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<document id="239CC973419711FC7EBACBACCB0BB2CB" ID-DOI="10.11646/zootaxa.4809.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3936907" ID-ZooBank="8FE54BBA-250D-4109-8023-1A28864A7D97" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594292004760" checkinUser="plazi" docAuthor="Hsueh, Pan-Wen" docDate="2020" docId="03D9878E833AFFE6FF18FBAF46969A87" docLanguage="en" docName="zootaxa.4809.3.7.pdf" docOrigin="Zootaxa 4809 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Garthasia ludaoensis Hsueh 2020, n. sp." docType="treatment" docVersion="3" lastPageNumber="540" masterDocId="FFE0FFF6833BFFE3FF8FFF95473A9F60" masterDocTitle="New species and record of xanthid crabs (Crustacean, Decapoda) from Taiwan" masterLastPageNumber="546" masterPageNumber="535" pageNumber="536" updateTime="1700277360130" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="327DFA2ABBE7136F6AAE0487DF72F89C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8E0552314C1F9F569AEA412B857B2B02">
<mods:title id="38A4B41385F5FC392A35B3DC6CAF73F0">New species and record of xanthid crabs (Crustacean, Decapoda) from Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="1D04529FF974B6148622C4E4AFF3222F" type="personal">
<mods:role id="19110D79DA02E22613539019C0B162FE">
<mods:roleTerm id="1C9550564C0C23E65446A7BDE38BB137">Author</mods:roleTerm>
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<mods:namePart id="975966FB63BE1D205BB3E5526B06EBAA">Hsueh, Pan-Wen</mods:namePart>
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<mods:typeOfResource id="3EF1AF4F25245CADA3C9BC5D46C1C1AA">text</mods:typeOfResource>
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<mods:title id="C9A8ADF9AAA4B68B2A6B620BF8428709">Zootaxa</mods:title>
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<mods:part id="C1D0F6AE521FB1F2A5BAFC533FD0CA21">
<mods:date id="F5B119DDC36BCBECF46365570FF124C9">2020</mods:date>
<mods:detail id="2EDAF4EB2F0736DE1D59B16A71BD06C5" type="pubDate">
<mods:number id="A9FDADA35B04D0D18E51BACF3BCE6E91">2020-07-08</mods:number>
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<mods:detail id="4F6219043F44254617B1A69C92DE68BB" type="volume">
<mods:number id="DC20F1F37914A401348AE108DFEE5879">4809</mods:number>
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<mods:detail id="76AC0992DD0214B3D26C4D7CB1117500" type="issue">
<mods:number id="0DD24A5F8C59B080B523CA24BC431C42">3</mods:number>
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<mods:classification id="FCE494AB04C024B53AB55B3DAD11F590">journal article</mods:classification>
<mods:identifier id="C7EA3199420E485F5D8CC380CD1264C2" type="DOI">10.11646/zootaxa.4809.3.7</mods:identifier>
<mods:identifier id="162B9A2A53CD360A0356E5E8E484F9BD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C6C9951A7FB4977BE7ABB8AE19278AAE" type="Zenodo-Dep">3936907</mods:identifier>
<mods:identifier id="69632267B97C0A75F24A8DE3B51B6A5D" type="ZooBank">8FE54BBA-250D-4109-8023-1A28864A7D97</mods:identifier>
</mods:mods>
<treatment id="03D9878E833AFFE6FF18FBAF46969A87" LSID="urn:lsid:plazi:treatment:03D9878E833AFFE6FF18FBAF46969A87" httpUri="http://treatment.plazi.org/id/03D9878E833AFFE6FF18FBAF46969A87" lastPageId="5" lastPageNumber="540" pageId="1" pageNumber="536">
<subSubSection id="C36A6513833AFFE2FF18FBAF46D89B35" box="[151,482,1082,1109]" pageId="1" pageNumber="536" type="nomenclature">
<paragraph id="8BCF3698833AFFE2FF18FBAF46D89B35" blockId="1.[151,482,1082,1145]" box="[151,482,1082,1109]" pageId="1" pageNumber="536">
<heading id="D08781F4833AFFE2FF18FBAF46D89B35" bold="true" box="[151,482,1082,1109]" fontSize="11" level="1" pageId="1" pageNumber="536" reason="1">
<emphasis id="B904EA8A833AFFE2FF18FBAF46D89B35" bold="true" box="[151,482,1082,1109]" pageId="1" pageNumber="536">
<taxonomicName id="4C704D1B833AFFE2FF18FBAF46A29B35" ID-CoL="84GTY" authority="Hsueh, 2020" authorityName="Hsueh" authorityYear="2020" box="[151,408,1082,1109]" class="Malacostraca" family="Xanthidae" genus="Garthasia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="536" phylum="Arthropoda" rank="species" species="ludaoensis" status="sp. nov.">
<emphasis id="B904EA8A833AFFE2FF18FBAF46A29B35" bold="true" box="[151,408,1082,1109]" italics="true" pageId="1" pageNumber="536">Garthasia ludaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23757F1833AFFE2FE10FBAE46D89B35" box="[415,482,1083,1109]" pageId="1" pageNumber="536" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A6513833AFFE2FF18FBCA468F9B19" box="[151,437,1119,1145]" pageId="1" pageNumber="536" type="description">
<paragraph id="8BCF3698833AFFE2FF18FBCA468F9B19" blockId="1.[151,482,1082,1145]" box="[151,437,1119,1145]" pageId="1" pageNumber="536">
<figureCitation id="134B2A1D833AFFE2FF18FBCA47ED9B19" box="[151,215,1119,1145]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="1" pageNumber="536">Figs 1</figureCitation>
A–G, 2A–E, 3A–G
