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<document id="739173A4214B388CD40DA04302D5CCA2" ID-DOI="10.5281/zenodo.184596" ID-GBIF-Dataset="690a17b6-8d89-439d-b24c-ba3efc6ee87d" ID-ISSN="1175-5326" ID-Zenodo-Dep="184596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459989687148" checkinUser="plazi" docAuthor="Anker, Arthur &amp; Tóth, Eva" docDate="2008" docId="03D987B85855FFA1FF1AFEAEFC38FDF2" docLanguage="en" docName="zt01915p028.pdf" docOrigin="Zootaxa 1915" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Synalpheus bocas Anker &amp; Tóth, 2008, n. sp." docType="treatment" docVersion="10" lastPageNumber="18" masterDocId="FFE0FFC05845FFB0FF8DFFD3FFCCFFB0" masterDocTitle="A preliminary revision of the Synalpheus paraneptunus Coutière, 1909 species complex (Crustacea: Decapoda: Alpheidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="17" updateTime="1698232578265" updateUser="plazi">
<mods:mods id="891F57EC998321BA54AEFA3AD5BE8B53" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="98DDB1E746E1702DB24CC3B7943E80D5">
<mods:title id="78AC068A8F7AA1E67A6EFC65F0DF02FC">A preliminary revision of the Synalpheus paraneptunus Coutière, 1909 species complex (Crustacea: Decapoda: Alpheidae)</mods:title>
</mods:titleInfo>
<mods:name id="028FB1817F6BC08A4C8FD7FE20C05BA0" type="personal">
<mods:role id="D3E80CE22B56621616AF11F4E75950E3">
<mods:roleTerm id="B03457B6B31A2CBE7565000EF6848837">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="62889CA460BDDCA22E5F286BB364CADC">Anker, Arthur</mods:namePart>
</mods:name>
<mods:name id="4A84F25532993C36E1937B9637E988DD" type="personal">
<mods:role id="FA834F601B53FA889A97F7ACD7F2D5CC">
<mods:roleTerm id="6E28D3D1652FCCB8E0DD65BF887380C9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="69AADA9DF1B5D25142C916DF4609D782">Tóth, Eva</mods:namePart>
</mods:name>
<mods:typeOfResource id="83625D9881D5C65B1CE20289FFDF1A42">text</mods:typeOfResource>
<mods:relatedItem id="329536B4CDE821528B8C6C679ADDA545" type="host">
<mods:titleInfo id="FF7A91AFF842F2590B134397587559D5">
<mods:title id="99E0716BD03CE05746F5C1DD4B95DFFA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2D0F84936EE175D1FEB16D8F8D88EFAF">
<mods:date id="C0BEA55AF1A394B2B388850893CBEEF3">2008</mods:date>
<mods:detail id="D82F485F7AC9B15F07F006F65F2DD821" type="volume">
<mods:number id="C9DFCF142A7D3EAA56BCFEDD3BB78A63">1915</mods:number>
</mods:detail>
<mods:extent id="7B492A3176A186A40D1E799070275C3E" unit="page">
<mods:start id="45AD790B22D9064FC01389858B17066F">1</mods:start>
<mods:end id="C9EF5ADA1D7644ACCAE60F45F302C3A9">28</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="62D97746B12407B581C48E5FFDC7CFED">journal article</mods:classification>
<mods:identifier id="B6755BCDCE8DBC36A8A7338F1B674880" type="DOI">10.5281/zenodo.184596</mods:identifier>
<mods:identifier id="A03E2BCFAC55DF91EBFE55158F0DCBC1" type="GBIF-Dataset">690a17b6-8d89-439d-b24c-ba3efc6ee87d</mods:identifier>
<mods:identifier id="4F58C9549F203CE78DE02264C8A51E68" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FD180CE9D683D93857B8ECA1B09889DE" type="Zenodo-Dep">184596</mods:identifier>
</mods:mods>
<treatment id="03D987B85855FFA1FF1AFEAEFC38FDF2" ID-DOI="http://doi.org/10.5281/zenodo.5694279" ID-GBIF-Taxon="119379847" ID-Zenodo-Dep="5694279" LSID="urn:lsid:plazi:treatment:03D987B85855FFA1FF1AFEAEFC38FDF2" httpUri="http://treatment.plazi.org/id/03D987B85855FFA1FF1AFEAEFC38FDF2" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="C36A65255855FFA0FF1AFEAEFE50FE0A" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BCF36AE5855FFA0FF1AFEAEFE74FE27" blockId="16.[151,440,381,442]" box="[151,440,381,407]" pageId="16" pageNumber="17">
<heading id="D08781C25855FFA0FF1AFEAEFE74FE27" bold="true" box="[151,440,381,407]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B904EABC5855FFA0FF1AFEAEFE74FE27" bold="true" box="[151,440,381,407]" pageId="16" pageNumber="17">
<taxonomicName id="4C704D2D5855FFA0FF1AFEAEFEA2FE27" ID-CoL="85YVF" box="[151,366,381,407]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">
<emphasis id="B904EABC5855FFA0FF1AFEAEFEA2FE27" bold="true" box="[151,366,381,407]" italics="true" pageId="16" pageNumber="17">Synalpheus bocas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23757C75855FFA0FEFBFEAEFE74FE27" box="[374,440,381,407]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36AE5855FFA0FF13FE73FE50FE0A" blockId="16.[151,440,381,442]" box="[158,412,416,442]" pageId="16" pageNumber="17">
(
