="[306,336,1607,1631]" captionStart="FIGURES 10 13" captionStartId="5.[151,264,1474,1497]" captionTargetBox="[152,1430,375,1452]" captionTargetId="figure@5.[151,1436,375,1452]" captionTargetPageId="5" captionText="FIGURES 10 13. Pronotum. 10, Tropiorhynchus orientis; 11, T. podagricus; 12, T. umbrinus; 13, T. annandalei (holotype)." httpUri="https://zenodo.org/record/1034260/files/figure.png" pageId="5" pageNumber="514">12</figureCitation>
,
<figureCitation id="134B2A60ED49FFB3AC7DF984FEF0F9EC" box="[349,379,1606,1631]" captionStart="FIGURES 14 17" captionStartId="6.[151,264,1832,1855]" captionTargetBox="[152,1433,227,1811]" captionTargetId="figure@6.[151,1436,227,1811]" captionTargetPageId="6" captionText="FIGURES 14 17. Elytra. 14, Tropiorhynchus orientis; 15, T. podagricus; 16, T. umbrinus; 17, T. annandalei (holotype)." httpUri="https://zenodo.org/record/1034262/files/figure.png" pageId="5" pageNumber="514">16</figureCitation>
,
<figureCitation id="134B2A60ED49FFB3ACA6F985FE2FF9EC" box="[390,420,1607,1631]" captionStart="FIGURES 18 21" captionStartId="7.[151,264,1156,1179]" captionTargetBox="[151,1432,193,1136]" captionTargetId="figure@7.[151,1436,193,1136]" captionTargetPageId="7" captionText="FIGURES 18 21. Pygidium. 18, Tropiorhynchus orientis; 19, T. podagricus; 20, T. umbrinus; 21, T. annandalei (holotype)." httpUri="https://zenodo.org/record/1034264/files/figure.png" pageId="5" pageNumber="514">20</figureCitation>
,
<figureCitation id="134B2A60ED49FFB3AC90F985FE72F9EC" box="[432,505,1607,1631]" captionStart="FIGURES 28 30" captionStartId="10.[151,264,1816,1839]" captionTargetBox="[151,1436,195,1734]" captionTargetId="figure@10.[151,1436,195,1734]" captionTargetPageId="10" captionText="FIGURES 28 30. Tropiorhynchus umbrinus. 28, profemur; 29, mesofemur; 30, metafemur." httpUri="https://zenodo.org/record/1034266/files/figure.png" pageId="5" pageNumber="514">2830</figureCitation>
, 39, 43, 47, 51)
</paragraph>
</subSubSection>
<subSubSection id="C36A656EED49FFB3ADB7F94CFC7BF8B3" pageId="5" pageNumber="514" type="reference_group">
<paragraph id="8BCF36E5ED49FFB3ADB7F94CFB9FF951" blockId="5.[151,1436,1678,1823]" pageId="5" pageNumber="514">
<treatmentCitationGroup id="AB6011CBED49FFB3ADB7F94CFB9FF951" infered="true" pageId="5" pageNumber="514">
<taxonomicName id="4C704D66ED49FFB3ADB7F94CFD18F917" authority="Machatschke, 1954: 62" authorityName="Machatschke" authorityPageNumber="62" authorityYear="1954" box="[151,659,1678,1700]" class="Insecta" family="Rutelidae" genus="Tropiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="514" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B904EAF7ED49FFB3ADB7F94CFE16F917" box="[151,413,1678,1700]" italics="true" pageId="5" pageNumber="514">Tropiorhynchus umbrinus</emphasis>
<treatmentCitation id="0AD110F4ED49FFB3AC83F94CFD18F917" author="Machatschke" box="[419,659,1678,1700]" page="62" pageId="5" pageNumber="514" year="1954">
<bibRefCitation id="EFE14B14ED49FFB3AC83F94CFDE7F917" author="Machatschke" box="[419,620,1678,1700]" pageId="5" pageNumber="514" refString="Machatschke, J. W. (1954) Ein neuer Tropiorhynchus, gleichzeitig Revision der Gattung. Beitragezur Entomologie, 4, 57 - 70." type="journal article" year="1954">Machatschke, 1954</bibRefCitation>