</paragraph>
</subSubSection>
<subSubSection id="C36A6513833AFFE2FF18FB3346319A69" pageId="1" pageNumber="536" type="materials_examined">
<paragraph id="8BCF3698833AFFE2FF18FB3346319A69" blockId="1.[151,1437,1190,2045]" pageId="1" pageNumber="536">
<emphasis id="B904EA8A833AFFE2FF18FB3346AA9BA1" bold="true" box="[151,400,1190,1217]" pageId="1" pageNumber="536">Material Examined.</emphasis>
<materialsCitation id="3B183CC5833AFFE2FE10FB3246319A69" collectingDate="2011-08-08" collectionCode="T" country="Taiwan" county="Hong" elevation="2" latitude="22.675833" location="Gongguan Port" longLatPrecision="12" longitude="121.49367" municipality="Ludao Township" pageId="1" pageNumber="536" specimenCode="NMNS 8194-001" specimenCount="1" stateProvince="Taitung" typeStatus="holotype">
<typeStatus id="54CB883A833AFFE2FE10FB32453D9BA1" box="[415,519,1191,1217]" pageId="1" pageNumber="536" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DBD69EE3833AFFE2FD90FB3245CA9BA1" box="[543,752,1191,1217]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="1" pageNumber="536">NMNS 8194-001</specimenCode>
), male, 12.8 by
<quantity id="4C889B7D833AFFE2FC49FB3243099BA1" box="[966,1075,1191,1217]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="1" pageNumber="536" unit="mm" value="10.6">10.6 mm</quantity>
,
<location id="8EAF6043833AFFE2FBC5FB3243C79BA1" LSID="urn:lsid:plazi:treatment:03D9878E833AFFE6FF18FBAF46969A87:8EAF6043833AFFE2FBC5FB3243C79BA1" box="[1098,1277,1191,1217]" country="Taiwan" county="Hong" latitude="22.675833" longLatPrecision="12" longitude="121.49367" municipality="Ludao Township" name="Gongguan Port" pageId="1" pageNumber="536" stateProvince="Taitung">Gongguan Port</location>
(
<geoCoordinate id="EE44505F833AFFE2FA9BFB3342A29BA1" box="[1300,1432,1190,1217]" degrees="22" direction="north" minutes="40.55" orientation="latitude" pageId="1" pageNumber="536" precision="9" value="22.675833">22°40.55'N</geoCoordinate>
,
<geoCoordinate id="EE44505F833AFFE2FF18FB5E461F9B86" box="[151,293,1227,1254]" degrees="121" direction="east" minutes="29.62" orientation="longitude" pageId="1" pageNumber="536" precision="9" value="121.49367">121°29.62'E</geoCoordinate>
),
<collectingMunicipality id="6BABACE2833AFFE2FEB5FB5E46C09B85" box="[314,506,1227,1253]" pageId="1" pageNumber="536">Ludao Township</collectingMunicipality>
,
<collectingRegion id="49B4F87A833AFFE2FD8AFB5E458D9B85" box="[517,695,1227,1253]" country="Taiwan" name="Taitung" pageId="1" pageNumber="536">Taitung County</collectingRegion>
,
<collectingCountry id="F3677608833AFFE2FD4DFB5E442D9B85" box="[706,791,1227,1253]" name="Taiwan" pageId="1" pageNumber="536">Taiwan</collectingCountry>
, subtidal hard bottom,
<quantity id="4C889B7D833AFFE2FB93FB5E43729B85" box="[1052,1096,1227,1253]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="536" unit="m" value="2.0">
<elevation id="005DD1AB833AFFE2FB93FB5E43729B85" box="[1052,1096,1227,1253]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="536" unit="m" value="2.0">2 m</elevation>
</quantity>
deep, coll. H.-
<collectionCode id="ED61AE5D833AFFE2FB7FFB5E423E9B85" box="[1264,1284,1227,1253]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="536">T</collectionCode>
.
<collectingCounty id="62AE4E14833AFFE2FA80FB5E426A9B85" box="[1295,1360,1227,1253]" pageId="1" pageNumber="536">Hong</collectingCounty>
,
<date id="FFCE1058833AFFE2FAD3FB5E463D9A69" pageId="1" pageNumber="536" value="2011-08-08">
<collectingDate id="EF8AE9B0833AFFE2FAD3FB5E463D9A69" pageId="1" pageNumber="536" value="2011-08-08">8 August 2011</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A6513833AFFE2FF48FA874210992D" pageId="1" pageNumber="536" type="diagnosis">
<paragraph id="8BCF3698833AFFE2FF48FA874210992D" blockId="1.[151,1437,1190,2045]" pageId="1" pageNumber="536">
<emphasis id="B904EA8A833AFFE2FF48FA8746789A4D" bold="true" box="[199,322,1298,1325]" pageId="1" pageNumber="536">Diagnosis.</emphasis>
Carapace oval, broader than long, length to width ratio about 1.21; dorsal surface evenly convex, region inconspicuous defined, granulose entirely except interspaces between regions; front bi-lobed with arched margins; anterolateral margin distinctly shorter than posterolateral margin, with small granules; frontal, anterolateral and anterior part of posterolateral margins with many long setae; posterolateral margin gently convex. Orbits as slits in dorsal view, eye small. Merus of third maxilliped rectangular, width to length ratio about 1.29. Chelipeds subequal in size, armed with many small teeth on dorsal margin of propodus and dactylus and outer surface of propodus; fingers subequal in length, strongly curved downward distally, gaping in a broad oval when closed. Sternum slightly elongates, without suture between sternites 1 and 2, surfaces of sternites 1 and 2 smooth. Surface of male abdomen smooth, pleonites 1 and 2 visible dorsally, telson semicircular, reaching posterior of sternum 4.