<figureCitation id="134B2A2B5855FFA0FF2AFE73FEC5FE0A" box="[167,265,416,442]" captionStart="FIGURE 10" captionStartId="17.[151,255,1907,1931]" captionTargetBox="[178,1406,627,1877]" captionTargetId="figure@17.[169,1413,616,1902]" captionTargetPageId="17" captionText="FIGURE 10. Synalpheus bocas n. sp., paratype, male from Bocas del Toro, Panama (USNM 1116692): A, frontal region, dorsal view; B, same, lateral view; C, third maxilliped, tip of ultimate segment, lateral view; D, second pereiopod, lateral view; E, third pereiopod, lateral view; F, abdomen, lateral view; G, telson and uropods, dorsal view. Scale bar: 0.5 mm (A – E, G), 1 mm (F)." httpUri="https://zenodo.org/record/184606/files/figure.png" pageId="16" pageNumber="17">Figs. 10</figureCitation>
,
<figureCitation id="134B2A2B5855FFA0FE98FE73FEFFFE0A" box="[277,307,416,442]" captionStart="FIGURE 11" captionStartId="18.[151,255,1826,1850]" captionTargetBox="[167,1420,478,1792]" captionTargetId="figure@18.[151,1436,461,1802]" captionTargetPageId="18" captionText="FIGURE 11. Synalpheus bocas n. sp., paratype, male from Bocas del Toro, Panama (USNM 1116692) [A – E]; paratype, male (USNM 1116690) [F]: A, major cheliped, lateral view; B, same, chela, mesial view; C, same, distal portion with opened fingers, lateral view; D, minor cheliped, lateral view; E, same, chela with opened fingers, mesial view, F, minor chela of young male with opened fingers, dorsolateral aspect. Scale bar: 0.5 mm (C – F), 1 mm (A, B)." httpUri="https://zenodo.org/record/184607/files/figure.png" pageId="16" pageNumber="17">11</figureCitation>
,
<figureCitation id="134B2A2B5855FFA0FECDFE73FE90FE0A" box="[320,348,416,442]" captionStart="FIGURE 14" captionStartId="21.[151,255,1824,1848]" captionTargetBox="[152,1434,195,1798]" captionTargetId="figure@21.[151,1436,194,1800]" captionTargetPageId="21" captionText="FIGURE 14. Life habitus and colour patterns. Synalpheus duffyi n. sp., Isla Grande, Panama [A, B]: A, larger queen and two smaller non-breeding colony members from the same colony, dorsal view; B, queen with embryos, lateral view. Synalpheus brevidactylus n. sp., Isla Grande, Panama [C, D]: C, breeding female and male (mated pair), dorsal view; D, breeding female, lateral view. Synalpheus bocas n. sp., Bocas del Toro, Panama [E – H]: E, male, dorsal view; F, breeding female; G, H, another breeding female, lateral and dorsal views on different backgrounds. Synalpheus belizensis n. sp., Carrie Bow Cay, Belize [I]: I, breeding female, dorsal view. Scale bars: ~ 2 mm." httpUri="https://zenodo.org/record/184610/files/figure.png" pageId="16" pageNumber="17">14</figureCitation>
E–H)
</paragraph>
</subSubSection>
<subSubSection id="C36A65255855FFA0FF1AFE3CFE56FC2A" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BCF36AE5855FFA0FF1AFE3CFE56FC2A" blockId="16.[151,1437,495,2002]" pageId="16" pageNumber="17">
<emphasis id="B904EABC5855FFA0FF1AFE3CFE89FDB9" bold="true" box="[151,325,495,521]" pageId="16" pageNumber="17">
<typeStatus id="54CB880C5855FFA0FF1AFE3CFF18FDB9" box="[151,212,495,521]" pageId="16" pageNumber="17">Type</typeStatus>
material
</emphasis>
.
<collectingCountry id="F367763E5855FFA0FEDEFE23FE7FFDBA" box="[339,435,496,522]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
, Caribbean coast.
<typeStatus id="54CB880C5855FFA0FD0AFE23FD3BFDBA" box="[647,759,496,522]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
, breeding female,
<collectionCode id="ED61AE6B5855FFA0FC40FE23FBEEFDBA" LSID="urn:lsid:biocol.org:col:34871" box="[973,1058,496,522]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
1116687,
<taxonomicName id="4C704D2D5855FFA0FB16FE23FB2FFDBA" box="[1179,1251,496,522]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">Bocas</taxonomicName>
del Boro,
<collectionCode id="ED61AE6B5855FFA0FAD1FE23FA50FDBA" box="[1372,1436,496,522]" pageId="16" pageNumber="17">STRI</collectionCode>
Bay, in sponge
<taxonomicName id="4C704D2D5855FFA0FEDBFDCBFD4CFD81" box="[342,640,536,561]" class="Demospongiae" family="Petrosiidae" genus="Xestospongia" kingdom="Animalia" order="Haplosclerida" pageId="16" pageNumber="17" phylum="Porifera" rank="species" species="rosariensis">
<emphasis id="B904EABC5855FFA0FEDBFDCBFD4CFD81" box="[342,640,536,561]" italics="true" pageId="16" pageNumber="17">Xestospongia rosariensis</emphasis>
</taxonomicName>
,
<quantity id="4C889B4B5855FFA0FD1FFDCBFD3DFD82" box="[658,753,536,562]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="16" pageNumber="17" unit="m" value="10.0" valueMax="15.0" valueMin="5.0">5–15 m</quantity>
, coll. E. Tóth,
<date id="FFCE106E5855FFA0FC21FDCBFBF4FD82" box="[940,1080,536,562]" pageId="16" pageNumber="17" value="2003-09-03">3 Sep 2003</date>
[03-033a].