: 62
</treatmentCitation>
</taxonomicName>
, fig. 1b (lateral side of pronotum); fig. 2a (metathoracic leg), 4a b, (frontal view paramere, lateral view aedeagus), fig. 5 (endophallus);
<treatmentCitation id="0AD110F4ED49FFB3AE0FF96FFBAFF970" author="Machatschke" box="[815,1060,1709,1731]" page="180" pageId="5" pageNumber="514" year="1957">
<bibRefCitation id="EFE14B14ED49FFB3AE0FF96FFC79F970" author="Machatschke" box="[815,1010,1709,1731]" pageId="5" pageNumber="514" refString="Machatschke, J. W. (1957) Rutelinae II. Genera Insectorum, 199 b, 1 - 219." type="journal article" year="1957">Machatschke 1957</bibRefCitation>
: 180
</treatmentCitation>
, plate 6, fig. 4a (habitus dorsal view, male), 4b (habitus ventral view, male), 4c (protarsi, male);
<treatmentCitation id="0AD110F4ED49FFB3AE3BF90EFB9BF951" author="Machatschke" box="[795,1040,1740,1762]" page="254" pageId="5" pageNumber="514" year="1972">
<bibRefCitation id="EFE14B14ED49FFB3AE3BF90EFC55F951" author="Machatschke" box="[795,990,1740,1762]" pageId="5" pageNumber="514" refString="Machatschke, J. W. (1972) Superfamilie Scarabaeoidae, Familie Melolonthidae, Subfamilie Rutelinae. Coleopterorum catalogus, 66 (1), 1 - 361." type="journal article" year="1972">Machatschke 1972</bibRefCitation>
: 254
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCF36E5ED49FFB3ADB7F928FC7BF8B3" blockId="5.[151,1436,1678,1823]" box="[151,1008,1770,1792]" pageId="5" pageNumber="514">
<treatmentCitationGroup id="AB6011CBED49FFB3ADB7F928FC7BF8B3" box="[151,1008,1770,1792]" pageId="5" pageNumber="514">
<taxonomicName id="4C704D66ED49FFB3ADB7F928FE17F8B3" authorityName="Machatschke" authorityYear="1954" box="[151,412,1770,1792]" class="Insecta" family="Rutelidae" genus="Tropiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="514" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B904EAF7ED49FFB3ADB7F928FE17F8B3" box="[151,412,1770,1792]" italics="true" pageId="5" pageNumber="514">Tropiorhynchus umbrinus</emphasis>
</taxonomicName>
form
<taxonomicName id="4C704D66ED49FFB3ACFAF929FDA5F8B3" box="[474,558,1771,1792]" class="Insecta" family="Rutelidae" genus="Tropiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="514" phylum="Arthropoda" rank="species" species="cruciata">
<emphasis id="B904EAF7ED49FFB3ACFAF929FDA5F8B3" box="[474,558,1771,1792]" italics="true" pageId="5" pageNumber="514">cruciata</emphasis>
</taxonomicName>
(unavailable name):
<treatmentCitation id="0AD110F4ED49FFB3AE24F928FC67F8B3" author="Machatschke" box="[772,1004,1770,1792]" page="62" pageId="5" pageNumber="514" year="1954">
<bibRefCitation id="EFE14B14ED49FFB3AE24F928FC4CF8B3" author="Machatschke" box="[772,967,1770,1792]" pageId="5" pageNumber="514" refString="Machatschke, J. W. (1954) Ein neuer Tropiorhynchus, gleichzeitig Revision der Gattung. Beitragezur Entomologie, 4, 57 - 70." type="journal article" year="1954">Machatschke 1954</bibRefCitation>
: 62
</treatmentCitation>
.