</paragraph>
</subSubSection>
<subSubSection id="C36A6513833AFFE6FF48F9C343B99C77" lastPageId="5" lastPageNumber="540" pageId="1" pageNumber="536" type="description">
<paragraph id="8BCF3698833AFFE2FF48F9C343BD989D" blockId="1.[151,1437,1190,2045]" pageId="1" pageNumber="536">
<emphasis id="B904EA8A833AFFE2FF48F9C346639911" bold="true" box="[199,345,1622,1649]" pageId="1" pageNumber="536">Description.</emphasis>
Carapace oval, broader than long, width to length ratio about 1.21 (
<figureCitation id="134B2A1D833AFFE2FBE5F9C243909911" box="[1130,1194,1623,1649]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="1" pageNumber="536">Fig. 1</figureCitation>
A–C); dorsal surface evenly convex, region inconspicuously defined, granulose entirely except interspaces between regions (
<figureCitation id="134B2A1D833AFFE2FA98F9EE42B599F5" box="[1303,1423,1659,1685]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="1" pageNumber="536">Fig. 1B, C</figureCitation>
). Front bi-lobed, separated medially by a long narrowed cleft, lobes arched, frontal margin about 0.4 times width of carapace, finely granulated with many forward-pointing long setae (
<figureCitation id="134B2A1D833AFFE2FC06F95644FC99BD" box="[905,966,1731,1757]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="1" pageNumber="536">Fig. 1</figureCitation>
A–C). Anterolateral margin arched, finely granulated with forward-pointing long setae, shorter than posterolateral margin. Anterior of posterolateral margin arched, finely granulated, converging towards gently convex posterior carapace margin with smooth surface (
<figureCitation id="134B2A1D833AFFE2FAE0F89E479A9829" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="1" pageNumber="536">Fig. 1</figureCitation>
A–C, E). Orbits poorly defined; eyes visible dorsally through an acute angular notch. Basal antennal segment rectangular, clearly separating antennular pits from orbits, flagellum short, project beyond front, antennules folding obliquely (
<figureCitation id="134B2A1D833AFFE2FE9EF8E2465D98F1" box="[273,359,1911,1937]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="1" pageNumber="536">Fig. 1E</figureCitation>
). Third maxilliped longitudinal, slender, length to width ratio about 2.70, fully covering buccal field when closed, outer margins of merus and carpus with forward-pointing long setae; ischium rectangular, length to width ratio about 1.25; merus rectangular, wider than long, width to length ratio about 1.29, outer and inner angles rounded; carpus slightly inflated, approximately one-fourth as large as merus (
<figureCitation id="134B2A1D833AFFE2FB8AF8764341989D" box="[1029,1147,2019,2045]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="1" pageNumber="536">Fig. 1F, G</figureCitation>
).
</paragraph>
<caption id="DF0F66108339FFE1FF18F8E547D69894" ID-DOI="http://doi.org/10.5281/zenodo.3936909" ID-Zenodo-Dep="3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="2" pageNumber="537" startId="2.[151,250,1904,1929]" targetBox="[165,1422,187,1878]" targetPageId="2">
<paragraph id="8BCF36988339FFE1FF18F8E547D69894" blockId="2.[151,1436,1904,2036]" pageId="2" pageNumber="537">
<emphasis id="B904EA8A8339FFE1FF18F8E5462F98E9" bold="true" box="[151,277,1904,1929]" pageId="2" pageNumber="537">FIGURE 1.</emphasis>
<taxonomicName id="4C704D1B8339FFE1FE91F8E546C398E8" authority="Hsueh, 2020" authorityName="Hsueh" authorityYear="2020" box="[286,505,1904,1928]" class="Malacostraca" family="Xanthidae" genus="Garthasia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="ludaoensis" status="sp. nov.">
<emphasis id="B904EA8A8339FFE1FE91F8E546C398E8" box="[286,505,1904,1928]" italics="true" pageId="2" pageNumber="537">Garthasia ludaoensis</emphasis>
</taxonomicName>
<emphasis id="B904EA8A8339FFE1FD8DF8E5450798E8" bold="true" box="[514,573,1904,1928]" pageId="2" pageNumber="537">
<taxonomicNameLabel id="A23757F18339FFE1FD8DF8E5450798E8" box="[514,573,1904,1928]" pageId="2" pageNumber="537" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm.