<typeStatus id="54CB880C5855FFA0FB45FDCBFAF3FD82" box="[1224,1343,536,562]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
: 2 nonbreeding specimens
<collectionCode id="ED61AE6B5855FFA0FE01FD93FE2DFDEA" LSID="urn:lsid:biocol.org:col:34871" box="[396,481,576,602]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
1116688, same collection data as for
<typeStatus id="54CB880C5855FFA0FC23FD93FBD9FDEA" box="[942,1045,576,602]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
[03-032]; 1 non-breeding specimen,
<collectionCode id="ED61AE6B5855FFA0FF55FDBBFEE1FD32" LSID="urn:lsid:biocol.org:col:34871" box="[216,301,616,642]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
1116689, same collection data as for
<typeStatus id="54CB880C5855FFA0FD65FDBBFC82FD32" box="[744,846,616,642]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
[03-033b];
<specimenCount id="9D76FD275855FFA0FC56FDBBFBE1FD32" box="[987,1069,616,642]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
,
<collectionCode id="ED61AE6B5855FFA0FBB4FDBBFB42FD32" LSID="urn:lsid:biocol.org:col:34871" box="[1081,1166,616,642]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
1116690, same collection data as for
<typeStatus id="54CB880C5855FFA0FEDDFD43FE7BFD1A" box="[336,439,656,682]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
[03-039];
<specimenCount id="9D76FD275855FFA0FDB8FD43FD58FD1A" box="[565,660,656,682]" pageId="16" pageNumber="17" type="male">2 males</specimenCount>
, 2 breeding females,
<collectionCode id="ED61AE6B5855FFA0FC1DFD43FC29FD1A" LSID="urn:lsid:biocol.org:col:34871" box="[912,997,656,682]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
1116691, same collection data as for
<typeStatus id="54CB880C5855FFA0FF1AFD6BFF32FD62" box="[151,254,696,722]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
[03-040];
<specimenCount id="9D76FD275855FFA0FEF3FD6BFE1EFD62" box="[382,466,696,722]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
, 1 breeding female, UP, same collection data as for
<typeStatus id="54CB880C5855FFA0FBCEFD6BFB66FD62" box="[1091,1194,696,722]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
[03-041]; 1 breding female,
<specimenCount id="9D76FD275855FFA0FF78FD33FE8BFD4A" box="[245,327,736,762]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
,
<collectionCode id="ED61AE6B5855FFA0FEDEFD33FE64FD4A" LSID="urn:lsid:biocol.org:col:34871" box="[339,424,736,762]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
1116692,
<taxonomicName id="4C704D2D5855FFA0FDACFD33FDA5FD4A" box="[545,617,736,762]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">Bocas</taxonomicName>
del Toro,
<collectionCode id="ED61AE6B5855FFA0FD6CFD33FCECFD4A" box="[737,800,736,762]" pageId="16" pageNumber="17">STRI</collectionCode>
Bay,
<quantity id="4C889B4B5855FFA0FCEEFD33FC7CFD4A" box="[867,944,736,762]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="16" pageNumber="17" unit="m" value="8.5" valueMax="9.0" valueMin="8.0">8–9 m</quantity>
, in sponge
<taxonomicName id="4C704D2D5855FFA0FBB9FD33FA96FD49" box="[1076,1370,736,761]" class="Demospongiae" family="Petrosiidae" genus="Xestospongia" kingdom="Animalia" order="Haplosclerida" pageId="16" pageNumber="17" phylum="Porifera" rank="species" species="rosariensis">
<emphasis id="B904EABC5855FFA0FBB9FD33FA96FD49" box="[1076,1370,736,761]" italics="true" pageId="16" pageNumber="17">Xestospongia rosariensis</emphasis>
</taxonomicName>
, coll. E. Tóth,
<date id="FFCE106E5855FFA0FE8DFCDBFE5DFC92" box="[256,401,776,802]" pageId="16" pageNumber="17" value="2006-01-15">15 Jan 2006</date>
[06-714, one specimen dissected]; 1 breeding female,
<collectionCode id="ED61AE6B5855FFA0FB90FCDBFBBEFC92" LSID="urn:lsid:biocol.org:col:34871" box="[1053,1138,776,802]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
116693,
<taxonomicName id="4C704D2D5855FFA0FB6CFCDBFAE4FC92" box="[1249,1320,776,802]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">Bocas</taxonomicName>
del Toro,
<collectingCountry id="F367763E5855FFA0FF1AFCE3FF0DFCFA" box="[151,193,816,842]" name="Iceland" pageId="16" pageNumber="17">Isla</collectingCountry>
Colón, Punta Caracol,
<quantity id="4C889B4B5855FFA0FE5BFCE3FDE8FCFA" box="[470,548,816,842]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="16" pageNumber="17" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">1–3 m</quantity>
, in sponges, coll. A. Anker, I. Marin and A. Baeza,
<date id="FFCE106E5855FFA0FB01FCE3FAEEFCFA" box="[1164,1314,816,842]" pageId="16" pageNumber="17" value="2007-04-28">28 Apr 2007</date>
[07-144b, photo voucher]; 1 non-breeding specimen,
<collectionCode id="ED61AE6B5855FFA0FD1AFC8BFD20FCC2" LSID="urn:lsid:biocol.org:col:34871" box="[663,748,856,882]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
1116694, same collection data as previous specimen [07- 141b, photo voucher].