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</paragraph>
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<treatmentCitationGroup id="AB6011CBED49FFB3ADB7F8CBFC44F8AC" box="[151,975,1801,1823]" pageId="5" pageNumber="514">
<taxonomicName id="4C704D66ED49FFB3ADB7F8CBFE17F8AC" authorityName="Machatschke" authorityYear="1954" box="[151,412,1801,1823]" class="Insecta" family="Rutelidae" genus="Tropiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="514" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B904EAF7ED49FFB3ADB7F8CBFE17F8AC" box="[151,412,1801,1823]" italics="true" pageId="5" pageNumber="514">Tropiorhynchus umbrinus</emphasis>
</taxonomicName>
form
<taxonomicName id="4C704D66ED49FFB3ACFAF8CBFD86F8AC" box="[474,525,1801,1823]" class="Insecta" family="Rutelidae" genus="Tropiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="514" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="B904EAF7ED49FFB3ACFAF8CBFD86F8AC" box="[474,525,1801,1823]" italics="true" pageId="5" pageNumber="514">flava</emphasis>
</taxonomicName>
(unavailable name):
<treatmentCitation id="0AD110F4ED49FFB3AFC2F8CBFC41F8AC" author="Machatschke" box="[738,970,1801,1823]" page="62" pageId="5" pageNumber="514" year="1954">
<bibRefCitation id="EFE14B14ED49FFB3AFC2F8CBFC2FF8AC" author="Machatschke" box="[738,932,1801,1823]" pageId="5" pageNumber="514" refString="Machatschke, J. W. (1954) Ein neuer Tropiorhynchus, gleichzeitig Revision der Gattung. Beitragezur Entomologie, 4, 57 - 70." type="journal article" year="1954">Machatschke 1954</bibRefCitation>
: 62
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36A656EED49FFB3ADB7F889FC05F83B" pageId="5" pageNumber="514" type="materials_examined">
<paragraph id="8BCF36E5ED49FFB3ADB7F889FC05F83B" blockId="5.[151,1296,1867,1928]" pageId="5" pageNumber="514">
<materialsCitation id="3B183CB8ED49FFB3ADB7F889FD17F8D7" ID-GBIF-Occurrence="1675496976" box="[151,668,1867,1892]" country="India" location="Maharashtra" pageId="5" pageNumber="514" specimenCount="1" stateProvince="Maharashtra">
<emphasis id="B904EAF7ED49FFB3ADB7F889FEF2F8D7" bold="true" box="[151,377,1867,1892]" pageId="5" pageNumber="514">Material examined</emphasis>
.
<collectingCountry id="F3677675ED49FFB3ACA7F889FE48F8D7" box="[391,451,1867,1892]" name="India" pageId="5" pageNumber="514">India</collectingCountry>
,
<collectingRegion id="49B4F807ED49FFB3ACEFF889FDD0F8D7" box="[463,603,1867,1892]" country="India" name="Maharashtra" pageId="5" pageNumber="514">Maharashtra</collectingRegion>
State
</materialsCitation>
,
<materialsCitation id="3B183CB8ED49FFB3AF87F889FE3CF83B" ID-GBIF-Occurrence="1675496975" collectionCode="ZSCI" country="India" location="Bombay" pageId="5" pageNumber="514" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F3677675ED49FFB3AF87F889FC8CF8D7" box="[679,775,1867,1892]" name="India" pageId="5" pageNumber="514">Bombay</collectingCountry>
,
<specimenCount id="9D76FD6CED49FFB3AE32F88EFCEBF8D7" box="[786,864,1867,1892]" pageId="5" pageNumber="514" type="male">1 male</specimenCount>
, (
<collectionCode id="ED61AE20ED49FFB3AE53F88EFC3AF8D7" box="[883,945,1868,1892]" pageId="5" pageNumber="514">ZSCI</collectionCode>
registration number: 2416/16).
<emphasis id="B904EAF7ED49FFB3ADE7F8ADFED2F83B" bold="true" box="[199,345,1903,1928]" pageId="5" pageNumber="514">Distribution</emphasis>
(
<figureCitation id="134B2A60ED49FFB3AC48F8ADFE26F83B" box="[360,429,1903,1928]" captionStart="FIGURE 5" captionStartId="3.[151,250,1316,1338]" captionTargetBox="[157,1436,155,1294]" captionTargetId="figure@3.[151,1436,147,1294]" captionTargetPageId="3" captionText="FIGURE 5. Map of India and adjacent countries showing distribution of the species belonging to the genus Tropiorhynchus." httpUri="https://zenodo.org/record/1034256/files/figure.png" pageId="5" pageNumber="514">Fig. 5</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3B183CB8ED49FFB3ACE2F8ADFC05F83B" ID-GBIF-Occurrence="1675496970" box="[450,910,1903,1928]" country="India" location="Maharashtra" pageId="5" pageNumber="514" specimenCount="1" stateProvince="Maharashtra">
<collectingCountry id="F3677675ED49FFB3ACE2F8ADFD8BF83B" box="[450,512,1903,1928]" name="India" pageId="5" pageNumber="514">India</collectingCountry>
:
<collectingRegion id="49B4F807ED49FFB3AF2AF8ADFD1DF83B" box="[522,662,1903,1928]" country="India" name="Maharashtra" pageId="5" pageNumber="514">Maharashtra</collectingRegion>
(Bombay, Khandala).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
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