</paragraph>
</caption>
<caption id="DF0F66108338FFE0FF18FA74461D9921" ID-DOI="http://doi.org/10.5281/zenodo.3936911" ID-Zenodo-Dep="3936911" httpUri="https://zenodo.org/record/3936911/files/figure.png" pageId="3" pageNumber="538" startId="3.[151,250,1505,1530]" targetBox="[171,1417,178,1480]" targetPageId="3">
<paragraph id="8BCF36988338FFE0FF18FA74461D9921" blockId="3.[151,1436,1505,1602]" pageId="3" pageNumber="538">
<emphasis id="B904EA8A8338FFE0FF18FA74462F9A9A" bold="true" box="[151,277,1505,1530]" pageId="3" pageNumber="538">FIGURE 2.</emphasis>
<taxonomicName id="4C704D1B8338FFE0FE92FA7746C29A9A" authority="Hsueh, 2020" authorityName="Hsueh" authorityYear="2020" box="[285,504,1506,1530]" class="Malacostraca" family="Xanthidae" genus="Garthasia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="ludaoensis" status="sp. nov.">
<emphasis id="B904EA8A8338FFE0FE92FA7746C29A9A" box="[285,504,1506,1530]" italics="true" pageId="3" pageNumber="538">Garthasia ludaoensis</emphasis>
</taxonomicName>
<emphasis id="B904EA8A8338FFE0FD8FFA7745019A9A" bold="true" box="[512,571,1506,1530]" pageId="3" pageNumber="538">
<taxonomicNameLabel id="A23757F18338FFE0FD8FFA7745019A9A" box="[512,571,1506,1530]" pageId="3" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(NMNS 8194-001): A, chelae, outer view; B, right chela, outer view; C, the fourth ambulatory leg; D, the sternum; E, line drawing of sternum, setae on surface of sternites 1–4 not shown. Scale: A–C, 2.0 mm; D, E, 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BCF36988338FFE0FF48F9E546BA9892" blockId="3.[151,1437,1647,2034]" pageId="3" pageNumber="538">
Chelipeds subequal in size, right chela slightly larger than left chela, right palm about 1.25 times length of left palm (
<figureCitation id="134B2A1D8338FFE0FF51F901466F99CE" box="[222,341,1684,1710]" captionStart="FIGURE 2" captionStartId="3.[151,250,1505,1530]" captionTargetBox="[171,1417,178,1480]" captionTargetId="figure@3.[171,1417,174,1480]" captionTargetPageId="3" captionText="FIGURE 2. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, chelae, outer view; B, right chela, outer view; C, the fourth ambulatory leg; D, the sternum; E, line drawing of sternum, setae on surface of sternites 1–4 not shown. Scale: A–C, 2.0 mm; D, E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936911" httpUri="https://zenodo.org/record/3936911/files/figure.png" pageId="3" pageNumber="538">Fig. 2A, B</figureCitation>
); merus rectangular, length to width ratio about 1.29, unarmed; carpus rectangular, anterior margin with a row of granules and forward-pointing long setae; propodus rectangular, anterior upheaved dorsally, dorsal margin with 2–3 rows of teeth, outer surface with scattered pinches of golden setae, anterior half of outer surface covered with rows of minute teeth, inner margin of outer surface with a row of small teeth running from posterior to anterior of propodus; fingers subequal in length, curved nearly 90° downward distally, leaving an oval gape when fully closed, 2 rows of small teeth present on dorsal margin of dactylus, forming a groove, cutting edge of right chela dactylus with a large triangular tooth proximally, a trace of a blunt tooth for left chela at same location, cutting edge of right chela pollex with 2 rectangular, offset, large teeth distally and 1 large triangular tooth proximally (
<figureCitation id="134B2A1D8338FFE0FAE0F805478198AE" captionStart="FIGURE 2" captionStartId="3.[151,250,1505,1530]" captionTargetBox="[171,1417,178,1480]" captionTargetId="figure@3.[171,1417,174,1480]" captionTargetPageId="3" captionText="FIGURE 2. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, chelae, outer view; B, right chela, outer view; C, the fourth ambulatory leg; D, the sternum; E, line drawing of sternum, setae on surface of sternites 1–4 not shown. Scale: A–C, 2.0 mm; D, E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936911" httpUri="https://zenodo.org/record/3936911/files/figure.png" pageId="3" pageNumber="538">Fig. 2B</figureCitation>
), pollex of left chela with a row of 3 small teeth at same location, distal one largest (
<figureCitation id="134B2A1D8338FFE0FBE2F82143F298AE" box="[1133,1224,1972,1998]" captionStart="FIGURE 2" captionStartId="3.[151,250,1505,1530]" captionTargetBox="[171,1417,178,1480]" captionTargetId="figure@3.[171,1417,174,1480]" captionTargetPageId="3" captionText="FIGURE 2. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, chelae, outer view; B, right chela, outer view; C, the fourth ambulatory leg; D, the sternum; E, line drawing of sternum, setae on surface of sternites 1–4 not shown. Scale: A–C, 2.0 mm; D, E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936911" httpUri="https://zenodo.org/record/3936911/files/figure.png" pageId="3" pageNumber="538">Fig. 2A</figureCitation>
). Colour of fingers whitish (
<figureCitation id="134B2A1D8338FFE0FF76F84D46499892" box="[249,371,2008,2034]" captionStart="FIGURE 2" captionStartId="3.[151,250,1505,1530]" captionTargetBox="[171,1417,178,1480]" captionTargetId="figure@3.[171,1417,174,1480]" captionTargetPageId="3" captionText="FIGURE 2. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, chelae, outer view; B, right chela, outer view; C, the fourth ambulatory leg; D, the sternum; E, line drawing of sternum, setae on surface of sternites 1–4 not shown. Scale: A–C, 2.0 mm; D, E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936911" httpUri="https://zenodo.org/record/3936911/files/figure.png" pageId="3" pageNumber="538">Fig. 2A, B</figureCitation>
).