</paragraph>
</subSubSection>
<subSubSection id="C36A65255855FFA0FF48FC74FB8CFA7A" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BCF36AE5855FFA0FF48FC74FB8CFA7A" blockId="16.[151,1437,495,2002]" pageId="16" pageNumber="17">
<emphasis id="B904EABC5855FFA0FF48FC74FEF1FC71" bold="true" box="[197,317,935,961]" pageId="16" pageNumber="17">Diagnosis</emphasis>
. Rostrum not lower than orbital hoods, rather broad at base, not particularly slender, subequal to orbital teeth, with subacute tip; orbital teeth triangular, broad at base, subacute; both rostrum and orbital teeth slightly up-turned distally; notches between orbital teeth and rostrum moderately deep, in-between V- and U-shaped. Antennular peduncles with second segment distinctly longer than wide; stylocerite overreaching distal margin of first segment. Antenna with basicerite bearing blunt, almost non-projecting distodorsal tooth; scaphocerite without blade. Third maxilliped with crown of eight-nine spine-like setae on tip of ultimate segment. Major chela with palm about 2.5 times as long as fingers; distodorsal tooth on palm not bulging, with anteriorly directed or slightly descendant acute point. Second pereiopod carpus with five segments, first carpal segment about four times length of second. Third pereiopod with propodus bearing eight spine-like setae (not including distal pair); dactylus stout. Uropodal exopod with two fixed teeth (including lateral tooth of diaeresis) and slender movable spine-like seta. Telson with longitudinal median depression and two pairs of very stout spine-like setae inserted anterior and posterior to mid-length, respectively; posterior margin broad, about half-length of anterior width, rounded (see
<figureCitation id="134B2A2B5855FFA0FD50FA5BFCF3FA12" box="[733,831,1416,1442]" captionStart="FIGURE 10" captionStartId="17.[151,255,1907,1931]" captionTargetBox="[178,1406,627,1877]" captionTargetId="figure@17.[169,1413,616,1902]" captionTargetPageId="17" captionText="FIGURE 10. Synalpheus bocas n. sp., paratype, male from Bocas del Toro, Panama (USNM 1116692): A, frontal region, dorsal view; B, same, lateral view; C, third maxilliped, tip of ultimate segment, lateral view; D, second pereiopod, lateral view; E, third pereiopod, lateral view; F, abdomen, lateral view; G, telson and uropods, dorsal view. Scale bar: 0.5 mm (A – E, G), 1 mm (F)." httpUri="https://zenodo.org/record/184606/files/figure.png" pageId="16" pageNumber="17">Figs. 10</figureCitation>
,
<figureCitation id="134B2A2B5855FFA0FCC1FA5BFCA7FA12" box="[844,875,1416,1442]" captionStart="FIGURE 11" captionStartId="18.[151,255,1826,1850]" captionTargetBox="[167,1420,478,1792]" captionTargetId="figure@18.[151,1436,461,1802]" captionTargetPageId="18" captionText="FIGURE 11. Synalpheus bocas n. sp., paratype, male from Bocas del Toro, Panama (USNM 1116692) [A – E]; paratype, male (USNM 1116690) [F]: A, major cheliped, lateral view; B, same, chela, mesial view; C, same, distal portion with opened fingers, lateral view; D, minor cheliped, lateral view; E, same, chela with opened fingers, mesial view, F, minor chela of young male with opened fingers, dorsolateral aspect. Scale bar: 0.5 mm (C – F), 1 mm (A, B)." httpUri="https://zenodo.org/record/184607/files/figure.png" pageId="16" pageNumber="17">11</figureCitation>
). For detailed description of general features of
<taxonomicName id="4C704D2D5855FFA0FF1AFA63FE99FA79" box="[151,341,1456,1481]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="paraneptunus">
<emphasis id="B904EABC5855FFA0FF1AFA63FE99FA79" box="[151,341,1456,1481]" italics="true" pageId="16" pageNumber="17">S. paraneptunus</emphasis>
</taxonomicName>
species complex see
<bibRefCitation id="EFE14B5F5855FFA0FDDCFA63FCC6FA7A" author="Dardeau" box="[593,778,1456,1482]" pageId="16" pageNumber="28" refString="Dardeau, M. R. (1984) Synalpheus shrimps (Crustacea: Decapoda: Alpheidae). I. The Gambarelloides group, with a description of a new species. Memoirs of the Hourglass Cruises, 7, 1 - 125." type="journal article" year="1984">Dardeau (1984)</bibRefCitation>
and
<bibRefCitation id="EFE14B5F5855FFA0FCCEFA63FBF1FA7A" author="Rios" box="[835,1085,1456,1482]" pageId="16" pageNumber="28" refString="Rios, R. and Duffy, J. E. (2007) A review of the sponge-dwelling snapping shrimp from Carrie Bow Cay, Belize, with description of Zuzalpheus, new genus, and six new species (Crustacea: Decapoda: Alpheidae). Zootaxa, 1602, 1 - 82." type="journal article" year="2007">Ríos &amp; Duffy (2007)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A65255855FFA0FF48FA04FAF8F90A" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BCF36AE5855FFA0FF48FA04FBC5F9AA" blockId="16.[151,1437,495,2002]" pageId="16" pageNumber="17">
<emphasis id="B904EABC5855FFA0FF48FA04FE86FA41" bold="true" box="[197,330,1495,1521]" pageId="16" pageNumber="17">Etymology</emphasis>
. The new species’ name was derived from the first word of the
<typeStatus id="54CB880C5855FFA0FBCFFA0BFBB9FA42" box="[1090,1141,1496,1522]" pageId="16" pageNumber="17">type</typeStatus>
locality,
<taxonomicName id="4C704D2D5855FFA0FB6FFA0BFAE6FA42" box="[1250,1322,1496,1522]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">Bocas</taxonomicName>
del Toro,
<collectingCountry id="F367763E5855FFA0FF1AF9D3FF34F9AA" box="[151,248,1536,1562]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
; “
<taxonomicName id="4C704D2D5855FFA0FE80F9D3FE99F9AA" box="[269,341,1536,1562]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bocas">bocas</taxonomicName>
” (mouths in Spanish) was here used as noun in opposition.
</paragraph>
<paragraph id="8BCF36AE5855FFA0FF48F9F4FAF8F90A" blockId="16.[151,1437,495,2002]" pageId="16" pageNumber="17">
<emphasis id="B904EABC5855FFA0FF48F9F4FE4EF9F1" bold="true" box="[197,386,1575,1601]" pageId="16" pageNumber="17">Colour pattern</emphasis>
. Whitish semitransparent, with reddish chromatophores distributed in diffuse bands over the entire body, more on abdomen; major chela pale beige, distal margin of palm and fingers pale brown (except for pale area surrounded by darker margin on dactylus), tips amber-yellow; ovaries or fresh brood green (
<figureCitation id="134B2A2B5855FFA0FF65F973FEF0F90A" box="[232,316,1696,1722]" captionStart="FIGURE 14" captionStartId="21.[151,255,1824,1848]" captionTargetBox="[152,1434,195,1798]" captionTargetId="figure@21.[151,1436,194,1800]" captionTargetPageId="21" captionText="FIGURE 14. Life habitus and colour patterns. Synalpheus duffyi n. sp., Isla Grande, Panama [A, B]: A, larger queen and two smaller non-breeding colony members from the same colony, dorsal view; B, queen with embryos, lateral view. Synalpheus brevidactylus n. sp., Isla Grande, Panama [C, D]: C, breeding female and male (mated pair), dorsal view; D, breeding female, lateral view. Synalpheus bocas n. sp., Bocas del Toro, Panama [E – H]: E, male, dorsal view; F, breeding female; G, H, another breeding female, lateral and dorsal views on different backgrounds. Synalpheus belizensis n. sp., Carrie Bow Cay, Belize [I]: I, breeding female, dorsal view. Scale bars: ~ 2 mm." httpUri="https://zenodo.org/record/184610/files/figure.png" pageId="16" pageNumber="17">Fig. 14</figureCitation>
E–H), exceptionally greenish yellow (one female from
<collectingCountry id="F367763E5855FFA0FC49F973FBA9F90A" box="[964,1125,1696,1722]" name="Saint Kitts and Nevis" pageId="16" pageNumber="17">San Cristobal</collectingCountry>
,
<taxonomicName id="4C704D2D5855FFA0FBFCF973FB75F90A" box="[1137,1209,1696,1722]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">Bocas</taxonomicName>
del Toro).
</paragraph>
</subSubSection>
<subSubSection id="C36A65255855FFA0FF48F914FAB9F952" box="[197,1397,1735,1762]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BCF36AE5855FFA0FF48F914FAB9F952" blockId="16.[151,1437,495,2002]" box="[197,1397,1735,1762]" pageId="16" pageNumber="17">
<emphasis id="B904EABC5855FFA0FF48F914FF3BF951" bold="true" box="[197,247,1735,1761]" pageId="16" pageNumber="17">Size</emphasis>
. – Largest male, CL
<quantity id="4C889B4B5855FFA0FE7DF91BFD81F952" box="[496,589,1736,1762]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="16" pageNumber="17" unit="mm" value="4.9">4.9 mm</quantity>
; largest breeding female, CL
<quantity id="4C889B4B5855FFA0FC2AF91BFBCCF952" box="[935,1024,1736,1762]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="16" pageNumber="17" unit="mm" value="5.8">5.8 mm</quantity>
(
<typeStatus id="54CB880C5855FFA0FB82F91BFBBBF952" box="[1039,1143,1736,1762]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
female, CL
<quantity id="4C889B4B5855FFA0FA87F91BFAA5F952" box="[1290,1385,1736,1762]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="16" pageNumber="17" unit="mm" value="5.7">5.7 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A65255855FFA0FF48F93CFD5AF8BA" box="[197,662,1775,1802]" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BCF36AE5855FFA0FF48F93CFD5AF8BA" blockId="16.[151,1437,495,2002]" box="[197,662,1775,1802]" pageId="16" pageNumber="17">
<emphasis id="B904EABC5855FFA0FF48F93CFEA8F8B9" bold="true" box="[197,356,1775,1801]" pageId="16" pageNumber="17">
<typeStatus id="54CB880C5855FFA0FF48F93CFECEF8B9" box="[197,258,1775,1801]" pageId="16" pageNumber="17">Type</typeStatus>
locality
</emphasis>
.
<taxonomicName id="4C704D2D5855FFA0FEFFF923FE75F8BA" box="[370,441,1776,1802]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">Bocas</taxonomicName>
del Toro,
<collectingCountry id="F367763E5855FFA0FDBFF923FD5FF8BA" box="[562,659,1776,1802]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A65255855FFA1FF48F8C4FBD3FEB2" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BCF36AE5855FFA0FF48F8C4FC15F882" blockId="16.[151,1437,495,2002]" box="[197,985,1815,1842]" pageId="16" pageNumber="17">
<emphasis id="B904EABC5855FFA0FF48F8C4FE92F881" bold="true" box="[197,350,1815,1841]" pageId="16" pageNumber="17">Distribution</emphasis>
. Presently known only from
<taxonomicName id="4C704D2D5855FFA0FD39F8CBFD30F882" box="[692,764,1816,1842]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">Bocas</taxonomicName>
del Toro,
<collectingCountry id="F367763E5855FFA0FCF9F8CBFC18F882" box="[884,980,1816,1842]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8BCF36AE5855FFA1FF48F8ECFBD3FEB2" blockId="16.[151,1437,495,2002]" lastBlockId="17.[151,1436,152,578]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B904EABC5855FFA0FF48F8ECFE97F8E9" bold="true" box="[197,347,1855,1881]" pageId="16" pageNumber="17">Life history</emphasis>
. This presumably pair-living species occurs in several sponges in
<taxonomicName id="4C704D2D5855FFA0FB2CF893FB25F8EA" box="[1185,1257,1856,1882]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">Bocas</taxonomicName>
del Toro. For instance, several sexual pairs of
<taxonomicName id="4C704D2D5855FFA0FD9AF8BBFDB6F831" box="[535,634,1896,1921]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">
<emphasis id="B904EABC5855FFA0FD9AF8BBFDB6F831" box="[535,634,1896,1921]" italics="true" pageId="16" pageNumber="17">S. bocas</emphasis>
</taxonomicName>
<emphasis id="B904EABC5855FFA0FD0EF8B4FD0AF831" bold="true" box="[643,710,1895,1921]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A23757C75855FFA0FD0EF8B4FD0AF831" box="[643,710,1895,1921]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
frequently inhabit
<taxonomicName id="4C704D2D5855FFA0FC24F8BBFF1EF81A" authority="Zea and Rutzler, 1983" authorityName="Zea and Rutzler" authorityYear="1983" class="Demospongiae" family="Petrosiidae" genus="Xestospongia" kingdom="Animalia" order="Haplosclerida" pageId="16" pageNumber="17" phylum="Porifera" rank="species" species="rosariensis">
<emphasis id="B904EABC5855FFA0FC24F8BBFB1CF831" box="[937,1232,1896,1921]" italics="true" pageId="16" pageNumber="17">Xestospongia rosariensis</emphasis>