</paragraph>
<caption id="DF0F6610833FFFE7FF18F8DA43359897" ID-DOI="http://doi.org/10.5281/zenodo.3936913" ID-Zenodo-Dep="3936913" httpUri="https://zenodo.org/record/3936913/files/figure.png" pageId="4" pageNumber="539" startId="4.[151,250,1871,1896]" targetBox="[151,1436,186,1845]" targetPageId="4">
<paragraph id="8BCF3698833FFFE7FF18F8DA43359897" blockId="4.[151,1437,1871,2040]" pageId="4" pageNumber="539">
<emphasis id="B904EA8A833FFFE7FF18F8DA46289808" bold="true" box="[151,274,1871,1896]" pageId="4" pageNumber="539">FIGURE 3.</emphasis>
<taxonomicName id="4C704D1B833FFFE7FE96F8C546CB9808" authority="Hsueh, 2020" authorityName="Hsueh" authorityYear="2020" box="[281,497,1872,1896]" class="Malacostraca" family="Xanthidae" genus="Garthasia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="539" phylum="Arthropoda" rank="species" species="ludaoensis" status="sp. nov.">
<emphasis id="B904EA8A833FFFE7FE96F8C546CB9808" box="[281,497,1872,1896]" italics="true" pageId="4" pageNumber="539">Garthasia ludaoensis</emphasis>
</taxonomicName>
<emphasis id="B904EA8A833FFFE7FE77F8C5450A9808" bold="true" box="[504,560,1872,1896]" pageId="4" pageNumber="539">
<taxonomicNameLabel id="A23757F1833FFFE7FE77F8C5450A9808" box="[504,560,1872,1896]" pageId="4" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(NMNS 8194-001): A, the male abdomen; B, line drawing of the male abdomen, setae on margins of all pleonites and telson not shown; C, left G1, arrow points the lappet next to the distal opening; D, line drawing of left G1, setae and spines not shown; E, the distal area of left G1, arrows point the lappet and minute spines on surface of the distal opening; F, left G2; G, close-up of left G2; H, close-up of the tip of the distal segment on left G2, arrows point minute and small spines. Scale: A, B, 2.0 mm; C, D, 1.0 mm; E, H, 0.1 mm; F, 0.5 mm; G, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BCF3698833EFFE6FF48FF0C44B99E07" blockId="5.[151,1437,153,1511]" pageId="5" pageNumber="540">
Ambulatory legs slender, first leg longest, fourth leg shortest with relatively robust merus (length to width ratio on 1–3 legs ranging about 1.96–2.45 versus that of 1.76 on fourth leg); merus of fourth leg unarmed, outer surface of merus, carpus and propodus smooth; 1–3 rows of small tooth present on outer margins of carpus, propodus and dactylus; a chitinous nail present at distal of dactylus; basis, ischium, carpus and propodus of fourth leg with a tuft of setae at each inner anterior corner, a row of long setae at outer margins of merus, carpus, propodus and inner margin of merus, short setae present at both margins of dactylus (
<figureCitation id="134B2A1D833EFFE6FC92FED8444C9E07" box="[797,886,333,359]" captionStart="FIGURE 2" captionStartId="3.[151,250,1505,1530]" captionTargetBox="[171,1417,178,1480]" captionTargetId="figure@3.[171,1417,174,1480]" captionTargetPageId="3" captionText="FIGURE 2. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, chelae, outer view; B, right chela, outer view; C, the fourth ambulatory leg; D, the sternum; E, line drawing of sternum, setae on surface of sternites 1–4 not shown. Scale: A–C, 2.0 mm; D, E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936911" httpUri="https://zenodo.org/record/3936911/files/figure.png" pageId="5" pageNumber="540">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8BCF3698833EFFE6FF48FEE443CE9D5F" blockId="5.[151,1437,153,1511]" pageId="5" pageNumber="540">
Sternum slightly elongates, surface of sternites covered with many setae, suture between sternites 1 and 2 absent, sternites 2 and 3 separated by straight groove, sternites 3 and 4 separated by shallow grooves curving towards abdominal cavity, a minute longitudinal slit present at center between sternites 4 and 5 (
<figureCitation id="134B2A1D833EFFE6FB08FE2C423F9EB3" box="[1159,1285,441,467]" captionStart="FIGURE 2" captionStartId="3.[151,250,1505,1530]" captionTargetBox="[171,1417,178,1480]" captionTargetId="figure@3.[171,1417,174,1480]" captionTargetPageId="3" captionText="FIGURE 2. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, chelae, outer view; B, right chela, outer view; C, the fourth ambulatory leg; D, the sternum; E, line drawing of sternum, setae on surface of sternites 1–4 not shown. Scale: A–C, 2.0 mm; D, E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936911" httpUri="https://zenodo.org/record/3936911/files/figure.png" pageId="5" pageNumber="540">Fig. 2D, E</figureCitation>
). Male abdomen with 6 somites, pleonites 1 and 2 visible in dorsal view, pleonite 2 semicircular, pleonite 3 widest, 3–5 pleonites fused, pleonite 6 rectangular, telson semicircular, reaching posterior of sternum 4, dorsal surface of abdomen smooth, margins of all pleonites and telson covered with short to medium-length setae (
<figureCitation id="134B2A1D833EFFE6FBE2FDB043DC9D5F" box="[1133,1254,549,575]" captionStart="FIGURE 3" captionStartId="4.[151,250,1871,1896]" captionTargetBox="[151,1436,186,1845]" captionTargetId="figure@4.[151,1436,181,1846]" captionTargetPageId="4" captionText="FIGURE 3. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, the male abdomen; B, line drawing of the male abdomen, setae on margins of all pleonites and telson not shown; C, left G1, arrow points the lappet next to the distal opening; D, line drawing of left G1, setae and spines not shown; E, the distal area of left G1, arrows point the lappet and minute spines on surface of the distal opening; F, left G2; G, close-up of left G2; H, close-up of the tip of the distal segment on left G2, arrows point minute and small spines. Scale: A, B, 2.0 mm; C, D, 1.0 mm; E, H, 0.1 mm; F, 0.5 mm; G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936913" httpUri="https://zenodo.org/record/3936913/files/figure.png" pageId="5" pageNumber="540">Fig. 3A, B</figureCitation>
).