Zea and Rützler, 1983
</taxonomicName>
(
<figureCitation id="134B2A2B5855FFA0FF6FF843FEFBF81A" box="[226,311,1936,1962]" captionStart="FIGURE 15" captionStartId="22.[151,255,1475,1499]" captionTargetBox="[151,1434,195,1451]" captionTargetId="figure@22.[151,1436,194,1451]" captionTargetPageId="22" captionText="FIGURE 15. Sponge hosts of some species from the Synalpheus paraneptunus Coutière, 1909 species complex: A, several members of a colony of Synalpheus duffyi n. sp. in their host sponge, possibly Neopetrosia subtriangularis, Isla Grande, Panama; B, same, close-up of a non-breeding colony member; C, Xestospongia rosariensis, main host of Synalpheus bocas n. sp., Bocas del Toro, Panama; D, same, longitudinal section showing details of canals; E, Xestospongia sp., occasional host of S. bocas n. sp., Bocas del Toro, Panama. Scale bars: ~ 3 mm (A, B), ~ 5 cm (C – E)." httpUri="https://zenodo.org/record/184611/files/figure.png" pageId="16" pageNumber="17">Fig. 15</figureCitation>
C, D), often together with congeners, such as the pair-living
<taxonomicName id="4C704D2D5855FFA0FC73F843FA50F81A" authority="Rios and Duffy, 2007" authorityName="Rios and Duffy" authorityYear="2007" box="[1022,1436,1936,1962]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dardeaui">
<emphasis id="B904EABC5855FFA0FC73F843FB49F819" box="[1022,1157,1936,1961]" italics="true" pageId="16" pageNumber="17">S. dardeaui</emphasis>
(
<bibRefCitation id="EFE14B5F5855FFA0FB19F843FA58F81A" author="Rios" box="[1172,1428,1936,1962]" pageId="16" pageNumber="28" refString="Rios, R. and Duffy, J. E. (2007) A review of the sponge-dwelling snapping shrimp from Carrie Bow Cay, Belize, with description of Zuzalpheus, new genus, and six new species (Crustacea: Decapoda: Alpheidae). Zootaxa, 1602, 1 - 82." type="journal article" year="2007">Ríos and Duffy, 2007</bibRefCitation>
)
</taxonomicName>
and the eusocial
<taxonomicName id="4C704D2D5855FFA0FED1F86BFD67F862" authority="Coutiere, 1909" authorityName="Coutiere" authorityYear="1909" box="[348,683,1976,2002]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rathbunae">
<emphasis id="B904EABC5855FFA0FED1F86BFE3DF861" box="[348,497,1976,2001]" italics="true" pageId="16" pageNumber="17">S. rathbunae</emphasis>
<bibRefCitation id="EFE14B5F5855FFA0FE74F86BFD67F862" author="Coutiere" box="[505,683,1976,2002]" pageId="16" pageNumber="28" refString="Coutiere, H. (1909) The American species of snapping shrimps of the genus Synalpheus. Proceedings of the United States National Museum, 36 (1659), 1 - 93." type="journal article" year="1909">Coutière, 1909</bibRefCitation>
</taxonomicName>
. It also occurs in single sexual pairs in the sponge
<taxonomicName id="4C704D2D5855FFA0FB73F86BFA50F861" box="[1278,1436,1976,2001]" class="Demospongiae" family="Petrosiidae" genus="Xestospongia" kingdom="Animalia" order="Haplosclerida" pageId="16" pageNumber="17" phylum="Porifera" rank="genus">
<emphasis id="B904EABC5855FFA0FB73F86BFA50F861" box="[1278,1436,1976,2001]" italics="true" pageId="16" pageNumber="17">Xestospongia</emphasis>
</taxonomicName>
sp. (
<figureCitation id="134B2A2B5854FFA1FF47FF4BFEECFF02" box="[202,288,152,178]" captionStart="FIGURE 15" captionStartId="22.[151,255,1475,1499]" captionTargetBox="[151,1434,195,1451]" captionTargetId="figure@22.[151,1436,194,1451]" captionTargetPageId="22" captionText="FIGURE 15. Sponge hosts of some species from the Synalpheus paraneptunus Coutière, 1909 species complex: A, several members of a colony of Synalpheus duffyi n. sp. in their host sponge, possibly Neopetrosia subtriangularis, Isla Grande, Panama; B, same, close-up of a non-breeding colony member; C, Xestospongia rosariensis, main host of Synalpheus bocas n. sp., Bocas del Toro, Panama; D, same, longitudinal section showing details of canals; E, Xestospongia sp., occasional host of S. bocas n. sp., Bocas del Toro, Panama. Scale bars: ~ 3 mm (A, B), ~ 5 cm (C – E)." httpUri="https://zenodo.org/record/184611/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
E), which superficially is intermediate between
<taxonomicName id="4C704D2D5854FFA1FCD6FF4BFED7FF6A" authority="Duchassaing and Michelotti, 1864" authorityName="Duchassaing and Michelotti" authorityYear="1864" class="Demospongiae" family="Petrosiidae" genus="Xestospongia" kingdom="Animalia" order="Haplosclerida" pageId="17" pageNumber="18" phylum="Porifera" rank="species" species="proxima">
<emphasis id="B904EABC5854FFA1FCD6FF4BFBAFFF01" box="[859,1123,152,177]" italics="true" pageId="17" pageNumber="18">Xestospongia proxima</emphasis>
(Duchassaing and Michelotti, 1864)
</taxonomicName>
and
<taxonomicName id="4C704D2D5854FFA1FEDAFF13FCE3FF6A" authority="Duchassaing, 1850" authorityName="Duchassaing" authorityYear="1850" box="[343,815,192,218]" class="Demospongiae" family="Petrosiidae" genus="Neopetrosia" kingdom="Animalia" order="Haplosclerida" pageId="17" pageNumber="18" phylum="Porifera" rank="species" species="subtriangularis">
<emphasis id="B904EABC5854FFA1FEDAFF13FDE3FF69" box="[343,559,192,217]" italics="true" pageId="17" pageNumber="18">X. subtriangularis</emphasis>
(Duchassaing, 1850)
</taxonomicName>
, and perhaps also in
<taxonomicName id="4C704D2D5854FFA1FBA1FF13FB2AFF69" box="[1068,1254,192,217]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="17" pageNumber="18" phylum="Porifera" rank="species" species="podatypa">
<emphasis id="B904EABC5854FFA1FBA1FF13FB2AFF69" box="[1068,1254,192,217]" italics="true" pageId="17" pageNumber="18">Calyx podatypa</emphasis>
</taxonomicName>
(E. Tóth, pers. obs.). The larvae of
<taxonomicName id="4C704D2D5854FFA1FE07FF3BFE27FEB1" box="[394,491,232,257]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">
<emphasis id="B904EABC5854FFA1FE07FF3BFE27FEB1" box="[394,491,232,257]" italics="true" pageId="17" pageNumber="18">S. bocas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23757C75854FFA1FE7EFF34FDFAFEB1" box="[499,566,231,257]" pageId="17" pageNumber="18" rank="species">
<emphasis id="B904EABC5854FFA1FE7EFF34FDFAFEB1" bold="true" box="[499,566,231,257]" pageId="17" pageNumber="18">n. sp.</emphasis>
</taxonomicNameLabel>
appear to be pelagic (E. Tóth, pers. obs.).
</paragraph>
</subSubSection>
<subSubSection id="C36A65255854FFA1FF48FEDCFC38FDF2" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BCF36AE5854FFA1FF48FEDCFC38FDF2" blockId="17.[151,1436,152,578]" pageId="17" pageNumber="18">
<emphasis id="B904EABC5854FFA1FF48FEDCFEFBFE99" bold="true" box="[197,311,271,297]" pageId="17" pageNumber="18">Remarks</emphasis>
.
<taxonomicName id="4C704D2D5854FFA1FECBFEC3FDDAFE99" box="[326,534,272,297]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">
<emphasis id="B904EABC5854FFA1FECBFEC3FDDAFE99" box="[326,534,272,297]" italics="true" pageId="17" pageNumber="18">Synalpheus bocas</emphasis>
</taxonomicName>
<emphasis id="B904EABC5854FFA1FD93FEDCFDACFE99" bold="true" box="[542,608,271,297]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23757C75854FFA1FD93FEDCFDACFE99" box="[542,608,271,297]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from all previous species, viz.
<taxonomicName id="4C704D2D5854FFA1FB92FEC3FB12FE99" box="[1055,1246,272,297]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="paraneptunus">
<emphasis id="B904EABC5854FFA1FB92FEC3FB12FE99" box="[1055,1246,272,297]" italics="true" pageId="17" pageNumber="18">S. paraneptunus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C704D2D5854FFA1FB61FEC3FA80FE99" box="[1260,1356,272,297]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="duffyi" status="sp. nov.">
<emphasis id="B904EABC5854FFA1FB61FEC3FA80FE99" box="[1260,1356,272,297]" italics="true" pageId="17" pageNumber="18">S. duffyi</emphasis>
</taxonomicName>
<emphasis id="B904EABC5854FFA1FADEFEDCFA59FE99" bold="true" box="[1363,1429,271,297]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23757C75854FFA1FADEFEDCFA59FE99" box="[1363,1429,271,297]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C704D2D5854FFA1FF1AFEEBFF20FEE1" box="[151,236,312,337]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="riosi" status="sp. nov.">
<emphasis id="B904EABC5854FFA1FF1AFEEBFF20FEE1" box="[151,236,312,337]" italics="true" pageId="17" pageNumber="18">S. riosi</emphasis>
</taxonomicName>
<emphasis id="B904EABC5854FFA1FF78FEE4FEF5FEE1" bold="true" box="[245,313,311,337]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23757C75854FFA1FF78FEE4FEF5FEE1" box="[245,313,311,337]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C704D2D5854FFA1FEF0FEEBFDF5FEE1" box="[381,569,312,338]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brevidactylus" status="sp. nov.">
<emphasis id="B904EABC5854FFA1FEF0FEEBFE47FEE1" box="[381,395,312,337]" italics="true" pageId="17" pageNumber="18">S</emphasis>
.