</paragraph>
<paragraph id="8BCF3698833EFFE6FF48FDDC43B99C77" blockId="5.[151,1437,153,1511]" pageId="5" pageNumber="540">
G1 long, slender, mid-region nearly straight, slightly curved outwards subdistally, distal sharply tapering to sharp point, a lappet present next to distal opening, a tuft of long setae present near distal opening, subdistal region lined with few medium-length to long setae, surfaces of distal to mid-region covered with dense minute spines and some relatively large spines (
<figureCitation id="134B2A1D833EFFE6FE6FFD2045189DAF" box="[480,546,693,719]" captionStart="FIGURE 3" captionStartId="4.[151,250,1871,1896]" captionTargetBox="[151,1436,186,1845]" captionTargetId="figure@4.[151,1436,181,1846]" captionTargetPageId="4" captionText="FIGURE 3. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, the male abdomen; B, line drawing of the male abdomen, setae on margins of all pleonites and telson not shown; C, left G1, arrow points the lappet next to the distal opening; D, line drawing of left G1, setae and spines not shown; E, the distal area of left G1, arrows point the lappet and minute spines on surface of the distal opening; F, left G2; G, close-up of left G2; H, close-up of the tip of the distal segment on left G2, arrows point minute and small spines. Scale: A, B, 2.0 mm; C, D, 1.0 mm; E, H, 0.1 mm; F, 0.5 mm; G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936913" httpUri="https://zenodo.org/record/3936913/files/figure.png" pageId="5" pageNumber="540">Fig. 3</figureCitation>
C–E); G2 short, basal segment distal end round, upper margin truncate without upright protrusion, distal segment elongate, slightly curved, sharply tapering to sharp point at distal area, tip spatuliform, surfaces of both segments covered with dense minute to small spines (
<figureCitation id="134B2A1D833EFFE6FC71FD6843049C77" box="[1022,1086,765,791]" captionStart="FIGURE 3" captionStartId="4.[151,250,1871,1896]" captionTargetBox="[151,1436,186,1845]" captionTargetId="figure@4.[151,1436,181,1846]" captionTargetPageId="4" captionText="FIGURE 3. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, the male abdomen; B, line drawing of the male abdomen, setae on margins of all pleonites and telson not shown; C, left G1, arrow points the lappet next to the distal opening; D, line drawing of left G1, setae and spines not shown; E, the distal area of left G1, arrows point the lappet and minute spines on surface of the distal opening; F, left G2; G, close-up of left G2; H, close-up of the tip of the distal segment on left G2, arrows point minute and small spines. Scale: A, B, 2.0 mm; C, D, 1.0 mm; E, H, 0.1 mm; F, 0.5 mm; G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936913" httpUri="https://zenodo.org/record/3936913/files/figure.png" pageId="5" pageNumber="540">Fig. 3</figureCitation>
F–H).
</paragraph>
</subSubSection>
<subSubSection id="C36A6513833EFFE6FF48FCB446BA9CC7" pageId="5" pageNumber="540" type="etymology">
<paragraph id="8BCF3698833EFFE6FF48FCB447D89C3F" blockId="5.[151,1437,153,1511]" pageId="5" pageNumber="540">
<emphasis id="B904EA8A833EFFE6FF48FCB446749C5B" bold="true" box="[199,334,801,827]" pageId="5" pageNumber="540">Etymology.</emphasis>
The specific name is derived from the name of the Ludao Township, where the crab was collected.
</paragraph>
<paragraph id="8BCF3698833EFFE6FF48FCFD46BA9CC7" blockId="5.[151,1437,153,1511]" pageId="5" pageNumber="540">
<emphasis id="B904EA8A833EFFE6FF48FCFD461A9CE3" bold="true" box="[199,288,872,899]" pageId="5" pageNumber="540">Colour.</emphasis>
In fresh specimen, carapace surface light greenish with scatted brown speckles, becoming beige in alcohol (
<figureCitation id="134B2A1D833EFFE6FF76FC1846499CC7" box="[249,371,909,935]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="5" pageNumber="540">Fig. 1A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6513833EFFE6FF48FC2443389CAB" box="[199,1026,944,971]" pageId="5" pageNumber="540" type="materials_examined">
<paragraph id="8BCF3698833EFFE6FF48FC2443389CAB" blockId="5.[151,1437,153,1511]" box="[199,1026,944,971]" pageId="5" pageNumber="540">
<materialsCitation id="3B183CC5833EFFE6FF48FC2444C49CAB" box="[199,1022,944,971]" country="Taiwan" location="Gongguan Port" municipality="Ludao Township" pageId="5" pageNumber="540" specimenCount="1" stateProvince="Taitung" typeStatus="holotype">
<emphasis id="B904EA8A833EFFE6FF48FC24465C9CAB" bold="true" box="[199,358,944,971]" pageId="5" pageNumber="540">
<typeStatus id="54CB883A833EFFE6FF48FC2446389CAB" box="[199,258,945,971]" pageId="5" pageNumber="540">Type</typeStatus>
locality.