<emphasis id="B904EABC5854FFA1FE11FEEBFDF5FEE1" box="[412,569,312,337]" italics="true" pageId="17" pageNumber="18">brevidactylus</emphasis>
</taxonomicName>
<emphasis id="B904EABC5854FFA1FDCEFEE4FD4BFEE1" bold="true" box="[579,647,311,337]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23757C75854FFA1FDCEFEE4FD4BFEE1" box="[579,647,311,337]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the consistent absence of a blade on the antennal scaphocerite (
<figureCitation id="134B2A2B5854FFA1FF12FEB3FF39FECA" box="[159,245,352,378]" captionStart="FIGURE 10" captionStartId="17.[151,255,1907,1931]" captionTargetBox="[178,1406,627,1877]" captionTargetId="figure@17.[169,1413,616,1902]" captionTargetPageId="17" captionText="FIGURE 10. Synalpheus bocas n. sp., paratype, male from Bocas del Toro, Panama (USNM 1116692): A, frontal region, dorsal view; B, same, lateral view; C, third maxilliped, tip of ultimate segment, lateral view; D, second pereiopod, lateral view; E, third pereiopod, lateral view; F, abdomen, lateral view; G, telson and uropods, dorsal view. Scale bar: 0.5 mm (A – E, G), 1 mm (F)." httpUri="https://zenodo.org/record/184606/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
A); the presence of only two teeth on the uropodal exopod (
<figureCitation id="134B2A2B5854FFA1FC35FEB3FBC3FECA" box="[952,1039,352,378]" captionStart="FIGURE 10" captionStartId="17.[151,255,1907,1931]" captionTargetBox="[178,1406,627,1877]" captionTargetId="figure@17.[169,1413,616,1902]" captionTargetPageId="17" captionText="FIGURE 10. Synalpheus bocas n. sp., paratype, male from Bocas del Toro, Panama (USNM 1116692): A, frontal region, dorsal view; B, same, lateral view; C, third maxilliped, tip of ultimate segment, lateral view; D, second pereiopod, lateral view; E, third pereiopod, lateral view; F, abdomen, lateral view; G, telson and uropods, dorsal view. Scale bar: 0.5 mm (A – E, G), 1 mm (F)." httpUri="https://zenodo.org/record/184606/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
G); the rostrum being subequal in length to the orbital teeth (
<figureCitation id="134B2A2B5854FFA1FE55FE5BFDE3FE12" box="[472,559,392,418]" captionStart="FIGURE 10" captionStartId="17.[151,255,1907,1931]" captionTargetBox="[178,1406,627,1877]" captionTargetId="figure@17.[169,1413,616,1902]" captionTargetPageId="17" captionText="FIGURE 10. Synalpheus bocas n. sp., paratype, male from Bocas del Toro, Panama (USNM 1116692): A, frontal region, dorsal view; B, same, lateral view; C, third maxilliped, tip of ultimate segment, lateral view; D, second pereiopod, lateral view; E, third pereiopod, lateral view; F, abdomen, lateral view; G, telson and uropods, dorsal view. Scale bar: 0.5 mm (A – E, G), 1 mm (F)." httpUri="https://zenodo.org/record/184606/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
A); and the third maxilliped distally armed with a crown of eight or nine spine-like setae (vs. at most six in the other species) (
<figureCitation id="134B2A2B5854FFA1FC9BFE63FCA1FE7A" box="[790,877,432,458]" captionStart="FIGURE 10" captionStartId="17.[151,255,1907,1931]" captionTargetBox="[178,1406,627,1877]" captionTargetId="figure@17.[169,1413,616,1902]" captionTargetPageId="17" captionText="FIGURE 10. Synalpheus bocas n. sp., paratype, male from Bocas del Toro, Panama (USNM 1116692): A, frontal region, dorsal view; B, same, lateral view; C, third maxilliped, tip of ultimate segment, lateral view; D, second pereiopod, lateral view; E, third pereiopod, lateral view; F, abdomen, lateral view; G, telson and uropods, dorsal view. Scale bar: 0.5 mm (A – E, G), 1 mm (F)." httpUri="https://zenodo.org/record/184606/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
C). In life,
<taxonomicName id="4C704D2D5854FFA1FC7FFE63FB98FE79" box="[1010,1108,432,457]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bocas" status="sp. nov.">
<emphasis id="B904EABC5854FFA1FC7FFE63FB98FE79" box="[1010,1108,432,457]" italics="true" pageId="17" pageNumber="18">S. bocas</emphasis>
</taxonomicName>
<emphasis id="B904EABC5854FFA1FBD0FE7CFB6DFE79" bold="true" box="[1117,1185,431,457]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23757C75854FFA1FBD0FE7CFB6DFE79" box="[1117,1185,431,457]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C704D2D5854FFA1FF5AFE0BFEFBFE41" box="[215,311,472,497]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="duffyi" status="sp. nov.">
<emphasis id="B904EABC5854FFA1FF5AFE0BFEFBFE41" box="[215,311,472,497]" italics="true" pageId="17" pageNumber="18">S. duffyi</emphasis>
</taxonomicName>
<emphasis id="B904EABC5854FFA1FEB3FE04FE4CFE41" bold="true" box="[318,384,471,497]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23757C75854FFA1FEB3FE04FE4CFE41" box="[318,384,471,497]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C704D2D5854FFA1FE35FE0BFDBEFE41" box="[440,626,472,497]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brevidactylus" status="sp. nov.">
<emphasis id="B904EABC5854FFA1FE35FE0BFDBEFE41" box="[440,626,472,497]" italics="true" pageId="17" pageNumber="18">S. brevidactylus</emphasis>
</taxonomicName>
<emphasis id="B904EABC5854FFA1FDF4FE04FD77FE41" bold="true" box="[633,699,471,497]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23757C75854FFA1FDF4FE04FD77FE41" box="[633,699,471,497]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(the only two species with known colour patterns) by the presence of diffuse bands of reddish chromatophores on the abdomen and in some areas of the carapace. It also differs from
<taxonomicName id="4C704D2D5854FFA1FEA7FDFBFE47FDF1" box="[298,395,552,577]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="duffyi" status="sp. nov.">
<emphasis id="B904EABC5854FFA1FEA7FDFBFE47FDF1" box="[298,395,552,577]" italics="true" pageId="17" pageNumber="18">S. duffyi</emphasis>
</taxonomicName>
<emphasis id="B904EABC5854FFA1FE1FFDF4FE18FDF1" bold="true" box="[402,468,551,577]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23757C75854FFA1FE1FFDF4FE18FDF1" box="[402,468,551,577]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by its much larger size in adults (see
<tableCitation id="C6F203155854FFA1FC00FDFBFC2FFDF2" box="[909,995,552,578]" captionStart="TABLE 1" captionStartId="25.[151,239,151,175]" captionTargetBox="[151,1436,220,664]" captionTargetPageId="25" captionText="TABLE 1. Characters useful for species separation in the Synalpheus paraneptunus Coutière, 1909 complex." httpUri="http://table.plazi.org/id/DF0F6626585CFFA9FF1AFF44FAEFFF1F" pageId="17" pageNumber="18" tableUuid="DF0F6626585CFFA9FF1AFF44FAEFFF1F">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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