</emphasis>
<location id="8EAF6043833EFFE6FEE3FC2445209CAB" LSID="urn:lsid:plazi:treatment:03D9878E833AFFE6FF18FBAF46969A87:8EAF6043833EFFE6FEE3FC2445209CAB" box="[364,538,945,971]" country="Taiwan" municipality="Ludao Township" name="Gongguan Port" pageId="5" pageNumber="540" stateProvince="Taitung">Gongguan Port</location>
,
<collectingMunicipality id="6BABACE2833EFFE6FDAAFC2445DE9CAB" box="[549,740,945,971]" pageId="5" pageNumber="540">Ludao Township</collectingMunicipality>
,
<collectingRegion id="49B4F87A833EFFE6FD61FC2444A49CAB" box="[750,926,945,971]" country="Taiwan" name="Taitung" pageId="5" pageNumber="540">Taitung County</collectingRegion>
,
<collectingCountry id="F3677608833EFFE6FC27FC2444C49CAB" box="[936,1022,945,971]" name="Taiwan" pageId="5" pageNumber="540">Taiwan</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6513833EFFE6FF48FC4145CB9C8F" box="[199,753,980,1007]" pageId="5" pageNumber="540" type="distribution">
<paragraph id="8BCF3698833EFFE6FF48FC4145CB9C8F" blockId="5.[151,1437,153,1511]" box="[199,753,980,1007]" pageId="5" pageNumber="540">
<emphasis id="B904EA8A833EFFE6FF48FC41465B9C8F" bold="true" box="[199,353,980,1007]" pageId="5" pageNumber="540">Distribution.</emphasis>
Only known from the
<typeStatus id="54CB883A833EFFE6FDEFFC4045AA9C8F" box="[608,656,981,1007]" pageId="5" pageNumber="540">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C36A6513833EFFE6FF48FC6C46969A87" pageId="5" pageNumber="540" type="discussion">
<paragraph id="8BCF3698833EFFE6FF48FC6C43859BA7" blockId="5.[151,1437,153,1511]" pageId="5" pageNumber="540">
<emphasis id="B904EA8A833EFFE6FF48FC6C46019B73" bold="true" box="[199,315,1017,1043]" pageId="5" pageNumber="540">Remarks.</emphasis>
<taxonomicName id="4C704D1B833EFFE6FECDFC6C45099B73" authority="Hsueh, 2020" authorityName="Hsueh" authorityYear="2020" box="[322,563,1017,1043]" class="Malacostraca" family="Xanthidae" genus="Garthasia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="540" phylum="Arthropoda" rank="species" species="ludaoensis" status="sp. nov.">
<emphasis id="B904EA8A833EFFE6FECDFC6C45099B73" box="[322,563,1017,1043]" italics="true" pageId="5" pageNumber="540">Garthasia ludaoensis</emphasis>
</taxonomicName>
<emphasis id="B904EA8A833EFFE6FDB5FC6C45409B73" bold="true" box="[570,634,1017,1043]" pageId="5" pageNumber="540">
<taxonomicNameLabel id="A23757F1833EFFE6FDB5FC6C45409B73" box="[570,634,1017,1043]" pageId="5" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
shares several characters with
<taxonomicName id="4C704D1B833EFFE6FC58FC6C42289B73" authority="(Garth, 1939)" baseAuthorityName="Garth" baseAuthorityYear="1939" box="[983,1298,1017,1043]" class="Malacostraca" family="Xanthidae" genus="Kraussia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="540" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B904EA8A833EFFE6FC58FC6C44D19B73" box="[983,1003,1017,1043]" italics="true" pageId="5" pageNumber="540">G</emphasis>
.
<emphasis id="B904EA8A833EFFE6FC75FC6F43489B73" box="[1018,1138,1018,1043]" italics="true" pageId="5" pageNumber="540">americana</emphasis>
(
<bibRefCitation id="EFE14B69833EFFE6FB0EFC6C42339B73" author="Garth, J. S." box="[1153,1289,1017,1043]" pageId="5" pageNumber="540" pagination="9 - 29" refId="ref5610" refString="Garth, J. S. (1939) New brachyuran crabs from the Galapagos Islands. Allan Hancock Pacific Expeditions, 5 (2), 9 - 29, pls. 1 - 10." type="journal article" year="1939">Garth, 1939</bibRefCitation>
)
</taxonomicName>
on: 1) ovalshaped carapace with many forward-pointing long setae on margins of front, anterolateral and anterior portion of posterolateral of the carapace; 2), front separated medially by a long narrowed cleft into two lobes; 3) whitish chelipeds with distal portion of fingers strongly bend, leaving a broad oval gap when closing; 4) poorly developed orbit region; 5) slightly elongated sternum without suture between sternites 1 and 2; and 6) telson inserted to posterior of sternum 4 (
<bibRefCitation id="EFE14B69833EFFE6FE9AFB3846F99BA7" author="Garth, J. S." box="[277,451,1197,1223]" pageId="5" pageNumber="540" pagination="9 - 29" refId="ref5610" refString="Garth, J. S. (1939) New brachyuran crabs from the Galapagos Islands. Allan Hancock Pacific Expeditions, 5 (2), 9 - 29, pls. 1 - 10." type="journal article" year="1939">Garth 1939: 20</bibRefCitation>
, pl. 7, figs 1, 3;
<bibRefCitation id="EFE14B69833EFFE6FDF7FB3844769BA7" author="Ng, P. K. L." box="[632,844,1196,1223]" pageId="5" pageNumber="540" pagination="133 - 157" refId="ref5882" refString="Ng, P. K. L. (1993) Kraussiinae, a new subfamily for the genera Kraussia Dana, 1852, Palapedia, new genus, and Garthasia, new genus (Crustacea: Decapoda: Brachyura: Xanthidae), with descriptions of two new species from Singapore and the Philippines. The Raffles Bulletin of Zoology, 41 (1), 133 - 157." type="journal article" year="1993">Ng 1993: 153–154</bibRefCitation>
, fig. 8A, D, E;
<figureCitation id="134B2A1D833EFFE6FC78FB3843899BA7" box="[1015,1203,1197,1223]" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="5" pageNumber="540">Fig. 1A, B, E, H</figureCitation>
).
</paragraph>
<paragraph id="8BCF3698833EFFE6FF48FB4446969A87" blockId="5.[151,1437,153,1511]" pageId="5" pageNumber="540">
However,
<taxonomicName id="4C704D1B833EFFE6FEB3FB4446ED9B8B" authority="Hsueh, 2020" authorityName="Hsueh" authorityYear="2020" box="[316,471,1233,1259]" class="Malacostraca" family="Xanthidae" genus="Garthasia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="540" phylum="Arthropoda" rank="species" species="ludaoensis" status="sp. nov.">
<emphasis id="B904EA8A833EFFE6FEB3FB44466A9B8B" box="[316,336,1233,1259]" italics="true" pageId="5" pageNumber="540">G</emphasis>
.
<emphasis id="B904EA8A833EFFE6FED0FB4446ED9B8B" box="[351,471,1233,1259]" italics="true" pageId="5" pageNumber="540">ludaoensis</emphasis>
</taxonomicName>
<emphasis id="B904EA8A833EFFE6FE50FB4445249B8B" bold="true" box="[479,542,1233,1259]" pageId="5" pageNumber="540">
<taxonomicNameLabel id="A23757F1833EFFE6FE50FB4445249B8B" box="[479,542,1233,1259]" pageId="5" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C704D1B833EFFE6FCDBFB4444D59B8B" baseAuthorityName="Garth" baseAuthorityYear="1939" box="[852,1007,1233,1259]" class="Malacostraca" family="Xanthidae" genus="Kraussia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="540" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B904EA8A833EFFE6FCDBFB4444529B8B" box="[852,872,1233,1259]" italics="true" pageId="5" pageNumber="540">G</emphasis>
.
<emphasis id="B904EA8A833EFFE6FCF8FB4744D59B8B" box="[887,1007,1234,1259]" italics="true" pageId="5" pageNumber="540">americana</emphasis>
</taxonomicName>
by having: 1) smaller width to length ratio (1.21 versus 1.35) of the carapace; 2) larger width to length ratio (3.2 versus 2.4) of the third maxilliped; 3) smaller width to length ratio of merus of the third maxilliped (1.3 versus 2.2); 4) no distinct longitudinal keel on surfaces of sternites 1 and 2; 5) smaller length to width ratio of merus (1.8 versus 2.7) of the last ambulatory leg; 6) smaller width to length ratio (2.1 versus 3.0) of pleonite 2 of the male abdomen; 7) pleonite
<quantity id="4C889B7D833EFFE6FB03FAF4438C9A1B" box="[1164,1206,1377,1403]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="540" unit="in" value="2.0">2 in</quantity>
semicircular-shaped (versus roughly rectangular); 8) a tuft of long setae (versus bared) near distal opening of G1; and 9) distal end of basal segment without upright protrusion (versus with upright protrusion) of G2 (
<bibRefCitation id="EFE14B69833EFFE6FBAEFA3C43849AA3" author="Ng, P. K. L." box="[1057,1214,1448,1475]" pageId="5" pageNumber="540" pagination="133 - 157" refId="ref5882" refString="Ng, P. K. L. (1993) Kraussiinae, a new subfamily for the genera Kraussia Dana, 1852, Palapedia, new genus, and Garthasia, new genus (Crustacea: Decapoda: Brachyura: Xanthidae), with descriptions of two new species from Singapore and the Philippines. The Raffles Bulletin of Zoology, 41 (1), 133 - 157." type="journal article" year="1993">Ng 1993: 153</bibRefCitation>
,155, fig. 8E–I;
<figureCitation id="134B2A1D833EFFE6FAE3FA3C479A9A87" captionStart="FIGURE 1" captionStartId="2.[151,250,1904,1929]" captionTargetBox="[165,1422,187,1878]" captionTargetId="figure@2.[165,1422,181,1878]" captionTargetPageId="2" captionText="FIGURE 1. Garthasia ludaoensis n. sp. (NMNS 8194-001): A, fresh animal; B, whole animal, dorsal view; C, close-up of the carapace; D, whole animal, ventral view; E, antennae and antennules areas, arrows pointed antennal segments; F, left third maxilliped; G, line drawing of left third maxilliped, setae on margins of the endopod not shown. Scale: A–D, F, G, 2.0 mm; E, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3936909" httpUri="https://zenodo.org/record/3936909/files/figure.png" pageId="5" pageNumber="540">Figs 1</figureCitation>
A–C, 2D, E, 3A, E–H).
</paragraph>
</subSubSection>
</treatment>
</document>
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